A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO

FAMILY: PSEPHENIDAE

ah52941

/^S^. UNITED STATES AGRICULTURE PREPARED BY (kki^ DEPARTMENT OF HANDBOOK AGRICULTURAL ^^^^ AGRICULTURE NUMBER 529-41 RESEARCH SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO

Fascicle ' Family Year issued Fascicle ' Family Year issued Fascicle ' Family Year issued

1_. . _ - 1979 45— .- —. 98—.-- ._ .... 2_-. Micromalthidae . 1982 46- Callirhipidae -.. 100— ..Lathridiidae 3_. .-.Carabidae 47__ — .. 1978 102—.— 4_. 48- ..Limnichidae 103—... 5- —Amphizoidae .._ — - 49— .. 1983 104—.. . 6_. -__Haliplidae 50- .. 105... „ 1982 ... 51.. .. 107... — 9_.— 52.- Cebrionidae 109...—Colydiidae 10_.—Gyrinidae 53__ Elateridae ..Monoflimatidae 13_-...Sphaeriidae 54.. —Throscidae 110- 14_.. Hydroscaphidae 55.. -- -_. 111—. --Cephaloidae 15....- 56.. ..Perothopidae .. 112—.— 16- ... .. 57— -- — 115-. —Tenebrionidae -_ 17_. Georyssidae 58 « «-Telegeusidae __. 116-. --Alleculidae 18.....Sphaeritidae 61 — — -- 117—..-Lagriidae 20-. 62. --Lampyridae 118 21.. ...Ptílüdae 63.- --Cantharidae 119-. -- 22.,—Limulodidae 64— .: 120—.—Pyrochroidae -— 1983 23..—Dasyceridae 65— -. — 121-. —Othniidae 24-.—-Micropeplidae — 66— -- - 122— --Inopeplidae 25 ...Leptinidae 67.- -- 123—.— ... 26..--. 69- —Ptinidae 124—, — .. 27.. —Scydmaenidae _. 70- --Anobiidae 1982 125...— 28-.— 71.. 126... --Rhipiphoridae _. 29..—Scaphidiidae 72.. —Lyctidae 127__._ Meloidae 30- —Staphylinidae .— 74.. -.Trogositidae 128... .- 31- Pselaphidae 76-— 129... —Pedilidae 32-. —-Lucanidae 78.. .- 130... ..Euglenidae 33--.. 79.. --Lymexylidae — 131... .-Cerambycidae .- 34- .-. __> 81.. 132—.--Bruchidae 35-, .-- 82_. Nitidulidae 133-. -.Chrysomelidae .. 36- —Helodidae 83.- --Rhizophagidae _ 134-. — .. 37-, .-- 86- -. 135—. .. 38-.-.-Dascillidae 90— — 138-..-.Allocorynidae _. .... 39- —Rhipiceridae 92.. —Languriidae 1983 140—. -. 40- — 93.. .. 141-. —Platypodidae — 1979 41- ..-Psephenidae 1983 94.. .- 142—..-Scolytidae 42- —- 95.- — 1982 143—.-- _- 43. ...Artematopidae .- 96- -.Corylophidae ._ 144.- —Stylopidae .—^_ 44. — - 97- — 145... --Fossil Coleóptera

' Missing numbers are those assigned in the computer program to families not found in the United States and Canada.

For sale by the Superintendent of Documents. U.S. Government Printing Office. Washington, D.C. 20402 y ^\ CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO FAMILY: PSEPHENIDAE i I

BY HARLEY P.[BROWN'.[BROV |0^ DEPARTMENT OF ZOOLOGY UNIVERSITY OF OKLAHOMA NORMAN, OK

,íA:. UNITED STATES AGRICULTURE PREPARED BY L|Ji; DEPARTMENT OF HANDBOOK AGRICULTURAL '^^ AGRICULTURE NUMBER 529-41 RESEARCH SERVICE

June 1983

FOREWORD

Many species of are important pests of agricultural crops, stored food products, forests, wood products and structures, and fabrics. Many other species, in contrast, are beneficial in the biological sup- pression of pest and weeds, as well as in the decomposition of plant detritus, carcasses, and dung. Part of our national responsibility to American agriculture is to provide correct identification of species of American beetles so that appropriate controls can be applied. Most information about animal species, whether agricultural, biological, or experimental, is filed under the species' scientific names. These names are therefore the keys to retrieval of such information. Because some species have been known by several names, a complete listing of these names for each species is neces- sary. For the user of scientific names, an up-to-date taxonomic catalog providing currently accepted names and pertinent bibliographic and distributional data is an indispensable tool. Although taxonomic literature is constantly changing to reflect current work, the traditional published taxonomic catalog remains static with updating left to the individual user until it is revised. Production of catalogs in the past has been laborious with long printing delays resulting in data that are obsolete before being published. However, the computer now provides the capability of storing, updating, and retrieving taxonomic data; rapid pubUca- tion through computer-driven typesetting machinery; and a greater degree of currentness and flexibility. All 124 fascicles in this catalog of the beetles of America north of Mexico are produced by an original group of computer programs, designed and written during a pilot project by personnel of the Systematic Entomology Laboratory, Agricultural Research Service, and the Communications and Data Services Divi- sion, Science and Education Management Staff. The published information is stored on computer tape, is updated periodically to reflect taxonomic progress in the family, and is available in a data base for computer searching.

