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11 April 2000 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON 113(1): 1-12. 2000. Austin Beatty Williams (17 October 1919-27 October 1999). Biographical summary

Rafael Lemaitre and Bruce B. CoUette (RL) Department of Invertebrate , National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560-0163, U.S.A.; (BBC) Bruce B. CoUette, National Marine Fisheries Service Systematics Laboratory, National Museum of Natural History, Smithsonian Institution, Washington, D.C. 20560-0153, U.S.A.

Abstract.—The professional career and aspects of the life of Austin Beatty Williams (17 October 1919-27 October 1999), Systematic Zoologist for the Systematics Laboratory of the U.S. National Marine Fisheries, are summarized. Included is a bibliography with the 118 papers published by Williams, and a list of all the new names he proposed along with the holotype repository and catalogue number of species and subspecies.

Austin B. Williams (Fig. 1) was System- ter for Bulletin No. 6: "The hydrothermal atic Zoologist at the Systematics Laborato- vents of the eastern Pacific: An overview" ry, National Marine Fisheries Service (Williams 1985b). (NMFS), based at the National Museum of Austin had a distinguished career span- Natural History, Smithsonian Institution, ning five decades during which he pub- Washington, D.C. After a valiant fight with lished 118 papers (see bibliography). Bom cancer, Austin passed away at his home in in Plattsburg, Missouri in 1919, he was the Falls Church, Virginia. Shortly after his first child of Oliver Perry Williams and death, the Council of the Biological Society Lucy Sell; his siblings are brothers Hillis of Washington unanimously voted to dedi- and Oliver. He married Jean McNicol with cate this issue, the first of the new millen- whom he had their only child, David (mar- ium, to his memory. This dedication is most ried to Anita Kyle, with two children, Lau- fitting as Austin generously served the So- ren and Kyle). His family had only modest ciety in many capacities during the last means so he had to work to support his ed- quarter of the 20th century, and gave luster ucation, first at McPherson College (A.B. to the Proceedings by using it to publish 1943), and then at the University of Kansas many of his important papers. He served (Ph.D. 1951), where he studied Ozark cray- the Society as a Editor of the Proceedings fishes. His studies on these re- (1974-1977), Vice-president (1983-1986), main among the key references to identify President (1986-1988), Past-President these decapods in the region. From 1951 to (1989-1999), Custodian of Publications 1955 he was with the University of North (1989-1995), and contributed significantly Carolina Institute of Fisheries Research, to its financial soundness as a member of studying the life history and ecology of the Finance Committee (1995-1999). He penaeid . He then worked at the was editor of Bulletin No. 3: "Symposium University of Illinois from 1956 to 1963, on the Composition and Evolution of Crus- after which he returned to the North Caro- taceans in the Cold and Temperate Waters lina Institute of Fisheries Research to con- of the World " (1979), based on the tinue his studies on marine and estuiirine results of a U.S.-U.S.S.R. Cooperative Pro- decapods. In the mid 1960s, Donald E gram. He also provided the sununary chap- Squires, Chairman of the Department of In- PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

Fig. I. Aiisliii B. Williams. Clockwi.sc from upper left: I'roni church directory. I99S: with tamily and friends at Heart Tail Ranch, Bulle County, South Dakota, one of his favorite fossil collecting sites, 31 July 1997. in back is son David, in front left to right are David's wife Anita with grandchildren Kyle and Lauren, and Nornia Samuels (Nornia Samuels); recording observations in his notebook on fossil decapcxis collected I'roni the Pierre Shale at Heart Tail Ranch, Butle County, South Dakota, 6 August 1997 (Gale A. Bishop); at Mount Rushmorc. South Dakota, August 1997 (Norina Samuels). and then Deputy Direc- down. This was a loss for the Museum but tor of the National Museum of Natural His- fortunately he was hired by the NMFS' tory, Smithsonian Institution, considered Systematics Laboratory in 1971, him for a job; however, his interview with In addition to his work, Aus- Secretary Dillon Ripley did not go well tin served NMFS and the Museum in a (most probably because of Austin's non-as- number of different ways. For example, he suming personality), and he turned him represented the Allied Agencies (NMFS, VOLUME 113, NUMBER 1

