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Vertebrates, Overview
VERTEBRATES, OVERVIEW Carl Gans* and Christopher J. Bell† *Department of Integrative Biology, University of Texas at Austin and †Department of Geological Sciences, University of Texas at Austin I. Introduction neurectoderm An embryonic tissue that gives rise to II. General Vertebrate Characteristics the central tube of the nervous system. III. Early Chordate and Vertebrate History notochord A stiff, flexible, longitudinal rod running IV. Vertebrate Classification along the middorsal portion of the chordate body. V. Definitions and Diagnoses of Major Chordate It is situated dorsal to the coelom and ventral to the Groups central tube of the nervous system. pharynx The anterior portion of the alimentary canal, characterized by lateral buds that provide skeletal GLOSSARY support for the gill region. tuberculum interglenoideum An anterior projection of chordate A member of the group Chordata. The the first (cervical) vertebra in salamanders. The tu- Chordata includes the most recent common ancestor berculum interglenoideum bears articular facets that of tunicates and cephalochordates and all of that insert into the foramen magnum of the skull and ancestor’s descendants. Tunicates, lancelets, hag- provide additional articulation points between the fishes, and vertebrates are all chordates. skull and the vertebral column. ectoderm An embryonic tissue that provides the future outside layer of the animal. ectothermy A method of body temperature control in which the animal utilizes external sources for gaining VERTEBRATES INCLUDE ALL the fishes, amphibians, and giving up heat, thus achieving temperature con- reptiles, birds, and mammals. These animals are united trol without affecting metabolic rate. in a more inclusive group, the Chordata, that includes endothermy A method of body temperature control in the closest living relatives of vertebrates, the hagfishes, which the animal modifies its metabolic rate to lancelets, and tunicates. -
The Ventricles
Guest Editorial Evolution of the Ventricles Solomon Victor, FRCS, FRCP We studied the evolution of ventricles by macroscopic examination of the hearts of Vijaya M. Nayak, MS marine cartilaginous and bony fish, and by angiocardiography and gross examination of Raveen Rajasingh, MPhil the hearts of air-breathing freshwater fish, frogs, turtles, snakes, and crocodiles. A right-sided, thin-walled ventricular lumen is seen in the fish, frog, turtle, and snake. In fish, there is external symmetry of the ventricle, internal asymmetry, and a thick- walled left ventricle with a small inlet chamber. In animals such as frogs, turtles, and snakes, the left ventricle exists as a small-cavitied contractile sponge. The high pressure generated by this spongy left ventricle, the direction of the jet, the ventriculoarterial ori- entation, and the bulbar spiral valve in the frog help to separate the systemic and pul- monary circulations. In the crocodile, the right aorta is connected to the left ventricle, and there is a complete interventricular septum and an improved left ventricular lumen when compared with turtles and snakes. The heart is housed in a rigid pericardial cavity in the shark, possibly to protect it from changing underwater pressure. The pericardial cavity in various species permits move- ments of the heart-which vary depending on the ventriculoarterial orientation and need for the ventricle to generate torque or spin on the ejected blood- that favor run-off into the appropriate arteries and their branches. In the lower species, it is not clear whether the spongy myocardium contributes to myocardial oxygenation. In human beings, spongy myocardium constitutes a rare form of congenital heart disease. -
Supplementary Tales
