In-Class Active Learning Exercise: Seedless and Seed Plants

Total Page:16

File Type:pdf, Size:1020Kb

In-Class Active Learning Exercise: Seedless and Seed Plants ALE 4 Plants – BSC2011, Spring 2020 In-Class Active Learning Exercise: Seedless and Seed Plants Introduction: Land plants acquired traits that made it possible to colonize land and survive out of the water. All land plants share the following characteristics: alternation of generations, with the haploid plant called a gametophyte, and the diploid plant called a sporophyte; formation of haploid spores in a sporangium; formation of gametes in a gametangium; protection of the embryo; and an apical meristem. Nonvascular seedless plants do not possess vascular conductive tissue to transport water and glucose, therefore they are very small and live in moist environments. Vascular seedless plants have an additional adaptation of conductive vascular tissue, xylem and phloem. Vascular plants are not limited in size and can inhabit drier environments. All land plants possess a waxy cuticle that prevents desiccation, and a tough sporopollenin layer that protects the spores. Land plants appeared about 500 million years ago in the Ordovician period. Objectives: • Learn vocabulary pertinent to plant structure • Understand Alternation of Generations in both seedless and seed plants • Distinguish between gametophytes and sporophytes in both seedless and seed plants • Describe plant stem and root anatomy • Describe differences between gymnosperm and angiosperm life cycles • Explain double fertilization in angiosperms Preparation related to the two assignments on this worksheet: • Read Chapters 25, 26, 30, and 32 • Answer the pre-lecture questions on Chapters 25, 26, 30, and 32 • Attend the lectures on Chapters 25, 26, 30 and 32 and actively participate in iClicker questions Grading: You have ~25 minutes to work through this worksheet in small groups. Afterwards, you’ll get 10 iClicker questions that you will have to answer individually. Each question is worth 1 point: 0.5 point participation + 0.5 point correct answer. These points contribute to your overall iClicker score, which is 10% of your final grade. De Bekker - 1 - ALE 4 Plants – BSC2011, Spring 2020 Assignment 1: Bryophyte Structure (~7 mins) Bryophytes are small, seedless, nonvascular plants. They are the closest living relatives of the earliest land plants. They undergo Alternation of Generations with a dominant gametophyte stage. As nonvascular plants, they do not have true leaves or stems and they do not have conductive vascular tissue. Bryophytes include liverworts, hornworts, and mosses. 1A Bryophyte Alternation of Generations Bryophytes have an alternation of generations life cycle in which the haploid gametophyte is the dominant stage. The gametophyte produces gametes which form a sporophyte upon fertilization. The sporophyte produces haploid spores which germinate into a gametophyte. Label each box as mitosis or meiosis. Label each of the following as diploid (2n) or haploid (n): gametophytes, sperm, egg, zygote, sporophytes, and spores. 1. In seedless plants, a fertilized egg will develop into a. A gametophyte b. A fruit c. A sporophyte d. Gametes e. Spores 2. The diploid generation of the plant life cycle always a. Is larger and more conspicuous than the haploid generation b. Is called the gametophyte c. Develops from a spore d. Produces spores e. Produces eggs and sperm De Bekker - 2 - ALE 4 Plants – BSC2011, Spring 2020 1B Life Cycle of Mosses Arrange the events in the life cycle of mosses in the flowchart below. Start the life cycle with the mature sporophyte stage in target 1. Not all labels will be used. 1C Terrestrial Plant Adaptations 1. Which of the following traits was the most important in enabling the first plants to move onto land? a. Rings of cellulose synthesizing complexes b. Alternation of Generations c. Apical meristems d. Development of sporopollenin to prevent desiccation of the zygote e. Peroxisome enzymes that minimize losses from photorespiration 2. What evolutionary development allowed terrestrial plants to grow tall? a. Sporophylls b. Waxy cuticle c. Rhizoids d. Leaves e. Lignified vascular tissue De Bekker - 3 - ALE 4 Plants – BSC2011, Spring 2020 Assignment 2: Plant Structure (~5 mins) 2A Building vocabulary Learning common prefixes and root words assists learning complex plant vocabulary. De Bekker - 4 - ALE 4 Plants – BSC2011, Spring 2020 2B Stems Vascular plants have conductive vascular tissue, xylem and phloem. Xylem cells transports water and dissolved minerals upward from the roots to the leaves of the plant. Phloem cells transport glucose from the leaves to various parts of the plants. Monocots are the grasses; they have