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INSECTA MUNDI, Vol. 15, No.1, March, 2001 35

Preliminary checklist of the orthopteroid (, Mantodea, ,) of Texas

John A. Stidham 301 Pebble Creek Dr., Garland, TX 75040

and

Thomas A. Stidham Department of Integrative Biology, Museum of Paleontology, and Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720,

Abstract: Texas has one of the most diverse orthopteroid assemblages of any state in the , reflecting the varied habitats found in the state. Three hundred and eighty-nine species and 78 subspecies of orthopteroid insects (Blattodea, Mantodea, Phasmatodea, and Orthoptera) have published records for the state of Texas. This is the first such comprehensive checklist for Texas and should aid future work on these groups in this area.

Introduction (Flook and Rowell, 1997). The species are listed alphabetically within families or subfamilies, and Overall, the orthopteroid fauna of North Amer­ therefore the alphabetical taxonomic listing does ica is fairly well known. However, details of the not imply any phylogenetic relationships within geographic distribution of many orthopteroid spe­ the subfamily or family. The reference for the cies in the U.S. are not well documented. We have published record is given after the species name compiled the published records of Texas ortho­ and authority. The alpha is still unclear pteroids, as a basis for future study. Most of these for a few species included in this list, and therefore records have been confirmed through collections additional species may be added in the future as the made by the authors throughout Texas over the last number of taxa is clarified. This checklist is meant 20 years. The study of Texas orthopteroids began in to act as an aid to collectors, collection managers, the mid-nineteenth century and continues today field workers, and other researchers working on with 389 species currently recorded in the state. Texas orthopteroids. This places the Texan orthopteroid fauna as one of the most diverse of any state. There are 197 species The Orthopteroidea of Texas of known from Texas, compared to the over 200 species from California (Strohecker et Order Blattodea al., 1968), though other orthopteroid groups are more diverse in Texas. The large number of species Farrrily Blattelidae is most likely related to the great variety of habitats found in Texas, from desert in the West, to high Blattella germanica (Linnaeus) (Hebard, 1943b) plains in the North, and subtropical climates in the Blattella uaga Hebard (Atkinson et al., 1991) South. Combined with failure of Pleistocene gla­ Cariblatta lutea lutea (Saussure and Zehntner) ciers to reach Texas, the state harbors factors (Atkinson et al., 1991) necessary for long term orthopteroid diversifica­ Cariblatta lutea minima Hebard (Atkinson et al., tion, as seen in its speciose fauna. However, the 1991) number of endemic species is low compared to Chorisoneura texensis Saussure and Zehntner (He­ California, since many of the habitats (and hence bard, 1943b) species ranges) in Texas extend into the adjacent Euthlastoblatta abortiua Caudell (Hebard, 1943b) states and into Mexico. Euthlastoblatta gemma (Hebard) (Atkinson et al., Supraspecific taxonomy follows Otte (1981, 1991) 1984) and Helfer (1987). Tridactylids are placed in Isch.noptera deropeltiformis (Brunner) (Hebard, following tradition, though recent molec­ 1943b) ular data suggest their placement within 36 Vohrme 15, No.1, March, 2001, INSECTA MUNDI

Ischnoptera rufa occidentalis Saussure (Hebard, Order Mantodea 1943b) Parcoblaita bolliana (Saussure and Zehntner) (He­ Family Mantidae bard, 1943b) caudelli Hebard (Hebard, 1943b) Brunnea borealis Scudder (Hebard, 1943b) Parcoblatta desertae (Rehn and Hebard) (Hebard, Litaneutria minor (Scudder) (Hebard, 1943b) 1943b) Oligonicellamexicanus (Saussure) (Hebard, 1943b) Parcoblatta divisa (Saussure and Zehntner) (He­ Oligonicella scudderi (Saussure) (Hebard, 1943b) bard, 1943b) Phyllovates chlorophaea (Blanchard) (Hebard, Parcoblatta fulvescens (Saussure and Zehntner) 1943b) (Hebard, 1943b) Stagmomantis californica Rehn and Hebard (He­ Parcoblatta lata (Brunner) (Hebard, 1943b) bard, 1943b) Parcoblatta pennsylvanica (DeGeer) (Hebard, Stagmomcaitis carolina (Johannson) (Hebard, 1943b) 1943b) Parcoblatia virginica (Brunner) (Hebard, 1943b) Stagmotnantie limbata (Hahn) (Hebard, 1943b) Parcoblatta zebra Hebard (Hebard, 1943b) Yersiniops solitarium. (Scudder) (Hebard, 1943b) Plectoptera picta Saussure and Zehntner (Atkin- son et al., 1991) Order Phasmatodea Pseudomops septentrionalis Hebard (Hebard, 1943b) Family Pseudophasmatidae Supella longipalpa (Fabricius) (Hebard, 1943b) Anieomorphaferruginea (Beauvois) (Hebard, 1943b) Family Blattidae Family Heteronemiidae Blatta lateralis (Walker) (Atkinson et al., 1991) Blatta orientalis Linnaeus (Hebard, 1943b) Diapheromera covillea (Rehn and Hebard) (He- Neostylopyga rhotnbifolea (Stoll) (Atkinson et al., bard, 1943b) 1991) Diapheromera [emorata (Say) (Hebard, 1943b) Periplaneta americana (Linnaeus) (Hebard, 1943b) Diapheromera persimilis Caudell (Hebard, 1943b) Periplaneta australasiae (Fabricius) (Hebard, Diapheromera tamaulipennis (Rehn) (Hebard, 1943b) 1943b) Periplaneta brunnea Burmeister (Hebard, 1943b) Diapheromera torquato Hebard (Hebard, 1943b) Periplaneta fuliginosa (Serville) (Hebard, 1943b) Diapheromera velii eucnetnis Hebard (Hebard, 1943b) Family Blaberidae Diapheromera velii velii Walsh (Hebard, 1943b) Megaph.asma denticrus (Sti'tl) (Hebard, 1943b) Panchlora cubensis Saussure (Hebard, 1943b) Parabacillus coloradus (Scudder) (Hebard, 1943b) Pycnoscelus surinamensis (Linnaeus) (Hebard, Pseudosennylestraminea (Scudder) (Hebard, 1943b) 1943b) Pseudosennyle strigata (Scudder) (Hebard, 1943b) Sermyle mexicana (Saussure) (Hebard, 1943b) Family Polyphagidae Order Orthoptera Arenivaga bolliana (Saussure) (Hebard, 1943b) Arenivaga erraiica Rehn (Hebard, 1943b) Suborder Caelifera Arenivaga grata Hebard (Hebard, 1943b) Arenivaga tonhauia Hebard (Hebard, 1943b) Family Attaphila fungicola Wheeler (Hebard, 1943b) Compsodes cucullatus Saussure and Zehntner (At- Neotettix [emoratus (Scudder) (Rehn and Grant, kinson et al., 1991) 1961) Compsodes schwarzi (Caudell) (Atkinson et al., Neotettix nullisinus (Hancock) (Rehn and Grant, 1991) 1961) Eremoblatia subdiaphana (Scudder) (Hebard, Nomotettix cristatus compreseue Morse (Rehn and 1943b) Grant, 1961) INSECTA MUNDI, Vol. 15, No.1, March, 2001 37