T. B. Kinney, Jr. Administrator Agricultural Research Service

PREFACE The Coleóptera, or beetles, are represented in the world by about 220,000 described species, of which about 24,000 occur in the United States and Canada. A comprehensive taxonomic catalog of beetles for this area has not been available except the series of world-based "Coleopterorum Catalogus" volumes (1909-present, Junk, Berlin). The Leng "Catalogue of the Coleóptera of America North of Mexico" (J. D. Sherman, Jr., Mt. Vemon, NY), which was published in 1920 with supplements to the end of 1947, is a checklist. However, it has served professional and amateur alike for nearly 60 years as the principal source of scientific names of beetles. Since 1947, many new taxa have been described and many changes in status and nomenclature have appeared in numerous scattered publications, but little effort has been made to summarize these changes. This catalog will supplant the Leng catalog and supply additional essential information. It is produced by an original suite of storage, retrieval, and printing programs written especially for automated taxonomic catalogs. The catalog for each family is published as a separate fascicle with its introductory text, bibliography, and index. Each family is numbered as listed, but the order of issuance of fascicles is not necessarily in numerical sequence. The publishing of separate fascicles makes data available shortly after they are assembled. Computer tapes for each fascicle are maintained for updating and necessary reprinting. The information on each family is the responsibility of the respective author or authors. The editors modify it only to correct obvious errors and to make it conform to the requirements of the computer programs. No original proposal for a new name, taxon, status, or classification is given, such data having been previously published, but new host and distributional data are often listed. The rules of "The International Code of Zoological Nomenclature" are followed. The geographic scope of this catalog includes the continental United States, Canada, Alaska, Greenland, and the associated continental islands. Names of taxa found only in other regions are excluded. If the range of a species extends outside these geographic limits, this fact is indicated. Inside the back cover is a map of the 12 faunal regions based on historical and faunal criteria to simplify distribution recordings. Two-letter Postal Service style abbreviations are used for States and Provinces, and faunal regions are indicated in each distribution record by a diagonal line between groups of abbreviations. It is not the purpose of this catalog to present a complete scheme of higher classification within the order. The familial makeup is somewhat intermediate between that of R. H. Arnett in "The Beetles of the United States" (1960-62, Catholic University Press, Washington, DC) and that of R. A. Crowson in "The Natural Classification of the Families of Coleóptera" (1967, Biddies Ltd., Guildford, England). Modifications of these two systems are largely those advocated by J. F. Lawrence based in part on suggestions by taxonomic specialists for certain families. Generic groups and higher categories within the family are arranged phylogenetically as indicated by the author of the particular fascicle, and species group names with their respective synonyms are arranged alphabetically. Names referable to incertae sedis and nomen dubium are listed separately at the end of the nearest applicable taxon with notations as to their status. Each available name is followed by its author, date proposed, and page number referring to the complete bibliographic citation containing the original description. Following each generic name are the type-species and method of its designation, necessary expianator>' notes, and pertinent references on immature stages, , redescription, ecology, and keys. After the specific name entry are the original genus (if different from the present placement), type-locality, geographical distribution by State, Province, and broad extralimital units, explanatory notes, pertinent references to immature stages, taxonomy, redescription, and ecology, depository of type-specimen and its sex, and hosts. In addition to the list under the map of faunal regions (inside back cover), the following abbrevia- tions are used in this catalog:

ABBREVIATIONS, GENERAL

Amer. Bor.—America Borealis Mus.—Museum Amer. Sept.—America Septentrionalis N. Amer.—North America Autom.—Automatic Orig. des.—Original designation C. Amer.—Central America Preocc.—Preoccupied Co.—County S. Amer.—South America Cosmop.—Cosmopolitan Sp.—Species Design.—Designated Subseq. monot.—Subsequent monotypy F.—Female Subsp.—Subspecies Holarc.—Holarctic Taut.—Tautonymy Isl.—Island Univ.—^University M.—Male USA—United States of America Mex.—Mexico Var.—Variety Monot.—Monotypy W. Ind.—West Indies

