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Papilio (Achillides) Dehaanii C Евразиатский энтомол. журнал 20(2): 82 –85 © EUROASIAN ENTOMOLOGICAL doi: 10.15298/euroasentj.20.2.04 JOURNAL, 2021 Papilio (Achillides) dehaanii C. et R. Felder, 1864 (Lepidoptera, Papilionidae) — íîâûé âèä äëÿ ìàòåðèêîâîé ÷àñòè Ðîññèéñêîãî Äàëüíåãî Âîñòîêà New records of Papilio (Achillides) dehaanii C. et R. Felder, 1864 (Lepidoptera, Papilionidae) from the continental part of the Russian Far East Â.Â. Äóáàòîëîâ*, **, Þ.À. ×èñòÿêîâ*** V.V. Dubatolov*, **, Yu.A. Tshistjakov*** * ФГУ «Заповедное Приамурье», ул. Юбилейная 8, Хабаровский край пос. Бычиха 680502 Россия. E-mail: [email protected]. * Federal State Institution «Zapovednoe Priamurye», Yubileinaya Str. 8, Khabarovskii Krai, Bychikha 680502 Russia. ** Институт систематики и экологии животных СО РАН, ул. Фрунзе 11, Новосибирск 630091 Россия. ** Institute of Systematics and Ecology of Animals, Siberian Branch of Russian Academy of Sciences, Frunze Str. 11, Novosibirsk 630091 Russia. *** Федеральный научный центр Биоразнообразия наземной биоты Восточной Азии Дальневосточного отделения Россий- ской академии наук, пр. 100-летия Владивостока 159, Владивосток 690022 Россия. E-mail: [email protected]. *** Federal Scientific Center of Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Prosp. 100- letiya Vladivostoka 159, Vladivostok 690022 Russia. Ключевые слова: парусники, Papilio, Achillides, Приморский край, Российский Дальний Восток, новые находки. Key words: swallow-tails, Papilio, Achillides, Primorskii Krai, Russian Far East, new records. Резюме. Приводятся достоверные находки Papilio лись [Kurentzov, 1970; Beljaev, Dubatolov, 1997]. (Achillides) dehaanii C. et R. Felder, 1984 из Хасанского Х. Ёсимото [Yoshimoto, 1998] высказал предполо- района Приморского края, где вид стал постоянно регис- жение, что под названием Papilio bianor скрывает- трироваться с 2004 года. Ранее этот вид считался подви- ся два близких, но самостоятельных вида. Номина- дом P. (A.) bianor Cr. тивный P. bianor обитает в Центральном и Южном Abstract. Confirmed records of Papilio (Achillides) de- Китае, а также на юге Японии, распространяясь на haanii C. et R. Felder, 1864, formerly considered as subspe- запад до Гималаев (P. b. polyctor Boisduval, 1836, cies of P. (A.) bianor Cramer, 1777, from Khasansky District который иногда трактуется в качестве самостоятель- of continental Russian Far East, regularly recorded since ного вида). На севере ареала, включая материковые 2004, are presented. районы севернее Пекина в Китае, в Корее и на боль- ших Японских островах (а также в России на юге Введение Сахалина и Южных Курилах) встречается самостоя- тельный вид Papilio dehaanii C. et R. Felder, 1864. Во второй половине XX века на территории СССР/ Это мнение впоследствии полностью подтвердилось России было известно два вида подрода Achillides филогенией подрода Achillides, построенной на ана- Hübner, 1819 рода Papilio Linnaeus, 1758: широко лизе ДНК [Condamine et al., 2013]. В упомянутой ра- распространённый в Приамурье, Приморье, на Са- боте, кроме того, было показано, что филогенети- халине и Южных Курилах Papilio maackii Ménétriès, чески близкие друг к другу P. bianor и