Paratettix aztecus (Saussure) (Rehn and Grant, Mertneria bivitatta bivitatta (Serville) (Knutson, 1961) 1940) brevipennis (Hancock) (Rehn and Grant, Mertneria bivitatta maculipennis Bruner (Knut- 1961) son, 1940) Paratettix cucullatue (Burmeister) (Rehn and Grant, Mertneria picta neomexicana (Thomas) (Otte, 1981) 1961) Mermeria picta picta (Walker) (Tinkham, 1948) Paratettix mexicanus (Saussure) (Rehn and Grant, Mermeria texana Bruner (Tinkham, 1948) 1961) obscura (Thomas) (Otte, 1981) Paratettix rugosu.s (Scudder) (Rehn and Grant, Orpliulella olivacea (Morse) (Gurney, 1940) 1961) olivacea halophila Rehn and Hebard Parateitix toltecus (Saussure) (Rehn and Grant, (Gurney, 1940) 1961) (Burmeister) (Otte, 1981) Tetrix arenosa angusta Hancock (Rehn and Grant, Orphulella speciosa (Scudder) (Otte, 1981) 1961) Parapomala pallida Bruner (Otte, 1981) Tetrix arenosa arenosa Burmeister (Rehn and Parapomala virgata Scudder (Otte, 1981) Grant, 1961) Parapomala wyomingensis (Thomas) (Otte, 1981) Tettigidea armata Morse (Rehn and Grant, 1961) Phlibostroma quadrimaculatum. (Thomas) (Otte, Tettigidea lateralis cazieri Rehn and Grant (Rehn 1981) and Grant, 1961) brach.yptera (Scudder) (Otte, 1981) Tettigidea lateralis lateralis (Say) (Rehn and Grant, delicatula delicatula (Scudder) (Rehn, 1961) 1942) pawnee Rehn (Rehn, 1942) Family Psoloessa texana pusilla (Scudder) (Rehn, 1942) Psoloessa texana texana Scudder (Rehn, 1942) Subfamily Rhammatocerue viatorius (Saussure) (Tinkham, 1948) Ach.urum hilliardi Gurney (Helfer, 1987) admirabilis (Uhler) (Otte, 1981) tninimipenne Caudell (Otte, 1981) Syrbula montezuma (Saussure) (Otte, 1981) Achuru.m. sumichrasti (Saussure) (Otte, 1981) Aerolophitus hirtipes hirtipes (Say) (Otte, 1981) Subfamily Aerolophitus hirtipes variegatus Bruner (Helfer, 1987) Metaleptea breoicornis (Johannson) (Otte, 1981) Aerolophitus maculipennis (Scudder) (Otte, 1981) breoipennis (Bruner) (Otte, 1981) Subfamily Oedipodinae Ageneotettix deorurn (Scudder) (Otte, 1981) Amphitornie coloradus (Thomas) (Otte, 1981) Anconia hebardi Rehn (Otte, 1984) elliotti (Thomas) (Otte, 1981) conepersa Scudder (Otte, 1984) Aulocara femoraturn. (Scudder) (Otte, 1981) Arphia pecos Otte (Otte, 1984) aurioentris McNeill (Otte, 1981) Arphia pseudonietana (Thomas) (Otte, 1984) Boopedon gracile Rehn (Otte, 1981) Arphia simplex Scudder (Otte, 1984) . (Say) (Otte, 1981) Arphia sulphurea (Fabricius) (Otte, 1984) argeniatiis Bruner (Otte, 1981) Arphia xanthoptera (Burmeister) (Otte, 1984) Cibolaeris parviceps (Walker) (Otte, 1981) Chortophaga australior Rehn and Hebard (Otte, Cibolaeris samalayucae Tinkham (Otte, 1981) 1984) Cordillocris erenulata (Bruner) (Otte, 1981) Chortophaga oiridifasciata (DeGeer) (Otte, 1984) occipitalis (Thomas) (Otte, 1981) rabula Rehn and Hebard (Otte, 1984) elegans (Morse) (Otte, 1981) Conozoa texana (Bruner) (Otte, 1984) Dichrotnorpha viridis (Scudder) (Otte, 1981) Derottnena haydenii (Thomas) (Otte, 1984) abortious (Bruner) (Otte, 1981) Derotmena laticinctum Scudder (Otte, 1984) (Scudder) (Otte, 1981) Dissosteira carolina (Linnaeus) (Otte, 1984) Heliaula rufa (Scudder) (Otte, 1981) Dissosteim longipennis (Thomas) (Otte, 1984) planum (Bruner) (Otte, 1981) 38 Volume 15, No.1, March, 2001, INSECTA MUNDI