MUSEUMS IN THE CONTINENTAL UNITED STATES AND CANADA

AMNH—American Museum of Natural History, FSCA—Florida State Collection, Gainesville New York HAHC—H. & A. Howden Collection, Ottawa, ANSP—Academy of Natural Sciences, Phila- Canada delphia, PA ICCM—Carnegie Museum, Pittsburgh, PA BYUC—Brigham Young University, Provo, UT INHS—Illinois Natural History Survey, Urbana CASC—California Academy of Sciences, San JGEC—J. G. Edwards Collection, San Jose, CA Francisco KMFC—K. M. Fender Collection, McMinnville, CISC—University of California, Berkeley OR CNCI—Canadian National Collections, Ottawa KSUC—Kansas State University, Manhattan CUIC—Cornell University, Ithaca, NY LACM—Los Angeles County Museum, CA CWOB—C. W. O'Brien Collection, Tallahassee, LSUC—Louisiana State University, Baton Rouge FL MCZC—Museum of Comparative Zoology, Har- DHKC^D. H. Kistner Collection, Chico State vard University, Cambridge, MA College, CA MSUC—Michigan State University, East Lansing ELSC—E. L. Sleeper Collection, Long Beach, NCSM—North Carolina State University, Raleigh CA NYSM—New York State Museum, Albany FMNH—Field Museum of Natural History, Chi- OSEC—Oklahoma State University, Stillwater cago, IL OSUC—Ohio State University, Columbus OSUO—Oregon State University, Corvallis

'Abbreviations for U.S. and Canadian museums abridged from Arnett, R. H., Jr., and Samuelson, G. A., 1969, "Di- rectory of Coleóptera Collections of North America (Canada Through Panama)," Cushing-Malloy, Ann Arbor, MI, 123 pp. vni PMNH—Peabody Museum, Yale University, New SMSH—Stovall Collection, University of Okla- Haven, CT homa, Norman PSUC—Pennsylvania State Museum, University TAMU—Texas A. & M. University, CoUege Park Station PURC—Purdue University, West Lafayette, IN UCDC—^University of California, Davis RUIC—Rutgers University, New Brunswick, NJ UMMZ—University of Michigan, Ann Arbor SEMC—Snow Museum, University of Kansas, UMRM—University of Missouri, Columbia Lawrence USNM—U.S. National Museum of Natural His- SJSC—San Jose State College, CA tory, Washington, DC SLWC—S. L. Wood Collection, Provo, UT WSUC—Washington State University, Pullman

MUSEUMS IN FOREIGN COUNTRIES

BMNH—British Museum (Natural History), NHRS—Naturhistoriske Riksmuseet, Stockholm London NMPC—^Narodni Museum, Prague, Czechoslo- BPBM—Bernice P. Bishop Museum, Honolulu vakia GUHC—Glasgow University, Hunterian College, SCUT—Spinola College, University of Turin, Scotland Italy HMOX—Hope Museum, Oxford, England SMTD—Staatliches Museum für Tierkunde, HNHM—Hungarian Natural History Museum, Dresden, East Germany Budapest UNAM—Universidad Nacional Autónoma, Mexi- IPZE—Institut Pflanzenschutzforschung Zweig- co City stelle, Eberswalde, East Germany UZMC—University Zoological Museum, Copen- IRSB—Institut Royal Sciences Belgique, Brus- hagen, Denmark sels UZMH—University Zoological Museum, Hel- MFNB—Museum für Naturkunde (Humboldt), sinki, Finland Berlin ZMAS—Zoological Museum, Academy of Sci- MGFT—Museum G. Prey, Tutzing, Munich, ences, Leningrad West Germany ZMPA—Zoological Museum, PoHsh Academy of MHNL—Museum d'Histoire Naturelle, Lyon, Sciences, Warsaw France ZMUL—Zoological Museum, University of Lund, MNHP—Museum National d'Histoire Naturelle, Sweden Paris ZMUM—Zoological Museum, University of MNSL—Museum of Natural Sciences, Leipzig, Moscow East Germany ZSBS—Zoologische Sammlung Bayerischen MZBS—Museum Zoología, Barcelona, Spain Staates, Munich, West Germany IX ACKNOWLEDGMENTS We are indebted to many individuals who contributed to the planning and development of this ca- talog. We are especially grateful to the following specialists who helped to make it as complete and accurate as possible: Richard H. Foote, Systematic Entomology Laboratory (SEL), Agricultural Research Service (ARS), for his suggestions, guidance, and encouragement; C. W. Sabrosky, SEL, for valuable counsel on nomenclatural problems; J. F. Lawrence, Division of Entomology, Com- monwealth Scientific and Industrial Research Organization, Canberra, Australia, for his recommenda- tions on higher categories; and more than 50 coleopterists in Canada, the United States, and Mexico for voluntarily contributing information about their specially groups. We thank the following members of the Communications and Data Services Division, SEMS: San- dra Strauss and Marianne Kingston for designing and writing the computer programs, and Margaret Seldin for developing the editing system.