Agriculture, and what was then the Fish and and usefulness in the identification of spec- Wildlife Service) on the Senate of Scientists imens while at the same time giving insight in the Museum. During his tenure in this into phylogenetic relationships. No major position, a question arose as to whether or group of decapods escaped Austin's atten- not the administrative staff of the Museum tion. He published important works on had increased significantly in the several crayfishes, peneaeoids, carideans, thalassin- preceding years. Discussion went back and ideans, , anomurans, and brachyu- forth between the Senate and the Director rans. His landmark studies on swimming of the Museum. Finally, Austin volunteered of the Callinectes, mud shrimps to get some real data on the issue. In a typ- of the family Upogebiidae, commercial lob- ical A. B. Williams way, he systematically sters, xanthid crabs, and deep-sea hydro- went through the entire telephone directory thermal vent decapods, among others, have and counted museum administrators at 5- earned him a place in the history of Zool- year intervals, and demonstrated that the ogy. His contributions to the systematics of type of positions that the Senate considered hydrothermal vent decapods inspired other as "administrative" had in fact increased colleagues, and one genus and species of significandy. the family Bythograeidae Williams, He was the acknowledged expert on and 1980, was named after him {Austinograea leader in studies of the systematics of east- williamsi Hessler & Martin, 1989, Journal em American decapod . He is of Crustacean Biology 9(4):645-661). In probably best known for his widely used March of 1996 he traveled to Kumamoto, monograph "Shrimps, lobsters, and crabs Japan, on a Japanese fellowship program to of the Atlantic coast of the eastern United join Keiji Baba (Kumamoto University Fac- States" published by the Smithsonian in ulty of Ecucation) in the study of galatheids 1984. His earlier study on the decapods of and other vent decapods from hydrother- the Carolinas published in 1965, a precursor mally active sites in the western Pacific. Al- to his 1984 monograph, was selected as a though his work concentrated on aspects of Science Citation Classic in 1983, a rare systematics and evolution, he also pub- honor for a systematist. His invaluable pa- lished key studies on the biology of com- per (Williams 1987a) on the identification mercial penaeid shrimps, ecology of mero- of spiny lobsters by color patterns of the plankton, larval genetics, and crustacean tails grew into a book co-authored with I. fisheries and mariculture. At the time of his Dore, entitled "Lobsters of the world—an death he had completed work but unfortu- illustrated guide" (Williams & Dore nately left unpublished, an important revi- 1988e); these two publications are indis- sion of the crab family Latreilliidae. pensable for anybody interested in this Austin's impact on carcinology is not group of economically significant decapods. limited to that derived from his publica- The primary focus of his research was tions. Throughout his exemplary career he the , systematics, biogeography actively participated in many professional and evolution of various decapod groups, societies in addition to the Biological So- both fossil and Recent. He named 101 new ciety of Washington. He was president of decapod taxa (see list), including one su- the Atlantic Estuarine Research Society perfamily, 2 families, 16 genera, 80 species, (1960-1961); co-founder, secretary (1971- and 2 subspecies. Occasionally he also 1973), and president (1983-1985) of the worked on other groups such as cirripeds, Estuarine Research Federation; secretary mysids, amphipods, and euphausiids, and (1985-1988) for the Society of Systematic even bird ecology. His publications provide Biology; associate editor for The Crusta- us with a standard of excellence, and are cean Society (1986-1991); and vice-presi- well known for attention to detail, accuracy. dent (1990-1991) and president (1991- PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