P. dehaanii в 1859, а также P. bianor Cramer, 1777, который указы- то же время не близкородственны P. maackii, бли- вался во многих работах с юга острова Сахалин и жайшими родственниками которого, по мнению ав- Южных Курильских островов [Esaki, 1922; Matsumura, торов, являются южнокитайско-индокитайский 1928; Kurentzov, 1970; Korshunov, 1972, 2002; P. krishna Moore, [1858], а также южнокитайско-ин- Krivolutskaya, 1973; Korshunov, Gorbunov, 1995; докитайский P. arcturus Westwood, 1842 и тайваньс- Gorbunov, Kosterin, 2001, Lvovsky, Morgun, 2008; кий P. hopponis Matsumura, 1907. Напротив, китайс- Dubatolov et al., 2019]. На материке в качестве север- кий P. syfanius Oberthür, 1886 совершенно не ной границы распространения P. bianor в XX веке отличается по ДНК от P. maackii. Согласно этим отмечалась Северная Корея; на территории же При- исследованиям, группа видов P. bianor, помимо морского края эти бабочки никогда не фиксирова- P. dehaanii, включает ещё южнокитайско-индокитай- Papilio (Achillides) dehaanii — новый вид для Российского Дальнего Востока 83 ский P. dialis Leech, 1893 и филиппинский P. hermeli каталоге насекомых Дальнего Востока России (Nuyda, 1992). Таким образом, на территории Даль- [Streltzov, 2016], а также в Каталоге чешуекрылых него Востока России обитает единственный вид из России [Dubatolov et al., 2019] P. bianor был приве- группы P. bianor — P. dehaanii, встречающийся дён для территории Южного Приморья только под также в наиболее северных провинциях Китая (к се- вопросом. веру от полуострова Шаньдун), в Корее (включая её Тем не менее, за последние 4 года накопилось северо-восточные районы), а также в Японии. немало наблюдений и сведений, причём задокумен- На территории России P. (Achilides) dehaanii C. тированных как на фото, так и непосредственными et R. Felder (=Papilio bianor в понимании прежних находками этого вида на юге Приморского края. Эти авторов) долгое время был известен только с Южно- данные, а также личные сборы и наблюдения авто- го Сахалина и острова Кунашир [Esaki, 1922; ров суммированы ниже и позволяют по-новому оце- Matsumura, 1928; Kurentzov, 1970], позднее найден нить распространение вида на Дальнем Востоке Рос- на Итурупе [Korshunov, 1998], точнее только в юж- сии. ной его половине [Tshikolovets, Streltzov, 2019]. При подробных исследованиях Сахалина в последние де- Papilio (Achillides) dehaanii C. et R. Felder, 1864 сятилетия этот вид был отмечен ещё в центральной Рис. 1–5. части острова (Мгачи, Леонидово, Ельники, река Papilio (Achillides) dehaanii Felder, Felder: 323, 371. Поронай) [Asahi et al., 1999], верховьях реки Алек- Типовое место: «Japonia». сандровка в Макаровском районе [Klitin, 2001], и с Материал. Ðîññèÿ: Ïðèìîðñêèé êðàé, Õàñàíñêèé ð-í, ïîëóîñòðîâ Ãàìîâà, áóõòà Âèòÿçü, 15.06.2004 — 1# острова Монерон [Asahi et al., 1999]. Тем не менее, (Y. Nagahata), 22.08.2006 — 1#, 1$ (Y. Nagahata), 8– наиболее северные находки на Сахалине связаны со 12.08.2017 — 2##, 1$ (×èñòÿêîâ), 20.08.2017 — 1# случайными залётами бабочек, так как кормовое (Y.Okatsu, Y. Nagahata), 12.08–1.09. 2020 — 3## (×èñòÿ- êîâ); áóõòà Âèòÿçü, êîòòåäæ Âîñòî÷íûé, 42°35'56" ñ.ø., растение его гусениц, бархат сахалинский, в цент- 131°11'13" â.