J. M. Kingsolver, editor in chief Systematic Entomology Laboratory, Agricultural Research Service Washington, D.C.

Editorial Board

J. M. Campbell, Biosystematics Research Institute, Agriculture Canada, Ottawa, ON T. L. Erwin, Department of Entomology, Smithsonian Institution, Washington, DC H. F. Howden, Department of Biology, Carleton University, Ottawa, ON P. J. Spangler, Department of Entomology, Smithsonian Institution, Washington, DC T. J. Spilman, Systematic Entomology Laboratory, ARS, Washington, DC R. E. White, Systematic Entomology Laboratory, ARS, Washington, DC

CONTENTS

Page Family Psephenidae I Subfamily Eubrianacinae 2 Subfamily 2 Subfamily Psepheninae 3 Bibliography 5 Index g Family PSEPHENIDAE Lacordaire, 1854 By Harley P. Brown Psephenides Lacordaire, 1854: 497; Psephenidae LeConte, 1861: 115; Psepheninae Zaitzev, 1910: 4. Larvae of this family are aquatic, almost discoidal, and often golden brown, hence the name water penny beetles. Five of the world's 24 genera and 15 of its approximately 120 spe- cies have been reported from North America. Another undescribed genus is in the Southeast- ern United States, but it is known only from the . The larva of the common eastern herricki (DeKay, 1844) was described as an isopod crustacean and given the name Fluvicola, which was preoccupied. The generic name Eurypalpus, which LeConte created for the , was also preoccupied. The name Psephenus is generally accredited to Haldeman (1853), when he and LeConte revised Melsheimer's catalog, although LeConte apparently considered himself author of the name. The family name comes from Lacordaire's Psephenides (1854), emended by LeConte to Psephenidae in 1861. However, the group was widely treated as a subfamily of Dryopidae (previously Parnidae) until Hinton (1939) presented a convincing case for the separation and for including within the family the Eubrianacinae and Eubriinae. Leng's catalog (1920) placed Psephenus in a separate family Pse- phenidae within the superfamily Dryopoidea, but it combined Eubrianax with , Dieran- opselaphus, and in a subfamily Eubriinae of the Dascillidae, in the superfamily Dascil- loidea. Arnett (1963) followed the same scheme. Bertrand (1972) also placed Eubrianax in the family Dascillidae, but removed Acneus, Ectopria, and to a family Eubriidae. The taxonomy of this group remains unsettled. On a worldwide basis, Grouvelle and Pic have named most of the species. In this hemi- sphere, Casey, Darlington, Hinton, and West have contributed materially. Murvosh is studying psephenid ecology and behavior and is joined by Brown, who is concentrating on psephenid tax- onomy. Brigham is working on the genus Ectopria. A discussion of relationships to other families is difficult because there is no clear consen- sus of opinion as to whether the three most common genera-Psephenus, Ectopria, and Eu- brianax-represent one, two, or three families or even whether they all belong in the same su- perfamily. Those who consider them all as psephenids within the superfamily Dryopoidea should, to be consistent, agree that their closest relatives must be other dryopoids, such as the dryopids and elmids of the subfamily Larinae. In behavior and ecology, they closely resemble the Larinae. Such limnichids as are also very close to Psephenus in these respects. Psephenids are also reminiscent of helodids or cyphonids. All psephenid larvae are aquatic, but they differ considerably in their respiratory adapta- tions to aquatic life. Larvae of Psephenus and Eubrianax have paired trachéal gills arising from the ventral surface of four or more abdominal segments, whereas larvae of Ectopria, Acneus, and Dicranopselaphus have retractile cloacal gills very much like those of elmid larvae. The larvae typically inhabit stream riffles, spending most daylight hours on the underside of rocks but moving in subdued light to their upper surface to graze on encrusting algae. Pupation occurs on sheltered firm substrates near the water, beneath the testudinate larval skin. The adults are short lived and feed little, if any. Males of Psephenus spend the night in hiding and the day scurrying over exposed parts of rocks in riffles in frantic search for mates. They fly readily and rapidly. After mating, females forsake flight, crawl into the water, and spend their few remaining days in oviposition. Their yellow-orange eggs are laid in a flat cluster on the lower side of stones in riffles. Habits of Eubrianax are similar to those of Psephenus, but adults of the Eubriinae are neither perceptibly riparian nor aquatic. They are active at night; Ecto- pria is often taken at lights in considerable numbers. They spend the daylight hours amid fo- liage near the water, chiefly on the lower side of leaves of trees and shrubs. Their oviposition has not been described. This manuscript was received May 1976. Hinton, 1939: 133, 1955: 543, and 1966: 211; Brown, 1972. 2 COLEÓPTERA CATALOG Subfamily EUBRIANACINAE Boeving and Craighead, 1931 Placonychini (Dascyllidae) Horn, 1880: 110 {Placonychus only). Hinton, 1939: 133 and 1955: 543. Genus EUBRIANAX Kiesenwetter Eubrianax Kiesenwetter, 1874: 246. Tjrpe-species: Eubrianax ramicornis Kiesenwetter (monot.). Placonycha Horn, 1880: 111 (replacement name for Dicranopselaphus edwardsii LeConte). Type-species: Dicranopselaphus edwardsii LeConte (automat.). IMMATURE STAGES: Brown, 1972: 66 (larva); Leech and Chandler, 1956: 365; Hinton, 1955: 546; Bertrand, 1972: 450. TAXONOMY: Hinton, 1939 and 1955. ECOLOGY: Leech and Chandler, 1956: 365. KEYS: Brown, 1972. edwardsii (LeConte), 1874: 57 (Dicranopselaphus). CA: Santa Barbara; OR/ CA NV. TYPE DEPOSITORY: MCZC. IMMATURE STAGES: Leech and Chandler, 1956: 13, figs. 58-59; Blackwelder, 1930; Brown, 1972: figs. 195-196 (larva). Subfamily EUBRIINAE LeConte, 1861 Eubriini (Dascyllidae) LeConte, 1861: 180; Eubriinae (Dascillidae) Champion, 1897: 595. Horn, 1880: 76; Hinton, 1955: 543; Brown, 1972. Genus ACNEUS Horn Acneus Horn, 1880: 97, pi. I, fig. 10. Type-species: Acneus quadrimaculatus Horn (monot.). Acna£us Fender, 1962: 44 (unjustified emendation). Type-species: Acneus quadrimaculatus Horn (monot.). IMMATURE STAGES: Bertrand, 1972: 429 (larva); Brown, 1972: 66 (larva). TAXONOMY: Fender, 1962; Leech and Chandler, 1956; Brown, 1972. REDESCRIPTION: Fender, 1962. ECOLOGY: Leech and Chandler, 1956: 365. KEYS: Fender, 1962. beeri Hatch, 1961: 98, pi. XXV, fig. 4. OR: Linn Co., 5-15 mi E. Cascadia; OR. TYPE DEPOSITORY: USNM. SEX OF TYPE: F. REDESCRIPTION: Fender, 1962. burnelli (Fender), 1962: 48, figs. 4, 7, 11 (Acnaeus), OR: Coos Co., Upper Middle Cr. 15 mi S.W. Powers; OR. TYPE DEPOSITORY: CASC?. SEX OF TYPE: M. oregonensis Fender, 1951: 271, figs. 1, 5, 9. OR: Multnomah Co., Multnomah Falls; OR. TYPE DEPOSITORY: OR State Univ., Corvallis. SEX OF TYPE: M. quadrimaculatus Horn, 1880: 98, pi. I, fig. 10. CA: Sauzalito?; OR/ CA. TYPE DEPOSITORY: MCZC. SEX OF TYPE: F. IMMATURE STAGES: Brown, 1972: figs. 189-190. REDESCRIPTION: Fender, 1962. Genus DICRANOPSELAPHUS Guerin-Meneville Dicranopselaphus Guerin-Meneville, 1861: 531, pi. 17. Type-species: Not yet designated. IMMATURE STAGES: Brown, 1972: figs. 191-192 (larva). KEYS: Champion, 1897. variegatus Horn, 1880: 97. IL and MD; IL/ NY PA MD. TYPE DEPOSITORY: MCZC. SEX OF TYPE: M. PSEPHENIDAE 3