1992) of the American Association for Zoo- knowledge of decapods made him an im- logical Nomenclature. He was also a mem- posing figiure to both students and junior ber of the American Association for the colleagues; however, his modesty was such Advancement of Science, American Fish- that he made sure to treat everyone as a eries Society, American Institute of Biolog- friend or colleague of similar stature. He ical Sciences, American Institute of Fishery accepted life-time honors bestowed upon Research Biologists, American Society of him by his peers only hesitantly, and main- Limnology and Oceanography, Society for tained until his end that he was undeserving Integrative and Comparative Biology (for- of such attention. His personality, working merly American Society of Zoologists), As- habits, discipline, and inspirations date back sociation of Systematic Collections, Ecolog- to his early life experiences which he often ical Society of America, Kansas Academy mentioned to friends during casual conver- of Science, and Society for the Study of Evo- sations. One of his first jobs prior to enter- lution. His editorial activities with various ing college was at his family's farm in Ster- journals and symposia proceedings pro- ling, Colorado, where he helped string fenc- duced many significant contributions, and es, some of which had to be modified to his involvement with doctoral and master- "first class communications grade" by in- level students at various academic institu- sulating the top strand of barbed wire to tions led to the development of outstanding carry telephone signals. This worked fine carcinologists who now must continue his until it rained and the circuits became legacy. His skills as reviewer were highly grounded by water. During his Sterling days regarded by editors and grant program man- he also taught high school. One of Austin's agers who were assured of a detailed, un- scientific strengths was his observational biased evaluation. and note taking abilities. He had been Several of Austin's papers won important trained at Kansas to write reflective notes awards or honorable mentions. His each night as if they were to be published. identification paper (Williams 1987a), for Those who have examined his field and of- example, won the highly regarded "Nation- fice notebooks are struck by how remark- al Marine Fisheries Service Outstanding ably clear and detailed they are. Publication Award" for best paper in the Austin often mentioned the impact of the 1997 Marine Fisheries Review. In recogni- Great Plains of Kansas and Colorado on his tion of Austin's life-time work. The Crus- psyche. One of his closest friends. Gale A. tacean Society presented him in 1997 with Bishop (Georgia Southern University), has their "Excellence in Research Award" dur- said that he was impressed with Austin's ing a ceremony at the National Museum of collegiality when he first met him during a Natural History, Smithsonian Institution, visit to the Smithsonian to study fossil Washington, D.C. [see Lemaitre, R., 1998, decapods. Gale suggested that he might Journal of Crustacean Biology 18(3):619- want to join him in the field in South Da- 620]. He was also honored with the "1999 kota to collect fossils. Austin did so with Elton Sette Award" from the Marine Fish- much enthusiasm, and the two worked to- eries Section of the Americjui Fisheries So- gether almost every summer from 1980 un- ciety. til the year of his death. Nancy Braimen Austin will be remembered not only for Marsh (Science Department, Portal High his impressive scientific accomplishments School, Georgia) also joined them, and the but also for his human qualities. Unselfish three collaborated in studies of decapods of almost to a fault, he made every effort to the Western Interior Cretaceous, collecting help colleagues and students alike. His numerous fossil crabs, lobsters, and shrimps height of 1.85 m (6'1") gave him a towering from the Carlile Shale and the Pierre Shale physique which combined with his deep of South Dakota, Wyoming and Colorado. VOLUME 113, NUMBER 1

These fossil collections have been donated choral music. He actively participated in to the Museum of Geology at the South Da- several choral groups, including the Wash- kota School of Mines, Rapid City. He con- ington Cathedral Choral Society which per- fessed to Gale that the timing of their col- formed in the National Cathedral, Washing- laboration was most appropriate as his wife ton, D.C. One of his performances is pre- Jean had passed away (1983) after a diffi- served on an audio CD-ROM entitled "Mil- cult illness (with Austin as major care giv- lenium; Russian Choral Music" (1990 er), and his return to the Great Plains was Centaur Record Inc.). He worshipped, sang just what he needed