Nomination of Dr. Eric Hoberg to Membership on Science Steering Committee Study of Environmental Arctic Change (SEARCH)

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25 February 2015

Re: Nomination of Dr. Eric Hoberg to Membership on Science Steering Committee Study of Environmental Arctic Change (SEARCH)

Dear Review Committee Members,

This letter is written in support of Dr. Eric Hoberg's appoint to the SEARCH Science Steering Committee. I have observed Dr. Hoberg in laboratory, field, scientific meetings and policy conferences over the past 25 years and I know he would make insightful and carefully considered contributions to the SEARCH agenda. In the early 1990’s, we were collaborators on a joint Russian/US research consortium (PanArctic Biota) initiated between the Russian Academy of Sciences and Smithsonian Institution. We first met in St Petersburg, Russia and subsequently put together an international (Finland, Russia, Canada, US, Sweden, Iceland) set of collaborations called the Beringian Coevolution Project that has produced more than 200 publications related to impact of arctic climate change on various aspects of biodiversity (e.g., emerging pathogens, conservation biology, community turnover). In addition to funding from a variety of natural resource agency sources (USDA, USGS, NPS), we have been co-funded through NSF (3 consecutive grants from 1999 to the present) to work on mammal and emerging pathogen questions related to the nexus between Asia and North America, Beringia. It is on the basis of this association that I make the following recommendation.

Dr. Hoberg is an extremely motivated and focused individual who has demonstrated considerable productivity throughout his career. He would bring substantial expertise and new perspectives on the effects of climate change on emerging pathogens, zoonotic disease, and subsistence food security of northern peoples, as well as a deeper time perspective on our dynamic northern environments. Significantly, he understands the critical role that natural history collections and associated big data play as scientific infrastructure that can stimulate interdisciplinary approaches to questions of environmental change across time and space.

Dr. Eric Hoberg is now Chief Curator and Zoologist at the US National Parasite Collection, among the largest archives of specimens and information documenting global parasite biodiversity. His undergraduate work was done at the University of Alaska Fairbanks, where he studied biology, ornithology and parasitology. Evolution of complex parasite faunas among seabirds became the focus for an MSc (Univ. Saskatchewan 1979), and then PhD (Univ. Washington-1984) with Dr. Robert Rausch. As a field biologist and biogeographer, he has traversed regions of Alaska, Canada, Siberia, Scandinavia and Antarctica. He has authored or coauthored over 250 publications, emphasizing the role of episodic events, climate, environmental perturbation and patterns of geographic colonization as determinants of diversity and emergent diseases in evolutionary and ecological time.

As reflected by the number of invited presentations he has already delivered and papers published, Dr. Hoberg’s research in parasitology, biogeography, and conservation biology is highly regarded by his peers. His contributions are numerous, have high impact and are published in high impact journals like the Global Change Biology, Proceedings Royal Society of London, Trends in Parasitology, Journal of Wildlife Diseases and others. Dr. Hoberg interacts well within the academic, agency and policy spheres. He has mentored three of my graduate students directly (one for over a year in the Beltsville USDA Lab) and has been heavily involved in other graduate student committees elsewhere. He provides strong and solid guidance, demonstrating the level of Dr. Hoberg's energy and commitment to climate change research at high latitudes.

Dr. Hoberg was a founding member of the Research Group for Arctic Parasitology, which drove the expansion of studies of parasite diversity in the north, becoming a powerful consortium linking the US National Parasite Collection, University of Calgary, University of Saskatchewan and research groups in Alaska, NWT, Yukon and Nunavut, Finland and Russia. RGAP complemented the Beringian Coevolution Project in working to develop geographically extensive baselines (from field sampling through agencies and Native communities) to understand environmental change and perturbation in northern systems and the consequences of emerging infectious diseases. Both RGAP and BCP were instrumental in the development of concepts for diversity addressed in the Arctic Biodiversity Assessment published last year.

In summary, Dr. Hoberg is a clear, original thinker, good listener, and consensus builder. He has developed a solid scientific reputation among our national and international colleagues. He has a quick wit and his sense of humor is a welcome addition to group interactions. He is a trustworthy and compassionate human being who is respected for his good taste, judgment and sensitivity to cultural issues. You will find him to be a pleasant, positive force that will be fully committed to SEACRH efforts. This letter is meant to be a strong recommendation for Dr. Eric Hoberg’s nomination to the Science Steering Committee for SEARCH.

Please do not hesitate to call me if you have further questions.

Sincerely,

Joseph A. Cook Professor of Biology Director, Museum of Southwestern Biology, University of New Mexico

United States National Parasite Collection

Agricultural Research Service, USDA

Current as of February 2015

1. NAME: Eric P. Hoberg

2. PERSONAL DATA: Born: 18 October 1953

3. ADDRESSES: Business Address: US National Parasite Collection Animal Parasitic Diseases Laboratory Beltsville Agricultural Research Center Agricultural Research Service United States Department of Agriculture BARC East, Bldg, 1180, 10300 Baltimore Avenue Beltsville, Maryland 20705-2350 USA Telephone: (301) 504-8588/ Fax: (301) 504-8979 [email protected]

Home Address: 16321 Oxford Court Bowie, Maryland 20715

4. ACADEMIC QUALIFICATIONS:

Degrees University Date Area

Ph.D. U. of Washington 1984 Systematics, biogeography, evolution of parasites M.Sc.. U. of Saskatchewan 1979 Parasitology B.Sc. U. of Alaska 1975 Biological Sciences

5. EMPLOYMENT HISTORY: Nov. 2000 - Present Research Zoologist, GS-15. Chief Curator, US National Parasite Collection. Unit Leader- Biosystematic Parasitology, Animal Parasitic Diseases Laboratory, USDA, ARS.

Sept. 1999 - Nov. 2000 Supervisory Zoologist, GS-15. Chief Curator, US National Parasite Collection. Research Leader, Biosystematics & National Parasite Collection Unit, USDA, ARS.

Sept. 1990 - Aug. 1999 Research Zoologist, GS-13/15. Assoc. Curator, US National Parasite Collection. Biosystematics & National Parasite Collection Unit, USDA, ARS.

Concurrent Appointment, Nov 2013 – Nov 2018 Adjunct Scientist, Research Associate, National Museum of Natural History, Smithsonian Institution

Concurrent Appointment, Nov 2013 – present Adjunct Professor, Department of Biology, Northern Michigan University, Marquette

Concurrent Appointment, June 2009- 2015 Adjunct Professor, Faculty of Veterinary Medicine, University of Calgary.

Concurrent Appointment, April 2004- present Adjunct Professor, Department of Biology, University of New Mexico, Albuquerque.

Concurrent Appointment, June 1995 - present Adjunct Professor, Department of Veterinary Microbiology, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon.

Concurrent Appointment, June 2000- 2007 Affiliate Associate Professor, Department of Biology and Wildlife, University of Alaska, Fairbanks.

Concurrent Appointment Sept. 2000- August 2003 Affiliate Faculty, Department of Biology, Idaho State University, Pocatello.

Jan. 1998 - June 1998 Administrative Detail to Office of Deputy Administrator, National Program Staff, Agricultural Research Service.

Jul. 1989 - Sept. 1990 Assistant Professor. Veterinary Parasitology. Atlantic Veterinary College, University of Prince Edward Island.

Jan. 1985 - June 1989 Research Associate. Veterinary Parasitology. College of Veterinary Medicine, Oregon State University, Corvallis.

Jul. 1984 - Nov. 1984 Field Biologist. College of Forest Resources, University of Washington, Seattle.

Jul. 1984 - June 1985 Post-Doctoral Senior Fellow. Department of Pathobiology, University of Washington, Seattle.

Jan. 1981 - Mar. 1981 Teaching Assistant. Parasitology. Department of Zoology, University of Washington, Seattle.

Oct. 1979 - Jun. 1984 Research Assistant. Department of Pathobiology, University of Washington, Seattle.

2 Sept. 1978 - Dec. 1978 Instructor, Parasitology. Department of Veterinary Microbiology and Department of Biology, University of Saskatchewan, Saskatoon, Canada.

Sept. 1977 - Dec. 1977 Teaching Assistant. Department of Veterinary Microbiology, University of Saskatchewan, Saskatoon, Canada.

May 1977 - Aug. 1977 Biological Technician. United States Fish and Wildlife Service, Anchorage, Alaska.

Jan. 1976 - Aug. 1976 Biological Technician. United States Fish and Wildlife Service, Anchorage, Alaska.

Apr. 1975 - Oct. 1975 Biological Technician. United States Fish and Wildlife Service, Aleutian Islands, NWR, Alaska.

Jan. 1974 - Dec. 1974 Histological Technician. University of Alaska, Fairbanks.

Jun. 1973 - Aug. 1973 Fisheries Technician. Alaska Department of Fish and Game, Bethel, Alaska.

6. TEACHING RESPONSIBILITIES: Western College of Veterinary Medicine, University of Saskatchewan: 1977 Helminthology (laboratory) 1978 Helminthology

University of Washington: 1981 General Parasitology (laboratory)

College of Veterinary Medicine, Oregon State University: 1986 VM 505 Parasitological Techniques 1986-88 VM 461 Parasitic Diseases of Domestic and Wild Animals

Atlantic Veterinary College, University of Prince Edward Island: 1989 VPM 450 Clinics in Diagnostic Services (Parasitology) 1990 VPM 422 Wildlife Disease (Helminthology) 1990 VPM 212 Parasitology (Helminths, Arthropods) 1990 VPM 450 Clinics in Diagnostic Services (Parasitology)

Western College of Veterinary Medicine, University of Saskatchewan: 1996 VTMC 839.3 Topics in Advanced Parasitology (invited lectures) 1996 BIOL 498.3 Parasitism (invited lectures)

WCVM, & Research Group for Arctic Parasitology (RGAP): 2000 Wildlife Parasitology & Disease, Short Course 17-20 April 2000; Ft. Smith, Northwest Territories

3 7. GRADUATE COMMITTEES/ STUDENTS:

Susan J. Kutz. Department of Veterinary Microbiology, Western College of Veterinary Medicine, University of Saskatchewan. PhD. (1995-1999). (CoAdvisor) [Thesis: The biology of Umingmakstrongylus pallikuukensis, a lung nematode of muskoxen in the Canadian Arctic: Field and Laboratory Studies].

Monica Santín-Durán. Universidad Complutense, Madrid. Facultad Patologia Animale I. PhD. (Co-Advisor) (1998-2002). [Thesis: Helmintofauna de cérvidos en la zona centro de España: Epidemiología e interés sanitario.].

Kurt E. Galbreath. (1) Department of Biology & Wildlife, and University of Alaska Museum, University of Alaska. MSc. (2000-2002). [Thesis: Genetic consequences of ice ages for a Holarctic rodent: Phylogeography and post-glacial colonization of the tundra vole, Microtus oeconomus, in Beringia]. (2) Department of Ecology & Evolutionary Biology, Cornell University. PhD (2003- 2008). [Thesis: Of Pikas and Parasites: Historical Biogeography of an Alpine Host-Parasite Assemblage]

Emily Jenkins. Department of Veterinary Microbiology, Western College of Veterinary Medicine, University of Saskatchewan. PhD. (Co-Advisor) (1999-2005). [Thesis: Ecological investigation of a new host-parasite relationship: Parelaphostrongylus odocoilei in thinhorn sheep (Ovis dalli)]. [Arctic Research Consortium of the US- ARCUS Award for Research Excellence 2003; WDA Outstanding Graduate Student Award 2004; WDA Outstanding Graduate Research Paper, Annual Meeting, Perth Australia, 2005]

Anson Koehler, Department of Biological Sciences, University of New Mexico. MSc. (2003- 2006). [Thesis: Systematics, phylogeography, distribution and lifecycle of Soboliphyme baturini].

Nathan deBruyn, Department of Ecosystem and Public Health, Veterinary Medicine, University of Calgary. MSc. (2007- 2010). [Thesis: Gastrointestinal nematodes of western Canadian Cervids: Molecular diagnostics, faunal baselines and management considerations].

Guilherme G. Verocai, Department of Ecosystem and Public Health, Veterinary Medicine, University of Calgary. PhD. (2009- present).

Kayce Bell, Department of Biology, University of New Mexico. PhD. (2010- present).

Heather Toman., Department of Biology, Northern Michigan University. MSc. (2014- present)

External Examiner, PhD:

Gregory John Klassen. Thesis: Phylogeny and biogeography of ostraciin boxfishes (Tetraodontiformes: Ostraciidae) and their gill parasitic Haliotrema species (Monogenea: Ancyrocephalidae): A study in host-parasite coevolution. University of Toronto. 21 January 1992.

4 Lotta M. Wickström. Thesis: Phylogeny, phyletic coevolution and phylogeography of Anoplocephaline cestodes in mammals. Metsäntutkimuslaitoksen Tiedonantoja 918, 2004. The Finnish Forest Research Institute, Research Papers. Vantaa Research Centre. University of Helsinki. 27 May 2004. (“Opponent” invited Faculty of Biosciences).

8. PUBLICATIONS:

(a) Papers in Refereed Journals

1. Hoberg EP. A redescription of Aporchis continuus McCauley and Pratt 1960 (Trematoda: Echinostomatidae) with new host and geographic records. Canadian Journal of Zoology 1981; 59:19-21.

2. Hoberg EP. Pseudogymnophallus alcae gen. n. et sp. n. (Trematoda: Gymnophallidae) from alcids (Charadriiformes) in subarctic seas. Proceedings of Helminthological Society of Washington 1981; 48:190-194.

3. Hoberg EP, McGee SG. Helminth parasitism in raccoons, Procyon lotor hirtus Nelson and Goldman, in Saskatchewan. Canadian Journal of Zoology 1982; 60:53-57.

4. Hoberg EP, Wehle DHS. Host and geographic records of ectoparasites of Alaskan seabirds (Charadriiformes: Alcidae and Laridae). Canadian Journal of Zoology 1982; 60:472-475.

5. Hoberg EP. Diorchis pelagicus sp. nov. (Cestoda: Hymenolepididae) from the whiskered auklet, Aethia pygmaea, and the crested auklet, A. cristatella, in the western Aleutian Islands, Alaska. Canadian Journal of Zoology 1982; 60:2198-2202.

6. Rausch RL, Maser C, Hoberg EP. Gastrointestinal helminths of the cougar, Felis concolor L., in northeastern Oregon. Journal of Wildlife Diseases 1983; 19:14-19.

7. Hoberg EP. Preliminary comments on parasitological collections from seabirds at Palmer, Antarctica. Antarctic Journal of the United States 1983; 18:206-208.

8. Hoberg EP. Trematode parasites of birds in Antarctica: the distribution of Gymnophallus deliciosus (Olsson, 1893). Antarctic Journal of the United States 1984; 19:159-160.

9. Hoberg EP. Alcataenia pygmaeus sp. n. (Cestoda: Dilepididae) from the whiskered auklet, Aethia pygmaea (Gmelin) in the western Aleutian Islands, Alaska. With a comment on the genera Alcataenia Spasskaia, 1971 and Rissotaenia Spasskaia and Kolitolova, 1972. Proceedings of the Helminthological Society of Washington 1984; 51:315-319.

10. Hoberg EP. Alcataenia fraterculae sp. n. from the horned puffin, Fratercula corniculata (Naumann), Alcataenia cerorhincae sp. n. from the rhinoceros auklet, Cerorhinca monocerata (Pallas), and Alcataenia larina pacifica ssp. n. (Cestoda: Dilepididae) in the North Pacific basin. Annales de Parasitologie Humaine et Comparee 1984; 59:335-351.

11. Hoberg EP. Alcataenia longicervica sp. n. from murres, Uria lomvia (Linnaeus) and Uria aalge (Pontoppidan) in the North Pacific basin, with redescriptions of Alcataenia armillaris (Rudolphi, 1810) and A. meinertzhageni (Baer, 1956) (Cestoda: Dilepididae). Canadian Journal of Zoology 1984; 62:2044-2052.