Genus ECTOPRIA LeConte Ectopria LeConte, 1853: 351. Type-species: thoracica Ziegler (subseq. monot.: Horn, 1880: 95) nervosa (Melsheimer). Horn (1880) placed all North American species names in E, nervosa, Eurea LeConte, 1853: 352. Type-species: Eubria nervosa Melsheimer (monot.). IMMATURE STAGES: Brown, 1972: 66; Leech and Chandler, 1956: 366; West, 1929b: 722 (larva, as Helichus). TAXONOMY: Horn, 1880: 94. REDESCRIPTION: Horn, 1880: 95. ECOLOGY: Brown, 1972. nervosa (Melsheimer), 1845: 222 (Eubria), PA; ON PQ/ KS lA MO IL IN OH KY/ NY PA NJ DE MD DC WV VA/ ME NH VT MA CT/ OK/ AR MS AL TN GA SC NC FL. thoracica Ziegler, 1845: 269 (EubriaXsynonymußa by Horn, 1880: 96). PA. tarsalis LeConte, 1853: 352 (synonymized by Horn, 1880: 96). GA. tibialis LeConte, 1853: 352 (synonymized by Horn, 1880: 96). PA. IMMATURE STAGES: Brown, 1972: figs. 193-194 (larva). TAXONOMY: Horn, 1880; LeConte, 1853: 351. REDESCRIPTION: Horn, 1880: 95.