Metabarcoding reveals different zooplankton communities in northern and southern areas of the North Sea Jan Niklas Macher, Berry B. van der Hoorn, Katja T. C. A. Peijnenburg, Lodewijk van Walraven, Willem Renema Supplementary tables 1-5 Table S1: Sampling stations and recorded abiotic variables recorded during the NICO 10 expedition from the Dutch Coast to the Shetland Islands Sampling site name Coordinates (°N, °E) Mean remperature (°C) Mean salinity (PSU) Depth (m) S74 59.416510, 0.499900 8.2 35.1 134 S37 58.1855556, 0.5016667 8.7 35.1 89 S93 57.36046, 0.57784 7.8 34.8 84 S22 56.5866667, 0.6905556 8.3 34.9 220 S109 56.06489, 1.59652 8.7 35 79 S130 55.62157, 2.38651 7.8 34.8 73 S156 54.88581, 3.69192 8.3 34.6 41 S176 54.41489, 4.04154 9.6 34.6 43 S203 53.76851, 4.76715 11.8 34.5 34 Table S2: Species list and read number per sampling site Class Order Family Genus Species S22 S37 S74 S93 S109 S130 S156 S176 S203 Copepoda Calanoida Acartiidae Acartia Acartia clausi 0 0 0 72 0 170 15 630 3995 Copepoda Calanoida Acartiidae Acartia Acartia tonsa 0 0 0 0 0 0 0 0 23 Hydrozoa Trachymedusae Rhopalonematidae Aglantha Aglantha digitale 0 0 0 0 1870 117 420 629 0 Actinopterygii Trachiniformes Ammodytidae Ammodytes Ammodytes marinus 0 0 0 0 0 263 0 35 0 Copepoda Harpacticoida Miraciidae Amphiascopsis Amphiascopsis cinctus 344 0 0 992 2477 2500 9574 8947 0 Ophiuroidea Amphilepidida Amphiuridae Amphiura Amphiura filiformis 0 0 0 0 219 0 0 1470 63233 Copepoda Calanoida Pontellidae Anomalocera Anomalocera patersoni 0 0 586 0 0 0 0 0 0 Bivalvia Venerida -
Skates and Rays Diversity, Exploration and Conservation – Case-Study of the Thornback Ray, Raja Clavata
UNIVERSIDADE DE LISBOA FACULDADE DE CIÊNCIAS DEPARTAMENTO DE BIOLOGIA ANIMAL SKATES AND RAYS DIVERSITY, EXPLORATION AND CONSERVATION – CASE-STUDY OF THE THORNBACK RAY, RAJA CLAVATA Bárbara Marques Serra Pereira Doutoramento em Ciências do Mar 2010 UNIVERSIDADE DE LISBOA FACULDADE DE CIÊNCIAS DEPARTAMENTO DE BIOLOGIA ANIMAL SKATES AND RAYS DIVERSITY, EXPLORATION AND CONSERVATION – CASE-STUDY OF THE THORNBACK RAY, RAJA CLAVATA Bárbara Marques Serra Pereira Tese orientada por Professor Auxiliar com Agregação Leonel Serrano Gordo e Investigadora Auxiliar Ivone Figueiredo Doutoramento em Ciências do Mar 2010 The research reported in this thesis was carried out at the Instituto de Investigação das Pescas e do Mar (IPIMAR - INRB), Unidade de Recursos Marinhos e Sustentabilidade. This research was funded by Fundação para a Ciência e a Tecnologia (FCT) through a PhD grant (SFRH/BD/23777/2005) and the research project EU Data Collection/DCR (PNAB). Skates and rays diversity, exploration and conservation | Table of Contents Table of Contents List of Figures ............................................................................................................................. i List of Tables ............................................................................................................................. v List of Abbreviations ............................................................................................................. viii Agradecimentos ........................................................................................................................ -
ESS 345 Ichthyology
ESS 345 Ichthyology Evolutionary history of fishes 12 Feb 2019 (Who’s birthday?) Quote of the Day: We must, however, acknowledge, as it seems to me, that man with all his noble qualities... still bears in his bodily frame the indelible stamp of his lowly origin._______, (1809-1882) Evolution/radiation of fishes over time Era Cenozoic Fig 13.1 Fishes are the most primitive vertebrate and last common ancestor to all vertebrates They start the branch from all other living things with vertebrae and a cranium Chordata Notochord Dorsal hollow nerve cord Pharyngeal gill slits Postanal tail Urochordata Cephalochordata Craniates (mostly Vertebrata) Phylum Chordata sister is… Echinodermata Synapomorphy – They are deuterostomes Fish Evolutionary Tree – evolutionary innovations in vertebrate history Sarcopterygii Chondrichthyes Actinopterygii (fish) For extant fishes Osteichthyes Gnathostomata Handout Vertebrata Craniata Figure only from Berkeley.edu Hypothesis of fish (vert) origins Background 570 MYA – first large radiation of multicellular life – Fossils of the Burgess Shale – Called the Cambrian explosion Garstang Hypothesis 1928 Neoteny of sessile invertebrates Mistake that was “good” Mudpuppy First Vertebrates Vertebrates appear shortly after Cambrian explosion, 530 MYA – Conodonts Notochord replaced by segmented or partially segmented vertebrate and brain is enclosed in cranium Phylogenetic tree Echinoderms, et al. Other “inverts” Vertebrate phyla X Protostomes Deuterostomes Nephrozoa – bilateral animals First fishes were jawless appearing -