to gain closure and and was an active member of the New York healing from the loss. This theme, Gale Avenue Presbyterian Church, of Washing- says, "came up many times and we con- ton, D.C, where well-attended and emo- cluded that getting back to our roots was an tional services were held for him on Octo- extremely healing process, both for Austin's ber 30, officiated by The Rev. Robert H. loss and for the loss of both of my parents; Craig. it was an annual 'rehealing' as we came Austin's remains are buried in Marion, back into harmony with our roots and Kansas, alongside those of his wife. Mother Earth. When working with Austin on fossils in the Western Interior his intel- Bibliography lect and collegiality were always apparent. (excluding abstracts and technical reports) His interests spanned the sciences, arts, ed- ucation, and humanities. He often would 1952a. Williams, A. B. Six new crayfishes of the genus Orconecles (: ) from Arkan- visit the Rapid City Astronomy club to par- sas. Missouri and Oklahoma.—Transactions of ticipate in telescopic observations, take us the Kansas Academy of Science 55(2):330-35l. all out dancing at the Broken Boot Saloon 1952b. , & A. B. Leonard The crayfishes of in Rapid City, climb Bear Butte or Hamey Kansas.—University of Kansas Science Bulle- Peak, or head us up to Rushmore for the tin 34, part II. (15):96I-I0I2. 1953. . Identification of juvenile (Pen- evening patriotic lighting program. While aeidae) in North Carolina.—Journal of the Eli- in the field, he brought new insights to pa- sha Mitchell Scientific Society 69(2): 156-160. leontology, often seeing things we took for 1954a. . Speciation and distribution of the cray- granted or forcing clearer explanation of fishes of the Ozark Plateaus and Ouachita Prov- our mutual deductions. Our collaboration inces.—University of Kansas Science Bulletin 36. part II, (12):803-918. was clearly very beneficial to Austin as 1954b. . An explanation for the distribution of well as to paleontologists Nancy Brennan a North American .—Ecology 35(4): Marsh, the late Reinhard Forster of Munich, 573-575. and Georgia Southern students Mike Klug, 1955a. . The genus Ogyrides (Crustacea: Car- Mehmet Samiratedu, and Amy Samiratedu. idea) in North Carolina.—Journal of the Wash- ington Academy of Science 45(2):56-59. These insights were carried over into the 1955b. A contribution to the life histories of laboratory and into collaborations on papers commercial .shrimps (Penaeidae) in North Car- and research comparing Recent and fossil olina.—Bulletin of Marine Science of the Gulf decapods." and Caribbean 5(2): 116-146. The multi-faceted personality of Austin 1955c. . A survey of North Carolina shrimp nursery grounds.—Journal of the Elisha Mitch- included a deep appreciation of the simple ell Scientific Society 7l(2):2(X)-207. things of life, his family and friends. During 1956. Darnell, R. M., & . 1956. A note on the the last decade or so of his life he was for- occurrence of the pink shrimp, Penaeus duor- tunate to share many moments with Norma ariim. in Louisiana waters.—Ecology 37(4): Samuels, of Fairfax, Virginia, whose com- 844-846. 1958. . Substrates as a factor in shrimp distri- panionship undoubtedly enriched his life. bution.—Limnology and Oceanography 3(3): Austin developed a passion for ballroom 283-290. and international dancing, and a love for 1959. . Spotted and brown shrimp postlarvae PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

(Penaeus) in North Carolina.—Bulletin of Ma- Mexico, 12-21 June 1967, FAO Fisheries Re- rine Science of the Gulf and Caribbean 9(3): ports, No. 57, vol. 3, E/33:643-656. 281-290. 1970. . Review: Crustacea Decapoda Brach- 1960. . The influence of temperature on os- yura. By M. E. Christiansen, 1969, Marine In- motic regulation in two species of estuarine vertebrates of Scandinavia, No. 2, Universitets- shrimps (Penaeus).—Biological Bulletin forlaget, Oslo, 143 pp.—Chesapeake Science 119(3):560-571. ll(l):69. 1962. . A re-examination of Ovalipes species 1971a. , & H. J. Porter. A ten-year study of in the Carolinas (Decapoda: Portunldae).— meroplankton in North Carolina estuaries: Oc- Crustaccana 4(1):39-41. currence of postmetamorphal bivalves.—Ches- 1963. Cerame-Vivas, M. J., , & I. E. Gray. New apeake Science 12(l):26-32. decapod crustacean records for the coast of 1971b. . A ten-year study of meroplankton in North Carolina.—Crustaceana 5(2): 157-159. North Carolina estuaries: Annual occurrence of 1964. , & H. J. Porter. An unusually large turtle some brachyuran developmental stages.