12. Hoberg EP. Alcataenia campylacantha (Krabbe, 1869) from pigeon guillemots, Cepphus columba Pallas, and black guillemots, Cepphus grylle (Linnaeus), and Alcataenia sp. indet. from Kittlitz's murrelets, Brachyramphus brevirostris (Vigors) in Alaska. Canadian Journal of 5 Zoology 1984; 62:2297-2301. 13. Hoberg EP. Nearshore food-webs and the distribution of acanthocephalan parasites in Antarctic seabirds. Antarctic Journal of the United States 1985; 19:161-162.

14. Hoberg EP. Reticulotaenia gen. n. for Lateriporus australis Jones and Williams, 1967 and L. mawsoni Prudhoe, 1969 (Cestoda: Dilepididae) from sheathbills, Chionis spp. in Antarctica, with a consideration of infraspecific variation and speciation. Journal of Parasitology 1985; 71:319- 326.

15. Hoberg EP, Zimmerman GL, Lichtenfels JR. First report of Nematodirus battus from North America: Redescription and comparison to other species. Proceedings of the Helminthological Society of Washington 1986; 53:80-88.

16. Hoberg EP. Aspects of ecology and biogeography of Acanthocephala in Antarctic seabirds. Annales de Parasitologie Humaine et Comparee 1986; 61:199-214.

17. Hoberg EP. Evolution and historical biogeography of a parasite-host assemblage: Alcataenia spp. (Cyclophyllidea:Dilepididae) and Alcidae (Charadriiformes). Canadian Journal of Zoology 1986; 64:2576-2589.

18. Hoberg EP. Eulimdana rauschorum sp. n. a filarioid nematode (Lemdaninae) from Larus dominicanus in Antarctica with comments on evolution and biogeography. Journal of Parasitology 1986; 72:755-761.

19. Zimmerman GL, Wallace DH, Schons DJ, Hoberg EP. Efficacy of Clorsulon against mature naturally acquired Fasciola hepatica infections in cattle and sheep. American Journal of Veterinary Research 1986; 47-1665-1667.

20. Hoberg EP, Zimmerman GL. Detection of ensheathed third-stage larvae of Haemonchus contortus (Trichostrongylidae) in sheep: Delayed exsheathement. Proceedings of the Helminthological Society of Washington 1987; 54:150-153.

21. Rickard LG, Hoberg EP, Zimmerman GL, Erno JK. Late fall transmission of Nematodirus battus (Trichostrongyloidea) in western Oregon. Journal of Parasitology 1987; 73:244-247.

22. Hoberg EP. Recognition of larvae of the Tetrabothriidae: Implications for the origin of tapeworms in marine homeotherms. Canadian Journal of Zoology. 1987; 65:997-1000.

23. Hoberg EP. Reighardia sternae (Diesing, 1864) (Pentastomida) from seabirds in Antarctica. Canadian Journal of Zoology. 1987; 65:1289-1291.

24. Richards LS, Zimmerman GL, Hoberg EP, Schons DJ, Dawley SW. The anthelmintic efficacy of netobimin against naturally acquired gastrointestinal nematodes in sheep. Veterinary Parasitology 1987; 26:87-94.

25. Bondarenko SK, Kondratieva L, Hoberg EP. Parazit chaek Hymenolepis (s.l.) haldemani Schiller, 1951: ego zhiznennyii tsikl. [Gulls' parasite Hymenolepis (s.l.) haldemani Schiller, 1951 and its life cycle.] Acta Parasitologica Lithuanica 1987; 22:82-91.

26. Hoberg EP. Tetrabothrius shinni sp.n. (Eucestoda), from Phalacrocorax atriceps bransfieldensis (Pelecaniformes) in Antarctica, with comments on morphological variation and host-parasite biogeography and evolution. Canadian Journal of Zoology 1987; 65:2969-2975.

27. Zimmerman GL, Hoberg, EP. Controlled release devices for the delivery of anthelmintics in cattle. Parasitology Today 1988; 4:55-56. 6

28. Hoberg EP, Rickard LG. Morphology of the synlophe of Nematodirus maculosus (Trichostrongyloidea) with comments on the evolution of Nematodirus spp. among the Caprinae (Artiodactyla). Proceedings of the Helminthological Society of Washington. 1988; 55:160- 164.

29. Hoberg EP, Rickard LG, Zimmerman GL. Synlophe of Nematodirus neotoma (Trichostrongyloidea). Journal of Parasitology. 1988; 74:686-691.

30. Hoberg, EP, Zimmerman, Rickard, LG, Schons DJ. Anthelmintic efficacy of febantel against natural gastrointestinal nematodes in calves, and recognition of Oesophagostomum venulosum in Oregon Cattle. American Journal of Veterinary Research. 1988; 49:1540-1542.

31. Zimmerman GL, Hoberg EP, Rickard LG, Bishop JK. Anthelmintic efficacy of ivermectin, levamisole, and fenbendazole against Nematodirus battus in sheep. American Journal of Veterinary Research. 1988; 49:2094-2095.

32. Poinar GO, Hoberg EP. Mermis nigrescens (Mermithidae) recovered from the mouth of a child. American Journal of Tropical Medicine and Hygiene. 1988. 39: 478-479.

33. Hoberg EP, Ryan PG. Ecology of helminth parasitism in Puffinus gravis (Procellariiformes) on the breeding grounds at Gough Island. Canadian Journal of Zoology. 1989; 67:220-225.

34. Rickard LG, Zimmerman GL, Hoberg EP, Lockwood PW, Weber DW, Miller RJ. Efficacy of morantel sustained release trilaminate matrix against gastrointestinal nematodes in beef calves. Veterinary Parasitology. 1989; 33:125-133.

35. Hoberg EP, Fruetel M, Rickard LG. Synlophe of Nematodirus odocoilei (Trichostrongyloidea) from deer and caribou in North America with comments on the evolution of Nematodirus spp. among the Cervidae. Canadian Journal of Zoology. 1989; 67:1489-1494.

36. Johnson MR, Hoberg EP. Differentiation of Moniezia expansa and M. benedeni (Eucestoda: Cyclophyllidea) by isoelectric focusing. Canadian Journal of Zoology. 1989; 67:1471-1475.

37. Hoberg EP, Miller GS, Wallner-Pendleton E, Hedstrom OR. Helminth parasites of northern spotted owls, Strix occidentalis caurina, from Oregon. Journal of Wildlife Diseases. 1989; 25:246-251.

38. Hoberg EP. Phylogenetic relationships among genera of the Tetrabothriidae (Eucestoda). Journal of Parasitology. 1989; 75:617-626.

39. Rickard LG, Hoberg EP, Bishop JK, Zimmerman GL. Epizootiology of Nematodirus battus (Trichostrongyloidea) in western Oregon. Proceedings of the Helminthological Society of Washington. 1989; 56:104-115.

40. Rickard LG, Hoberg EP. Evaluation of the synlophe of Nematodirus urichi and N. lamae (Trichostrongyloidea) from Mazama americana and camelids (Artiodactyla) in the neotropics. Canadian Journal of Zoology. 1990; 68:1005-1011.

41. Hoberg EP. Trigonocotyle sexitesticulae sp. n. (Tetrabothriidae) from pygmy killer whales (Feresa attenuata). Canadian Journal of Zoology. 1990; 68:1835-1838.

42. Hoberg EP, Aubry KB, Brittell, J.D. Helminth parasitism in pine martens (Martes americana) and 7 ermines (Mustela erminea) from the Pacific Northwest, with comments on the distribution of Trichinella spiralis. Journal of Wildlife Diseases. 1990; 26:447-452.

43. Pence DB, Hoberg EP. Thalassornectes (Alcidectes) aukletae subgen. et sp. n. (Acari: Hypoderatidae) from the crested and parakeet auklets (Aves: Charadriiformes; Alcidae). Journal of Medical Entomology. 1991; 28:205-209.

44. Brooks DR, Hoberg EP, Weekes PJ. Preliminary phylogenetic systematic analysis of the major lineages of the Eucestoda Southwell, 1930 (Platyhelminthes: Cercomeria). Proceedings of the Biological Society of Washington. 1991; 104: 651-668.

45. Hoberg EP. Alcataenia atlantiensis n. sp. (Dilepididae) from the razorbill (Alca torda Linnaeus) in the eastern North Atlantic basin. Systematic Parasitology. 1991; 20: 83-88.

46. Hoberg EP, Adams AM, Rausch RL. Revision of the genus Anophryocephalus Baylis, 1922 from pinnipeds in the Holarctic with descriptions of Anophryocephalus nunivakensis sp. n. and A. eumetopii sp. n. (Tetrabothriidae) and evaluation of host records from the Phocidae. Canadian Journal of Zoology. 1991; 69: 1653-1668.

47. Rickard LG, Zimmerman GL, Hoberg EP, Wallace DH. Effect of ivermectin in a sustained release bolus on the productivity of beef cattle in Oregon. Veterinary Parasitology. 1991; 39: 267-277.

48. Hoberg EP, Lichtenfels JR. Morphology of the synlophe and genital cone of Parostertagia heterospiculum (Trichostrongylidae), with comments on the subfamilial placement of the genus. Systematic Parasitology. 1992; 22: 1-16.

49. Hoberg EP, Adams AM. Phylogeny, historical biogeography and ecology of Anophryocephalus spp. (Tetrabothriidae) among pinnipeds of the Holarctic during the late Tertiary and Pleistocene. Canadian Journal of Zoology. 1992; 70: 703-719.

50. Zimmerman GL, Hoberg EP, Pankavich JA. Efficacy of orally administered moxidectin against naturally acquired gastrointestinal nematodes in cattle. American Journal of Veterinary Research. 1992; 53: 1409-1410.

51. Hoberg EP. Congruent and synchronic patterns in biogeography and speciation among seabirds, pinnipeds and cestodes. Von Ihering Centenary Symposium in Biogeography and Coevolution. Journal of Parasitology. 1992; 78: 601-615.

52. Lichtenfels JR, Pilitt PA, Hoberg EP. The National Parasite Collection- a century of service. Journal of Parasitology. 1992; 78: 924-929.

53. Hoberg EP, Kondratiev AI, Kitaysky AS. Larids, scolopacids and passerines exploiting ephemeral prey at Talan Island, USSR. Northwestern Naturalist. 1992; 73: 57-58.

54. Rickard LG, Zimmerman GL, Hoberg EP, Bishop JK, Pettit RJ. Influence of ivermectin and clorsulon in a cow-calf herd on the southern Oregon coast. Veterinary Parasitology. 1992; 41:45-55.

55. Lichtenfels JR, Hoberg EP. The systematics of nematodes that cause ostertagiasis in domestic and wild ruminants in North America: An update and a key to species. 2nd Ostertagia Workshop, University of Maryland, March 1992. Veterinary Parasitology. 1993; 46: 33-53.

56. Lichtenfels JR, Hoberg EP, Pilitt PA, Belem AMG. Cuticular ridge patterns in Mazamastrongylus 8 odocoilei and Mazamastrongylus pursglovei (Nematoda: Trichostrongyloidea) from white-tailed deer (Odocoileus virginianus). Systematic Parasitology. 1993; 24: 1-15.

57. Hoberg EP, Lichtenfels JR, Pilitt PA. Comparative morphology of Ostertagia mossi and Ostertagia dikmansi (Trichostrongylidae) from Odocoileus virginianus with comments on other Ostertagiinae from the Cervidae. Systematic Parasitology. 1993; 24: 111-127. 58. Hoberg EP, Lichtenfels JR, Pilitt PA. Synlophe of Cooperia neitzi (Trichostrongylidae: Cooperiinae) with comments on vulval inflations and hypertrophy of cuticular ridges among the trichostrongylids. Journal of the Helminthological Society of Washington. 1993; 60: 153-161.

59. Hoberg EP, Daoust P-Y, McBurney S. Bolbosoma capitatum (von Linstow, 1880) and Bolbosoma sp. (Acanthocephala) from sperm whales (Physeter macrocephalus L.) stranded on Prince Edward Island, Canada. Journal of the Helminthological Society of Washington. 1993; 60: 205-210.

60. Kritsky DC, Hoberg EP, Aubry KB. Lagarocotyle salamandrae gen. et sp. n. (Monogenoidea, Polyonchoinea, Lagarocotylidea ord. n.) from the cloaca of Rhyacotriton cascadae (Good and Wake) (Caudata: Rhyacotritonidae) in Washington State. Journal of Parasitology. 1993; 79: 322-330.

61. Rickard, LG, Hoberg EP, Allen NM, Zimmerman GL, Craig TM. Spiculopteragia spiculoptera and S. asymmetrica (Nematoda: Trichostrongyloidea) from red deer (Cervus elaphus) in Texas. Journal of Wildlife Diseases. 1993; 29: 512-515.

62. Brooks DR, Hoberg EP, Houtman A. Some platyhelminths inhabiting white-throated sparrows, Zonotrichia albicollis (Gmelin) (Aves: Emberizidae: Emberizinae), from Algonquin Park, Ontario, Canada. Journal of Parasitology. 1993; 79: 610-612.

63. Hoberg EP, Cawthorn RC, and Hedstrom OR. Enteric coccidia (Apicomplexa) in the small intestine of the northern spotted owl (Strix occidentalis caurina). Journal of Wildlife Diseases. 1993; 29: 495-497.

64. Hoberg EP, Lichtenfels JR, Pilitt PA. Affiliation of Hyostrongylus rubidus (Nematoda: Trichostrongylidae) with the Ostertagiinae, and evaluation of the synlophe and other structural characters. Journal of the Helminthological Society of Washington. 1993. 60: 219-233.

65. Hoberg EP, Miller S, Brown MA. Echinococcus granulosus (Taeniidae) and autochthonous echinococcosis in a North American horse. Journal of Parasitology. 1994; 80: 141-144.

66. Lichtenfels JR, Pilitt PA, Hoberg EP. New morphological characters for identifying individual specimens of Haemonchus spp. and a key to species in ruminants from North America. Journal of Parasitology. 1994; 80: 107-119.

67. Huss BT, Miller MA, Corwin RM, Hoberg EP, Obrien DP. Fatal cerebral coenurosis in a cat. Journal of the American Veterinary Medical Association. 1994; 205: 69-71.

68. Hoberg EP, Lichtenfels JR. Phylogenetic systematic analysis of the Trichostrongylidae (Nematoda) with an initial assessment of coevolution and biogeography. Journal of Parasitology. 1994; 80: 976-996.

69. Hoberg EP, Lloyd S, Omar HM. Libyostrongylus dentatus sp. n. (Nematoda: Trichostrongylidae) from ostriches in North America, with comments on the genera Libyostrongylus and Paralibyostrongylus. Journal of Parasitology. 1995; 81: 85-93.

70. Hoberg EP, Measures LN. Anophryocephalus inuitorum sp. nov. and A. arcticensis sp. nov. 9 (Eucestoda: Tetrabothriidae) in ringed seals, Phoca hispida and harp seals, P. groenlandica from high latitude seas of eastern Canada and the Arctic basin. Canadian Journal of Zoology. 1995; 73: 34-44.

71. Hoberg, E.P. Historical biogeography and modes of speciation across high latitude seas of the Holarctic: concepts for host-parasite coevolution among the Phocini (Phocidae) and Tetrabothriidae (Eucestoda): Canadian Journal of Zoology. 1995; 73: 45-57.

72. Hoberg EP, Sims DE, Odense PH. Comparative morphology of the scolex and microtriches among some species of Tetrabothrius (Eucestoda: Tetrabothriidae). Journal of Parasitology. 1995; 81: 475-481.

73. Khrustalev AV, Hoberg EP. Silver staining for elucidation of the synlophe in trichostrongyle nematodes. Journal of Parasitology. 1995; 81: 1016-1018.

