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New Poecilanthrax, with Notes on Described Species (Diptera: Bombylidae) D Great Basin Naturalist Volume 17 Article 1 Number 1 – Number 2 7-31-1957 New Poecilanthrax, with notes on described species (Diptera: Bombylidae) D. Elmer Johnson University of Nevada, Reno Lucile Maughan Johnson Follow this and additional works at: https://scholarsarchive.byu.edu/gbn Recommended Citation Johnson, D. Elmer and Johnson, Lucile Maughan (1957) "New Poecilanthrax, with notes on described species (Diptera: Bombylidae)," Great Basin Naturalist: Vol. 17 : No. 1 , Article 1. Available at: https://scholarsarchive.byu.edu/gbn/vol17/iss1/1 This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. LIBRARY SEP 10 1957 HARVARD The Great Basin Naturalist UNIVERSITY purlished by the Department of Zoology and Entomology Brigham Young University, Provo, Utah Volume XVII JULY 31, 1957 Nos. 1-2 NEW POECILANTHRAX, W^ITH NOTES ON DESCRIBED SPECIES (DIPTERA: BOMBYLIDAE) D. Elmer' and Lucile Maughan Johnson In his Notes on Osten Sacken's Group Poecilanthrax. Cole'-' re- viewed most of the previous work on this genus and described several new species and varieties. A number of these forms were figured, and some of Osten Sacken's illustrations were also included. He did not, however, construct a key. Apparently having at hand representatives of all the forms described from North America, as well as a number of forms we consider to be undescribed, we present below descrip- tions of most of the latter, notes on some of the species heretofore de- scribed, as well as an analytical table which we hope will be usefid to others in the identification of species. Our study of the genus has indicated the urgent need for much more collecting of long series from all parts of the country. The pres- ent study is, of necessity, limited to our own collection, to those speci- mens from other miscellaneous collections that have passed through our hands during the past several years, and to a few specimens loaned to us by the United States National Museum. In all we have examined between seven and eight hundred specimens, mostly from the Middle and Far Western United States, but including a few from Mexico and Canada. The minor differences between forms, though consistent, are often subtle and difficult to describe, and in some instances require close familiarity with the gi'oup to be apparent. In a few instances the intra-specific variation is greater than the usual inter-specific variation. Of the characters we have used for the differentiation of species one of the most useful has been the venation of the medial area of the wing, particularly the courses of the medial cross-vein and the second medial vein, with the resultant effect on the shape of the discal cell. The presence or absence of an appendiculate spur on the medial cross-vein, though not entirely constant within a species, has. never- theless, been very useful. The color pattern of the wing has. with- in fairly narrow limits, been constant in most species. Ground 1. Assistant Entomologist, Ecological Research, University of Utah, Dugway, Utah. 2. Cole, F. R., Notes on Osten Sacken's Group Poecilanthrax. with descriptions of new species. Jour. New York Ent. Soc. Vol. XXV. pp. 1.7. Mil. ", ],l,,u-.. M.inh. I'il7 The Great Basin Naturalist 2 JOHNSON AND JOHNSON Vol. XVII. NoS. 1-2 color of legs, antennae, and abdomen has also been useful; however, in some instances, as in the abdomen of alcyon and willistoni the intra-specific variation has been so great that this character has been rendered almost useless in these species. The color, nature and distri- bution of the vestiture has been useful; in some instances it has been the most convenient means of differentiation between forms, but its fragility limits its usefulness and wherever possible we have used other more durable characters. Because of the extreme similarity of some forms, certainly there will be flies that cannot be placed by means of our key. even by one familiar with the group. Vhe accompanying text figures have been prepared with as much care as possible and frequent reference is made to them in an attempt to form a basis for comparison, and to clarify the key as much as possible. All drawings were made with the use of a camera lucida. We wish here to express our gratitude to the following men and institutions for the loan of material: Illinois State Natural History Survey, United States National Museum. Colorado Agricultural and Mechanical College, Utah State University, Brigham Young Univer- sity, the Universities of Arizona, Colorado, Idaho, Minnesota and Utah, Dr. D. Geo. Butler, Jr., and Mr. Joe Schuh. We wish also to express gratitude to