parallel leaf venation and fibrous roots. Cross sections of monocot stems and roots show scattered vascular bundles. All other plants are Eudicots (sometimes called dicots); they have branched leaf venation and a main taproot. Cross sections of eudicots show vascular bundles that form a ring. 1. Label the parts of the stems 2. The primary growth of a plant adds ________ and the secondary growth adds ______. a. Height, branching b. Branching, girth c. Girth, height d. Branching, flowers e. Height, girth 3. Label the areas of the root De Bekker - 5 - ALE 4 Plants – BSC2011, Spring 2020 4. Cell division in the roots occurs in the _______________. a. Root cap b. Area of Elongation c. Apical Meristem d. Area of Maturation Assignment 3: Seed Plants (~10 mins) Seed plants are vascular plants with a dominant diploid sporophyte stage. Two major innovations of seed plants are seeds and pollen. Seeds protect the embryo from desiccation and provide it with a store of nutrients to support the early growth of the sporophyte. Seeds are also equipped to delay germination until growth conditions are optimal. Pollen allows seed plants to reproduce in the absence of water. In the gymnosperms pollen is dispersed by wind, and their naked seeds developed in the sporophylls of a strobilus. Angiosperms bear both flowers and fruit. Flowers expand the possibilities for pollination, especially by insects. Fruits offer additional protection to the embryo during its development. Angiosperms have become the dominant plant life in terrestrial habitats. 3A Gymnosperm Life Cycle Gymnosperms have male cones and female cones. The male cones contain microsporangia that produce haploid microspores via meiosis. Each microspore undergoes mitosis to produce two nuclei, the generative nucleus and the tube nucleus, which is pollen, the male gametophyte. The female cones contain megasporangia that contain megaspore mother cells which produce four haploid megaspores via meiosis. One megaspore divides via mitosis to produce the female gametophyte, the other three divide to produce the archegonia. Label the diagram De Bekker - 6 - ALE 4 Plants – BSC2011, Spring 2020 3B Gymnosperm Reproduction 1. In pine trees, the embryo develops within the _______. a. Macrogametophyte b. Pollen cone c. Female gametophyte d. Male gametophyte e. Microsporophyll 3C Angiosperm Life Cycle 2. Angiosperms are different from all other plants because only they have _______. a. A vascular system b. A life cycle that involves alternation of generations c. Seeds d. Flowers e. A sporophyte phase 3. The major difference between angiosperms and gymnosperms come from the a. Production of microspores versus megaspores b. Presence or absence of a protective covering over the ovule c. Presence or absence of vascular structures d. Dominance of sporophyte in angiosperms and gametophyte in gymnosperms e. Presence or absence of alternation of generations 3D Flower Structure Label the parts of the flower De Bekker - 7 - ALE 4 Plants – BSC2011, Spring 2020 3E Angiosperm Life Cycle Some angiosperm flowers contain both female and male reproductive structures. The male anther contains microsporangia which produce microspore mother cells. These mother cells produce haploid microspores (pollen) via meiosis. Then each pollen divides via mitosis to produce a pollen tube cell and a generative cell. The female carpel contains megasporangia which produce megaspore mother cells. These mother cells produce 4 haploid megaspores via meiosis. One lives and the other three die. That one then undergoes mitosis without cytokinesis to produce a gametophyte with seven cells and eight nuclei. Two of the nuclei fuse to form a single diploid polar body. Angiosperms undergo double fertilization to produce the zygote and the endosperm which provides nutrition to the zygote. When the pollen lands on the stigma, one of the cell, the pollen tube cell germinates and grows through the style to reach the ovule. The other pollen cell, the generative cell, divides to form two sperm cells. One sperm cell fertilizes the egg to form the zygote, the other sperm cell joins with two polar nuclei in the ovule to form the endosperm. 4. Which of these is unique to flowering plants? a. Pollen production b. Dominant sporophyte c. Double fertilization d. Embryo surrounded by nutritive tissue e. Haploid gametophytes 5. In angiosperms, each pollen grain produces two sperm. What do these sperm do? a. Each one fertilizes a separate egg cell b. One fertilizes an egg, the other combines with two polar nuclei which develop into the endosperm c. One fertilizes an egg, the other fertilizes the fruit d. Both sperm fertilize a single egg e. One fertilizes and egg, the other is kept in reserve De Bekker - 8 - .