List of Animal Species with Ranks October 2017
Washington Natural Heritage Program List of Animal Species with Ranks October 2017 The following list of animals known from Washington is complete for resident and transient vertebrates and several groups of invertebrates, including odonates, branchipods, tiger beetles, butterflies, gastropods, freshwater bivalves and bumble bees. Some species from other groups are included, especially where there are conservation concerns. Among these are the Palouse giant earthworm, a few moths and some of our mayflies and grasshoppers. Currently 857 vertebrate and 1,100 invertebrate taxa are included. Conservation status, in the form of range-wide, national and state ranks are assigned to each taxon. Information on species range and distribution, number of individuals, population trends and threats is collected into a ranking form, analyzed, and used to assign ranks. Ranks are updated periodically, as new information is collected. We welcome new information for any species on our list. Common Name Scientific Name Class Global Rank State Rank State Status Federal Status Northwestern Salamander Ambystoma gracile Amphibia G5 S5 Long-toed Salamander Ambystoma macrodactylum Amphibia G5 S5 Tiger Salamander Ambystoma tigrinum Amphibia G5 S3 Ensatina Ensatina eschscholtzii Amphibia G5 S5 Dunn's Salamander Plethodon dunni Amphibia G4 S3 C Larch Mountain Salamander Plethodon larselli Amphibia G3 S3 S Van Dyke's Salamander Plethodon vandykei Amphibia G3 S3 C Western Red-backed Salamander Plethodon vehiculum Amphibia G5 S5 Rough-skinned Newt Taricha granulosa -
Vii Fishery-At-A-Glance Night Smelt Scientific Name: Spirinchus Starksi Range
Fishery-at-a-Glance Night Smelt Scientific Name: Spirinchus starksi Range: Night Smelt are distributed coast-wide from southeast Alaska to Point Arguello, Santa Barbara County. Habitat: Night Smelt occur in the surf and in depths from the surface to approximately 400 feet (122 meters). Size (length and weight): Night Smelt measure less than 6 inches total length (140 millimeters) weighing to 11 grams. Males are slightly longer and heavier than females. Life span: Night Smelt are short lived and believed to reach a maximum of 2 to 3 years. Reproduction: Spawning occurs in the surf along open coast coarse sand beaches from January to September. Eggs are fertilized in the wash of the surf, adhere to sand grains, and sink. Hatching occurs in approximately 2 weeks. Prey: Night Smelt feed on small crustaceans—primarily gammarid amphipods and mysid shrimp. Predators: Night Smelt provide forage for a wide range of predators, including Striped Bass, Redtail Surfperch, salmon, Harbor Seals, California Sea Lions, terns, gulls, and cormorants. Fishery: Commercial and recreational fisheries are shore-based. Area fished: Historically, fishing occurred from Moss Landing, Monterey County to the Oregon border. Currently, fishing occurs from San Mateo County to Del Norte County. Fishing season: Fishing occurs during the spawning season—January to September. Fishing gear: Fishermen fish from shore using A-frame dip nets. Market(s): Landed fish are sold for human consumption and aquarium food. Current stock status: No formal stock assessments exist for Night Smelt. Although catch rates have increased on average since the early 2000s, it is undetermined if this increase in the index is due to increased abundance or changes in fishermen behavior. -
A Framework of Ichthyofaunal Ecostratigraphy of the Oligocene–Early Miocene Strata of the Polish Outer Carpathian Basin
Annales Societatis Geologorum Poloniae (2006), vol. 76: 1–111. A FRAMEWORK OF ICHTHYOFAUNAL ECOSTRATIGRAPHY OF THE OLIGOCENE–EARLY MIOCENE STRATA OF THE POLISH OUTER CARPATHIAN BASIN Janusz KOTLARCZYK1, Anna JERZMAÑSKA2, Ewa ŒWIDNICKA2 & Teresa WISZNIOWSKA2 1 Department of General and Mathematical Geology, AGH University of Science and Technology, Al. Mickiewicza 30, 30-059 Kraków, Poland 2 Department of Palaeozoology, Institute of Zoology, University of Wroc³aw, Sienkiewicza, 21, 50-335 Wroc³aw, Poland Kotlarczyk, J., Jerzmañska, A., Œwidnicka, E. & Wiszniowska, T., 2006. A framework of ichthyofaunal ecostrati- graphy of the Oligocene–Early Miocene strata