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coursework that is integrated with the major BIOCHEMISTRY AND program. BIOPHYSICS We describe below only the four more popular programs of study within the concentration. Students Department Website: interested in other options, such as a concentration https://www.haverford.edu/biochemistry-biophysics in both biochemistry and biophysics, should consult with the faculty representatives listed above to The Concentration in Biochemistry and Biophysics design a course of study encompassing the required recognizes enduring trends in interdisciplinary courses and any proposed substitutions. However, science, by establishing in the curriculum a formal students may not obtain both a minor program of classroom and laboratory training at and a biochemistry concentration, or both a the interface between the physical, chemical and minor and a biophysics concentration. biological sciences. Learning Goals Concentration Requirements Biochemistry/Biophysics Core Curriculum (required of • Identify, formulate, and solve complex problems all): at the interface of and the physical sciences using state-of-the-art equipment and • BIOL H200 (, and Genomes) techniques. and BIOL H201 (, Cells, and ) • Apply knowledge of chemistry, biology, physics • CHEM H104, CHEM H112, CHEM H114 (Chemical and to develop a coherent Dynamics). understanding of biological processes and solve • MATH H118 (Calculus II) or a one semester problems in living organisms or in vitro systems mathematics or course with MATH H118 derived from biological specimens. as a prerequisite. • Learn to search, read and interpret original • PHYS H105 and PHYS H106, or PHYS H101 and scientific literature, both for research and for PHYS H102 (two semesters of Introductory ongoing learning. Physics), or the Bryn Mawr equivalents. • Recognize enduring trends in interdisciplinary Concentrators typically complete required science, while navigating the program of coursework at the 200-level and higher in classroom and laboratory training at the interface participating Haverford departments. One course between the physical and biological sciences. taken elsewhere may be substituted with prior • (For biochemistry) study the importance of approval of the relevant department and the biological at all levels of the Concentration coordinator; students seeking natural sciences, including the , the , the additional flexibility may petition the Concentration , and larger ecological systems. Committee in advance regarding their needs and • Examine and analyze natural phenomena at plans. the appropriate level(s) (molecular, cellular, organismal and/or ecological), using a variety of The advanced interdisciplinary coursework methods informed by evolutionary theory. requirements vary by major and desired • Communicate findings (either verbally and/or concentration: via written expression) effectively and clearly to Biology Major with a Biochemistry Concentration: diverse audiences. Biology majors seeking a biochemistry concentration Haverford’s Institutional Learning Goals are must complete the biochemistry/biophysics core available on the President’s website, at http://hav.to/ curriculum (see above) as well as the following learninggoals. additional requirements: Curriculum • CHEM H111 or CHEM H113 or CHEM H115 (Chemical Structure and Bonding), CHEM H222 All concentrators must complete a major in biology, and CHEM H225 (Organic Chemistry). chemistry or physics while taking additional • CHEM H304 (Statistical and coursework that spans mathematics and all of Kinetics) or CHEM H305 (). these disciplines. The concentration requirements provide guidance for students while • One semester of advanced biochemistry-related allowing considerable leeway for tailoring the laboratory coursework: CHEM H301 or CHEM H302 program to specific interests. All concentrators take (Laboratory in Chemical Structure and Reactivity) a required core curriculum as well as advanced or BIOL/CHEM H303 (Laboratory in Biochemical Research). For concentrators, this course will 2 Biochemistry and Biophysics

count toward the Biology major in lieu of either Biological Nano-Machines), BIOL H452 (Advanced BIOL H300 or BIOL H301. Topics in ), BIOL H454 (Advanced • Two half-semester advanced courses with Topics in ), and BIOL H457 (Topics in significant biochemistry content: CHEM H351 Science). (Bioinorganic Chemistry), CHEM H352 (Topics in Chemistry Major with a Biochemistry Area of ), and CHEM H357 (Topics in Bioorganic Chemistry); students may take topics Concentration: courses multiple times with different topics. Chemistry majors desiring a biochemistry area of • Two half-semester courses with significant concentration must complete the biochemistry/ biochemistry content: BIOL H311 (Advanced biophysics core curriculum (see above) as well as the Genetic Analysis), BIOL H313 (Structure and following additional requirements: of Macromolecules), BIOL H314 • One semester of advanced biochemistry-related (Biochemistry: Metabolic Basis of