Subfamily PSEPHENINAE LeConte and Horn, 1883 Eurypalpini LeConte, 1852: 41. Boeving and Craighead, 1931; Hinton, 1939: 133 and 1955: 543. Genus PSEPHENUS Haldeman Psephenus Haldeman, 1853: 34. Type-species: Fluvicola herricki DeKay (monot.).Although the name Psephenus is attributed to Haldeman, he did not describe it, and apparently LeConte considered himself author of the name. The name simply appears in the portion of Melsheimer's catalog (1853) prepared by LeConte and Haldeman, but with Haldeman's name affixed to it. Fluvicola DeKay, 1844: 53, pi. 10 (preoccupied Swainson, 1827). Type-species: Fluvicola herricki DeKay (monot.). Eurypalpus LeConte, 1852: 42 (preoccupied Macquart, 1835; nomen nudum in Dejean, 1837). Type-species: Eurypalpus lecontei LeConte (automat.) herricki (DeKay). IMMATURE STAGES: West, 1929a; Hinton, 1955 and 1966 (larvae, pupae); Brown and Arrington, 1967 (larva, pupa). TAXONOMY: Brown and Murvosh, 1974. REDESCRIPTION: Brown and Murvosh, 1974: 308. ECOLOGY: Murvosh, 1971. KEYS: Brown and Murvosh, 1974; Brown, 1972. arizonensis Brown and Murvosh, 1974: 303, figs. 14-16, 29, 32, 33. AZ: Cochise Co., Cave Creek near Portal; AZ. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. TAXONOMY: Brown and Murvosh, 1974. falli Casey, 1893: 579. CA: Los Angeles Co.; WA OR ID/ CA NV. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. veluticollis Casey, 1893: 579 (synonymized by Brown and Murvosh, 1974: 308). CA: Mendocino Co. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. lanei Blaisdell, 1923: 234 (synonymized by Brown and Murvosh, 1974: 308). ID: Juliaetta. TYPE DEPOSFTORY: CASC. SEX OF TYPE: M. 4 COLEÓPTERA CATALOG

calaveras Blaisdell, 1923: 236 (synonymized by Brown and Murvosh, 1974: 308). CA: Calaveras Co., Calaveras River. TYPE DEPOSITORY: CASC. SEX OF TYPE: F. TAXONOMY: Brown and Murvosh, 1974. REDESCRIPTION: Brown and Murvosh, 1974. herricki (DeKay), 1844: 53 (Fluvicola). NY: gorge of Niagara River; MN WI MI ON PQ/ NB NS/ KS MO IL IN OH KY/ NY PA NJ DE MD DC WV VA/ ME NH VT MA RI CT/ OK/ AR MS AL TN GA SC NC. TYPE DEPOSITORY: USNM (neotype). SEX OF TYPE: M. lecontei LeConte, 1852: 42 (EurypalpusXxiomen nudum listed by Dejean, 1837; validated by LeConte, 1852; synonymized by Leng, 1920). 'Western NY and PA I trentonensis Zimmermann, 1869: 259 (synonymized by Horn, 1870: 30). NY: Trenton Falls. TYPE DEPOSITORY: USNM. IMMATURE STAGES: West, 1929a (eggs, larvae, pupae). TAXONOMY: Brown and Murvosh, 1974. REDESCRIPTION: Brown and Murvosh, 1974. ECOLOGY: Murvosh, 1971. minckleyi Brown and Murvosh, 1974: 322, figs. 10, 11, 22-24, 34. AZ: Yavapai Co., West Clear Creek S.E. Camp Verde; AZ NM. TYPE DEPOSITORY: USNM. SEX OF TYPE: M. montanus Brown and Murvosh, 1974: 328, figs. 12, 13, 19-21, 30. AZ: Apache Co., E. Fork of Black R. in White Mtns.; AZ. TYPE DEPOSFFORY: USNM. SEX OF TYPE: M. murvoshi Brown, 1970: 34, figs. 1-5. AZ: Gila Co., East Verde River near Payson; AZ NM/ Mex. TYPE DEPOSITORY: SMSH. SEX OF TYPE: M. REDESCRIPTION: Brown and Murvosh, 1974 (female). texanus Brown and Arrington, 1967: 86, figs. 1-3, 9-11. TX: Val Verde Co., San Felipe Cr., Del Rio, 300 m; TX/ Mex. TYPE DEPOSITORY: SMSH. SEX OF TYPE: M. IMMATURE STAGES: Brown and Arrington, 1967: figs. 9, 10 (larva), fig. 11 (pupa); Brown and Murvosh, 1974: ñg. 62. BIBLIOGRAPHY