ä., 6.09.2020 — 1$, 9.09.2020 — 1# (Äóáàòî- ральной части Сахалина не растёт. Таким образом, ëîâ); ïîñ. Âèòÿçü, ñåâåðíûé áåðåã áóõòû, 42°35'25" ñ.ø., северная граница зоны обитания (а не только встреч!) 131°10'30" â.ä., 9.09.2020 — 2## (Äóáàòîëîâ, âèçóàëüíî); вида на Сахалине отмечалась по перешейку Поясок Õàñàíñêèé ðàéîí, ïîñ. Áåçâåðõîâî, 16.08.2020 — 1$ (Ë. Ïåò- ðîâà-Åëàíñêàÿ); Ïðèìîðñêèé êðàé, Ëàçîâñêèé ãîñóäàðñòâåí- [Asahi et al., 1999; Klitin, 2001]. В пределах материко- íûé çàïîâåäíèê, áóõòà Ïåòðîâà, 28–29.08.2017 — 1$ (Â. Ìåä- вой части ареала А.И. Куренцов [Kurentzov, 1970: âåäåâ), 7.09.2017 — 1$ (Â. Ìåäâåäåâ). fig. 6, p. 9] экстраполировал северную границу рас- Распространение в Приморском крае России. В по- пространения этого вида на Корейском полуостро- сёлке Витязь P. dehaanii впервые достоверно зарегистри- ве южнее 41-й параллели и не рассматривал возмож- рован в июне 2004 года, а также два года спустя в августе ность его обитания на территории Южного 2006 г. Затем, в течение последующих 10 лет эти бабочки не попадались. Но в последние годы они здесь отмечают- Приморья, хотя для двух других видов, обитающих в ся почти ежегодно, регулярно появляясь в начале авгус- Корее и Северо-Восточном Китае — P. (Menelaides) та. Отдельные экземпляры этого вида также были отмече- protenor demetrius Cramer, [1782]. и P. (Menelaides) ны Л. Петровой-Еланской несколько севернее в macilentus Janson, 1877 он допускал вероятность их Безверхово [https://yadi.sk/i/GQCUrmMFphkCVw], в нахождения в Южном Приморье в будущем, хотя бы Хасанском районе. Единственной достоверной наиболее в качестве залётных. Тем не менее, эти два вида на северной точкой, где был найден A. dehaanii, является юге Приморского края до сих пор так и не найдены, бухта Петрова в Лазовском государственном заповедни- хотя они и указывались для Северной Кореи (КНДР) ке [https://m.facebook.com/story.php?story_fbid= [Chu, Im, 1987]. На территории Северной Кореи 1088681854597477&id=100003670188830], где этот вид P. dehaanii также обитает и, судя по изображению был сфотографирован В. Медведевым (рис. 5). После- дняя находка может служить косвенным подтверждением бабочек [Chu, Im, 1987], северокорейские «P. bianor» достоверности и его нахождения на острове Попова следует относить именно к P. dehaanii. С материко- [Martynenko, Chichvarkhin, 1997]. вой части Дальнего Востока России какие-либо дос- Замечания по систематике. А. Коттон [Cotton, 2017] товерные материалы по P. dehaanii до недавнего признаёт существование единственного, номинативного времени отсутствовали [Dubatolov et al., 2019], за подвида A. dehaanii, распространённого в континенталь- исключением указания о находке P. bianor на остро- ной части ареала, а также на больших островах Японии. ве Попова по единственному самцу, собранному Отдельные подвиды, по его мнению, встречаются только 31 августа 1995 года [Martynenko, Chichvarkhin, 1997]. на изолированных островах на юге Японии (архипелаг Это указание ошибочно было изменено на остров Рюкю) и на острове Улунгдо в Корее. Значительную из- Рейнеке и иную дату поимки — 30 августа 1995 года менчивость в размерах он не выделяет из сезонной измен- чивости. Тем не менее, на островах Сахалинской области
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