Encoptolophus sordidus costalis (Scudder) (Knut­ Subfamily son, 1940) subgracilis texensis Bruner (Knut- elegans (Scudder) (Tinkham, 1948) son, 1940) bruneri (Caudell) (Wallace, Eximacris superburn (Hebard) (Otte, 1984) 1955) magnifica (Rehn) (Otte, 1984) Agroecotettix modestus aristus Hebard (Hebard, (Say) (Otte, 1984) 1943a) Heliastus benjamini Caudell (Otte, 1984) Agroecotettix modestus crypsidomus Hebard (He­ Heliastus eubroeeus Caudell (Otte, 1984) bard, 1943a) Heliastus sumichrasti (Saussure) (Otte, 1984) Campylacantha lamprotata Rehn and Hebard (Rehn Hippiscus ocelote (Saussure) (Otte, 1984) and Hebard, 1909) Hippopedon capito (Stal) (Otte, 1984) Campylacantha olivacea olivacea (Scudder) (Knut­ Lepru.s intermedius Saussure (Otte, 1984) son, 1940) Leprus ioheeleri (Thomas) (Otte, 1984) Campylacantha olivacea vivax (Scudder) (Helfer, Mestobregma plattei corrugata (Scudder) (Otte, 1987) 1984) Chloropus cactocaetes Hebard (Helfer, 1987) Mestobregma terricolor Rehn (Otte, 1984) Clematodes larrae Scudder (Tinkham, 1948) pardalinus (Saussure) (Otte, 1984) Dactylotutn. bicolor piciurn. (Thomas) (Knutson, Pardalophora haldemanii (Scudder) (Otte, 1984) 1940) Pardalophora phoenicoptera (Burmeister) (Otte, Dactylotum bicolor oariegatum (Scudder) (Helfer, 1984) 1987) Pardalophora saussurei (Scudder) (Otte, 1984) Dendrotettix quercus Packard (Helfer, 1987) Platylactista aztecus (Saussure) (Tinkham, 1948) Hesperotettix speciosus (Scudder) (Knutson, 1940) Psinidia cunplicornis Caudell (Otte, 1984) Hesperotettix viridis pratensis Scudder (Knutson, Psinidia fenestralis fenestralis (Serville) (Otte, 1984) 1940) Psinidia fenestralis fraier Rehn (Rehn, 1919) Hesperotettix viridis viridis (Thomas) (Knutson, Spharagemon. bolli Scudder (Otte, 1984) 1940) Spharagemoti bolli inornatum. Morse (Tinkham, Hypochlora alba (Dodge) (Knutson, 1940) 1948) alexanderi Hilliard (Hilliard, 2001) Spharagemon. campestrie (McNeill) (Otte, 1984) Melanoplus angularis Little (Knutson, 1940) Spharagemoti collare (Scudder) (Otte, 1984) Melanoplus angustipennis angustipennis (Dodge) Spharagemoti crietatum (Scudder) (Otte, 1984) (Helfer, 1987) Spharagemon. equale (Say) (Otte, 1984) Melanoplus angustipennis itnpiger Scudder (Knut- Spharagemon. marmorata picta (Scudder) (Otte, son, 1940) 1984) Melanoplus aridus (Scudder) (Spooner, 1989) Trachyrhachis aspera Scudder (Otte, 1984) Melanoplus arizonae Scudder (Helfer, 1987) Trachyrhachis coronata Scudder (Otte, 1984) Melanoplus bispinosus Scudder (Knutson, 1940) Trachyrhachis kiowa [uecifrons (Stal) (Otte, 1984) Melanoplus bivittatus (Say) (Knutson, 1940) Trachyrhachis kiowa kiowa (Thomas) (Otte, 1984) Melanoplus bowditchi Scudder (Rehn and Hebard, citrina Scudder (Otte, 1984) 1909) Trimerotropis cyaneipennis Bruner (Otte, 1984) Melanoplus confusus Scudder (Knutson, 1940) Trimerotropis latifasciata Scudder (Otte, 1984) Melanoplus dakini Hilliard (Hilliard, 2001) Trimerotropis melanoptera McNeill (Otte, 1984) Melanoplus desultorius Rehn (Spooner, 1989) Trimerotropis pallidipennis (Burmeister) (Otte, Melanoplus differentialis nigricans Cockerell (Rob- 1984) erts, 1942) Trimerotropis pistrinaria Saussure (Otte, 1984) Melanoplus discolor (Scudder) (Knutson, 1940) Trimerotropis salina McNeill (Otte, 1984) Melanoplus eumera Hebard (Tinkham, 1948) Trimerotropis saxatilis McNeill (Isely, 1935) Melanoplus [emur-rubru.m. (DeGeer) (Knutson, Trimerotropis schaefferi Caudell (Otte, 1984) 1940) Tritnerotropis strenua McNeill (Otte, 1984) Melanoplus flabellatus Scudder (Knutson, 1940) Tropidolophus [ormosue (Say) (Otte, 1984) Melanoplus flavidus Scudder (Knutson, 1940) Xanthippus corallipes pantherinus (Scudder) Melanoplus foedus fluviatilis Brunner (Knutson, (Helfer, 1987) 1940) INSECTA MUNDI, Vol. 15, No.1, March, 2001 39