Recommended publications
  • Tansley Review Evolution of Development of Vascular Cambia and Secondary Growth
    New Phytologist Review Tansley review Evolution of development of vascular cambia and secondary growth Author for correspondence: Rachel Spicer1 and Andrew Groover2 Andrew Groover 1The Rowland Institute at Harvard, Cambridge, MA, USA; 2Institute of Forest Genetics, Pacific Tel: +1 530 759 1738 Email: [email protected] Southwest Research Station, USDA Forest Service, Davis, CA, USA Received: 29 December 2009 Accepted: 14 February 2010 Contents Summary 577 V. Evolution of development approaches for the study 587 of secondary vascular growth I. Introduction 577 VI. Conclusions 589 II. Generalized function of vascular cambia and their 578 developmental and evolutionary origins Acknowledgements 589 III. Variation in secondary vascular growth in angiosperms 581 References 589 IV. Genes and mechanisms regulating secondary vascular 584 growth and their evolutionary origins Summary New Phytologist (2010) 186: 577–592 Secondary growth from vascular cambia results in radial, woody growth of stems. doi: 10.1111/j.1469-8137.2010.03236.x The innovation of secondary vascular development during plant evolution allowed the production of novel plant forms ranging from massive forest trees to flexible, Key words: forest trees, genomics, Populus, woody lianas. We present examples of the extensive phylogenetic variation in sec- wood anatomy, wood formation. ondary vascular growth and discuss current knowledge of genes that regulate the development of vascular cambia and woody tissues. From these foundations, we propose strategies for genomics-based research in the evolution of development, which is a next logical step in the study of secondary growth. I. Introduction this pattern characterizes most extant forest trees, significant variation exists among taxa, ranging from extinct woody Secondary vascular growth provides a means of radially lycopods and horsetails with unifacial cambia (Cichan & thickening and strengthening plant axes initiated during Taylor, 1990; Willis & McElwain, 2002), to angiosperms primary, or apical growth.