Drs. Willis W. Wirth and Paul H. Arnaud, United States Department of Agriculture, for their efforts in our be- half in making comparisons with types in the U. S. National Mu- seum. Except where otherwise noted types of the species herein de- scribed are in our collection, presently located at the Brigham Young University, Provo. Utah. Table of Species 1. Wings distinctly mottled hyaline or subhyaline and brown or black, at least in area of discal cell; veins in posterodistal half usually more or less distinctly bordered with brown. (Figs. 3, 4, 7-12, 14-22) 9 Anteroproximal half of wings brown, posterodistal half hya- line or subhyaline; wings evenly brown; or wings with color gradually evanescent from anteroproximal to postercclista!, with at most faint borders of darker along veins, but without distinct mottling in the discal cell area. (Figs. 1, 2, 5, 6, 13) 2 2. Anteroproximal half of wing almost evenly brown, posterodis- tal half evenly subhyaline. (Fig. 2) 3 Wings not as above 4 3. No black hairs in lateral fringe of abdominal segments anterior to sixth; very few if any black hairs and very little or no black tomentum on hind margins of tergites sackenii sackenii (Coq.) Black hairs in lateral fringe usually of most abdominal seg- ments anterior to sixth; distinct fasciae of black pile and tomentum on hind margins of most or all tergites after the first (Fig. 2) sackenii monticola, n. sub-sp. 4. Wings evenly brown, only slightly paler behind (Fig 6) 5 Wings definitely darker along anteroproximal portion 6 July 31. 1957 the bee-fly genus peocilanthrax 3 5. Abdomen with distinct fasciae of black pile and tomentum along hind margins of segments caudad of the first; black pile in lateral fringe of at least two or three abdominal segments, relatively slender species lucifer (Fab.) Abdomen without fasciae of black pile and tomentum, relatively more robust species (Fig. 6) tegminipenn\s (Say) 6. Medial cross-vein not angled near distal third or fourth, regu- larly without short spur extending into cell 2M:, color of wings and general appearance blackish gray; legs mostly black; first joint of antennae, only, yellow (Fig. 1) painteri Maughan Medial cross-vein angled near distal third or fourth, usually with a spur extending into 2M, or at least a thickening of the vein at this point ; wing color and general appearance usually brownish 7 7. Abdomen without red on sides; second and third antennal seg- ments black; tarsi and most of tibiae black flaviceps (Lw.) Sides of at least second abdominal segment with spots of red; tibiae mostly red 8 8. To the naked eye discal cell appears to be paler than rest of posterodistal half of wing; third antennal segment very long-conical, the bulbous base not well defined; lateral fringe markedly tufted with black pile, a band of black pile and to- mentum across hind margins of all abdominal segments after the first (Fig. 5) fuliginosa (Lw.) To the naked eye the discal cell concolorous with rest of postero- distal portion of wing; hind angles of abdominal segments well defined; black pile and tomentum usually confined to hind margins of second and third abdominal segments or less (Fig. 13.) pilosa (Coq.) 9. Regularly cell R, divided into two by a supernumerary cross- vein connecting veins R, and Rr, (Fig. 18) 10 Regularly without cell Rj divided by a supernumerary cross- vein 11 10. Sides of abdomen narrowly red; wings blackish brown alpha alpha (O.S.) Sides of abdomen broadly red; wings more reddish brown (Fig. 18) alpha zionensis, n. sub-sp. 11. Wings normally with supernumerary cross-vein in cell 2M2 con- necting vein M, + Cu, to the medial cross-vein (Figs. 4, 16, 20) 12 Wings normally without supernumerary cross-vein and extra cell 14 12. No free stump of vein projecting into cell 2M,; sides of abdomen broadly red; Mid-western and Western species (Figs. 4, 23c). arethitsa (O.S.) Free stump of vein projecting into cell 2M;; abdomen usually more narrowly red at sides 13 13. Free stump of vein which projects into cell 2M2 is attached to medial cross-vein between attachment of supernumerary cross-vein and vein M, (Figs. 16, 23a) alcyon (Say) Free stump of vein projecting into cell 2M, is attached to super- numerary cross-vein (Figs. 20, 23b) nigripennis (Cole) 14. Dark of wings grayish black: at least fore femora dark basally; pile of middle of face black; at most, spots of red on sides of second and third segments (Fig. 21) californicus (Cole) Dark of wings reddish brown 15 1.5. Abdomen broadly red at sides 16 At most, sides of abdomen narrowly red, usually all black or at most small spots on sides of second to fourth segments 22 The Great Basin Naturalist 4 JOHNSON AND JOHNSON Vol.
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