Melanoplue [oedus foedus Scudder (Hebard, 1936) Schistocerca albolineata (Thomas) (Hubbell, 1960) Melanoplus [oedus iselyi Hebard. (Knutson, 1940) Schistocerca alutacea (Harris) (Hubbell, 1960) Melanoplus gladstoni Scudder (Helfer, 1987) Schistocerca americana (Drury) (Knutson, 1940) Melanoplus glaucipes (Scudder) (Knutson, 1940) Schistocerca damnified (Saussure) (Knutson, 1940) Melanoplus herbaceous Bruner (Rehn and Hebard, Schistocerca emarginata (Scudder) (Knutson, 1940) 1909) Schistocerca nitens (Thunberg) (Helfer, 1987) Melanoplus inconepicuus Caudell (Hebard, 1917a) Schistocerca obscura (Fabricius) (Helfer, 1987) Melanoplue heeleri (Thomas) (Knutson, 1940) Schistocerca rubiginosa (Harris) (Helfer, 1987) Melanoplus lahinus (Scudder) (Knutson, 1940) Schistocerca shoshone (Thomas) (Hubbell, 1960) Melanoplus mastigiphallus Strohecker (Stroheck- er, 1941) Subfamily Leptysminae Melanoplue mexicanus (Saussure) (Helfer, 1987) Melanoplus occidentalis (Thomas) (Helfer, 1987) Lepiysma marginicollis (Serville) (Knutson, 1940) Melanoplus packardii Scudder (Knutson, 1940) Stenacris uitreipennis (Marschall) (Hilliard, 1982) Melanoplus plebejus (Stal) (Helfer, 1987) Melanoplus ponderoeus (Scudder) (Knutson, 1940) Subfamily Melanoplue ponderosue uiola (Thomas) (Hebard, 1941) (Girard) (Rehn and Grant, 1961) Melanoplus punctulatue arboreus Scudder (Rehn, Phrynotettix robuetus occultus Rehn and Grant 1946) (Rehn and Grant, 1961) Melanoplus regalis (Dodge) (Helfer, 1987) Phrynotettix robuetus robustue (Bruner) (Rehn and Melanoplus rusticue obouatipennis Blatcheley (Cop­ Grant, 1961) pock, 1962) Phrynotettix tschiuauensis tschiuauensis (Halde­ Melanoplus eanguinipee defectus Scudder (Helfer, man) (Rehn and Grant, 1961) 1987) Phrynotettix tschiuauensis oerruculatus Glover Melanoplue sanguiiiipes uulturnis Gurney and (Rehn and Grant, 1961) Brooks (Gurney and Brooks, 1959) Rotnalea gutiatus (Stoll) (Rehn and Grant, 1961) Melanoplus scudderi latue Morse (Knutson, 1940) Taeniopoda equues (Burmeister) (Rehn and Grant, Melanoplue scudderi texensis Hart (Knutson, 1940) 1961) Melanoplue terininalis Scudder (Scudder, 1897) Melanoplue texanus (Scudder) (Knutson, 1940) Suborder Ensifera Melanoplue ihomasi Scudder (Helfer, 1987) Melanoplus tuberculatus Morse (Helfer, 1987) Family Gryllacrididae Melanoplu.s ioarneri Little (Knutson, 1940) Netrosoma nigropleura Scudder (Helfer, 1987) Paraidemona fratercula Hebard (Hebard, 1918) Subfamily Rhaphidophorinae Paraidetnotia latifurcula Hebard (Hebard, 1918) Paraidemana mimica (Scudder) (Hebard, 1918) Ammobaenetes phrixocnemoides (Caudell) (Helfer, Paraidemona nuttingi Yin and Smith (Yin and 1987) Smith, 1989) Ceuthophilus carlsbadensis Caudell (Hubbell, 1936) Paroidemona olsoni Yin and Smith (Yin and Smith, Ceuthophilus conicaudu.s Hubbell (Hubbell, 1936) 1989) Ceuthophilus cunicularis Hubbell (Hubbell, 1936) Paraidemona punctata (Stal) (Hebard, 1918) Ceuthophilus dioergens Scudder (Hubbell, 1936) Paratylotropidea brunneri Scudder (Knutson, 1940) Ceuthophilus isletae Hubbell (Hubbell, 1936) Paroxya atlantica Scudder (Knutson, 1940) Ceuthophilus nodulosus Brunner (Hubbell, 1936) Paroxya claouliger (Serville) (Helfer, 1987) Ceuthophilus pallidue Thomas (Hubbell, 1936) Phaedrotettix accola (Scudder) (Hebard, 1917b) Ceuthophilus polingi Hubbell (Hubbell, 1936) Phaedrotettix dumicola (Scudder) (Hebard, 1917b) Ceuthophilus rehni Hubbell (Hubbell, 1936) Phaedrotettix palmeri (Scudder) (Hebard, 1917b) Ceuthophilus secretus Scudder (Hubbell, 1936) Phaulotettix compreeeus Scudder (Helfer, 1987) Ceuthophilus umbratilis Hubbell (Hubbell, 1936) Phaulotettix eurycercus Hebard (Helfer, 1987) Ceuthophilus utaheneis Thomas (Hubbell, 1936) Philoeleon nigrouittata (Stal) (Tinkham, 1948) Ceuthophilus uariegatus Scudder (Hubbell, 1936) Photaliotes nebrascensis (Thomas) (Knutson, 1940) Daihinia breuipes Haldeman (Helfer, 1987) 40 Volume 15, No.1, March, 2001, INSECTA MUNDI