    [Show full text]
  • Chapter 5: the Shoot System I: the Stem
    Chapter 5 The Shoot System I: The Stem THE FUNCTIONS AND ORGANIZATION OF THE SHOOT SYSTEM PRIMARY GROWTH AND STEM ANATOMY Primary Tissues of Dicot Stems Develop from the Primary Meristems The Distribution of the Primary Vascular Bundles Depends on the Position of Leaves Primary Growth Differs in Monocot and Dicot Stems SECONDARY GROWTH AND THE ANATOMY OF WOOD Secondary Xylem and Phloem Develop from Vascular Cambium Wood Is Composed of Secondary Xylem Gymnosperm Wood Differs from Angiosperm Wood Bark Is Composed of Secondary Phloem and Periderm Buds Are Compressed Branches Waiting to Elongate Some Monocot Stems Have Secondary Growth STEM MODIFICATIONS FOR SPECIAL FUNCTIONS THE ECONOMIC VALUE OF WOODY STEMS SUMMARY ECONOMIC BOTANY: How Do You Make A Barrel? 1 KEY CONCEPTS 1. The shoot system is composed of the stem and its lateral appendages: leaves, buds, and flowers. Leaves are arranged in different patterns (phyllotaxis): alternate, opposite, whorled, and spiral. 2. Stems provide support to the leaves, buds, and flowers. They conduct water and nutrients and produce new cells in meristems (shoot apical meristem, primary and secondary meristems). 3. Dicot stems and monocot stems are usually different. Dicot stems tend to have vascular bundles distributed in a ring, whereas in monocot stems they tend to be scattered. 4. Stems are composed of the following: epidermis, cortex and pith, xylem and phloem, and periderm. 5. Secondary xylem is formed by the division of cells in the vascular cambium and is called wood. The bark is composed of all of the tissues outside the vascular cambium, including the periderm (formed from cork cambium) and the secondary phloem.
    [Show full text]
  • SECONDARY GROWTH in PLANTS Compiled and Circulated by Arpita Chakraborty, Govt.Approved Part-Time Teacher, Narajole Raj College, Narajole
    COMPILED AND CIRCULATED BY ARPITA CHAKRABORTY, GOVT. APPROVED PART TIME TEACHER, DEPARTMENT OF BOTANY, NARAJOLE RAJ COLLEGE. SECONDARY GROWTH IN PLANTS compiled and circulated by Arpita Chakraborty, Govt.approved Part-time teacher, Narajole Raj College, Narajole. BOTANY: SEM- IV, PAPER: GE4T:PLANT ANATOMY AND EMBRYOLOGY:UNIT-3:SECONDARY GROWTH COMPILED AND CIRCULATED BY ARPITA CHAKRABORTY, GOVT. APPROVED PART TIME TEACHER, DEPARTMENT OF BOTANY, NARAJOLE RAJ COLLEGE. •CHAPTER OUT LINE- • 1. Overview of secondary growth • 2. Growth patterns in wood and bark • 3. Commercial Uses of wood and bark BOTANY: SEM- IV, PAPER: GE4T:PLANT ANATOMY AND EMBRYOLOGY:UNIT-3:SECONDARY GROWTH COMPILED AND CIRCULATED BY ARPITA CHAKRABORTY, GOVT. APPROVED PART TIME TEACHER, DEPARTMENT OF BOTANY, NARAJOLE RAJ COLLEGE. CHAPTER OBJECTIVES- Students should have an idea of; 1. How wood and bark develop 2. How stems and roots become thicker and stronger 3. Commercial benefits of wood and bark of a plant with secondary growth BOTANY: SEM- IV, PAPER: GE4T:PLANT ANATOMY AND EMBRYOLOGY:UNIT-3:SECONDARY GROWTH COMPILED AND CIRCULATED BY ARPITA CHAKRABORTY, GOVT. APPROVED PART TIME TEACHER, DEPARTMENT OF BOTANY, NARAJOLE RAJ COLLEGE. SECONDARY GROWTH- Cambial 1.Vascular cambium a)Fusiform Initials (Vertically oriented) Secondary Xylem Secondary Phloem b)Ray Initials (Horizontally oriented) Vascular Rays Xylem rays Phloem ray 2.Cork cambium (Phellogen) Periderm Phellem (Cork cells) Phelloderm (Cork Parenchyma) BOTANY: SEM- IV, PAPER: GE4T:PLANT ANATOMY AND EMBRYOLOGY:UNIT-3:SECONDARY
    [Show full text]
  • Secondary Tree Growth Increments 11-07
    Secondary Tree Growth Increments: Ring Development & Forms by Dr. Kim D. Coder, Professor of Tree Biology & Health Care, University of Georgia Trees attempt to occupy more space and control available resources through cell divisions and cell expansion. From the end of a shoot tip to the end of a root tip, trees elongate. Along this axis of growth, trees also expand radially, which is termed “secondary growth.” Tree growth is initiated in the shoot tips (growing points / buds), root tips, vascular cambium (i.e. or simply cambium), and phellogen which generates the periderm. The first two meristems are primary generation systems and the second two are termed secondary meristems. Cambial activity and xylem formation, visible as increases in growth increment volume, is radial growth, and reviewed here. The secondary meristem phellogen, sometimes generically referred to as the cork cambium, will not be discussed here although it does increase tree diameter. Cambium The cambium is a