Arnett, R. H., Jr. 1963 The beetles of the United States (A manual for identification). Part III. The Catho- lic University of America Press, Washington, D.C., pp. 369-526, illus. Bertrand, H. P. 1. 1972 Larves et nymphes des Coléoptères aquatiques du globe, avec tableaux de determi- nation des genres et 561 figures. F. Paillart, Abbeville, France, 804 pp., illus. Blackwelder, R. E. 1930 The larva of Eubrianax edwardsi (LeC.) (Coleóptera: Psephenidae). The Pan-Pacific Entomologist, vol. 6, no. 3, pp. 139-142, illus. Blaisdell, F. E., Sr. 1923 Two new species of Psephenus Hald., with a note on Narpus angustus Casey (Coleop.). Entomological News, vol. 34, pp. 234-238. Boeving, A. 6. and F. C. Craighead 1931 An illustrated synopsis of the principal larval forms of the order Coleóptera. Ento- mológica Americana, n. ser., vol. 11, pp. 1-351, illus. Brown, H. P. 1970 A new species of Psephenus from Arizona (Coleóptera, Psephenidae). The Coleop- terists' Bulletin, vol. 24, no. 2, pp. 34-38, illus. Brown, H. P. 1972 Aquatic dryopoid beetles (Coleóptera) of the United States. Biota of Freshwater Ecosystems Identification Manual No. 6, Water Pollution Control Research Series, U. S. Environmental Protection Agency, Washington, D. C, ix and 82 pp., illus. Brown, H. P. and R. Arrington, Jr. 1967 A new species of Psephenus from Mexico and Texas (Coleóptera, Dryopoidea, Pse- phenidae). The Coleopterists' Bulletin, vol. 21, no. 3, pp. 86-91, illus. Brown, H. P. and C. M. Murvosh 1974 A revision of the genus Psephenus (water-penny beetles) of the United States and Canada (Coleóptera, Dryopoidea, Psephenidae). Transactions of the American Entomological Society, vol. 100, pp. 289-340, illus. Casey, T. L. 1893 Coleopterological notices. V. Annals of the New York Academy of Science, vol. 7, pp. 281-606. Champion, G. C. 1897 Rhipiceridae, Dascillidae and supplement. In Biologia Centrali-Americana, Insecta. Coleóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 3, pt. 1, pp. 585-662, illus. Dejean, P. F. M. A. 1837 Catalogue des Coléoptères de la collection de M. le compte Dejean. Troisième edi- tion, revue, corrigée et augmentée. (Ed. 4). Paris, France, Livre 5, pp. 385-503. DeKay, J. E. 1844 Zoology of New-York, or the New-York fauna; comprising detailed descriptions of all the hitherto observed within the state of New-York, with brief no- tices of those occasionally found near its borders, and accompanied by appropri- ate illustrations. Part VI, Crustacea. Albany, New York, pp. 1-70, illus. Fender, K. M. 1951 A new species of Acneus (Coleóptera, Dascillidae). Proceedings of the Entomologi- cal Society of Washington, vol. 53, no. 5, pp. 271-272. 6 COLEÓPTERA CATALOG Fender, K. M. 1962 A review of the genus Acnaeus (Coleoptera-Dascillidae). Northwest Science, vol. 36, no. 2, pp. 44-49, illus. Guerin-Meneville, F. E. 1861 Monographie du nouveau genre Dicranopselaphus et description de quelques autres insectes Coléoptères appartenant aussi a la famille des Dascillides. Revue et Magasin de Zoologie Pure et Appliquée, vol. 13, pp. 531-547, illus. Haldeman, S. S. 1853 In Melsheimer, F. E. Catalogue of the described Coleóptera of the United States. Washington, D. C, 174 pp. Hatch, M. H. 1961 The beetles of the Pacific Northwest. University of Washington Publications in Bi- ology, vol. 16., pt. 3, ix and 503 pp., illus. Hinten, H. E. 1939 An inquiry into the natural classification of the Dryopoidea, based partly on a study of their internal anatomy (Col.). Transactions of the Entomological Society of London, vol. 89, pp. 133-184, illus. Hinton, H. E. 1955 On the respiratory adaptations, biology, and taxonomy of the Psephenidae, with notes on some related families (Coleóptera). Proceedings of the Zoological Society of London, vol. 125, pp. 543-568, illus. Hinton, H. E. 1966 Respiratory adaptations of the pupae of beetles of the family Psephenidae. Philo- sophical Transactions of the Royal Society of London, ser. B, vol. 25L no 771 pp. 211-245, illus. Horn, G. H. 1870 Synopsis of the Parnidae of the United States. Transactions of the American Ento- mological Society, vol. 3, pp. 29-42. Horn, G. H. 1880 Synopsis of the Dascyllidae of the United States. Transactions of the American En- tomological Society, vol. 8, pp. 76-114, illus. Von Kiesenwetter, E. A. H. 1874 Die Malacodermen Japans nach dem Ergebnisse der Sammlungen des Herrn G. Lewis waehrend der Jahre 1869-1871. Berliner Entomologische Zeitschrift, vol 18, pp. 241-288. Lacordaire, J. T. 1854 Histoire naturelle des Insectes. Genera des Coléoptères ou expose méthodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Roret, Paris, vol. 2, 548 pp. LeConte, J. L. 1852 Synopsis of the Parnidae of the United States. Proceedings of the Academy of Nat- ural Sciences of Philadelphia, vol. 6, pp. 41-45. LeConte, J. L. 1853 Synopsis of the Atopidae, Rhipiceridae and Cyphonidae of the United States. Pro- ceedmgs of the Academy of Natural Sciences of Philadelphia, vol. 6, pp. 350-357. LeConte, J. L. 1861 Classification of the Coleóptera of North America. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collections, no. 136, pt. 1, pp. 1-208. LeConte, J. L. 1874 Descriptions of new Coleóptera chiefly from the Pacific slope of North America Transactions of the American Entomological Society, vol. 5, pp. 43-72. PSEPHENIDAE í