Daihiniella bellicosus (Hubbell) (Hubbell, 1936) Subfamily Pseudophyllinae Daihiniodes hausteriferum (Rehn) (Hubbell, 1936) Udeopsylla robusta (Haldeman) (Helfer, 1987) Paracyrtoph.yllus excelsus (Rehn and Hebard) (He­ bard, 1941) Subfamily Stenopelmatinae Paracyrtoph.yllue robustus Caudell (Hebard, 1941) Pterophylla camellifolia camellifolia (Fabricius) Stenopelmatu.s fuscus Haldeman (Hebard, 1916a) (Hebard, 1941) Pterophylta camellifolia intennedia (Caudell) (He­ Family bard, 1941) Pterophyllafurcata furcata (Caudell) (Hebard, 1941) Subfamily Pieroph.ylla furcata laletica Hebard (Hebard, 1941)

Bucrates malivolens (Scudder) (Helfer, 1987) Subfamily brevipennis (Scudder) (Rehn and Hebard, 1915a) Amblycorypha floridana Rehn and Hebard (Rehn Conocephalus fasciatus (DeGeer) (Rehn and He­ and Hebard, 1914c) bard, 1915a) Amblycoryplui hausteca (Saussure) (Rehn and He­ Conocephalus hygrophilus Rehn and Hebard (Rehn bard, 1914c) and Hebard, 1915a) Amblycorypha insolita Rehn and Hebard (Rehn Conocephalus resacensis Rehn and Hebard (Rehn and Hebard, 1914c) and Hebard, 1915a) Amblycorypha oblongifolia (DeGeer) (Rehn and Conocephalus saltans (Scudder) (Rehn and He­ Hebard, 1914c) bard, 1915a) Amblycoryph.a rotundifolia parvipennis Stal (Rehn Conocephalus spartinae (H. Fox) (Rehn and He­ and Hebard, 1914c) bard, 1915a) AmblycoryphauhleriStal(RehnandHebard,1914c) Conocephalus strictus (Scudder) (Rehn and He­ Arethaea ambulaior Hebard (Rehn and Hebard, bard, 1915a) 1914a) exiliscanorus (Davis) (Helfer, Arethaea arachnopyga Rehn and Hebard. (Rehn 1987) and Hebard, 1914a) Neoconocephalus nebrascensis (Brunner) (Walker, Arethaea constricta comanche Hebard (Hebard, 1998) 1936b) Neoconocephalus robustus crepitans (Scudder) Arethaea constricta constricta Brunner (Hebard, (Tinkham, 1944) 1936b) Neoconocephalus triops (Linnaeus) (Helfer, 1987) Arethaea gracilipes cerciata Hebard. (Rehn and Neoconocephalus velox Rehn and Hebard (Walker, Hebard, 1914a) 1998) Arethaea gracilipes gracilipes (Thomas) (Rehn and Orchelitnum agile (DeGeer) (Morris and Walker, Hebard, 1914a) 1976) Arethaea grallator (Scudder) (Rehn and Hebard, Orchelimurn. bullatum. Rehn and Hebard (Rehn 1914a) and Hebard, 1915b) Arethaea phantasm a Rehn and Hebard (Rehn and . calcaratum Rehn and Hebard (Rehn Hebard, 1914a) and Hebard, 1915b) Arethaea semialata Rehn and Hebard (Rehn and Orchelimum. concinnum concinnum Scudder (Rehn Hebard, 1914a) and Hebard, 1915b) Dichopetala brevihastata Morse (Rehn and He­ Orchelitnum concinnum delicatulum Bruner (Tho­ bard, 1914b) mas and Alexander, 1962) Dichopetala castanea Rehn and Hebard (Rehn and Orchelimum. nigripes Scudder (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala catinata Rehn and Hebard (Rehn and Orchelimum. vulgare Harris (Rehn and Hebard, Hebard, 1914b) 1915b) Dichopetala emarginata Brunner (Rehn and He­ Pyrgocorypha uncuinata (Harris) (Helfer, 1987) bard, 1914b) INSECTA MUNDI, Vol. 15, No.1, March, 2001 41