74. Hoberg EP, Polley L, Gunn A, Nishi J. Umingmakstrongylus pallikuukensis gen. n. et sp. n. (Nematoda: Protostrongylidae) from muskoxen Ovibos moschatus, in the central Canadian Arctic with comments on biology and biogeography. Canadian Journal of Zoology. 1995; 73: 2266- 2282.

75. Lichtenfels JR, Hoberg EP, Pilitt PA, Craig TM. The synlophe and other structural characters of Sarwaria bubalis (Nematoda: Trichostrongyloidea) from cattle in Guyana. Journal of Parasitology. 1996; 82: 146-154.

76. Khrustalev AV, Hoberg EP. Carmine-propionic acid stain for elucidation of fine cellular structure in nematodes. Journal of Parasitology. 1996; 82: 176-178.

77. Hoberg EP. Emended description of Mazamastrongylus peruvianus (Nematoda: Trichostrongylidae), with comments on the relationships of the genera Mazamastrongylus and Spiculopteragia. Journal of Parasitology. 1996; 82: 470-477.

78. Van Baren D, Hoberg EP, Botzler RG. Abomasal parasites in tule elk (Cervus elaphus nannodes) from Grizzly Island, California. Journal of the Helminthological Society of Washington. 1996; 63: 222-225.

79. Hoberg EP, Khrustalev AV. Re-evaluation of Mazamastrongylus dagestanica (Nematoda: Trichostrongylidae) with analysis of the synlophe, genital cone and other structural characters. Journal of Parasitology. 1996; 82: 778-787.

80. Hoberg EP. Faunal diversity among avian parasite assemblages: the interaction of history, ecology and biogeography. Bulletin of the Scandinavian Society of Parasitology. 1996; 6: 65- 89.

81. Lichtenfels JR, Hoberg EP, Zarlenga DS. Systematics of gastrointestinal nematodes of domestic ruminants: Advances 1992-1995. Veterinary Parasitology. 1997; 72: 225-245.

82. Hoberg EP, Henny CJ, Hedstrom OR, Grove RA. Intestinal helminths of river otters (Lutra canadensis) from the Pacific Northwest. Journal of Parasitology. 1997; 83: 105-110.

83. McAlpine D, Murison LD, Hoberg EP. New records for the pygmy sperm whale, Kogia breviceps, (Physeteridae) from Atlantic Canada with notes on diet and parasites. Marine Mammal Science. 1997; 13: 701-704.

84. Hoberg EP, Gardner SL, Campbell RA. Paradigm shifts and tapeworm systematics. Parasitology Today. 1997; 13: 161-162. 10

85. Rickard LG, Hoberg EP, Mulrooney DM, Zimmerman GL. Isoelectric focusing of soluble proteins in the characterization of species and isolates of Nematodirus (Nematoda: Trichostrongyloidea). Journal of Parasitology. 1997; 83: 895-901.

86. Hoberg EP, Mariaux J, Justine J-L, Brooks DR, Weekes P. Phylogeny of the orders of the Eucestoda (Cercomeromorphae) based on comparative morphology: Historical perspectives and a new working hypothesis. Journal of Parasitology. 1997; 83: 1128-1147.

87. Adams A, Hoberg EP, McAlpine DF, Clayden S. Campula oblonga (Trematode: Campulidae) from an atypical host, the thresher shark, Alopias vulpinus (Bonnaterre). Journal of Parasitology. 1998; 84: 435-438.

88. Rego AA, de Chambrier A, Hanzelova V, Hoberg E, Scholz T, Weekes P, Zehnder M. Preliminary phylogenetic analysis of subfamilies of the Proteocephalidea (Eucestoda). Systematic Parasitology. 1998; 40: 1-19.

89. Hoberg EP, Brooks DR, Molina-Urena H, Erbe E. Echinocephalus janzeni sp. n. (Nematoda: Gnathostomatidae) in Himantura pacifica (Chondrichthyes: Myliobatiformes) from the Pacific coast of Costa Rica and Mexico, with historical biogeographic analysis of the genus. Journal of Parasitology. 1998; 84: 571-581.

90. Zarlenga D, Hoberg EP, Stringfellow F, Lichtenfels JR. Comparisons of two polymorphic species of Ostertagia and phylogenetic relationships within Ostertagiinae (Trichostrongylidae) inferred from ribosomal DNA repeat and mitochondrial DNA sequences. Journal of Parasitology. 1998; 84: 806-812.

91. Mignucci-Giannoni AA, Hoberg EP, Siegel-Causey D, Williams EH. Helminth fauna of cetaceans from the Carribean. Journal of Parasitology. 1998; 84: 939-946.

92. Hoberg EP, Gardner SL, Campbell RA. Systematics of the Eucestoda: Advances toward a new phylogenetic paradigm, and observations on the early diversification of tapeworms and vertebrates. Systematic Parasitology. 1999; 42: 1-12.

93. Bray RA, Jones A, Hoberg EP. Observations on the phylogeny of the cestode order Pseudophyllidea. Systematic Parasitology. 1999; 42: 13-20.

94. Ivanov V, Hoberg EP. Preliminary comments on a phylogenetic study of the order Diphyllidea van Beneden in Carus, 1863. Systematic Parasitology. 1999; 42: 21-27.

95. Hoberg EP, Jones A, Bray, RA. Phylogenetic analysis of the families of the Cyclophyllidea based on comparative morphology with new hypotheses for coevolution in vertebrates. Systematic Parasitology. 1999; 42: 51-73.

96. Carreno RA, Hoberg EP. Evolutionary relationships among the Protostrongylidae (Nematoda: Metastrongyloidea) as inferred from morphological characters, with consideration of parasite-host coevolution. Journal of Parasitology.1999; 85: 638-648.

97. Gray ML, Rogers R, Little S, Puette M, Ambrose D, Hoberg EP. Sparganoisis in feral hogs (Sus scrofa) from Florida. Journal of the American Veterinary Medical Association. 1999; 215: 204-208.

98. Hoberg EP, Monsen KJ, Kutz S, Blouin MS. Structure, biodiversity and historical biogeography of 11 nematode faunas in Holarctic ruminants: Morphological and molecular diagnoses for Teladorsagia boreoarcticus n. sp. (Nematoda: Ostertagiinae) a dimorphic cryptic species in muskoxen (Ovibos moschatus). Journal of Parasitology. 1999; 85: 910-934.

99. Hoberg EP, Ebinger W, Render JA. Fatal cysticercosis by Taenia crassiceps (Cyclophyllidea: Taeniidae) in a presumed immunocompromised canine host. Journal of Parasitology.1999; 85: 1174-1178.

100. Kutz SJ, Hoberg EP, Polley L. Experimental infections in muskoxen (Ovibos moschatus) and domestic sheep (Ovis aries) with Umingmakstrongylus pallikuukensis (Nematoda: Protostrongylidae): Parasite development, population structure and pathology. Canadian Journal of Zoology. 1999; 77: 1562-1572.

101. Hoberg EP, Jones A, Rausch RL, Eom KS, Gardner SL. A Phylogenetic hypothesis for species of the genus Taenia (Eucestoda: Cyclophyllidea). Journal of Parasitology. 2000; 86: 89-98.

102. Rickard L, Hoberg EP. Reassignment of the genus Lamanema (Nematoda: Trichostrongyloidea) from Nematodirinae to Molineinae. Journal of Parasitology. 2000; 86: 647- 650.

103. Brooks DR, Hoberg EP. Triage for the biosphere: The need and rationale for taxonomic inventories and phylogenetic studies of parasites. Comparative Parasitology. 2000; 67: 1-25. (order of authorship is arbitrary).

104. Kutz SJ, Hoberg EP, Polley L. Emergence of third stage larvae of Umingmakstrongylus pallikuukensis from three gastropod intermediate host species. Journal of Parasitology. 2000; 86: 743-749.

105. Nadler SA, Hoberg EP, Hudspeth DSS, Rickard LG. Relationships of Nematodirus species and Nematodirus battus isolates (Nematoda: Trichostrongyloidea) based on nuclear ribosomal DNA sequences. Journal of Parasitology. 2000; 86: 588-601.

106. Santin-Duran M, Alunda JM, San Miguel JM, Hoberg EP, de la Fuente C. Elaeophorosis in red deer (Cervus elaphus) from central Spain. Journal of Wildlife Diseases. 2000; 36: 779- 782.

107. Hoberg EP, Adams A. Phylogeny, history and biodiversity: Understanding faunal structure and biogeography in the marine realm. Bulletin of the Scandinavian Society of Parasitology. 2000; 10: 19-37.

108. Kutz SJ, Hoberg EP, Polley L. Umingmakstrongylus pallikuukensis (Nematoda: Protostrongylidae) in gastropods: Larval morphology, morphometrics, and development rates. Journal of Parasitology. 2001; 87: 527-535.

109. Kutz SJ, Veitch A, Hoberg EP, Elkin B, Jenkins EJ, Polley L. New host and geographic records for the protostrongylid nematodes Protostrongylus stilesi and Parelaphostrongylus odocoilei in Dall’s sheep (Ovis dalli dalli). Journal of Wildlife Diseases. 2001; 37: 761-774.

110. Zamparo D, Brooks DR, Hoberg EP, McLennan, DA. Phylogenetic analysis of the Rhabdoceola (Platyhelminthes) with emphasis on the Neodermata and relatives. Zoologica Scripta. 2001; 30: 59-77.

111. Kutz S, Hoberg EP, Polley L. A new lungworm in muskoxen: An exploration in Arctic parasitology. Trends in Parasitology. 2001; 17: 276-280. 12

112. Hoberg EP, Alkire NL, de Queiroz A, Jones A. Out of Africa: Origins of the Taenia tapeworms in humans. Proceedings Royal Society of London, Series B. 2001; 268: 781-787.

113. Carreno R, Durden L, Brooks DR, Abrams A, Hoberg EP. Parelaphostrongylus tenuis (Nematoda: Protostrongylidae) and other parasites of white-tailed deer (Odocoileus virginianus) in Costa Rica. Comparative Parasitology. 2001; 68: 177-184.

114. Brooks DR, Hoberg EP. Parasite systematics in the 21st Century: Opportunities and obstacles. Trends in Parasitology. 2001; 17: 273-275.

115. Santin-Duran M, Alunda JM, de la Fuente C, Hoberg EP. Onchocercosis in red deer (Cervus elaphus) from Spain. Journal of Parasitology. 2001; 87: 1213-1215.

116. Hoberg EP, Abrams A. Synlophe in Ostertagia cf. kasakhstanica (Nematoda: Ostertagiinae), the minor morphotype of O. bisonis in western North America. Journal of Parasitology. 2001; 87: 1181-1184.

117. Santin-Duran M, Alunda JM, de la Fuente C, Rosenthal B, Hoberg EP. Identical ITS-1 and ITS-2 sequences indicate that Spiculopteragia asymmetrica and S. quadrispiculata (Nematoda: Trichostrongylidae) constitute morphologically distinct variants of a single species. Journal of Parasitology. 2002; 88: 417-418.

118. Hoberg EP, Kutz S, Nagy J, Jenkins E, Elkin B, Branigan M, Cooley D. Protostrongylus stilesi (Protostrongylidae): Ecological isolation and putative host switching between muskoxen and Dall’s sheep in a contact zone. Comparative Parasitology. 2002; 69: 1-9.

119. Hoberg EP. Foundations for an integrative parasitology: collections archives and biodiversity informatics. Comparative Parasitology. 2002; 69: 124-131.

120. Hoberg EP. Taenia tapeworms: biology, evolution and socioeconomic significance. Microbes and Infection. 2002; 4: 859-866.

121. Hoberg EP, Klassen GJ. Revealing the faunal tapestry: Coevolution and historical biogeography of hosts and parasites in marine systems. Parasitology. 2002; 124 (Supplement): S3-S22.

122. Lichtenfels JR, Pilitt PA, Gibbons LM, Hoberg EP. Redescriptions of Haemonchus mitchelli and Haemonchus okapiae (Nematoda: Trichostrongyloidea). Journal of Parasitology. 2002; 88: 947-960.

123. Hoberg EP. Colonization and diversification: historical and coevolutionary trajectories among cestodes, cetaceans and pinnipeds. In Symposium- Parasites of Marine Mammals, ICOPA X. Monduzzi Editore. Bologna, Italy. 2002. pp. 65-69.

124. Hoberg EP, Abrams A, Carreno R, Lichtenfels JR. Ashworthius patriciapilittae n. sp. (Trichostrongyloidea: Haemonchinae), an abomasal nematode in Odocoileus virginianus from Costa Rica, and a first record for the genus in the Western Hemisphere. Journal of Parasitology. 2002; 88: 1187-1199.

125. Kutz SJ, Hoberg EP, Nishi J, Polley L. Development of the muskox lungworm, Umingmakstrongylus pallikuukensis (Protostrongylidae) in gastropods in the Arctic. Canadian Journal of Zoology. 2002; 80: 1972-1985. 13

126. Wünschmann A, Garlie V, Averbeck G, Kurtz H, Hoberg EP. Cerebellar cysticercosis by Taenia crassiceps in a domestic cat. Journal of Veterinary Diagnostic Investigation. 2002; 19: 240-253.

127. Winker K, Gibson DD, Sowls A, Lawhead BE, Martin PD, Hoberg EP, Causey D. The birds of St. Matthew Island, Bering Sea, at the end of the millennium. Wilson Bulletin. 2002; 114: 491-509.

128. Hoberg EP, Kutz SJ, Galbreath KE, Cook J. Arctic biodiversity: From discovery to faunal baselines- revealing the history of a dynamic ecosystem. In Plenary of the ICOPA X: Parasitology and Biodiversity in an Age of Discovery. Journal of Parasitology. 2003; 89: (Supplement): S-84- S-95.

129. Dowling APG, van Veller MGP, Hoberg EP, Brooks DR. A priori and a posteriori methods in comparative evolutionary studies of host-parasite associations. Cladistics. 2003; 19: 240-253.

130. Brooks DR, Dowling APG, van Veller MGP, Hoberg EP. Ending a decade of deception: A valiant failure, a not-so-valiant failure, and a success story. Cladistics. 2004; 20: 32-46.

131. Hoberg EP, Lichtenfels JR, Gibbons L. Phylogeny for species of the genus Haemonchus (Nematoda: Trichostrongyloidea): Considerations of their evolutionary history and global biogeography among Camelidae and Pecora (Artiodactyla). Journal of Parasitology 2004; 90: 1085-1102.

132. Kutz SJ, Hoberg EP, Nagy J, Polley L, Elkin B. “Emerging” parasitic infections in Arctic ungulates. Integrative and Comparative Biology 2004; 44: 109-118.

133. Santín-Durán M, Alunda JM, Hoberg EP, de la Fuente C. Abomasal parasites in wild sympatric cervids, red deer Cervus elaphus and fallow deer Dama dama, at three locations from Central and Western Spain: Relationship to host density and land management. Journal of Parasitology 2004; 90: 1378-1386.

134. Jenkins EJ, Hoberg EP, Veitch A, Schwantje H, Wood M, Toweill D, Kutz SJ, Polley L. Parasite fauna of mountain goats (Oreamnos americana) in the Northwest Territories, British Columbia and Idaho. Proceedings of the Northern Wild Sheep and Goat Council. 2004; 14: 141-158.

135. Hoberg EP, Lichtenfels JR, Rickard L. Phylogeny for genera of Nematodirinae (Nematoda: Trichostrongylina). Journal of Parasitology. 2005; 91: 382-389.

136. Hoberg EP. Coevolution and biogeography among Nematodirinae (Nematoda: Trichostrongylina), Lagomorpha and Artiodactyla (Mammalia): Exploring determinants of history and structure for the northern fauna across the Holarctic. Journal of Parasitology. 2005; 91: 358-369.