cell generation zone which blends (within several cell thicknesses) inward into the xylem (wood) and outward into the phloem (inner bark). The cambium occupies the circumference of the wood cylinder which comprises the bulk of a tree. The cambium is responsible for modifying cell divisions, cell forms, and cell wall materials in response to mechanical, biological and defensive stresses at each point along its surface. The cambium receives signals from sensors both internally (from divid- ing and expanding cells in the cambial zone) and externally (located locally farther inside and outside the cambium, and from shoot tips and root tips.) A tree radially expands tissues as shoots and roots elongate.
    [Show full text]
  • Tree Anatomy Stems and Branches
    Tree Anatomy Series WSFNR14-13 Nov. 2014 COMPONENTSCOMPONENTS OFOF PERIDERMPERIDERM by Dr. Kim D. Coder, Professor of Tree Biology & Health Care Warnell School of Forestry & Natural Resources, University of Georgia Around tree roots, stems and branches is a complex tissue. This exterior tissue is the environmental face of a tree open to all sorts of site vulgarities. This most exterior of tissue provides trees with a measure of protection from a dry, oxidative, heat and cold extreme, sunlight drenched, injury ridden site. The exterior of a tree is both an ecological super highway and battle ground – comfort and terror. This exterior is unique in its attributes, development, and regeneration. Generically, this tissue surrounding a tree stem, branch and root is loosely called bark. The tissues of a tree, outside or more exterior to the xylem-containing core, are varied and complexly interwoven in a relatively small space. People tend to see and appreciate the volume and physical structure of tree wood and dismiss the remainder of stem, branch and root. In reality, tree life is focused within these more exterior thin tissue sets. Outside of the cambium are tissues which include transport cells, structural support cells, generation cells, and cells positioned to help, protect, and sustain other cells. All of this life is smeared over the circumference of a predominately dead physical structure. Outer Skin Periderm (jargon and antiquated term = bark) is the most external of tree tissues providing protection, water conservation, insulation, and environmental sensing. Periderm is a protective tissue generated over and beyond live conducting and non-conducting cells of the food transport system (phloem).
    [Show full text]
  • Brassinosteroid Regulation of Wood Formation in Poplar
    Research Brassinosteroid regulation of wood formation in poplar Juan Du1,2,3*, Suzanne Gerttula3*, Zehua Li2, Shu-Tang Zhao2, Ying-Li Liu2, Yu Liu1, Meng-Zhu Lu2,4 and Andrew T. Groover3,5 1College of Life Sciences, Zhejiang University, 866 Yu Hang tang Road, Hangzhou 310058, China; 2State Key Laboratory of Tree Genetics and Breeding, Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, China; 3Pacific Southwest Research Station, US Forest Service, Davis, CA 95618, USA; 4State Key Laboratory of Subtropical Silviculture, School of Forestry and Biotechnology, Zhejiang Agriculture and Forest University, Hangzhou 311300, China; 5Department of Plant Biology, University of California Davis, Davis, CA 95616, USA Summary Authors for correspondence: Brassinosteroids have been implicated in the differentiation of vascular cell types in herba- Meng-Zhu Lu ceous plants, but their roles during secondary growth and wood formation are not well Tel: +1 86 10 62872015 defned. Email: [email protected] Here we pharmacologically and genetically manipulated brassinosteroid levels in poplar Andrew Groover trees and assayed the effects on secondary growth and wood formation, and on gene expres- Tel: +1 530 759 1738 sion within stems. Email: [email protected] Elevated brassinosteroid levels resulted in increases in secondary growth and tension wood Received: 6 March 2019 formation, while inhibition of brassinosteroid synthesis resulted in decreased growth and sec- Accepted: 30 April 2019 ondary vascular differentiation. Analysis of gene expression showed that brassinosteroid action is positively associated with genes involved in cell differentiation and cell-wall biosyn- New Phytologist (2020) 225: 1516–1530 thesis. doi: 10.1111/nph.15936 The results presented here show that brassinosteroids play a foundational role in the regula- tion of secondary growth and wood formation, in part through the regulation of cell differen- tiation and secondary cell wall biosynthesis.