Dichopetalagladiator Rehn and Hebard (Rehn and Family Tridactylidae Hebard, 1914b) Dichopetala oreoeca Rehn and Hebard (Rehn and Tridactylus apicalis Say (Helfer, 1987) Hebard, 1914b) Tridactylus minuiis Scudder (Helfer, 1987) Dichopetala pollicifera Rehn and Hebard (Rehn and Hebard, 1914b) Family Myrmecophilidae Dichopetala seeuersi Strohecker (Strohecker, 1941) Ineara elegans (Scudder) (Tinkham, 1944) Myrtnecophila nebrascensis Lugger (Helfer, 1987) Microcentrum latifrone Spooner (Spooner, 1989) Microcentrum minus Strohecker (Strohecker, 1952) Family Microcentrum rhombifoliurn. (Saussure) (Helfer, 1987) Montezumina modesta (Brunner) (Walker, 1998) Acheta domesticus (Linnaeus) (Blatchley, 1920) Scudderia curuicauda-laticauda Brunner (Rehn Allonemobius [aeciatus (DeGeer) (Hebard, 1913) and Hebard, 1914) Allonemobius fasciatus socius (Scudder) (Hebard, Scudderia [urcaia Brunner (Rehn and Hebard, 1913) 1914) Allonemobius griseus funeral is (Hart) (Hebard, Scudderia furcata [urcifera (Scudder) (Rehn and 1913) Hebard, 1914) Allonemobius sparsalus (Fulton) (Walker, 1998) Scudderia mexicana (Saussure) (Rehn and He­ Anaxipha exigua (Say) (Helfer, 1987) bard, 1914) Anurogryllus muticus (DeGeer) (Helfer, 1987) Scudderia texensis Saussure and Pictet (Rehn and Cycloptilium. antillarum (Redtenbacher) (Hebard, Hebard, 1914) 1916b) Cycloptilium. comprehendens copmprehendens He­ Subfamily Decticinae bard (Hebard, 1916b) Cycloptilium. comprehendens fortior Hebard (He­ bard, 1916b) Anabrus simplex Haldeman (Caudell, 1907) Cycloptilium kelainopum Love and Walker (Love Capnobotes fuliginosus (Thomas) (Tinkham, 1944) and Walker, 1979) Eremopedes couilleae Hebard (Tinkham, 1948) Cycloptilium pigriun Love and Walker (Love and Eremopedes scudderi Cockerell (Tinkham, 1948) Walker, 1979) Neobarrettia spinosus (Caudell) (Helfer, 1987) Cycloptilium slossoni Scudder (Love and Walker, Neobarrettia uictoriae (Caudell) (Helfer, 1987) 1979) Pediodectes daedalus (Rehn and Hebard) (Rehn Cycloptilium. squamosu.m. Scudder (Love and Walk­ and Hebard, 1920) er' 1979) Pediodectes grand is (Rehn) (Tinkham, 1944) Cycloptilium trigonipalpum Rehn and Hebard (Love Pediodectes haldemanii (Girard) (Tinkham, 1944) and Walker, 1979) Pediodectes nigromarginata (Caudell) (Rentz and Cyrtoxipha columbiana Caudell (Walker, 1969) Birchim, 1968) carolinus (Scudder) (Hebard, 1913) Pediodectes pratti (Caudell) (Rentz and Birchim, Eunemobiue carolinus neomexicanus (Scudder) 1968) (Hebard, 1913) Pediodectes steuensoni (Thomas) (Tinkham, 1944) Falcicula hebardi Rehn (Blatchley, 1920) Pediodectes tinkhami Hebard (Rentz and Birchim, Gryllodes sigillatus Walker (Alexander and Walker, 1968) 1962) firma Scudder (Alexander, 1968) Family Gryllotalpidae Gryllus fultoni Alexander (Alexander, 1968) Gryllus pennsyluanicus Burmeister (Alexander, Gryllotalpa cultriger Uhler (Frank et al., 1995) 1968) Neocurtilla hexadactyla (Perty) (Helfer, 1987) Grylluspersonatus Uhler (Rehn and Hebard, 1915c) Scapteriecus borelli Giglio-Tos (Frank et al., 1995) Gryllus rubens (Scudder) (Walker, 1998) Scapteriscus uicinus Scudder (Frank et al., 1995) Gryllue texensis Cade and Otte (Cade and Otte, 2000) Hapithus agitator Uhler (Helfer, 1987) 42 Volume 15, No.1, March, 2001, INSECTA MUNDI

Hoploephyru.m. boreale (Scudder) (Helfer, 1987) Flook, P.K. and C.H.F. Rowell. 1997. The phy­ Miogryllue lineatus (Scudder) (Hebard, 1915) logeny of the Caelifera (Insecta, Orthoptera) as Miogryllue uerticalis (Serville) (Hebard, 1915) deduced from mtrRNA gene sequences. Mol. Neonemobiue cubensis mormoniu.s (Scudder) (He- Phylog. Evol. 8: 89-103. bard, 1913) Frank, J.H., T.R. Fasulo, and D.E. Short. 1995. Neoxabea bipunctata (DeGeer) (Walker, 1963) McricketKnowledgebasehttp://gnv.lfas.ufl.edu/ Neoxabea formosa (F. Walker) (Walker, 1967) -entI/mcricket/. argentinue Saussure (Walker, 1962) Gurney, A.B. 1940. A revision of the grasshoppers Oecanthus califomicus Saussure (Walker, 1963) ofthe Orphulella from America North of Oecanthus celerinictus T.J. Walker (Walker, 1962) Mexico. Entomologica Am .. 20: 85-157. Oecanthus exclamationis Davis (Walker, 1963) Gurney, A.B. and A.R. Brooks. 1959. Grasshop­ Oecanthus fultoni T.J. Walker (Walker, 1963) pers of the Mexicanus group, genus Melano­ Oecanthus leptogrammus T.J. Walker (Walker, plus (Orthoptera: Acrididae). Proc. UB. Natl. 1963) Mus. 110: 1-93. Oecanthus niueus (DeGeer) (Walker, 1963) Hebard, M. 1913. A revision of the species of the Oecanthus pini Beutenmuller (Walker, 1962) genus Nemobius (Orthoptera: Gryllidae) found Oecanthus quadripunctatus Beutenmuller (Walk- in North America North of the Isthmus of er, 1962) Panama. Proc. Acad. Nat. Sci. Phila. 65: 394­ Orocharis saltator Uhler (Walker, 1969) 492. Phyllopalpus pulchellus (Uhler) (Walker, 1998) Hebard, M. 1915. The American species of the Trigonidomimus belfragei Caudell (Caudell, 1912) genus Miogryllus. J. New York Entomol. Soc. 23: 101-121. References Hebard, M. 1916a. A study of the species of the genus Stenopelmatus found in the United States. Alexander, R.D. 1968. Life cycle origins, specia­ J. New York Entomol. Soc. 24: 70-86. tion, and related phenomena in crickets. Quart. Hebard, M. 1916b. The Mogoplistinae (Ortho­ Rev. BioI. 43: 1-41. ptera, Gryllidae) of the United States. Trans. Alexander, R.D. and T.J. Walker. 1962. Two Am. Entomol. Soc. 42: 135-160. introd uced field crickets new to Eastern United Hebard, M. 1917a. New genera and species of the States (Orthoptera: Gryllidae). Ann. Entomol. Melanopli found within the United States and Soc. Am. 55: 90-94. Canada (Orthoptera: Acrididae). Parts X to Atkinson, T.A., P.G. Koehler, and R.S. Patter­ XIV. Trans. Am. Entomol. Soc. 43:147-163. son. 1991. Catalog and atlas of the cockroaches Hebard, M. 1917b. Notes on Mexican Melanopli (Dictyoptera) of North America North of Mexi­ (Orthoptera: Acrididae). Proc. Acad. Nat. Sci. co. Misc. Publ. Entomol. Soc. Amer. 78: 85 pp. Phila. 69: 251-275. Blatchley, W.S. 1920. Orthoptera of Northeastern Hebard, M. 1918. New genera and species of America. Nature Publ. Co., Indianapolis, Indi­ Melanopli found within the United States (01'­ ana. thoptera: Acrididae). Trans. Am. Entomol. Soc. Cade, W.H. and D. Otte. 2000. Gryllus texensis n. 44: 141-169. sp.: a widely studied field (Orthoptera; Hebard, M. 1936a. New genera and species of the Gryllidae) from the Southern United States. Melanopli found within the U.S. and Canada, Trans. Amer. Entomol. Soc. 126: 117-123. Part VIII. Trans. Am. Entomol. Soc. 62: 182­ Caudell, AX 1907. The Decticinae (a group of 190. Orthoptera) of North America. Proc. U.S. Natl. Hebard, M. 1936b. VI. A revision of the genus Mus. 32: 285-410. Arethaea (Tettigoniidae, Phaneropterinae). Caudell, A.N. 1912. A new genus and species of Trans. Am. Entomol. Soc. 62: 231-256. Gryllidae from Texas. Proc. Entomol. Soc. Wash. Hebard, M. 1941. The group Pterophyllae as found 14: 187-189. in the United States (Tettigoniidae: Pseudo­ Coppock, S., Jr. 1962. The grasshoppers of Okla­ phyllinae). Trans. Am. Entomol. Soc. 67: 197­ homa (Orthoptera: Acrididae) Oklahoma State 219. Univ. Agric. Exp. Sta. Report P-399: 143 pp. Hebard, M. 1943a. New genera and species of Melanoplifound within the UB., PartlV. Trans. Am. Entomol. Soc. 68: 49-66. INSECTA MUNDI, Vol. 15, No.1, March, 2001 43