of the Polish Outer Carpathian basin. Annales Societatis Geologorum Poloniae, 76: 1–111. Abstract: The paper presents the results of an analysis of ichthyofaunal variability throughout the section of the Menilite-Krosno Series (MKS) in the Outer Carpathians of Poland. The studied tanathocoenoses were formed at the bottom of a more than 2,000 m deep northern basin of the Tethys, being largely represented by the continental rise and bottoms of its narrow furrows, and – to a lesser degree – the continental slope and slopes of a submarine high. Lateral variability of statististically representative assemblages of tanathocoenoses hosted in thin, isochro- nous horizons is interpreted as a result of both local changes of ichthyocoenoses and the influence of post-mortem relocation of fishes that mainly dwelled the shelf and upper continental slope. Vertical variability, in turn, is considered as a resulting from changeable conditions of the ecological environment, the input and outflow of taxa whose evolution proceeded in the Indo-Pacific area, and the species extinction. Changeability of ichthyofauna within a ca. -
DEEP SEA LEBANON RESULTS of the 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project
DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project March 2018 DEEP SEA LEBANON RESULTS OF THE 2016 EXPEDITION EXPLORING SUBMARINE CANYONS Towards Deep-Sea Conservation in Lebanon Project Citation: Aguilar, R., García, S., Perry, A.L., Alvarez, H., Blanco, J., Bitar, G. 2018. 2016 Deep-sea Lebanon Expedition: Exploring Submarine Canyons. Oceana, Madrid. 94 p. DOI: 10.31230/osf.io/34cb9 Based on an official request from Lebanon’s Ministry of Environment back in 2013, Oceana has planned and carried out an expedition to survey Lebanese deep-sea canyons and escarpments. Cover: Cerianthus membranaceus © OCEANA All photos are © OCEANA Index 06 Introduction 11 Methods 16 Results 44 Areas 12 Rov surveys 16 Habitat types 44 Tarablus/Batroun 14 Infaunal surveys 16 Coralligenous habitat 44 Jounieh 14 Oceanographic and rhodolith/maërl 45 St. George beds measurements 46 Beirut 19 Sandy bottoms 15 Data analyses 46 Sayniq 15 Collaborations 20 Sandy-muddy bottoms 20 Rocky bottoms 22 Canyon heads 22 Bathyal muds 24 Species 27 Fishes 29 Crustaceans 30 Echinoderms 31 Cnidarians 36 Sponges 38 Molluscs 40 Bryozoans 40 Brachiopods 42 Tunicates 42 Annelids 42 Foraminifera 42 Algae | Deep sea Lebanon OCEANA 47 Human 50 Discussion and 68 Annex 1 85 Annex 2 impacts conclusions 68 Table A1. List of 85 Methodology for 47 Marine litter 51 Main expedition species identified assesing relative 49 Fisheries findings 84 Table A2. List conservation interest of 49 Other observations 52 Key community of threatened types and their species identified survey areas ecological importanc 84 Figure A1. -
First Records of the Night Smelt, Spirinchus Starksi, in the Salish Sea
First Records of the Night Smelt, Spirinchus starksi, in the Salish Sea, Washington Author(s): Melanie M Paquin , Anna N Kagley , Kurt L Fresh , and James W Orr Source: Northwestern Naturalist, 95(1):40-43. 2014. Published By: Society for Northwestern Vertebrate Biology DOI: http://dx.doi.org/10.1898/NWN13-05.1 URL: http://www.bioone.org/doi/full/10.1898/NWN13-05.1 BioOne (www.bioone.org) is a nonprofit, online aggregation of core research in the biological, ecological, and environmental sciences. BioOne provides a sustainable online platform for over 170 journals and books published by nonprofit societies, associations, museums, institutions, and presses. Your use of this PDF, the BioOne Web site, and all posted and associated content indicates your acceptance of BioOne’s Terms of Use, available at www.bioone.org/ page/terms_of_use. Usage of BioOne content is strictly limited to personal, educational, and non- commercial use. Commercial inquiries or rights and permissions requests should be directed to the individual publisher as copyright holder. BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. 40 NORTHWESTERN NATURALIST 95(1) NORTHWESTERN NATURALIST 95:40–43 SPRING 2014 FIRST RECORDS OF THE NIGHT SMELT, SPIRINCHUS STARKSI, IN THE SALISH SEA, WASHINGTON MELANIE MPAQUIN,ANNA NKAGLEY,KURT LFRESH, AND JAMES WORR Key words: COI, distribution, genetics, (frozen or preserved in ethanol) from 11 S. starksi Longfin Smelt, Night Smelt, Osmeridae, Spir- and 5 S. thaleichthys were obtained from the inchus starksi, Spirinchus thaleichthys, Washing- Northwest Fisheries Science Center (NWFSC) ton and the University of Washington Fish Collec- tion (UW). -