LeConte, J. L. and G. H. Horn 1883 Classification of the Coleóptera of North America. Prepared for the Smithsonian Institution. Smithsonian Miscellaneous Collections, no. 507, vol. 26, pt. 4, pp. i- xxxvii and 1-567. Leech, H. B. and H. P. Chandler 1956 Aquatic Coleóptera, pp. 293-371. In Usinger, R. L. (ed.). Aquatic of Califor- nia. University of California Press, Berkeley, i-ix and 508 pp., illus. Leng, C. W. 1920 Catalogue of the Coleóptera of America, north of Mexico. John D. Sherman, Jr., Mt. Vernon, New York, 470 pp. Leng, C. W. and A. J. Mutchler 1927 Supplement 1919-1924 (inclusive) to catalogue of the Coleóptera of America north of Mexico. John D. Sherman, Jr., Mt. Vernon, New York, 78 pp. Melsheimer, F. E. 1846 Descriptions of new species of Coleóptera of the United States. Proceedings of the Academy of Natural Sciences of Philadelphia, vol. 2, pp. 213-223. Melsheimer, F. E. 1853 Catalogue of the described Coleóptera of the United States. Washington, D. C, 174 pp. Murvosh, C. M. 1971 Ecology of the water penny beetle (DeKay). Ecological Mono- graphs, vol. 41, pp. 79-96, illus. West, L. S. 1929a Life history notes on Psephenus lecontei Lee. (Coleóptera; Dryopoidea; Psepheni- dae). Battle Creek College Bulletin, vol. 3, no. 1, pp. 3-20, illus. West, L. S. 1929b A preliminary study of larval structure in the Dryopidae. Annals of the Entomo- logical Society of America, vol. 22, no. 4, pp. 691-727, illus. Zaitzev, P. A. 1910 Dryopidae, Cyathoceridae, Georyssidae, Heteroceridae. In Coleopterorum catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. W. Junk, Berlin, pt. 17, (vol. 14), pp. 1-68. Ziegler, D. 1845 Descriptions of new North American Coleóptera. Proceedings of the Academy of Natural Sciences of Philadelphia, vol. 2, pp. 266-272. Zimmermann, C. C. A. 1869 Synonymical notes on Coleóptera of the United States, with descriptions of new species from the MSS of the late Dr. C. Zimmermann. Edited by John L. Le- Conte, M. D. Transactions of the American Entomological Society, vol. 2, pp. 243-259. INDEX

Names are indexed as follows: EUBRIINAE 2 CAPITALS: All names for taxa above the generic Eurea LeConte 3 level; Eurypalpus LeConte 3 Boldface: Valid generic and subgeneric names; falli Casey, Psephenus 3 Roman: Valid specific and subspecific names; Fluvicola DeKay 3 Italic: All invalid names such as synonyms, nomina herricki (DeKay), Psephenus 4 nuda, and extra-limital taxa even though valid. lanei Blaisdell, Psephenus 3 Parentheses around an author's name indicate that lecontei (LeConte), Psephenus 4 the specific name has been transferred from its original minckleyi Brown and Murvosh, Psephenus 4 genus. The generic name following the author's name in- montanus Brown and Murvosh, Psephenus 4 dicates the present placement of the species. Synonyms murvoshi Brown, Psephenus 4 of species-group names are listed with the original spell- nervosa (Melsheimer), Ectopria 3 ing. oregonensis Fender, Acneus 2 Acnaeus Fender 2 Placonycha Horn 2 Acneus Horn 2 PSEPHENINAE 3 arizonensis Brown and Murvosh, Psephenus 3 Psephenus Haldeman 3 beeri Hatch, Acneus 2 quadrimaculatus Horn, Acneus 2 burnelli (Fender), Acneus 2 tarsalis LeConte, Ectopria 3 calaveras Blaisdell, Psephenus 4 texanus Brown and Arrington, Psephenus 4 Dicranopselaphus Guerin-Meneville 2 thoracica (Ziegler), Ectopria 3 Ectopria LeConte 3 tibialis LeConte, Ectopria 3 edwardsii (LeConte), Eubrianax 2 trentonensis Zimmermann, Psephenus 4 EUBRIANACINAE 2 variegatus Horn, Dicranopselaphus 2 Eubrianax Kiesenwetter 2 veluticollis Casey, Psephenus 3 ->?'*iï?£t'=^^''

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