Hebard, M. 1943b. The Dermaptera and Ortho­ Rehn, J.A.G. 1942. On the locust genus Psoloessa pterous families Blattidae, Mantidae, and Phas­ (Orthoptera: Acrididae: Acridinae). Trans. Am. midae of Texas. Trans. Am. Entomol. Soc. 48: Entomol. Soc. 68: 167-237. 239-313. Rehn, J.A.G. 1946. On the Punctulatus species­ Helfer, J.R. 1987. How to know the grasshoppers, group of the genus Melanoplus (Orthoptera, crickets, cockroaches and their allies. Dover Acrididae, Cyrtacanthacridinae). Proc. Acad. Publications, Inc., New York. Nat. Sci. 98: 241-269. Hilliard, J.R., Jr. 1982. Endophytic oviposition by Rehn, J.A.G. and H.J. Grant Jr. 1961. A mono­ Lepiysma marginicollis marginicollis and Ste­ graph of the Orthoptera of North America (North nacris vitreipennis (Orthoptera: Acrididae: Lep­ of Mexico), Vol. 1. Monographs of the Acad. Nat. tysminae) with life history notes. Trans. Am. Sci. Phila., 12: 257 pp. Entomol. Soc. 108: 153-180. Rehn, J.A.G. and M. Hebard. 1909. An ortho­ Hilliard, J.R., Jr. 2001. Two new pterological reconnaisance of the Southwest­ species in the Texanus group of the genus ern United States. Part II: New Mexico and Melanoplus (Orthoptera: Acrididae: Molano­ Western Texas. Proc. Acad. Nat. Sci. Phila. 61: plinae) with biologicalnotes on the group. Trans. 111-175. Am. Entomol. Soc. 127: 31-68. Rehn, J.x.G. and M. Hebard. 1914a. A revision of Hubbell, T.H. 1936. A monographic revision ofthe the orthopterous group Insarae (Tettigoniidae, genus Ceuthophilue (Orthoptera, Gryllacrid­ Phaneropterinae), Trans. Am. Entomol. Soc. idae, Rhaphidophorinae). Univ. Fla. Pub. BioI. 40: 37-184. Sci. Series 2: 1-551. Rehn, J.A.G. and M. Hebard. 1914b. A study of Hubbell, T.H. 1960. The sibling species of the the species of the genus Dichopetala (Ortho­ Alutacea group of the bird-locust genus Schis­ ptera: Tettigoniidae). Proc. Acad. Nat. Sci. Phi­ iocerca (Orthoptera, Acrididae, Cyrtacan­ lao 66: 64-160. thacridinae). Misc. Publ. Mus. Zool., Univ. Mich. Rehn, J.A.G.and M. Hebard. 1914c. Studies in 116: 1-91. American Tettigoniidae (Orthoptera) I and II. Isely, F.B. 1935. Acridian researches within North­ Trans. Am. Entomol. Soc. 40: 271-344. eastern Texas (Orthoptera). Entomol. News 46: Rehn, J.A.G. and M. Hebard. 1915a. A synopsis 37-75. of the species of the genus Conocephalue Knutson, H. 1940. A key to the Acrididae (Ortho­ (Xiphidium of authors) found in North America ptera) of Northeastern Texas with ecological North of Mexico. Trans. Am. Entomol. Soc. 41: notes. Field and Laboratory 8: 33-58. 155-224. Love, R.E. and T.J. Walker. 1979. Systematics Rehn, J.A.G. and M. Hebard. 1915b. A synopsis and acoustic behavior of scaly crickets (Ortho­ of the species of the genus Orchelimum, Trans. ptera: Gryllidae: Mogoplistinae) of Eastern Am. Entomol. Soc. 41: 11-83. United States. Trans. Am. Entomol. Soc. 105: 1­ Rehn, J.A.G. and M. Hebard. 1915c. The genus 66. Gryllus as found in America. Proc. Acad. Nat. Morris, G.K. and T.J. Walker. 1976. Calling Sci. Phila. 77: 293-322. songs of Orchelimurn. meadow katydids (Tet­ Rehn, J.A.G. and M. Hebard. 1920. Description tigoniidae), 1. mechanism, terminology, and of new genera and species of North American geographic distribution. Can. Entomol. 108: Decticinae (Orthoptera: Tettigoniidae). Trans. 785-800. Am. Entomol. Soc. 46: 225-265. Otte, D. 1981. The North American grasshoppers, Rentz, D.C. and J.D. Birchim. 1968. Revisionary Volume I Acrididae, Gomphocerinae, and Acrid­ studies in the Nearctic Decticinae. Mem. Pacif­ inae. Harvard Univ. Press, Cambridge, Massa­ ic Coast Entomol. Soc. 3: 1-173. chusetts. Roberts, H.R. 1942. Two subspecies of Melano­ Otte, D. 1984. The North American grasshoppers, plus differentialis and related new subspecies Volume II Oedipodinae. Harvard Univ. Press, from Mexico with discussion of their variation. Cambridge, Massachusetts. Trans. Amer. Entomol. Soc. 68: 151-166. Rehn, J.A.G. 1919. Descriptions of new and criti­ Scudder, S.H. 1897. Revision of the orthopteran cal notes upon previously known forms of North group Melanopli (Acrididae), with special refer­ American Oedipodinae (Orthoptera: Acrididae). ence to North American forms. Proc. U.S. Natl. Trans. Am. Entomol. Soc. 45: 229-255. Mus. 20: 293-294. 44 Volume 15, No.1, March, 2001, INSECTA MUNDI