Updated Checklist of Marine Fishes (Chordata: Craniata) from Portugal and the Proposed Extension of the Portuguese Continental Shelf
European Journal of Taxonomy 73: 1-73 ISSN 2118-9773 http://dx.doi.org/10.5852/ejt.2014.73 www.europeanjournaloftaxonomy.eu 2014 · Carneiro M. et al. This work is licensed under a Creative Commons Attribution 3.0 License. Monograph urn:lsid:zoobank.org:pub:9A5F217D-8E7B-448A-9CAB-2CCC9CC6F857 Updated checklist of marine fishes (Chordata: Craniata) from Portugal and the proposed extension of the Portuguese continental shelf Miguel CARNEIRO1,5, Rogélia MARTINS2,6, Monica LANDI*,3,7 & Filipe O. COSTA4,8 1,2 DIV-RP (Modelling and Management Fishery Resources Division), Instituto Português do Mar e da Atmosfera, Av. Brasilia 1449-006 Lisboa, Portugal. E-mail: [email protected], [email protected] 3,4 CBMA (Centre of Molecular and Environmental Biology), Department of Biology, University of Minho, Campus de Gualtar, 4710-057 Braga, Portugal. E-mail: [email protected], [email protected] * corresponding author: [email protected] 5 urn:lsid:zoobank.org:author:90A98A50-327E-4648-9DCE-75709C7A2472 6 urn:lsid:zoobank.org:author:1EB6DE00-9E91-407C-B7C4-34F31F29FD88 7 urn:lsid:zoobank.org:author:6D3AC760-77F2-4CFA-B5C7-665CB07F4CEB 8 urn:lsid:zoobank.org:author:48E53CF3-71C8-403C-BECD-10B20B3C15B4 Abstract. The study of the Portuguese marine ichthyofauna has a long historical tradition, rooted back in the 18th Century. Here we present an annotated checklist of the marine fishes from Portuguese waters, including the area encompassed by the proposed extension of the Portuguese continental shelf and the Economic Exclusive Zone (EEZ). The list is based on historical literature records and taxon occurrence data obtained from natural history collections, together with new revisions and occurrences. -
New Zealand Fishes a Field Guide to Common Species Caught by Bottom, Midwater, and Surface Fishing Cover Photos: Top – Kingfish (Seriola Lalandi), Malcolm Francis
New Zealand fishes A field guide to common species caught by bottom, midwater, and surface fishing Cover photos: Top – Kingfish (Seriola lalandi), Malcolm Francis. Top left – Snapper (Chrysophrys auratus), Malcolm Francis. Centre – Catch of hoki (Macruronus novaezelandiae), Neil Bagley (NIWA). Bottom left – Jack mackerel (Trachurus sp.), Malcolm Francis. Bottom – Orange roughy (Hoplostethus atlanticus), NIWA. New Zealand fishes A field guide to common species caught by bottom, midwater, and surface fishing New Zealand Aquatic Environment and Biodiversity Report No: 208 Prepared for Fisheries New Zealand by P. J. McMillan M. P. Francis G. D. James L. J. Paul P. Marriott E. J. Mackay B. A. Wood D. W. Stevens L. H. Griggs S. J. Baird C. D. Roberts‡ A. L. Stewart‡ C. D. Struthers‡ J. E. Robbins NIWA, Private Bag 14901, Wellington 6241 ‡ Museum of New Zealand Te Papa Tongarewa, PO Box 467, Wellington, 6011Wellington ISSN 1176-9440 (print) ISSN 1179-6480 (online) ISBN 978-1-98-859425-5 (print) ISBN 978-1-98-859426-2 (online) 2019 Disclaimer While every effort was made to ensure the information in this publication is accurate, Fisheries New Zealand does not accept any responsibility or liability for error of fact, omission, interpretation or opinion that may be present, nor for the consequences of any decisions based on this information. Requests for further copies should be directed to: Publications Logistics Officer Ministry for Primary Industries PO Box 2526 WELLINGTON 6140 Email: [email protected] Telephone: 0800 00 83 33 Facsimile: 04-894 0300 This publication is also available on the Ministry for Primary Industries website at http://www.mpi.govt.nz/news-and-resources/publications/ A higher resolution (larger) PDF of this guide is also available by application to: [email protected] Citation: McMillan, P.J.; Francis, M.P.; James, G.D.; Paul, L.J.; Marriott, P.; Mackay, E.; Wood, B.A.; Stevens, D.W.; Griggs, L.H.; Baird, S.J.; Roberts, C.D.; Stewart, A.L.; Struthers, C.D.; Robbins, J.E.