137. Hoberg EP, Abrams A. Pseudostertagia bullosa (Nematoda: Trichostrongyloidea) in artiodactyl hosts from North America: Redescription and comments on systematics. Journal of Parasitology. 2005; 91: 370-381.

138. Jenkins EJ, Appleyard GD, Hoberg EP, Rosenthal BM, Kutz SJ, Veitch A, Schwantje H, Polley L. Geographic distribution of the muscle dwelling nematode Parelaphostrongylus odocoilei (Protostrongylidae: Elaphostrongylinae) in North America using molecular identification of first- stage larvae. Journal of Parasitology. 2005; 91: 574-584.

14 139. Pozio E, Pagani P, Marucci G, Zarlenga DS, Hoberg EP, La Rosa G, Rossi L. Trichinella britovi etiological agent of sylvatic trichinellosis in the Republic of Guinea (West Africa) and a re- evaluation of geographic distribution for encapsulated species in Africa. International Journal for Parasitology. 2005; 35: 955-960.

140. Jenkins EJ, Hoberg EP, Polley L. Development and pathogenesis of Parelaphostrongylus odocoilei in experimentally infected thinhorn sheep (Ovis dalli). Journal of Wildlife Diseases. 2005; 41:669-682.

141. Hoberg, EP, Jenkins EJ, Rosenthal BM, Wong M, Erbe E, Kutz SJ, Polley L. Caudal polymorphism and cephalic morphology among first-stage larvae of Parelaphostrongylus odocoilei (Protostrongylidae: Elaphostrongylinae) in Dall’s sheep from the Mackenzie Mountains, Canada. Journal of Parasitology. 2005; 91:1318-1325.

142. Kutz SJ, Hoberg EP, Polley L, Jenkins EJ. Global warming is changing the dynamics of Arctic host-parasite systems. Proceedings of the Royal Society London, B. 2005; 272: 2571- 2576.

143. Cook JA, Hoberg EP, Koehler A, Henttonen H, Wickström L, Haukisalmi V, Galbreath K, Chernyavski F, Dokuchaev N, Lahzuhtkin A, MacDonald SO, Hope A, Waltari E, Runck A, Veitch A, Popko R, Jenkins E, Kutz S, Eckerlin R. Beringia: Intercontinental exchange and diversification of high latitude mammals and their parasites during the Pliocene and Quaternary. Mammal Study. 2005; 30: S33-S44.

144. Muzaffar SB, Hoberg EP, Jones II. Possible recent range expansion for Alcataenia longicervica (Eucestoda: Dilepididae) parasitic in murres (Uria spp.) into the North Atlantic. Marine Ornithology. 2005; 33: 189-191.

145. Jenkins EJ, Kutz SJ, Hoberg EP, Polley L. Bionomics of larvae of Parelaphostrongylus odocoilei (Nematoda: Protostrongylidae) in experimentally infected gastropod intermediate hosts. Journal of Parasitology. 2006; 92: 298-305.

146. Hoberg EP. Phylogeny of Taenia: Defining species and origins of human parasites. Parasitology International. 2006; 50 (Supplement): S23-S30.

147. Mortenson JA, Abrams A, Rosenthal B, Dunams D, Hoberg EP, Bildfell RJ, Green RL. Parelaphostrongylus odocoilei in Oregon black-tailed deer. Journal of Wildlife Diseases. 2006; 42: 527-535. 148. Jenkins EJ, Veitch AM, Kutz SJ, Hoberg EP, Polley L. Climate change and the epidemiology of protostrongylid nematodes in northern ecosystems: Parelaphostrongylus odocoilei and Protostrongylus stilesi in Dall’s sheep (Ovis dalli dalli). Parasitology. 2006; 132: 387-401.

149. Brooks DR, Hoberg EP. Systematics and emerging infectious diseases: From management to solution. Journal of Parasitology. 2006; 92: 426-429.

150. Brooks DR, McLennan DA, León-Règagnon V, Hoberg EP. Phylogeny, ecological fitting and lung flukes: helping solve the problem of emerging infectious diseases. Revista Mexicana de Biodiversidad. 2006; 77: 225-233.

151. Zarlenga DS, Rosenthal B, Pozio E, La Rosa G, Hoberg EP. Post-Miocene expansion, colonization, and host switching drove speciation among extant nematodes of the archaic genus Trichinella. Proceedings of the National Academy of Sciences (US). 2006; 103: 7354-7359.

15 152. Jenkins EJ, Veitch AM. Kutz SJ, Bollinger T, Chirino-Trejo M, Elkin BT, West K, Hoberg EP, Polley L. Protostrongylid parasites and pneumonia in captive and wild thinhorn sheep (Ovis dalli). Journal of Wildlife Diseases. 2007; 43: 189-205.

153. Karpenko SV, Dokuchaev NE, Hoberg EP. Nearctic shrews, Sorex spp., as paratenic hosts of Soboliphyme baturini (Nematoda: Soboliphymidae). Comparative Parasitology. 2007; 74: 81-87.

154. Waltari E, Hoberg EP, Lessa EP, Cook JA. Eastward Ho: phylogeographic perspectives on colonization of hosts and parasites across the Beringian nexus. Journal of Biogeography. 2007; 34: 561-574.

155. Muzaffar SB, Hoberg EP, Jones IL. First record of Alcataenia larina larina (Cestoda: Dilepididae) in an Atlantic Puffin (Aves, Alcidae, Fratercula arctica) from Newfoundland, Canada. Comparative Parasitology. 2007; 74: 380-382.

156. Hoberg EP, Abrams A. Emended description and redetermination of Sarwaria caballeroi n. comb. (Nematoda: Ostertagiinae) with details of the synlophe and esophageal characters. Journal of Parasitology. 2007; 93: 1140-1150.

157. Koehler AVA, Hoberg EP, Dokuchaev NE, Cook JA. Geographic and host range of the nematode Soboliphyme baturini across Beringia. Journal of Parasitology. 2007; 93: 1070- 1083.

158. Brooks DR, Hoberg EP. How will climate change affect host-parasite assemblages? Trends in Parasitology. 2007; 23: 571-574.

159. Kutz SJ, Asmundsson IM, Hoberg EP, Appleyard GD, Jenkins EJ, Beckmen K, Branigan M, Butler L, Chilton NB, Cooley D, Elkin B, Huby-Chilton F, Johnson D, Kuchboev A, Nagy J, Oakley M, Polley L, Popko R, Scheer A, Simard M, Veitch A. Serendipitous discovery of a novel prostostrongylid (Nematoda: Metastrongyloidea) associated with caribou (Rangifer tarandus), muskoxen (Ovibos moschatus) and moose (Alces alces) from high latitudes of North America based on DNA sequence comparisons. Canadian Journal of Zoology. 2007; 85: 1143-1156.

160. Brooks DR, Hoberg EP. Darwin’s necessary misfit and the sloshing bucket: The evolutionary biology of emerging infectious diseases. Outreach and Education in Evolution. 2008; 1: 2-9.

161. Hoberg EP, Polley L, Jenkins EJ, Kutz SJ, Veitch A, Elkin B. Integrated approaches and empirical models for investigation of parasitic diseases in northern wildlife. Emerging Infectious Diseases. 2008; 14: 10-17.

162. Santin-Durán M, Alunda JM, Hoberg EP, de la Fuente C. Age distribution and seasonal dynamics of abomasal helminths in wild red deer from Central Spain. Journal of Parasitology. 2008; 94: 1031-1037. 163. Asmundsson IM, Mortenson JA, Hoberg EP. Muscleworms, Parelaphostrongylus andersoni (Nematoda: Protostrongylidae), discovered in Columbia white-tailed deer from Oregon and Washington: Implications for biogeography and host associations. Journal of Wildlife Diseases. 2008; 44: 16-27.

164. Hoberg EP, Abrams A, Ezenwa VO. An exploration of diversity among the Ostertagiinae (Nematoda: Trichostrongyloidea) in ungulates from Sub-Saharan Africa with a proposal for a new genus. Journal of Parasitology. 2008; 94: 230-251.

165. Hoberg EP, Abrams A. Hamulonema gen. nov. for Teladorsagia hamata and Ostertagia 16 kenyensis in the ostertagiine fauna (Nematoda: Trichostrongyloidea) from African ungulates. Journal of Parasitology. 2008; 94: 866-879.

166. Hoberg EP, Polley L, Jenkins EJ, Kutz SJ. Pathogens of domestic and free-ranging ungulates: Global climate change in temperate to boreal latitudes across North America. Revue Scientifique et Technique Office International des Épizooties. 2008; 27: 511-528.

167. Hoberg EP, Brooks DR. A macroevolutionary mosaic: Episodic host-switching, geographic colonization, and diversification in complex host-parasite systems. Journal of Biogeography. 2008; 35: 1533-1550.

168. Kutz SJ, Schock D, Brook R, Hoberg EP. Climate change and infectious diseases in wildlife. Wildlife Professional. 2008; Fall, 42-46.

169. Zarlenga DS, Rosenthal B, Hoberg E, Mitreva M. Integrating genomics and phylogenetics in understanding the history of Trichinella species. Veterinary Parasitology. 2008; 159: 210-213. (doi:10:1016/j.vetpar.2008.10.061)

170. Carta LK, Handoo ZA, Hoberg EP, Erbe EF, Wergin WP. Evaluation of some vulval appendages in nematode taxonomy. Comparative Parasitology. 2009; 76: 191-209.

171. Koehler VA, Hoberg EP, Torres-Pérez F, Cook JA. A molecular view of the superfamily Dioctophymatoidea (Nematoda). Comparative Parasitology. 2009; 76: 100-104.

172. Koehler AVA, Hoberg EP, Dokuchaev NE, Trabenkova NA, Whitman JS, Nagorsen DW, Cook JA. Phylogeography of a Holarctic nematode, Soboliphyme baturini among mustelids: Climate change, episodic colonization and diversification in a complex host-parasite system. Biological Journal of the Linnean Society. 2009; 96: 651-663.

173. Hoberg EP, Pilitt PA, Galbreath KE. Why museums matter: A tale of pinworms (Oxyuroidea: Heteroxynematidae) among pikas (Ochotona princeps and O. collaris) in the American west. Journal of Parasitology. 2009; 95: 490-501.

174. Pozio E, Hoberg EP, La Rosa G, Zarlenga DS. Molecular taxonomy and phylogeny of nematodes of the genus Trichinella. Infection Genetics and Evolution. 2009; 9: 606-616.

175. Hoberg EP, Abrams A, Pilitt P. Robustostrongylus aferensis gen. nov. et sp. nov. (Nematoda: Trichostrongyloidea) in kob (Kobus kob) and hartebeest (Alcelaphus buselaphus jacksoni) (Artiodactyla) from Sub-Saharan Africa, with further ruminations on the Ostertagiinae. Journal of Parasitology. 2009; 95: 702-717.

176. Hoberg EP, Abrams A, Pilitt PA. Synlophe structure for species of Longistrongylus (Nematoda: Trichostrongyloidea), abomasal parasites among ungulates from sub-Saharan Africa, with comparisons to the global ostertagiine fauna. Journal of Parasitology. 2009; 95: 1468- 1478.

177. Kutz SJ, Dobson A.P., Hoberg E.P. Where are the parasites? Science. 2009; 326: 1187- 1188. 178. Gasbarre L, Smith LL, Hoberg EP, Pilitt PA.. Further characterization of a cattle nematode population with demonstrated resistance to current anthelmintics. Veterinary Parasitology. 2009; 166: 275-280.

17 179. Reinstein SL, Lucio-Forster A, Bowman DD, Eberhard ML, Hoberg EP, Pot S, Miller PE. Surgical extraction of intraocular Parelaphostrongylus tenuis in a horse. Journal of the American Veterinary Medical Association. 2010; 237: 196-199.

180. Hoberg EP, Abrams A, Pilitt PA. A new species of trichostrongyloid in African buffalo (Syncerus caffer) from Uganda. Journal of Parasitology. 2010; 96: 129-136.

181. Hoberg EP, Kumsa B, Pilitt P and Abrams A. Synlophe structure in Pseudomarshallagia elongata (Nematoda: Trichostrongyloidea), abomasal parasites among Ethiopian ungulates, with consideration of other morphological attributes and differentiation within the Ostertagiinae. Journal of Parasitology. 2010; 96: 401-411.

182. Polley L, Hoberg EP, Kutz SJ. Climate change, parasites and shifting boundaries. Acta Veterinaria Scandinavica. 2010; 52: S1-S5.

183. Hoberg EP. Invasive processes, mosaics and the structure of helminth parasite faunas. Revue Scientifique et Technique Office International des Épizooties. 2010; 29: 255-272.

184. Ezenwa VO, Hines AM, Archie EA, Hoberg EP, Asmundsson I, Hogg JT. Muellerius capillaris dominates the lungworm community of bighorn sheep at the National Bison Range, Montana. Journal of Wildlife Diseases. 2010; 46: 988-993.

185. Durette-Desset M-C, Galbreath KE, Hoberg EP. Discovery of new Ohbayashinema spp. (Heligmosomidae: Nematoda) in Ochotona princeps and O. cansus (Lagomorpha: Ochotonidae) from western North America and central Asia, with consideration of historical biogeography. Journal of Parasitology. 2010; 96: 569-579.

186. Laaksonen S, Pusenious J, Kumpula J, Venäläinen A, Kortet R, Oksanen A, Hoberg EP. Climate change promotes the emergence of serious disease outbreaks of Filarioid nematodes. EcoHealth. 2010; 7: 7-13.

187. Weaver HJ, Hawdon J, Hoberg EP. Soil transmitted helminthiases: implications of climate change and human behaviour. Trends in Parasitology. 2010; 26: 574-581.

188. Rossin MA, Timi JT, Hoberg EP. An endemic Taenia from South America: Validation of T. talicei Dollfus, 1960 (Cestoda; Taeniidae) with characterization of metacestodes and adults. Zootaxa. 2010; 2636: 49-58.

189. Kuchboev AE, Hoberg EP. Morphological and ultrastructural changes in tissues of intermediate and definitive hosts infected by protostrongylid lungworms (Nematoda: Metastrongyloidea). 190. Turkish Journal of Zoology. 2011; 35: 1-7.

191. Binkiene R, Kontrimavichus V, Hoberg EP. Overview of the cestode fauna of European shrews of the genus Sorex, with an exploration of historical processes in post-glacial Europe. Helminthologia. 2011; 48: 201-228.

192. Galbreath KE, Hoberg EP. Return to Beringia: Parasites reveal cryptic biogeographic history of North American pikas. Proceedings of the Royal Society, B. 2012; 279: 371-378.

193. Hoberg EP, Abrams A, Pilitt PA, Kutz SJ. Discovery and description of the “davtiani” morphotype for Teladorsagia boreoarcticus (Trichostrongyloidea: Ostertagiinae) an abomasal parasite in muskoxen, Ovibos moschatus and caribou, Rangifer tarandus from the North 18 American Arctic. Journal of Parasitology. 2012; 98: 355-364.

194. Wagner BA, Hoberg EP, Somers CM, Soos C, Fenton H, Jenkins EJ. Gastrointestinal helminth parasites of double-crested cormorants (Phalacrocorax auritus) at four sites in Saskatchewan, Canada, 2006-2007. Comparative Parasitology. 2012; 79: 275-282.

195. Makarikov AA, Gardner SL, Hoberg EP. New species of Arostrilepis (Cyclophyllidea: Hymenolepididae) in members of Cricetidae and Geomyidae (Rodentia) from the western Nearctic. Journal of Parasitology. 2012; 98: 617-626.

196. Hoberg EP, Abrams A, Pilitt PA, Jenkins EJ. Discovery and description of a new trichostrongyloid species (Nematoda: Ostertagiinae), abomasal parasites in mountain goat, Oreamnos americanus from the western Cordillera of North America. Journal of Parasitology. 2012; 98: 817-846.