—Ches- (Chelonibia p. patula) on a blue crab apeake Science 12(2):53-61. from Delaware Bay.—Chesapeake Science 1972a. . Review: Guide to identification of ma- 5(3): 150-153. rine and estuarine invertebrates. Cape Hatteras 1965a. . Marine decapod crustaceans of the to the Bay of Fundy. By K. L. Gosner, 1971, Carolinas.—Fishery Bulletin 65(1): 1-298. Wiley-Intersciences, New York, 693 pp.— 1965b. . A new genus and species of snapping IVansactions of the American Fisheries Society shrimp (Decapoda: Alpheidae) from the south- 101(3):570-571. eastern United States.—Crustaceana 9(2): 192- 1972b. , & W. S. Brown. Notes on structure 198. and parasitism of iris A. Milne Ed- 1966a. . Technical note no. 2 - An inexpensive wards (Decapoda, Galatheidae) from North Car- scallop cleaner built from spare parts.—Com- olina, USA.—Crustaceana 22(3):303-308. mercial Fisheries Review 28(1): 12-13. 1972c. . A ten-year study of meroplankton in 1966b. . The western Atlantic swimming crabs North Carolina estuaries: Juvenile and adult Callinectes omatus, C. danae, and a new, re- Ogyrides (: Ogyrididae).—Chesapeake lated species (Decapoda: Portunidae).—^l\ilane Science 13(2): 145-148. Studies in Zoology 13(3):83-93. I972d. , & K. H. Bynum. A ten-year study of 1967a. , G. S. Posner, W. J. Woods, & E. E. meroplankton in North Carolina estuaries: Am- Deubler, Jr. A hydrographic atlas of larger North phipods.—Chesapeake Science 13(3):175-192. Carolina sounds.—U.S. Fish and Wildlife Ser- 1972e. . A ten-year study of meroplankton in vice Data Report 20:1-130. (Reprinted 1973 as North Carolina estuaries: Mysid shrimps.— Sea Grant Publication UNC-5G-73-10, 129 pp.) Chesapeake Science 13(4):254-262. 1967b. . Review: The crabs of Sagami Bay: 1972f. . Decapod crustaceans of the Chesa- Collected by His Majesty the Emperor of Japan. peake Bay.—Chesapeake Science 13 (Supple- By T. Sakai, 1965. East-West Center Press, ment):S119-S121. Honolulu, Hawaii, 206 pp.—Transactions 1974a. . Allactaea lithostrota. a new genus and American Fisheries Society 96:81-82. species of crab (Decapoda: ) from 1968a. , & E. E. Deubler, Jr. A ten-year study North Carolina, U.S.A.—Proceedings of the Bi- of meroplankton in North Carolina estuaries: ological Society of Washington 87(3): 19-26. Assessment of environmental factors and sam- 1974b. . Two new axiids (Crustacea: Decapoda: pling success among bothid flounders and pen- Thalassinidea: Calocaris) from North Carolina aeid shrimps.—Chesapeake Science 9(1):27- and the Straits of Florida.—Proceedings of the 41. Biological Society of Washington 87(39):451- 1968b. , L. R. McCloskey, & I. E. Gray. New 464. records of brachyuran decapod crustaceans 1974c. . A new species of Hypsophrys (Deca- from the continental shelf off North Carolina.— poda: Homolidae) from the Straits of Florida, Crustaceana 15(l):41-66. with notes on related crabs.—Proceedings of 1969a. . A ten-year study of meroplankton in the Biological Society of Washington 87(42): North Carolina estuaries: Cycles of occurrence 485-492. among penaeidean shrimps.—Chesapeake Sci- 1974d. , T. E. Bowman, & D. M. Damkaer. ence 10(l):36-47. Distribution, variation, and supplemental de- 1969b. . Penaeid shrimp catch and heat sum- scription of the opossum shrimp. Neomysis mation, an apparent relationship.—Proceedings americana (Crustacea: Mysidacea).—Fishery World Scientific Conference on Biology and Bulletin 72(3):835-842. Culture of Shrimps and Prawns, Mexico City, I974e. . The swimming crabs of the genus Cal- VOLUME 113, NUMBER 1

linectes (Decapoda: Portunidae).—Fishery Bul- 1981b. & D. McN. Williams. Carolinian re- letin 72(3):685-798. cords for American lobster, american- 1974f. . Marine flora and fauna of the north- us, and tropical swimming crab, Callinecles bo- eastern United States. Crustacea: Decapoda.— courti. Postulated means of dispersal.—Fishery NOAA Technical Report NMFS Circular 389: Bulletin 79(1): 192-198. 1-50. 1982a. . Revision of the genus Roux 1976a. . Distinction between a Gulf of Mexico (Brachyura: ).—Quademi del La- and a Carolinian Atlantic species of swimming boratorio de Tccnologia della Pesca, Ancona, It- crab Ovalipes (Decapoda: Portunidae).—Pro- aly 3(2-5):227-255. ceedings of the Biological Society of Washing- 1982b. , & E A. Chace, Jn A new caridean ton 89(14):205-214. shrimp of the family Bresiliidae from thermal 1976b. . Integumental organs of unknown func- vents of the Galapagos Rift.