Spooner, J.D. 1989. Microcentrum latifrone, a lidae: Oecanthinae). 1. The Genus Neoxabea new species of angular-winged katydid from and the Niueus and Varicornis Groups of the Southwestern United States (Orthoptera, Tet­ Genus Oecanthus. Ann. Entomol. Soc. Am. 55: tigoniidae, Phaneropterinae). Trans. Am. En­ 303-322. tomol. Soc. 114: 311-317. Walker, T.J.1963. The taxonomy and calling songs Strohecker, H.F. 1941. Two new species of Ortho­ of the United States tree crickets (Orthoptera: ptera from Texas. Ann. Entomol. Soc. Am. 34: Gryllidae: Oecanthinae). II. The Nigricornis 539-542. Group of the Genus Oecanthus. Ann. Entomol. Strohecker, H.F. 1952. Descriptions of new spe­ Soc. Am. 56: 772-789. cies and notes on North American Orthoptera. Walker, T.J. 1967. Revision of the Oecanthinae Am. Midland Naturalist 48: 683-688. (Gryllidae: Orthoptera) of America South of the Strohecker, H.F., W.W. Middlekauff, and D.C. United States. Ann. Entomol. Soc. Am. 60: 784­ Rentz. 1968. The grasshoppers of California 796. (Orthoptera: ). Bull. California In­ Walker, T.J. 1969. Systematics and acoustic be­ sect Survey 10: 177pp. havior of United States crickets of the genus Thomas, E.S. and R.D. Alexander. 1962. Sys­ Orocharis (Orthoptera: Gryllidae). Ann. Ento­ tematic and behavioral studies on the meadow mol. Soc. Am. 62: 752-762. grasshoppers of the Orchelimuiri concinnum Walker, T.J. 1969. Systematics and acoustic be­ group (Orthoptera: Tettigoniidae). Occ. Papers havior of United States crickets of the genus Mus. Zool., Univ. Mich. 626: 1-30. Cyrtoxipha (Orthoptera: Gryllidae). Ann. En­ Tinkham, E.R. 1944. Biological, taxonomic and tomol. Soc. Am. 62: 945-952. faunistic studies on the shield-back katydids of Walker, T.J. 1998. Handbook of katydids and the North American deserts. Am. Midland Nat­ crickets of America North of Mexico. http:// uralist 31: 257-328. cassrvr .entnem. ufl.ed u/ -walker/handbook/ Tinkham, E.R. 1948. Faunistic and ecological hdbkhmpg.htm. studies on the Orthoptera of the Big Bend Wallace, H.S. 1955. Revision of the genus Aeol­ Region of Trans-Pecos Texas, with especial oplides (Orthoptera, Acrididae), Ann. Entomol. reference to the orthopteran zones and faunae Soc. Am. 48: 453-480. of Midwestern North America. Am. Midland Yin, X. and R.L. Smith. 1989. Three new grass­ Naturalist 40: 521-663. hoppers from the Western United States (01'­ Walker, T.J.1962. The taxonomy and calling songs thoptera: Acrididae). Pan-Pacific Entomol. 65: ofUnited States tree crickets (Orthoptera: Gryl- 166-171.