197. Anh NTL, Madsen H, Thi TD, Hoberg E, Dalsgaard A, Murrell KD. Rats are not an important reservoir for fishborne zoonotic trematodes in Vietnamese aquaculture. Parasitology Research. 2012; 111: 1045-1048.

198. Miller DS, Hoberg E, Weiser G, Aune K, Atkinson M, Kimberling C. A review of determinants associated with bighorn sheep (Ovis canadensis) die-offs. Veterinary Medicine International. 2012; Article ID 796517. 19 pgs. doi:10.1155/2012/796517.

199. Hoberg EP, Galbreath KE, Cook JA, Kutz SJ, Polley L. Northern host-parasite assemblages: History and biogeography on the borderlands of episodic climate and environmental transition. D. Rollinson and S.I. Hays (eds.). Elsevier. Advances in Parasitology. 2012; 79: 1-97.

200. Kutz SJ, Ducrocq J, Verocai G, Hoar B, Colwell D, Beckmen K, Polley L, Hoberg E. Parasites in ungulates of Arctic North America and Greenland: A view of contemporary diversity, ecology, and impact in a world under change. D. Rollinson and S.I. Hays (eds.). Elsevier. Advances in Parasitology. 2012; 79: 99-252.

201. Galbreath KE, Cook JA, Hoberg EP. Climate’s role in polar bear past. Science. 2012; 336: 1230.

202. Verocai GG, Lejeune M, Beckmen KB, Kashivakura CK, Veitch AM, Popko RA, Fuentealba C, Hoberg EP, Kutz SK. Defining parasite biodiversity at high latitudes of North America: new host and geographic records for Onchocerca cervipedis (Nematoda: Onchocercidae) in moose and caribou. Parasites and Vectors. 2012; 5: 242. doi:10.1186/1756- 3305-5-242.

203. Jenkins E, Castrodale L, de Rosemond S, Dixon B, Elmore S, Gesy K, Hoberg E, Polley L, Schurer J, Simard M, Thompson RCA. Tradition and transition: Parasitic zoonoses of people and animals in Alaska, northern Canada and Greenland. D. Rollinson and S.I. Hays (eds.). Elsevier. Advances in Parasitology. 2013: 82: 33-204.

204. Makarikov AA, Galbreath KE, Hoberg EP. Parasite diversity at the Holarctic nexus: Species of Arostrilepis (Eucestoda: Hymenolepididae) in voles and lemmings (Cricetidae: Arvicolinae) from greater Beringia. Zootaxa. 2013: 3608: 401-439.

205. Lavikainen A, Haukisalmi V, Deksne G, Holmala K, Virapin ML, Isomursu M, Jokelainen P, Näreaho A, Laakkonen J, Hoberg E, Sukura A. Molecular identification of Taenia spp. in Eurasian lynx (Lynx lynx) from Finland. Parasitology. 2013: 140: 653-662.

206. Kutz SJ, Checkley S, Verocai GG, Dumond M, Hoberg EP, Peacock R, Wu J, Orsel K, Seegers K, Warren A, Abrams A. Invasion, establishment, and range expansion of two parasitic nematodes in the Canadian Arctic. Global Change Biology. 2013: 19: 3254-3262. DOI 10:1111/gcb.12315.

207. Galbreath KE, Ragaliauskaitė K, Kontrimavichus L, Makarikov A, Hoberg EP. A widespread distribution of Arostrilepis tenuicirrosa (Eucestoda: Hymenolepididae) in Myodes voles (Cricetidae: Arvicolinae) from the Palearctic based on molecular and morphological criteria: Historical and biogeographic implications. Acta Parasitologica. 2013: 58: 441-452.

208. Platt TR, Hoberg EP, Chisholm L. Critical Comment: On the morphology and taxonomy of Griphoibilharizia amoena Platt and Blair, 1991 (Schistosomatoidea), a dioecious digenetic trematode parasite of the freshwater crocodile, Crocodylus johnstoni, in Australia. Journal of Parasitology. 2013: 99: 888-891.

209. Steele J, Orsel K, Cuyler C, Hoberg EP, Kutz SJ. Divergent parasite faunas in adjacent populations of West Greenland caribou: Suggested natural and anthropogenic influences. International Journal for Parasitology- Parasites and Wildlife. 2013: 2: 197-202.

210. Justine J-L, Rahmouni C, Gey D, Schoelinck C, Hoberg EP. The monogenean which lost its clamps. PLos One. 2013: 8(11): e79155. Doi:10.1371/journal.pone.0079155.

211. Hoberg EP. Robert Lloyd Rausch- A life in nature and field biology. Journal of Parasitology. 2014: 100: 547-552.

212. Kutz S, Hoberg EP, Molnár PK, Dobson A, Verocai G. A walk on the tundra: Host- parasite interactions in an extreme environment. International Journal for Parasitology- Parasites and Wildlife. 2014: 3: 198-208.

213. Yanagida T, Carod J-F, Mirua T, Nakayama T, Sako Y, Nakao M, Hoberg EP, Ito A. Genetics of the pig tapeworm in Madagascar reveal a history of human dispersal and colonization. PLos One. 2014: 9(10):e109003.doi:10.1371/journal.pone.0109002.

214. Bronson E, Kelly K, Hoberg EP. Airsac nematode Monopetalonema alcedinis in the Belted Kingfisher (Megaceryle alcyon). Journal of Wildlife Disease. 2014: 50: 938-941.

215. Haukisalmi V, Hardman LM, Hoberg EP, Henttonen H. Phylogenetic relationships and taxonomic revision of Paranoplocephala Lühe, 1910 sensu lato (Cestoda, Cyclophyllidea, Anoplocephalidae). Zootaxa. 2014: 3873: 371-415.

216. Brooks, DR, Hoberg EP, Gardner SL, Boeger W, Galbreath KE, Herczeg D, Mejía-Madrid HH, Racz E, Tsogtsaikhan A. Finding them before they find us: Informatics, parasites and environments in accelerating climate change. Comparative Parasitology. 2014: 81: 155-164.

217. Zarlenga D, Hoberg EP, Rosenthal B, Mattiucci S, Nascetti G. Anthropogenics: human influence on global and genetic homogenization of parasite populations. Journal of Parasitology. 2014: 100: 756-772.

218. Blume GR, Reis JL, Gardiner CH, Hoberg EP, Pilitt PA , Silvestres R, de Sant'Ana FJF. Granulomatous pneumonia due to Spirocerca lupi in two free-ranging maned wolves (Chrysocyon brachyurus) from central Brazil. Journal of Veterinary Diagnostic Investigation. 2014: 26: 815- 817.

219. Verocai G, Kutz SJ, Simard M, Hoberg EP. Varestrongylus eleguneniensis sp. n. (Nematoda: Protostrongylidae): a widespread, multi-host lungworm of North American ungulates, with an emended diagnosis for the genus and explorations of biogeography. Parasites and Vectors. 2014. 7: 556. DOI 10.1186/s13071-014-0556-9.

220. Verocai GG, Hoberg EP, Vikoren T, Davidson RK, Rezansoff AM, Gilleard JS, Ytrehus B, Handeland K, Kutz SJ. Resurrection and redescription of Varestrongylus alces (Nematoda; Protostrongylidae) a lungworm of Eurasian elk (Alces alces), with a report of associated pathology. Parasites and Vectors. 2014. 7: 557. DOI 10.1186/s13071-014-0557-8

221. Hoberg EP, Brooks DR. Evolution in action: Climate change, biodiversity dynamics and emerging infectious disease. Theme Issue- Climate Change and Vector-Borne Diseases. 222. Philosophical Transactions of the Royal Society B. 2015: 370: 20130553. 223. dx.doi.org/10.1098/rstb.2013.0553

224. Brooks DR, Hoberg EP, Boeger WA. In the eye of the Cyclops- re-examining the classic case of cospeciation: Why paradigms are important. Comparative Parasitology. 2015. In Press.

225. Hoberg EP, Agosta SJ, Boeger WA, Brooks DR. An Integrated parasitology: Revealing the elephant through tradition and invention. Trends in Parasitology. 2015: In Press.

226. Davidson RK, Kutz S, Madslien K, Hoberg EP, Handeland K. Gastrointestinal parasites in an isolated population of wild red deer (Cervus elaphus) in Norway. Acta Veterinaria Scandinavica. In Press.

227. Galbreath KE, Hoberg EP. Host responses to historical climate change shape parasite communities in North America’s Intermountain West. Folia Zoologica. In Press.

228. Makarikov A, Nims T, Galbreath KE, Hoberg EP. Hymenolepis folkertsi n. sp. (Eucestoda: Hymenolepididae) from the oldfield mouse Peromyscus polionotus (Wagner) (Rodentia: Cricetidae: Neotominae) from the southeastern Nearctic with comments on tapeworm faunal diversity among deer mice. Folia Parasitologica. In Press.

229. Budischak SA, Hoberg, EP, Abrams A, Jolles AE, Ezenwa VO. Molecular barcoding enables accurate, non-invasive sampling of parasitic nematode communities in wild hosts. Molecular Ecology Resources. In Press.

230. Dudley JP, Hoberg EP, Jenkins EJ, Parkinson AJ. Climate change in the North American Arctic: A One Health perspective. EcoHealth. In Review.

231. Budischak SA, Hoberg, EP, Abrams A, Jolles AE, Ezenwa VO. Experimental insight into the process of parasite community assembly. Journal of Animal Ecology. In Review.

232. Araujo SBL, Braga MP, Brooks DR, Agosta S, Hoberg EP, von Hathental F, Boeger WA. Modeling the Stockholm Paradigm: Ecological fitting facilitates host switching in sloppy fitness space. Oikos. In Review.

233. Laaksonen S, Oksanen A, Hoberg E. A lymphatic dwelling filarioid nematode, Rumenfilaria andersoni (Filarioidea; Splendidofilariinae), is an emerging parasite in Finnish cervids. Parasites and Vectors. In Review.

234. Verocai GG, Hoberg EP, Kutz SJ. Phylogeny, history and biogeography among 21 Varestrongylus lungworms in ungulates, processes for host and geographic colonization. Folia Parasitologica.

235. Haukisalmi V, Hardman LM, Fedorov V, Hoberg EP, Henttonen H. Molecular systematics and phylogeography of cestodes of the genus Anoplocephaloides Baer, 1923 s. s. (Cyclophyllidea: Anoplocephalidae) in lemmings (Lemmus, Synaptomys).

(b) Thesis, Dissertation, Chapters in Books 1. Hoberg EP. Studies of seabird parasites from Ugaiushak Island. A preliminary review. Part 5. In: Population dynamics and trophic relationships of marine birds in the Gulf of Alaska and southern Bering Sea. NOAA-OCSEAP 01-022-2538. JC Bartonek, CJ Lensink, PJ Gould, RE Gill, GA Sanger co-principal investigators. U.S. Fish and Wildlife Service 1977; 1-14.

2. Wehle DHS, Hoberg EP. Studies on marine birds on Ugaiushak Island. Part 4. In: Population dynamics and trophic relationships of marine birds in the Gulf of Alaska and southern Bering Sea. NOAA-OCSEAP 01-022-2538. JC Bartonek, CJ Lensink, PJ Gould, RE Gill, GA Sanger co- principal investigators. U.S. Fish and Wildlife Service 1977; 1-78.

3. Nysewander D, Hoberg EP. The breeding biology of marine birds associated with Chiniak Bay, Kodiak Island, Alaska 1977. pp. 525-574. In: Environmental Assessment of the Alaskan Continental Shelf, Annual Reports of the Principal Investigators, Vol. 3 National Oceanographic Atmospheric Administration Environmental Research Laboratory, Boulder, Colorado 1978.

4. Hoberg, EP. Helminth parasites of marine birds (Charadriiformes: Alcidae and Laridae) occurring in the North Pacific Ocean and Gulf of Alaska. MSc.Thesis. University of Saskatchewan, Saskatoon, Canada, 1979. 1-121.

5. Hoberg, EP. Systematics, zoogeography, and ecology of platyhelminth parasites of the seabird family Alcidae (Charadriiformes: Suborder Alcae) PhD. Dissertation. University of Washington, Seattle, 1984. 1-324.

6. Hoberg EP. Ekologiia gel'mintov morskikh ptits ostrova Talan (predvaritel'ny obzor). [Ecology of helminth parasitism among seabirds at Talan Island, USSR, a preliminary overview.] In: Prebrezhnye Ekosistemy Severnogo Okhotomor'ia Ostrov Talan. [Coastal Ecosystems in the Northern Part of the Sea of Okhotsk]. F.B. Tschernyavsky and A.Ia. Kondratiev (eds); 239 p. Institute for Biological Problems of the North. Far Eastern Branch, Russian Academy of Sciences. Magadan, Russia. 1992. pp. 116-136.

7. Hoffmann R, Siegel-Causey D, Moore JH, Murray DF, Hoberg EP. The global scientific heritage of the FSU. In: JH Moore, RS Hoffmann (eds), Sustaining Excellence in Science and Engineering in the Former Soviet Union. National Academy Press, Washington D.C. 1993. pp. 43-56.

8. Hoberg EP. Order Tetrabothriidea. In: CIP Keys to the Cestode Parasites of Vertebrates. LF Kahlil, A Jones and R Bray (eds). CAB International Institute of Parasitology and British Museum (Natural History). 1994. pp. 295-304.

9. Hoberg EP. Phylogeny and historical reconstruction: host parasite systems as keystones in biogeography and ecology. In: Biodiversity II: Understanding and Protecting Our Resources, M Reaka-Kudla, EO Wilson and D Wilson (eds). Joseph Henry Press, National Academy of Sciences, Washington, D.C. 1997; pp. 243-261.

10. Hoberg EP, Brooks DR, Siegel-Causey D. Host-parasite cospeciation: History, principles and prospects. In: Host-Parasite Evolution: General principles and avian models, D Clayton and J 22 Moore (eds). Oxford University Press. 1997. pp. 212-235.

11. Hoberg EP. Parasite biodiversity and emerging pathogens: A role for systematics in limiting the impacts on genetic resources. In: Global Genetic Resources- Access, Ownership, and Intellectual Property Rights. KE Hoagland and AY Rossman (eds). Association of Systematics Collections, Washington, D.C. 1997. pp. 71-83. 12. Lichtenfels JR, Hoberg EP, Pilitt PA. Animal parasites: US National Parasite Collection. In: Systematic Collections of the Agricultural Research Service, JR Lichtenfels, JH Kirkbride, DJ Chitwood (eds.). US Dept. of Agriculture Misc. Publ. No. 1343. 1998. pp. 1-7.

13. Hoberg EP, Kocan A, Rickard LG. Gastrointestinal strongyles in wild ruminants. In: Parasitic Diseases of Wild Mammals. W Samuel, M Pybus and A Kocan (eds). Iowa State University Press. 2001. pp. 193-227.

14. Hoberg EP, Mariaux J, Brooks DR. Phylogeny among orders of the Eucestoda (Cercomeromorphae): Integrating morphology, molecules and total evidence. In: Interrelationships of the Platyhelminthes. DTJ Littlewood and RA Bray (eds). Systematics Association Special Vol. Ser. 60. Taylor and Francis. 2001. pp. 112-126.

15. Hoberg EP. Coevolution in Marine Systems. In: Marine Parasitology, (Chapter 8, Coevolution and Speciation). K. Rohde (ed.). CSIRO, Sydney Australia. 2005. pp. 327-339.

16. Hoberg EP. Marine birds and their helminth parasites. In: Marine Parasitology, (Chapter 10, Economic, environmental and medical importance). K. Rohde (ed.). CSIRO, Sydney, Australia. 2005. pp. 414-421.

17. Hoberg EP, Brooks DR. Beyond vicariance: Integrating taxon pulses, ecological fitting and oscillation in historical biogeography and evolution. In: The Geography of Host-Parasite Interactions. S. Morand and B. Krasnov (eds.). Oxford University Press. 2010. pp. 7-20.