    [Show full text]
  • Tree Anatomy Stems and Branches
    Tree Anatomy Series WSFNR14-15 Nov. 2014 COMPONENTSCOMPONENTS OFOF STEMSSTEMS by Dr. Kim D. Coder, Professor of Tree Biology & Health Care Warnell School of Forestry & Natural Resources, University of Georgia Trees attempt to occupy more space and control available resources through cell divisions and cell expansion. From the end of a shoot tip to the end of a root tip, trees elongate. Along this axis of growth, trees also expand radially, which is termed “secondary growth.” Tree growth is initiated in the shoot tips (growing points / buds), root tips, vascular cambium (i.e. or simply cambium), and phellogen which generates periderm. The first two meristems are primary generation systems and the second two are termed secondary meristems. Cambial activity and xylem formation, as seen in visible increases in growth increment volume is radial growth, and reviewed in the next two chapters. Cross-Section Tree stem anatomy is usually visualized in cross-section. In this form, two-demensional “layers” or “rings,” and tissue types can be identified. More difficult is to visualize these two-dimensional cross- sections and incorporate them into a three dimensional object which changes and grows over time. Moving from outside to inside a stem, tissues encountered include those associated with periderm, secondary cortex, phloem, xylem, and pith. A traditional means of describing a mature tree stem cross-section defines which tissue type is cut first, second and third using a handsaw. A list of generic tissues in stems from outside to inside could include: (Figure 1). epidermis and primary cortex = exterior primary protective tissues quickly rent, torn, and discarded with secondary growth (expansion of girth by lateral meristems – vascular cambium and phellogen).
    [Show full text]
  • Botany of Trees 2
    Botany of Trees California’s Part 2: How Wood Forms Iconic Flora North Coast California Na.ve Plant Society Free Webinar 7 PM Tomorrow Night Dr. Ma9 Ri9er h9ps://us04web.zoom.us/webinar/register/ WN_s4i_BmnlTryUe_7t45a9Qw Botany of Trees Tuesday, April 14th: IntroducTon to trees, growth, development, leaves, and morphology • Five part series -Tuesdays, 11 AM to 12:30 PM • Each will be ~1 hour with 30 minutes of ques.ons Tuesday, April 28th: Tree names, Tuesday, April 21st: Trunks, branches, diversity, and why names change tree form, branching pa9erns, and shape, and how wood forms Tuesday, May 5th: Water in trees, Tuesday, May 12th: ReproducTon, photosynthesis, and respiraTon flower formaTon, fruit, and seeds All aboveground plant structures are: Interpret the sharp structure: stems, leaves, or buds Modified leaf - spine bud becoming a new branch Pointy Structures on Plants Big Leaf Maple (Acer macrophyllum) Thorn Spine Prickle modified branch modified leaf epidermal that comes from that comes from outgrowths that an axillary bud below the occur at random on axillary bud the stem (not necessarily at nodes) Big Leaf Maple (Acer Pine Leaves macrophyllum) • Leaves of two kinds: primary scales and secondary needles • Primary leaves of pines are membranous scales • A set number of needle leaves are produced on short branches (fascicles) • Each bundle (fascicle) of needles is surrounded by membranous bud scales Leaf Leaf Leaf Stump SprouTng & Epicormic Growth Meristems: where growth occurs • Apical Meristems • Form primary .ssues • Increase in
    [Show full text]
  • Lecture 12: Gymnosperms and Angiosperms