Disease laboratory coursework: normally BIOL H300 and ), BIOL H316 (Inter- and or BIOL H301 (Advanced Lab in Biology) or Intra-Cellular Communication), BIOL H320 BIOL/CHEM H303 (Laboratory in Biochemical (Molecular ), BIOL H322 (Cell Research). Any of these courses, along with one Architecture), BIOL H324 (), semester of either CHEM H301 or CHEM H302, BIOL H326 (Biochemical ), BIOL H328 satisfy the advanced laboratory requirement for (Immunology), BIOL H334 (Biochemistry of Gene the Chemistry major. Expression), BIOL H451 (Molecular Motors and • Two half-semester courses with significant Biological Nano-Machines), BIOL H452 (Advanced biochemistry content: CHEM H351 (Bioinorganic Topics in Immunology), BIOL H454 (Advanced Chemistry), CHEM H352 (Topics in Biophysical Topics in Virology), and BIOL H457 (Topics in Chemistry), and CHEM H357 (Topics in Bioorganic Protein Science). Chemistry). Students may take topics courses Biology Major with a Biophysics Concentration: multiple times with different topics. Biology majors seeking a biophysics concentration • Two half-semester courses with significant must complete the biochemistry/biophysics core biochemistry content: BIOL H311 (Advanced curriculum (see above) as well as the following Genetic Analysis), BIOL H313 (Structure and additional requirements: Function of Macromolecules), BIOL H314 (Biochemistry: Metabolic Basis of Disease • MATH H121 (Calculus III) or MATH H216 and Adaptation), BIOL H316 (Inter- and (Advanced Calculus). Intra-Cellular Communication), BIOL H320 • PHYS H213 (Waves and ), PHYS H211 (Molecular Microbiology), BIOL H322 (Cell (Laboratory in Electronics, Waves and Optics); Architecture), BIOL H324 (Photosynthesis), half-credit course), and PHYS H301 (Advanced BIOL H326 (Biochemical Adaptations), BIOL H328 Laboratory in ). For concentrators, (Immunology), BIOL H334 (Biochemistry of Gene PHYS H301 will count toward the Biology major in Expression), BIOL H451 (Molecular Motors and lieu of either BIOL H300 or BIOL H301. Biological Nano-Machines), BIOL H452 (Advanced • PHYS H214 (Quantum ) or CHEM H305 Topics in Immunology), BIOL H454 (Advanced (Quantum Chemistry). Topics in Virology), and BIOL H457 (Topics in • PHYS H303 () or CHEM H304 Protein Science). (Statistical Thermodynamics and Kinetics). Physics Major with a Biophysics Area of Concentration: • PHYS H320 (Topics in Biological Physics) or a Physics majors desiring a biophysics area of similar course approved by the coordinating concentration must complete the biochemistry/ committee. biophysics core curriculum (see above) as well as the • Two half-semester courses with significant following additional requirements: biophysics content: BIOL H311 (Advanced Genetic Analysis), BIOL H313 (Structure and • An advanced biochemistry or biology laboratory Function of Macromolecules), BIOL H314 course: one semester of BIOL H300 or BIOL H301 (Biochemistry: Metabolic Basis of Disease (Advanced Lab in Biology) or BIOL/CHEM H303 and Adaptation), BIOL H316 (Inter- and (Laboratory in Biochemical Research). Intra-Cellular Communication), BIOL H320 • Either PHYS H320 (Topics in Biological Physics) (Molecular Microbiology), BIOL H322 (Cell or two half-semester courses with significant Architecture), BIOL H324 (Photosynthesis), biophysics content: BIOL H311 (Advanced BIOL H326 (Biochemical Adaptations), BIOL H328 Genetic Analysis), BIOL H313 (Structure and (Immunology), BIOL H451 (Molecular Motors and Function of Macromolecules), BIOL H314 Biochemistry and Biophysics 3

(Biochemistry: Metabolic Basis of Disease and Adaptation), BIOL H316 (Inter- and Intra-Cellular Communication), BIOL H320 (Molecular Microbiology), BIOL H322 (Cell Architecture), BIOL H324 (Photosynthesis), BIOL H326 (Biochemical Adaptations), BIOL H328 (Immunology), BIOL H451 (Molecular Motors and Biological Nano-Machines), BIOL H452 (Advanced Topics in Immunology), BIOL H454 (Advanced Topics in Virology), and BIOL H457 (Topics in Protein Science). The Biophysics Concentration may be elected with either the Interdisciplinary or Traditional Physics major. BIOL H200, BIOL H201 and the advanced biochemistry or biology laboratory course satisfy the Interdisciplinary Physics major requirement of two 200(+)-level courses in a related field as well as the upper level laboratory course requirement. The 300- level biology courses meeting concentration requirements may be used in lieu of one or two of the six 300-level physics courses required for the Traditional Physics major. Concentration Coordinating Committee Karin Åkerfeldt Professor Emeritus of Chemistry Professor of Physics and , Coordinator of Biochemistry and Biophysics Lou Charkoudian Associate Professor of Chemistry Amy Cooke Assistant Professor of Biology Rob Fairman Professor of Biology Karl Johnson Professor and Chair of Biology Casey Londergan Professor of Chemistry Robert Scarrow Professor and Chair of Chemistry Walter Smith The Paul and Sally Bolgiano Professor of Physics; Professor of Physics and Astronomy Kristen Whalen Assistant Professor of Biology