18. Polley L, Kutz SJ, Hoberg EP. Zoonotic parasites, our environment and change. In: Encyclopedia of Environmental Health, Volume 4. JO Nriagu (ed.). Elsevier. 2011. pp. 325-345.

19. Hoberg EP, Koehler AVA, Cook J. Complex host-parasite systems in Martes: Implications for conservation biology of endemic faunas. In: Biology and Conservation of Martens, Sables and Fishers: A New Synthesis. K.B. Aubry (ed.) Cornell University Press. 2012. pp. 39-57. [Awarded- Outstanding Academic Titles by Choice magazine for books published in 2012]

20. Hoberg EP, Brooks DR. Episodic processes, invasion, and faunal mosaics in evolutionary and ecological time. In: The Balance of Nature and Human Impact. K. Rohde (ed.). Cambridge University Press. 2013. pp. 199-213.

21. Brooks DR, Hoberg EP. The emerging infectious diseases crisis and pathogen pollution: A question of ecology and evolution. In: The Balance of Nature and Human Impact. K. Rohde (ed.). Cambridge University Press. 2013. pp. 215-229.

22. Hoberg EP, Kutz SJ, Cook JA, Galaktionov K, Haukisalmi V, Henttonen H, Laaksonen S, Makarikov A, Marcogliese DJ. Parasites in terrestrial, freshwater and marine systems. In: Arctic Biodiversity Assessment- Status and Trends in Arctic Biodiversity, H. Meltofte (ed.). Conservation of Arctic Floral and Fauna, Arctic Council, Akureyi, Iceland. 2013. pp. 476-505.

23. Cook J, Brochman C, Talbot SL, Fedorov V, Taylor E, Väinölä R, Hoberg EP, Kholodova M, Magnusun, M. Genetics. In: Arctic Biodiversity Assessment- Status and Trends in Arctic Biodiversity, H. Meltofte (ed.). Conservation of Arctic Floral and Fauna, Arctic Council, Akureyi, Iceland. 2013. pp. 514-539. 23

24. Meltofte H, Barry T, Berteaux D, Bueltmann H, Christiansen JS, Cook JA, Daniëls FJA, Dahlberg A, Friðriksson F, Ganter B, Gaston AJ, Gillespie L, Grenoble L, Henry G, Hoberg EP, Hodkinson I, Huntington HP, Ims RA, Josefson AB, Kutz SJ, Kuzmin SA, Laidre K, Lassuy DR, Lewis PN, Lovejoy C, Michel C, Mokievskiy V, Payer D, Reid D, Reist J, Tessler D, Wrona F. Status and Trends in Arctic Biodiversity- Synthesis: Implications for Conservation. In: Arctic Biodiversity Assessment- Status and Trends in Arctic Biodiversity, H. Meltofte (ed.). Conservation of Arctic Floral and Fauna, Arctic Council, Akureyi, Iceland. 2013. pp. 21-66.

25. Zarlenga D, Hoberg EP, Detwiler JT. Diversity and history as drivers of helminth systematics and biology. In: Helminth Infections and Their Impact on Global Public Health. F. Bruschi (ed.). Springer Verlag. 2014. pp. 1-28.

Hoberg EP, Zimmerman GL, Rickard LG. Nematodirus battus: A review of recent studies relative to the development of surveillance and control programs. Proceedings of the 89th Annual Meeting, United States Animal Health Association, 1985, Milwaukee, Wisconsin 1986; 420-431.

Zimmerman GL, Hoberg EP, Rickard LG, Erno JK, Wilson D, Strickland RK, Gerrish RR, Prichard WD, Knott E. Broadened geographic range and periods of transmission for Nematodirus battus in the United States. Proceedings of the 90th Annual Meeting of U.S. Animal Health Association 1986; 404-412.

(d) Other Papers, Book Reviews, Research Reports and Letters

Hoberg EP. Acceptance speech, Henry Baldwin Ward Medal. 67th Annual Meeting American Society of Parasitologists, Philadelphia, 7 August 1992. Journal of Parasitology. 1992; 78: 968-971.

American Society of Parasitologists. Results of the American Society of Parasitologists 1992 membership assessment by the prioities committee. (KD Murrell, SS Desser, TD Geary, R Grieve, EP Hoberg, P LoVerde, M Moser, DB Pence). Journal of Parasitology. 1993; 79: 798-807.

Miller S, Hoberg EP, Brown MA. Autochthonous echinococcosis in a North American horse. Journal of the American Veterinary Medical Association. 1993; 203: 1117.

Hoberg EP. Book Review: Parascript: Parasites and the Language of Evolution. D.R. Brooks and D.A. McLennan, Smithsonian Institution Press. 429 p. 1993. Journal of Parasitology. 1993; 79: 976-977.

Hoberg EP. Endorsement by Helminthological Society of Washington of Great Smokey Mountains National Park All-Taxa Biotic Inventory (ATBI). Journal of the Helminthological Society of Washington. 1998; 65: 269.

Hoberg EP, Hendrix S. A proposal: New name for the Journal of the Helminthological Society of Washington. Journal of the Helminthological Society of Washington. 1998; 65: 270.

Hoberg EP. Presentation of the ASP Distinguished Service Award to Robert L. Rausch 76th Annual Meeting, 2 July 2001, Albuquerque, New Mexico. Journal of Parasitology 2001; 87: 1259.

Galaktionov KV, Ryss AJ, Pugachev O, Pelgunov A, Kontrimavichus VL, Hoberg EP. In Memoriam- Mark Dmitrievich Sonin 1932-2010. Journal of Parasitology 2010; 96: 836-837.

Hoberg EP. Remembering Robert Rausch 1921-2012. Newsletter. Wildife Disease Association. January 2013. (http://www.wildlifedisease.org/wda/PUBLICATIONS/Newsletter.aspx).

24 Hoberg EP. Parasite biodiversity: Providing context in a changing world. The Circle. World Wildlife Fund, Global Arctic Program. 2014; Vol 2: 15-16.

Hoberg EP, Phillips, AJ. Transfer of the US National Parasite Collection. Comparative Parasitology. 2014; 81:300-301.

Hoberg EP, Phillips AJ. Transfer of the US National Parasite Collection. Systematic Parasitology. 2014. 89:1-2.

(e) Abstracts

Invited Symposia

Hoberg EP. Notes on the zoogeography of seabirds and their cestode parasites. Symposium on Significance of the Beringian Land in the Modern Biogeography of the Northern Part of the Pacific Region. Fourteenth Pacific Science Congress, Khabarovsk, Union of Soviet Socialist Republics. 1979.

Hoberg EP. Phylogeny of the Tetrabothriidae. Stunkard Centenary Session on Phylogenetic Systematics of Worms. Sixty-fourth Annual Meeting American Society of Parasitologists. Vancouver, British Columbia. 1989.

Hoberg EP. Congruent and synchronic patterns in biogeography and speciation among seabirds, pinnipeds and cestodes. Von Ihering Centenary Symposium in Parasite Biogeography and Coevolution. Sixty-sixth Annual Meeting American Society of Parasitologists. Madison, Wisconsin. 1991.

Lichtenfels JR, Hoberg EP. Systematics of nematodes that cause ostertagiasis: An update. Second Ostertagia Workshop. University of Maryland, College Park. March 1992.

Hoberg, EP. Acceptance speech, Henry Baldwin Ward Medal. American Society of Parasitologists, 67th Annual Meeting, Philadelphia, 7 August 1992.

Lichtenfels JR, Pilitt PA, Hoberg EP. Planning a second century for the U.S. Parasite Collection. Workshop on the Future of Parasite Collections. Sixty-seventh Annual Meeting American Society of Parasitologists. Philadelphia, Pennsylvania. August 1992.

Hoberg EP. Species and the phylogenetic perspective. Presidents Symposium, Southeastern Society of Parasitologists and Association of Southeastern Biologists. Virginia Beach, Virginia. April 1993.

Hoberg EP. Parasite-host systems: Foundations for comparative studies in biodiversity. Symposium on Comparative evolution in host-parasite systems (co-speciation, host switching, evolutionary rates). 4th Congress of the European Society of Evolutionary Biology. Montpellier, France. August 1993.(Symposium Co-Organizer/ Chair).

Hoberg EP. The roles of parasitology in biodiversity studies. Biodiversity Symposium. American Society of Parasitologists. Atlanta, Georgia. October 1993. (Symposium Co-Organizer/ Chair).

Hoberg EP. Phylogeny and historical reconstruction: host-parasite systems as keystones in biogeography and ecology. In: Biodiversity: From 1986 to the 21st Century. Inaugural Symposium, Consortium for Systematics and Biodiversity. Smithsonian Institution, Washington, D.C. November 1993.

Hoberg EP. Biodiversity and parasitology, new concepts and perspectives. Biodiversity Workshop, American Society of Parasitologists, 69th Annual Meeting, Fort Collins, Colorado. August 1994. (Workshop Organizer).

25 Hoberg EP. 1994. Systematics, coevolution and biogeography, unifying concepts in parasitology. Science Symposium, Scientist of the Year Awards. Agricultural Research Service. November 1994.

Lichtenfels JR, Hoberg EP, Zarlenga DS. Systematics of gastrointestinal nematodes of domestic ruminants: Advances 1992-1995. Fourth Ostertagia Workshop. University of Maryland. 5-8 March 1996.

Hoberg EP. Parasite biodiversity and emerging pathogens: A role for systematics in limiting the impacts on genetic resources. In: Global Genetic Resources- Access, Ownership, and Intellectual Property Rights. Beltsville Symposium XXI. Agricultural Research Service, US Department of Agriculture, Beltsville, Maryland, May 1996.

Hoberg EP. Faunal diversity among avian parasite assemblages: The interaction of history ecology and biogeography. In: Parasites and Ecology of Marine and Coastal Birds. Special Symposium, Scandinavian Society of Parasitology, Stykkishólmur, Iceland, June 1996.

Hoberg EP, Mariaux J, Justine J-L, Gulyaev V, Vaucher C. Phylogeny of the Eucestoda: Relationships among the orders. 2nd International Workshop for Tapeworm Systematics, University of Nebraska State Museum, Lincoln, Nebraska, October 1996.

Hoberg EP, Jones A, Gardner SL, Rausch RL, Eom KS. Phylogenetic relationships among species of Taenia. 2nd International Workshop for Tapeworm Systematics, University of Nebraska State Museum, Lincoln, Nebraska, October 1996.

Hoberg EP. Biodiversity and historical biogeography of nematode faunas in Holarctic ruminants. In: Holarctic Mammals, with R. Hoffmann and V. Neronov. US-Russia Area V. 25th Anniversary Conference, US Fish and Wildlife Service. Trinity College, Washington, D.C., June 1997.

Hoberg EP, Zarlenga D. The species question in Echinococcus and Taenia: Taxonomic controversies. Special Symposium, Advances in Cestode Research, American Association of Veterinary Parasitologists, Reno, Nevada. July 1997.

Hoberg EP, Mariaux J. Phylogeny of the Eucestoda, morphological and molecular perspectives. Zoologica and Botanica '98. Museum of Natural History, Geneva, Switzerland, February 1998.

Hoberg EP. Nematode phylogeny and comparative morphology: Lessons on the evolution of structure. Nematode Evolution Symposium. American Society of Parastiologists, 74th Annual Meeting and Society of Nematology, Monterey, California, July 1999.

Hoberg EP, Mariaux J, Brooks DR, Georgiev B. A phylogeny for the Eucestoda: Ordinal relationships integrating morphology, molecules and total evidence. In: Interrelationships of the Platyhelminthes. Linnean Society of London, Systematics Association, British Society of Parasitology, London, UK, July 1999.

Lichtenfels JR, Hoberg EP. Preliminary morphological phylogeny of Strongyloidea of horses. WAAVP Workshop- Systematics of the Cyathostominae of Horses. World Association for Advancement of Veterinary Parasitology, 17th International Conference, Copenhagen Denmark. 15-19 August 1999.

Hoberg EP. A parasite biosystematics perspective. Agricultural Research Service, USDA. Animal Parasite Genomics Workshop. Beltsville, MD. 23 May 2000.

Hoberg EP, Schauff M. USDA Collections at the interface of agriculture and the environment. Association of Systematics Collections, National Conference. Symposium, Information, Technology, and 26 Partnerships: Natural History Collections in the New Millennium. Baltimore, Maryland. 13-15 May 2000.

Hoberg EP. Phylogeny, history and biodiversity: Understanding faunal structure and biogeography in the marine realm. In: Ecological Parasitology on the Turn of the Millennium. Zoological Institute, Russian Academy of Sciences & Scandinavian Society for Parasitology, St. Petersburg, Russia. July 2000.

Hoberg EP. Biodiversity and Biogeography: Establishing the context and keystones for Arctic parasitology. General Introduction. International Workshop for Arctic Parasitology, Prince Albert National Park, Saskatchewan, Canada. October 2000. Kutz S, Hoberg EP, Polley L. Global warming and host-parasite systems in the Arctic: Should we be concerned? Issues Presentation. International Workshop for Arctic Parasitology, Prince Albert National Park, Saskatchewan, Canada. October 2000.

Jenkins E, Hoberg EP, Kutz S, Veitch A, Polley L. Challenges in investigating protostrongylids in northern hosts: Parelaphostrongylus odocoilei in Dall’s sheep (Ovis dalli dalli) of northern North America. Issues Presentation. International Workshop for Arctic Parasitology, Prince Albert National Park, Saskatchewan, Canada. October 2000.

Hoberg EP. Presentation of the ASP Distinguished Service Award to Robert L. Rausch. 76th Annual Meeting American Society of Parasitologists, 2 July 2001, Albuquerque, New Mexico.

Hoberg EP. Foundations for an integrative parasitology: collections archives and biodiversity informatics. In: Symposium- Parasitology in Science and Society. Helminthological Society of Washington, 676th Meeting, and Smithsonian Institution, Washington D.C. 27 October 2001.

Hoberg EP. Host-parasite systems and global climate change: models from the Arctic. Beltsville Area Research Center, Workshop on Global Climate Change. February 2002.

Cook J, Galbreath K, Runck A, Fedorov V, Hoberg EP. Beringian Coevolution Project: An interface between mammalogy and parasitology. In: Biogeography and Evolution in Northern Host-Parasite Systems: A Symposium Honoring Robert L. Rausch and Virginia R. Rausch. International Congress of Parasitology X. Vancouver, Canada. 4-9 August 2002.

Hoberg EP. Colonization and diversification: Historical and coevolutionary trajectories among cestodes, cetaceans and pinnipeds. In: Symposium for Parasites of Marine Mammals. International Congress of Parasitology X. Vancouver, Canada. 4-9 August 2002.

Hoberg EP, Kutz SJ, Cook J. Arctic biodiversity: From discovery to faunal baselines- revealing the history of a dynamic ecosystem. SubPlenary Lecture, In: Parasitology and Biodiversity in an Age of Discovery. International Congress of Parasitology X. Vancouver, Canada. 4-9 August 2002.

Hoberg EP. Robert and Virginia Rausch: Fifty years on the frontiers of northern parasitology. In: Biogeography and Evolution in Northern Host-Parasite Systems: A Symposium Honoring Robert L. Rausch and Virginia R. Rausch. International Congress of Parasitology X. Vancouver, Canada. 4-9 August 2002.

Hoberg EP, Alkire N, de Queiroz A, Jones A. Phylogeny of Taenia tapeworms: Insights about the origins of species in humans. In: Symposium on Taenia asiatica and cysticercosis. International Congress of Parasitology X. Vancouver, Canada. 4-9 August 2002.

Hoberg EP. Hominids, carnivores and Taenia tapeworms: Revealing historical associations for parasites and pathogens in humans.In: Biologie des cestodes: recherches actuelles. IIIe Cycle Romand en Sciences Biologiques. Muséum d’Histoire naturelle de Geneve. 1-3 October 2002.