    12: Gymnosperms and Angiosperms • Gymnosperm – Intro and evolution – Life cycle and reproduction – Uses and significance • Angiosperms: Flowering plants – Intro and evolution – Life cycle and reproduction – Uses and significance – Monocots vs. dicots Kingdom Plantae • Evolutionary tree of plants • From primitive more advanced traits Gymnosperms __________ _______ Bryophytes Flowers ________ Green alga Vascular ancestor Terrestrial GYMNOSPERMS • Introduction – Gymnosperm means “naked seed” (From the Greek: gymnos = naked; sperm = seed) • More advanced than ferns – do not have spores, they have seeds. • The seeds of the gymnosperms lack a protective enclosure (unlike flowering plants which have flowers and fruit). • Examples of gymnosperms: • Conifers (pine trees), cycads, ginkgo biloba Evolution of gymnosperms • Gymnosperms evolved from fern-like ancestors • Advancements of gymnosperms over ferns: • 1. __________ (plant embryo, food storage tissue, and seed coat) • 2. Gymnosperms do not depend on water for fertilization (have air-borne pollen) • 3. Have a more dominant _______________ generation • 4. Have a more efficient vascular system Gymnosperm life cycle • Exhibits alternation of generations • Sporophyte generation (2n) is dominant • Gametophyte generation (1n) is contained in and dependent on the sporophyte generation Gymnosperm lifecycle Sporophyte generation • Sporophyte produces two types of spores (heterosporous) • Megasporangium – undergoes meiosis to produce megaspores (female gametophyte) • ________sporangium – undergoes meiosis to produce haploid microspores, germinate to produce male gametophyte (pollen) • Many gymnosperms use wind for pollination and seed dispersal Wood produced by gymnosperms • Gymnosperms have a very efficient and effective vascular system • Usually woody plants • Xylem wood of a tree • Phloem bark of the tree • Wood is formed from secondary growth Primary vs. secondary growth • 1. Primary growth – occurs in apical meristems of shoots and roots • Results in increase in length • 2.
    [Show full text]
  • Anatomy of Flowering Plants
    84 BIOLOGY CHAPTER 6 ANATOMY OF FLOWERING PLANTS 6.1 The Tissues You can very easily see the structural similarities and variations in the external morphology of the larger living organism, both plants and 6.2 The Tissue animals. Similarly, if we were to study the internal structure, one also System finds several similarities as well as differences. This chapter introduces 6.3 Anatomy of you to the internal structure and functional organisation of higher plants. Dicotyledonous Study of internal structure of plants is called anatomy. Plants have cells and as the basic unit, cells are organised into tissues and in turn the tissues Monocotyledonous are organised into organs. Different organs in a plant show differences in Plants their internal structure. Within angiosperms, the monocots and dicots are also seen to be anatomically different. Internal structures also show 6.4 Secondary adaptations to diverse environments. Growth 6.1 THE TISSUES A tissue is a group of cells having a common origin and usually performing a common function. A plant is made up of different kinds of tissues. Tissues are classified into two main groups, namely, meristematic and permanent tissues based on whether the cells being formed are capable of dividing or not. 6.1.1 Meristematic Tissues Growth in plants is largely restricted to specialised regions of active cell division called meristems (Gk. meristos: divided). Plants have different kinds of meristems. The meristems which occur at the tips of roots and shoots and produce primary tissues are called apical meristems (Figure 6.1). 2021-22 ANATOMY OF FLOWERING PLANTS 85 Central cylinder Cortex Leaf primordium Protoderm Shoot apical Meristematic zone Initials of central cylinder Root apical and cortex Axillary bud meristem Differentiating Initials of vascular tissue root cap Root cap Figure 6.1 Apical meristem: (a) Root (b) Shoot Root apical meristem occupies the tip of a root while the shoot apical meristem occupies the distant most region of the stem axis.