Hoberg EP. Old worms in new hosts? Radiation of the Tetrabothriidea in marine birds and mammals. In: 27 Biologie des cestodes: recherches actuelles. IIIe Cycle Romand en Sciences Biologiques. Muséum d’Histoire naturelle de Geneve. 1-3 October 2002.

Kutz SJ, Hoberg EP. Stress in the Arctic- Emerging parasitic infections of arctic ungulates. In: Biology of the Canadian Arctic: A crucible for change in the 21st Century. Society of Integrative and Comparative Biology. Toronto, Canada. 4-9 January 2003.

Jenkins E, Veitch AM, Appleyard GD, Hoberg EP, Kutz SJ, Polley L. Geographic distribution and seasonal patterns of larval shedding of the muscle-dwelling nematode Parelaphostrongylus odocoilei in thinhorn sheep from northern North America. 15th Annual Arctic Forum, Arctic Research Consortium of the United States. 28-29 April 2003. (ARCUS Award for Research Excellence- Life Sciences).

Hoberg EP. Collections, archives and informatics: A new synergy for tracking history and biodiversity from species to populations. International Workshop for Arctic Parasitology II. Arctic Centre, University of Lapland, Rovaniemi, Finland. 1-7 September 2003.

Kutz SJ, Hoberg EP, Nagy J, Elkin B, Veitch A, Jenkins E, Polley L. Emerging parasitic infections and diseases of Arctic ungulates. International Workshop for Arctic Parasitology II. Arctic Centre, University of Lapland, Rovaniemi, Finland. 1-7 September 2003.

Koehler A, Hoberg EP. Telford S, Henttonen H, Galbreath K, Runck A, Kutz S, MacDonald S, Waltari E, Cook J. The Beringian Coevolution Project. International Workshop for Arctic Parasitology II. Arctic Centre, University of Lapland, Rovaniemi, Finland. 1-7 September 2003.

Hoberg EP. Structuring the biosphere: Episodic host switching and geographic colonization as determinants of diversification in complex host-parasite systems. International Biogeography Society- Symposium: Biogeography of Parasites and Pathogens. US NCTC, Shepherdstown, WVA. January 2005.

Hoberg EP. Phylogeny of Taenia: Defining species and understanding origins of human parasites. In “Taeniasis/Cysticercosis and Echinococcosis International Symposium with a Focus on Asia and the Pacific;” and Third Congress of Federation of Asian Parasitologists Focused on Cestode Zoonoses. Asahikawa, Japan. 5-8 July 2005.

Hoberg EP, Cook J, Kutz S, Jenkins E, Polley L. Beringian biogeography: Ungulates, parasites and taxon pulses. “Mammal parasite diversity- Aspects of Scale, Diversity and Coevolution of Host-Parasite Assemblages.” International Mammalogical Congress 9, Sapporo, Japan 31 July- 5 August 2005.

Cook J, Hoberg EP, Henttonen H, Wickström L, Haukisalmi V, Galbreath K. Reflections on a northern host/ parasite biota: Beringia. “Mammal parasite diversity- Aspects of Scale, Diversity and Coevolution of Host-Parasite Assemblages.” International Mammalogical Congress 9, Sapporo, Japan 31 July- 5 August 2005.

Kutz SJ, Hoberg EP. Northern host-parasite systems: Understanding invasions and emergence of ungulate parasites in space and time. Symposium-“Biology of the Canadian Arctic II Integrating Across Scales: From Site Specific to Regional and Global Scales. Canadian Society of Zoologists, 45th Annual Meeting, Edmonton, Alberta. 2-6 May 2006.

Hoberg EP, Cook J. A macroevolutionary mosaic: Episodic processes and history of northern faunas. In: Northern Parasite Faunas: Using the Past to Understand the Present and Future. International Workshop for Arctic Parasitology III. Calgary, Alberta, 6-10 November 2006. (keynote).

Hoberg, EP. Parasites, pathogens and model archives: Exploring ecosystems in dynamic change. Helminthological Society of Washington, 659th Annual Meeting. George Washington University. 4-5 May 28 2007. (keynote).

Hoberg EP, Rosenthal B. Climate change and ecological structure: Building baselines and biodiversity archives. Climate Change and Infectious Diseases of Wildlife in Arctic and Mountainous Regions (Workshop). National Veterinary Institute, Tromso, Norway. 23-24 April 2007.

Hoberg EP, Polley L, Kutz S, Jenkins E, Elkin B, Veitch A. Climate and pathogens: Consequences for human health and subsistence food chains across high latitudes of the North. Climate Change and Impacts on Human Health in the Arctic: An International Workshop on Emerging Threats and Response of Arctic Communities to Climate Change. Alaska Forum On the Environment. Anchorage, Alaska. 13-15 February 2008.

Kutz SJ, Hoberg EP, Veitch AM, Polley L. Arctic parasitology and climate change: Perspectives from the Canadian Arctic. Dutch Wildlife Disease Society. April 2008. (keynote).

Polley L, Hoberg EP, Kutz SJ. Climate change, parasites and shifting boundaries. NkVet Symposium 2008 (Nordic Committee for Veterinary Scientific Cooperation)- Parasitic Infections of Domestic Animals in the Nordic Countries – Emerging Threats and Challenges. Finnish Food Safety Authority Evira Headquarters, Mustialankatu, Finland. September 2008.

Kutz S, Curry P, deBruyn N, Ducrocq J, Dumond M, Elkin B, Hoberg E, Lair S, Orsel K, Polley L, Seegers K, Spans R, Torretti L, Veitch A. Climate change and parasitism in Arctic and Subarctic terrestrial mammals. Wildlife Society, Alaska Chapter, Annual Meeting. April 2009. (plenary).

Hoberg EP. Parasites in a changing landscape: A global perspective on history, connectivity and consequences. WAAVP 2009 Theme Symposium: One World, One Health: Parasites in a Changing Landscape. 22nd Conference, World Association for Advancement of Veterinary Parasitology. Calgary, Alberta. 8-13 August 2009. (invited theme lecture).

Hoberg EP, Koehler A, Cook JA. Exploring complex host-parasite systems in Martes: Historical biogeography and coevolutionary processes. Biology and Conservation of Martens, Sables and Fishers: A New Synthesis. 5th International Martes Symposium. University of Washington, Seattle. 8-12 Sept. 2009. (invited keynote).

Cook JA, Hoberg EP, MacDonald SO, Hope AG, Dawson SG, Fedorov V. Beringia: History and climate structure northern assemblages of mammals and parasites at the nexus of Asia and North America. American Society of Parasitologists, Special Symposium. Arctic Passages: Explorations of History, Diversity and Dynamic Change. American Society of Parasitologists, 86th Annual Meeting, Anchorage, Alaska. 1-4 June 2011.

Galbreath KE, Hoberg EP. A worm with a view: parasite perspectives on Beringian biogeography. American Society of Parasitologists, Special Symposium. Arctic Passages: Explorations of History, Diversity and Dynamic Change. ASP, 86th Annual Meeting, Anchorage, Alaska. 1-4 June 2011.

Hoberg EP, Adams A. Presentation of the ASP Eminent Parasitologist Award to Robert L. Rausch. ASP, 86th Annual Meeting, Anchorage, Alaska. 1-4 June 2011.

Hoberg EP. Arctic Passages: Explorations of History, Diversity and Dynamic Change. Closing Summary. American Society of Parasitologists, 86th Annual Meeting, Anchorage, Alaska. 1-4 June 2011.

Hoberg EP, Galbreath KE, Makarikov AA, Brooks DR. Revealing History: Integrated Pathways Exploring the Biosphere. Parascript in the 21st Century: Climate Change, Biodiversity and Emerging Disease. 29 International Symposium and Workshop sponsored by the HW Manter Laboratory/ Cedar Point Biological Station, University of Nebraska. 5-6 September 2012. (keynote)

Hoberg EP. What is the US National Parasite Collection, And Why is it Important? 43rd Rocky Mountain Conference for Parasitology. Cedar Point Biological Station, Nebraska. 7-8 September 2012.

Ito A, Nakao M, Yanagida T, Okamoto M, Sako Y, Carod J-F, Hoberg EP. Taenia solium: genetic diversity, history of its global expansion, and outcrossing. Australian Society for Parasitology, Canberra, Australia. 30 June- 3 July 2014. Hoberg EP, Kutz SJ, Galbreath KE, Cook JA. An Integrated Perspective on Pathogens of the Arctic. In: Symposium- Pathogen Diversity in the Arctic Arena: Implications for Ecosystem Sustainability and Human Health Under Accelerating Change. Arctic Biodiversity Congress/ 2-4 December 2014, Trondheim, Norway.

Kutz SJ, Branigan M, Checkley S, Dobson A, Elkin B, Forde T, Hoberg EP, Kafle P, Leclerc L, Stasiak I, Tomaselli M. Is changing pathogen biodiversity in muskoxen influencing human health, economies, socio- cultural well-being, and wildlife conservation? In: Symposium- Pathogen Diversity in the Arctic Arena: Implications for Ecosystem Sustainability and Human Health Under Accelerating Change. Arctic Biodiversity Congress/ 2-4 December 2014, Trondheim, Norway.

Cook JA, Hoberg EP, Galbreath K, Greiman S, Haukisalmi V, Hentonnen H, Hope A, Kutz S, Makarikov A, Talbot S, Tkach V, Tsevetkova A, Yanagihara R. The Beringian Coevolution Project: A case study for understanding the exchange and diversification of high latitude mammals and their pathogens In: Symposium- Pathogen Diversity in the Arctic Arena: Implications for Ecosystem Sustainability and Human Health Under Accelerating Change. Arctic Biodiversity Congress/ 2-4 December 2014, Trondheim, Norway.

Galbreath KE, Hoberg EP, Cook JA. Looking back to look forward: What Beringia’s past may tell us about its future. In: Symposium- Biodiversity of Beringia: past, present and future. Arctic Biodiversity Congress/ 2-4 December 2014, Trondheim, Norway.

Contributed Papers (with abstracts= 105)

Professional Presentations and Lectures, Invited- (without abstracts= 90)

9. CONSULTATION/ INVITED WORKSHOPS/ SYMPOSIA:

Merck, Sharp and Dohme AGVET, Third Parasite Control Workshop. Chateau Montebello, Quebec. 15- 16 January 1990.

Panarctic Biota Project, First planning session for Invertebrate Biodiversity Research: 2-8 February 1992, Russian Academy of Sciences, Zoological Institute, St. Petersburg, Russia.

Agricultural Research Service, USDA, Biosystematics and Biodiversity Team to Russia, for development and implementation of cooperative research programs. September 1992.

Comparative Evolution in Host-Parasite Systems, 4th Congress of the European Society for Evolutionary Biology, Montpellier, France. August 1993. Symposium Co-Organizer/Chair.

First International Workshop on Systematic Zoology of Tapeworms. Museum of Natural History, Geneva, Switzerland. 30-31 August 1993.

Biodiversity Symposium. American Society of Parasitologists. 68th Annual Meeting, Atlanta, 30 Georgia. October 1993. (Symposium Co-Organizer/ Chair).

Consortium for Systematics and Biodiversity. Inaugural Symposium, Biodiversity: From 1986 to the 21st Century. Smithsonian Institution, Washington, D.C. November 1993.

International Science Foundation. International Science Foundation and the National Academy of Sciences Commission on Life Sciences, Biology III (Zoology) Washington D.C. 14-16 December 1993; 19-22 July 1994.

Biodiversity Workshop, American Society of Parasitologists. 69th Annual Meeting, Fort Collins, Colorado. August 1994. (Workshop Organizer).

US-Russia Environmental Protection Agreement, Area V Working Group, US Fish and Wildlife Service, Arlington, Virginia, 7-11 February 1994; February 1996.

National Science Foundation. Systematic Biology, Special Competition: Partnerships for Enhancing Expertise in Taxonomy (PEET) Advisory Panel, Arlington, VA., 17-19 May 1995.

International Workshop for Tapeworm Systematics II. University of Nebraska, Lincoln. 2-6 October 1996. Organizer and Chair (Grant support provided through- American Society of Parasitologists, Agricultural Research Service, Foreign Agriculture Service, U. S. Civilian Research and Development Foundation).

Parasitic Nematodes of Domestic Animals, Diagnostics Workshop. Merck, Sharp and Dohme, AGVET, Animal Science Research. Rockville, Maryland. 11 December 1996.

US-Russia Joint Commission on Economic and Technical Cooperation (Gore-Primakov Commission) 10th Meeting Science and Technology Committee, US Delegate. 1999. [And Arctic Working Group for GPC.]

Interagency Arctic Research Policy Committee (IARPC) (USA). Representative for Agricultural Research Service. 1998- 2001.

WAAVP Workshop- Systematics of the Cyathostominae of Horses. World Association for Advancement of Veterinary Parasitology, 17th International Conference, Copenhagen, Denmark. 15-19 August 1999.

Agricultural Research Service, USDA, BARC. Beef Cattle Research Field Day. 6 October 1999.

National Science Foundation. Division of Environmental Biology: Biotic Surveys and Inventories Advisory Panel, Arlington, VA., 10-11 February 2000.

Agricultural Research Service, USDA. Invasive Species Symposium. Beltsville, MD. 15 February 2000.

U. S Civilian Research and Development Foundation (CRDF): 1999-2000 Cooperative Grants Program- Biological & Biomedical Sciences and Engineering Review Panel. Washington, D.C., 13-14 March 2000.

Association of Systematics Collections: Information, Technology, and Partnerships: Natural History Collections in the New Millennium. National Conference. 13-14 May 2000. Baltimore, Maryland. Meeting Planning Committee.

Agricultural Research Service, USDA. Animal Parasite Genomics Workshop Beltsville, MD. 23 May 2000.

31 National Science Foundation, Division of Environmental Biology, Systematic and Population Biology Cluster. Committee of Visitors (COVS). Arlington, Virginia. 19-22 July 2000.

1st International Workshop for Arctic Parasitology (IWAP): Prince Albert National Park, Canada. Co- Organizer. Issues Session Co-Chair; Summary of Issues. 1-4 October 2000.

U. S Civilian Research and Development Foundation (CRDF): 2001 Cooperative Grants Program- Biological & Biomedical Sciences and Engineering Review Panel. 26-27 September 2001.

Beltsville Area Research Center Workshop. Global Climate Change. February 2002.

International Congress of Parasitology X. Scientific Program Committee. Co-Organizer, Plenary- Parasitology & Biodiversity in an Age of Discovery. Organizer, and Co-Chair, Biogeography & Evolution in Northern Host-Parasite Systems, A Symposium Honoring Robert L. Rausch and Virginia R. Rausch. Vancouver Canada. August 2002.

Biologie des Cestodes: Recherches actuelles. IIIe Cycle Romand en Sciences Biologiques. Muséum d’Histoire naturelle de Geneve. 1-3 October 2002. (advanced course for graduate students).

Assembling the Tree of Life- Nematode Phylogeny Workshop. University of New Hampshire. July 2003. Chair- Strongyle Phylogeny Working Group in NEMATOL.

2nd International Workshop for Arctic Parasitology. Rovaniemi, Finland, 1-7 September 2003. Organizing Committee (2002-2003). Chair- Opening session. Chair- Discussion on Collections.

National Scientific Leadership Conference, Agricultural Research Service. New Orleans, 21-23 January 2004.

Federal Interagency Committee on Invasive Terrestrial Animals and Pathogens (ITAP). Planning Retreat. National Conservation Training Center. Shepherdstown, West Virginia. 22-24 June 2004. Chair- Systematics Subcommittee, 2004- 2006.

3rd International Workshop for Arctic Parasitology. Calgary, Alberta, 6-9 November 2006. Organizing Committee (2005-2006).Chair- Session II- Northern Parasite Faunas: What Can the Past Tell Us About the Present and Future?

Climate Change and Infectious Diseases of Wildlife in Arctic and Mountainous Regions. National Veterinary Institute, Tromso, Norway. 23-24 April 2007.