    [Show full text]
  • Are Secondary Growth.Wpd
    Topic 16. Secondary Growth Introduction: Secondary growth results from the cell division at lateral meristems called cambia.To properly understand secondary growth, one must first be familiar with primary structure of the stem and the root. Specifically you should have an understanding of the organization of the primary tissues in the two stems we have studied (Medicago and Coleus) and of the Ranunculus root. It may be a good idea to review both "Cells and Tissues of the Plant Body", “The Root”, and "The Shoot" before proceeding. Some Important Definitions: Primary tissues: Tissues generated from the growth of an apical meristem. Cambium: A lateral meristem consisting of a sheet of cells. Growth of these cells increases the girth of the plant organ involved. Secondary tissues: Tissues generated from the growth of a cambium. Vascular Cambium: A cambium that gives rise to secondary xylem to the inside, and to secondary phloem to the outside. Periderm: A structure that consists of a cork cambium (phellogen), producing cork tissue (phellem) to the outside, and in some cases, a layer of cells to the inside called phelloderm. Periderm functions to limit dehydration and block pathogens after the epidermis is disrupted by the onset of secondary growth. Cork (phellem, you need know only the term "cork"): Tissue, dead at maturity generated from a cork cambium. The cell walls of the tissue are impregnated with suberin. This waterproofs the tissue. The cork used to seal wine bottles is cork tissue harvested from a species of oak.The cell theory was first proposed by Robert Hooke in 1665 after microscopic examination of a slice of cork.
    [Show full text]
  • Secondary Growth in Vertebraria Roots from the Late Permian of Antarctica: a Change in Developmental Timing
    KU ScholarWorks | http://kuscholarworks.ku.edu Please share your stories about how Open Access to this article benefits you. Secondary Growth in Vertebraria Roots from the Late Permian of Antarctica: A Change in Developmental Timing by Anne-Laure Decombeix, Edith L. Taylor, and Thomas N. Taylor 2009 This is the published version of the article, made available with the permission of the publisher. The original published version can be found at the link below. [Citation] Published version: http://www.jstor.org/stable/10.1086/597784 Terms of Use: http://www2.ku.edu/~scholar/docs/license.shtml KU ScholarWorks is a service provided by the KU Libraries’ Office of Scholarly Communication & Copyright. Int. J. Plant Sci. 170(5):644–656. 2009. Ó 2009 by The University of Chicago. All rights reserved. 1058-5893/2009/17005-0007$15.00 DOI: 10.1086/597784 SECONDARY GROWTH IN VERTEBRARIA ROOTS FROM THE LATE PERMIAN OF ANTARCTICA: A CHANGE IN DEVELOPMENTAL TIMING Anne-Laure Decombeix,1 Edith L. Taylor, and Thomas N. Taylor Department of Ecology and Evolutionary Biology, and Natural History Museum and Biodiversity Research Center, University of Kansas, Lawrence, Kansas 66045-7534, U.S.A. Permineralized Vertebraria roots from the late Permian of the Central Transantarctic Mountains, Antarctica, are investigated to understand the unusual vascular anatomy of the genus. The specimens range from ;1mmto several centimeters in diameter and illustrate all the stages of secondary growth. Our observations confirm previous hypotheses on the development of these roots and suggest that their unique anatomy is the result of a change in developmental timing. Vertebraria is characterized by a vascular cambium that remains discontinuous through several growth seasons, leading to the formation of lacunae alternating in cross section with wedges of secondary vascular tissues.
    [Show full text]