CircumArctic Rangifer Monitoring & Assessment Network (CARMA). International Polar Year. Research Protocols and Planning Meeting. University of Calgary, Alberta, Canada. 7-8 June 2007.

Smithsonian Institution Global Earth Observatories (SEIGO). Center for Tropical Forest Science Workshop. Monitoring Vertebrates and Vertebrate-Ecosystem Interactions. Center for Species Survival and Conservation Research & Smithsonian National Zoological Park. Front Royal, Virginia. 29-31 October 2007.

Climate Change and Impacts on Human Health in the Arctic: An International Workshop on Emerging Threats and Response of Arctic Communities to Climate Change. Alaska Forum On the Environment. Anchorage, Alaska. 13-15 February 2008.

WAAVP, One World, One Health: Parasites in a Changing Landscape. 22nd Conference World Association for Advancement of Veterinary Parasitology. Calgary, Alberta. 8-13 August 2009.

Biology and Conservation of Martens, Sables and Fishers: A New Synthesis. 5th International 32 Martes Symposium. University of Washington, Seattle, WA. 8-12 September 2009.

Research Group for Arctic Parasitology, Research Workshop. Ressor Ranch, Cypress Hills Interprovincial Park, Saskatchwan/Alberta, Canada. 14-16 May 2010. Discussion Leader- invasive species and processes.

Advancing Integration of Museums into Undergraduate Programs (AIM UP). NSF Research Coordination Network. Santa Fe, NM. 15-16 October 2010.

American Society of Parasitologists, Special Symposium. Arctic Passages: Explorations of History, Diversity and Dynamic Change. ASP Annual Meeting, Anchorage, Alaska. 1-4 June 2011. Organizer and Chair.

Conservation of Arctic Flora and Fauna (CAFF). Arctic Biodiversity Assessment Authors Workshop. Biosphere Museum, Environment Canada, Montreal, Canada, 21-22 April 2012. Writing workshop in advance of the International Polar Year 2012 conference: “From Knowledge to Action.” Lead author for “Parasites in Terrestrial, Freshwater and Marine Systems” and contributor to “Genetics” and to “Arctic Biodiversity Synthesis and Recommendations.”

Parascript in the 21st Century: Climate Change, Biodiversity and Emerging Disease. An international Symposium and Workshop sponsored by the HW Manter Laboratory. Cedar Point Biological Station, University of Nebraska. 5-6 September 2012. Co-Organizer w/ S.L. Gardner and D.R. Brooks.

FDA, Center for Veterinary Medicine and ARS Joint Meeting on Anthelmintic Resistance. Invited Presentation: Hoberg EP. US National Parasite Collection- A Portal for Biodiversity Informatics. Beltsville, MD., 26 April 2013.

National Science Foundation, Collections Web RCN: Building a National Community for Natural History Collections. Collections Web Stakeholders Workshop. Invited representative of the US National Parasite Collection; presentations and discussions on the development of informatics resources and digitization of natural history collections. Smithsonian Institution NMNH, 1-3 May 2013.

Scientific Collections International: Engaging Scientific Collections in Emerging Infectious Disease Research. Invited Workshop. Prediction and Monitoring Panel. Smithsonian Insitution, National Museum of Natural History. 23-24 October 2014.

Arctic Biodiversity Congress, Conservation of Arctic Flora and Fauna (CAFF). Symposium: Pathogen Diversity in the Arctic Arena: Implications for Ecosystem Sustainability and Human Health Under Accelerating Change- Symposium Organizer, invited lectures. Symposium: Biodiversity of Beringia: Past, Present and Future- Symposium Co-organizer. Trondheim, Norway, 2-4 December 2014.

10. RESEARCH:

(a) Grants received:

Arctic Institute of North America. Grant in Aid for studies of helminth parasites of Charadriiformes in the Chaunsk Gulf Region, USSR. E.P. Hoberg. 1981. $1500.

Arctic Institute of North America. Grant in Aid for studies of helminth parasites of Alcidae in Alaska. E.P. Hoberg. 1982. $1500.

National Science Foundation. A study of ecological relationships, zoogeography and systematics of helminth parasites in antarctic seabirds (DPP 8115975). R.L. Rausch, E.P. Hoberg. 1982-1984. 33 $32,500.

Oregon State University, College of Veterinary Medicine, USDA Formula 1433 Funds. Culture and electrophoretic studies of Nematodirus battus. E.P. Hoberg, G.L. Zimmerman. 1985. $6,500.

Merck and Co. Clorsulon/Ivermectin clinical field trial/productivity trial/parasite epidemiology. G.L. Zimmerman, E.P. Hoberg. 1986. $31,777.

Merck and Co. Efficacy of Ivermectin against Nematodirus battus. G.L. Zimmerman, E.P. Hoberg. 1986. $5,000.

Merck and Co. Effects of Ivermectin sustained release bolus on production of cattle. G.L. Zimmerman, E.P. Hoberg. 1986. $26,071. Hoechst. Efficacy of Fenbendazole against Nematodirus battus. G.L. Zimmerman, E.P. Hoberg. 1986. $9,000.

American Cyanamid. Efficacy of Levamisole against Nematodirus battus. G.L. Zimmerman, E.P. Hoberg. 1986. $4,000.

Oregon State University, General Research Fund. Evolution and phylogenetic relationships of Nematodirus spp. (Trichostrongyloidea) in North America. E.P. Hoberg, G.L. Zimmerman. 1986-87. $3,184.

Merck and Co. Efficacy of sustained release Ivermectin against inhibited Ostertagia. G.L. Zimmerman, E.P. Hoberg. 1987. $24,600.

American Cyanamid. Anthelmintic efficacy of a new compound (nemadectin) against inhibited Ostertagia. G.L. Zimmerman, E.P. Hoberg. 1987. $26,000.

Merck and Co. Influence of sustained release Ivermectin on productivity in calves. G.L. Zimmerman, E.P. Hoberg. 1987. $51,281.

Agricultural Research Foundation. Prevention of losses to the sheep industry by Nematodirus battus, an agent of foreign animal disease. G.L. Zimmerman, E.P. Hoberg. 1987. $6,957.

Oregon State University, College of Veterinary Medicine, USDA Formula 1433 Funds. Epizootiology of Nematodirus battus. E.P. Hoberg, G.L. Zimmerman. 1987-1988. $10,000.

American Cyanamid. Efficacy of a new compound (injectable oxydectin) against inhibited Ostertagia. G.L. Zimmerman. E.P. Hoberg. 1988. $40,560.

Natural Sciences and Engineering Research Council of Canada. Population Biology. Helminth faunas of seabirds and marine mammals: model systems for phylogenetic reconstruction, coevolution and historical ecology. E.P. Hoberg. 1990- 1993. $75,000.

National Science Foundation. The Beringian Coevolution Project. (DEB BS&I 0196095) J. Cook, E.P. Hoberg, S. Telford III. 1999- 2003 (with extension). $240,000.

Civilian Research Development Foundation (CRDF) Cooperative Grants Program. Comparative Morphology, Phylogenetic Analyses, and Classification of Strongylid Nematode Parasites of Equidae. J.R. Lichtenfels, V. Kharchenko, E.P. Hoberg.2001-2003. $80,000.

ARS-Former Soviet Union Scientific Cooperation Program. Population Ecology and Systematics of the Family Protostrongylidae, the Parasites of Bovidae. Collaborative Partnership Grant: Institute of 34 Zoology, Academy of Sciences, Uzbekistan & US National Parasite Collection. 2002-2005. $340,151.

National Science Foundation. The Beringian Coevolution Project II (DEB BS&I 0415668) J. Cook, E.P. Hoberg. 2004-2007. $502,000.

Western Northwest Territories Biophysical Study. Defining Parasite Biodiversity in Barren-Ground Caribou (An apparently new nematode parasite: Identification and geographic distribution in the Northwest Territories). S.J. Kutz (PI., University of Saskatchewan)/ E. Jenkins, G. Appleyard, B. Elkin, E. Hoberg, L. Polley (CoPI’s.). 2004-2006. $80,000. National Science Foundation. (BRC). Integration and Curation of the Robert and Virginia Rausch Helminthological Collection-A Resource for Science and Society in the MSB Division of Parasitology. J.A. Cook, PI (University of New Mexico), G. Jarrell, S. Loker, E.P. Hoberg (Co-PIs). 2011-2014. $489,490. (Division of Biological Infrastructure, Biological Research Collections, 1057383).

National Science Foundation. Collaborative Research: RUI---- Integrated Inventory of Biomes of the Arctic. J.A. Cook (PI) (University of New Mexico), E.P. Hoberg (Co-PI) (USNPC, USDA), K.E. Galbreath (Co-PI) (Northern Michigan University), E. DeChaine (CoPI) (Western Washington University). 36 months. 2013-2016. $945,000. (DEB- Biodiversity Discovery and Analysis- 1258010-1256832- 1256493).

(b) Consultant/ Collaborator on Extramural Grants:

National Science Foundation. Molecular Systematics of Ascaridoid Nematodes: A Phylogenetic Framework for Testing Hypotheses of Life History and Morphological Evolution. S.A. Nadler, Principle Investigator (Department of Biological Sciences, Northern Illinois University)/ E.P. Hoberg, Consultant. 1992-1995. $180,000.

National Research Council, Canada. Climate Change Impacts & Adaptations Directorate. “An Evaluation of the Role of Climate Change in the Emergence of Pathogens and Diseases in Arctic and Subarctic Ungulate Populations.” S.J. Kutz (PI., University of Saskatchewan)/ E.P. Hoberg Consultant. 2004-2006. $179,700.

Alberta Ingenuity Fund. “Impacts of Climate Change on the Biodiversity, Epidemiology, and Impacts of Parasites in Caribou.” S.J. Kutz (PI, University of Calgary)/ E.P. Hoberg et al., collaborator and consultant. 2007-2009. $300,000.

US Geological Survey (Alaska Area Regional Office)/ Science on the DOI Landscape Initiative. “A Molecular Framework for Predicting the Responses of High Latitude Vertebrates and Associated Pathogens to Climate Change.” S. Talbot (PI., Alaska Science Center, USGS). 2009-2011. $300,000.

National Science Foundation, Research Coordination Network (RCG, UBE)- Advancing Integration of Museums into Undergraduate Programs (AIM UP!). J.A. Cook (PI), E.P. Hoberg and others (Senior Personnel). 2010, February, 60 months. $485,429.

Stockholm Paradigm- Integrating taxon pulses/ecological fitting/ explanations of faunal assembly/ history Parasite biodiversity assessment. Historical biogeography, host-parasite coevolution and systematics. Climate, episodic processes and diversity, Invasive processes in evolutionary and ecological time. Systematics and phylogeny of Stongylate nematodes in Artiodactyla. Integrated comparative methods (morphology/ molecular) for differentiation of cestodes and nematodes. Phylogeny and evolution of the Eucestoda (systematics of Tetrabothriidea & Cyclophyllidea).

(d) Field Research/ Collections/ Integrated Biodiversity Survey & Inventory: Parasitology, ornithology, mammalogy

Aleutian Islands and Gulf of Alaska. April-October 1975, January-August 1976; May-August 1977. Chaunsk Gulf, Chukotka, Russia. July-August 1981. Bering Sea (St. Matthew Island). July-August 1982. Antarctica (Palmer Station, Antarctic Peninsula). December 1982-February 1983. Washington State, Marine collections, September 1982; August, September, 1985; August 1987. Washington State, Amphibian collections, July-November 1984. Talan Island, Sea of Okhotsk, Russia. July-September, 1988. Coppermine, Northwest Territories, Canada. April 1994. Kugluktuk, Nunavut, Canada. May 1998. Norman Wells, Mackenzie Mountains, NWT, Canada. April 1999. Cantwell, Alaska. September 1999. Wrangell-St. Elias Mountains, Alaska (Beringian Coevolution Project). July-August 2001. Finnish Lapland, Pallasjärvi and Kilpisjärvi, Finland. May-June 2004. Southeastern Alaska (Beringian Coevolution Project). July 2006. Alaska, Yukon, Northwest Territories (Integrated Inventory of Biomes of the Arctic). July-August 2013. New Mexico and Idaho (Integrated Inventory of Biomes of the Arctic). June 2014.

Professional Affiliations:

Museum Research Associate (Zoology), Burke Memorial, Washington State Museum, July 1985- 2001.

International Panarctic Biota Project, Leader for the Working Group of Invertebrates, University of Kansas, Smithsonian Institution, Russian Academy of Sciences, October 1991-1995.

Museum Research Affiliate, Mammalogy, University of Alaska Museum, Fairbanks, November 1996- 2000.

Museum Research Affiliate, Parasitology, Harold W. Manter Laboratory, Nebraska State Museum, University of Nebraska, Lincoln, 1997- present.

Museum Research Associate, Mammalogy & Parasitology, Museum of Southwestern Biology, University of New Mexico, Albuquerque, 2004- present.

11. PROFESSIONAL AWARDS, HONORS, RECOGNITION:

Anniversary Award, Helminthological Society of Washington. October 2004.

Washington Biologists Field Club, inducted to membership October 2004.

Certificate of Merit, Agricultural Research Service, 2002. “For playing a key role on the team that planned and implemented the Systematics Summit, BARC.

36 Patronyms- Pararhinebothroides hobergi- Zamparo, Brooks & Barriga 1999, J. Parasitology 85:534- 539. Proteocephalus hobergi- de Chambrier & Vaucher, 1999, Rev. Suisse Zool. 106:165-240. Tetrabothrius hobergi- Nikolov et al., 2010, J. Parasitology 96:746-751. Neoskrjabinolepis hobergi- Kornienko & Dukochaev, 2012, Syst. Parasitol. 83:179-188.

Certificate of Merit, Agricultural Research Service, 1995 - 2012 (consecutive), “For outstanding contributions to the systematics of helminth parasites of ruminants.”

Outstanding Early Career Research Scientist, USDA, Agricultural Research Service, November 1994.

Early Career Scientist of the Year, Beltsville Area Research Center, USDA, Agricultural Research Service, October 1994.

Henry Baldwin Ward Medal, American Society of Parasitologists, 1992.

Robert and Virginia Rausch Visiting Professorship: University of Saskatchewan, Western College of Veterinary Medicine, Saskatoon, Canada, March 1992.

Certificate of Merit, Agricultural Research Service, 1991, 1992, 1993, 1994.

National Academy of Sciences, Interacademy Exchange Program: long term exchange to Russia, 1988.

RAS, Institute of Biological Problems of the North, Chukhotka, Sea of Okhotsk, Russia (invited visiting scientist, Russian Academy of Sciences) June-September 1988.

Xi Sigma Pi Society of Foresters, 1983.

Antarctica Service Medal, National Science Foundation, August 1983.

RAS, Institute of Biological Problems of the North, Magadan and Chaun, Russia (invited visiting scientist, Russian Academy of Sciences) July-August, 1981.

Graduate Teaching Fellowship, University of Saskatchewan, 1977, 1978.

12. EDITORIAL ACTIVITY:

Associate Editor: Journal of Parasitology: for Systematics and Phylogenetics, 1993-1996. Systematic Parasitology: Invited Editor, Vol. 42 (Eucestodes….) 1999. International Journal for Parasitology- Parasites and Wildlife, 2012- present.

Editorial Board: Journal of the Helminthological Society of Washington, 1992-1999 Zoology and Ecology (replacing) Acta Zoologica Lithuanica, 2005- present. Vestnik Zoologii (Ukraine), 2010- present.

Grants Reviewed for Granting Agencies: Natural Sciences and Engineering Research Council (Canada) National Science Foundation (USA) International Science Foundation (Soros Foundation) Australian Research Council 37 Hungarian Scientific Research Fund (OTKA). Grant Agency of the Czech Republic Swiss National Science Foundation U.S. Civilian Research and Development Foundation (CRDF) Leakey Foundation

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