—Journal of Crus- tion on chelipeds of deep-sea crabs, genus Hyp- tacean Biology 2(1): 136-147. sophrys.—^Journal of Morphology 150(4):889- 1983a. . Marine decapod crustaceans of the 899. Carolinas. Fishery Bulletin 65:1-298.—Current 1977a. . J. K. Shaw, & T. S. Hopkins. Stilbom- Contents 14(I7):20. astax, a new genus of spider crab (Majidae: Ty- 1983b. , & C. L. Van Dover. A new species of chinae) from the West Indies region, with notes Munidopsis from submarine thermal vents of on American relatives.—Proceedings of the Bi- the East Pacific Rise at 2I°N (: Gal- ological Society of Washington 90(4):884-893. atheidae).—Prtxieedings of the Biological So- 1977b. , & R. L. Wigley. Distribution of deca- ciety of Washington 96(3):481-488. pod Crustacea off northeastern United States 1984a. . The mud crab, herbstii, s.l. based on specimens at the Northeast Fisheries Partition into six species (Decapoda: Xanthi- Center, Woods Hole, Massachusetts.—NOAA dae).—Fishery Bulletin 81(4):863-882. Technical Report NMFS Circular 407:i-iii, 1- 1984b. Sullivan, B.. K. Miller, K. Singleton, A. G. 44. Scheer, & . Electrophoretic analyses of 1978a. . Transfer to Pseudomedaeus of the xan- hemocyanins from four species of mud crabs, thid crab Micropanope distinctus (Rathbun).— genus Punopeus, with observations on the ecol- Proceedings of the Biological Society of Wash- ogy of P. obesus.—Fishery Bulletin 81(4):883- ington 91(2):546-557. 885. 1978b. . True crabs. (Unpaginated) in W. Fi- 1984c. Reams, R. C, & . Mud crabs of the scher, ed., FAO Species Identification Sheets for Panopeus herbstii H. M. Edw., s.l., complex in Fishery Purposes. Western Central Atlantic Alabama, U.S.A.—Fishery Bulletin 81(4):885- (Fishing Area 31), vol. 6, Food and Agriculture 890. Organization of the United Nations, Rome. 1984d. Van Dover, C. L., , & J. R. Factor The 1979a. Herbst, G. N., , & B. B. Boothe, Jr. first zoeal stage of a hydrothermal vent crab Reassessment of northern geographic limits for (Decapoda: Brachyura: Bythograeidae).—Pro- decapod crustacean species in the Carolinian ceedings of the Biological Society of Washing- Province, USA; some major range extensions ton 97(2):4I3-418. itemized.—Proceedings of the Biological Soci- 1984e. Millikin, M. R., & . Synopsis of bio- ety of Washington 9l(4):989-998. logical data on the blue crab, Catlinectes sapi- 1979b. , & T. W. Duke. Chapter 6. Crabs (Ar- dus Rathbun.—NOAA Technical Report NMFS thropoda: Crustacea: Decapoda: Brachyura). Pp. 1, FAO Fisheries Synopsis 138:1-39. 171-233 in C. W. Hart and S. M. N. Fuller, eds.. I984f. . Shrimps, lobsters, and crabs, of the At- Pollution ecology of estuarine invertebrates. lantic coast of the eastern United States, Maine Academic Press. N.Y., 406 pp. to Florida. Smithsonian Institution Press. 550 1979c. . A new crab family fi-om shallow wa- pp. ters of the West Indies (Crustacea: Decapoda: 1984g. Review: Shallow-water crabs. By R. Brachyura).—Proceedings of the Biological So- W. Ingle. 1983, Linnean Society of London and ciety of Washington 92(2):399-413. the Estuarine and Brackish-Water Sciences As- 1980. . A new crab family from the vicinity of sociation, Cambridge University Press, London, submarine thermal vents on the Galapagos Rift 206 pp.—Estuaries 7(3):266-267. (Crustacea: Decapoda: Brachyura).—Proceed- 1984h. . Review: Crustacean Phytogeny. F. R. ings of the Biological Society of Washington Schram, ed., 1983, Crustacean Issues 1. Based 93(2):443-472. on a symposium, Dallas, Dec. 1981, Balkema, 1981a. . Western Atlantic species of the cari- Rotterdam, 372 pp.—American Scientist 72(5): dean shrimp genus Ogyrides.—^Journal of Crus- 514-515. tacean Biology 1(1): 143-147. 1985a. Van Dover, C. L., J. R. Factor, , & C. 8 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

J. Berg, Jr. Reproductive patterns of decapod 1988b. -. Notes on decapod and euphausiid crustaceans from hydrothermal vents of the crustaceans, continental margin, western Atlan- Eastern Pacific. In M. L. Jones, ed.. The hydro- tic, Georges Bank to western Florida, USA.— thermal vents of the Eastern Pacific: an over- Fishery Bulletin 86(l):67-76. view.—Bulletin of the Biological Society of 1988c. . New marine decapod crustaceans from Washington 6:223-227. waters influenced by hydrothermal discharge, 1985b. . Summary comments. In M. L. Jones, brine, and hydrocarbon seepage.—Fishery Bul- ed.. The hydrothermal vents of the Eastern Pa- letin 86(2):263-287. cific: An overview.—Bulletin of the Biological 1988d. . Conjoined twin adult shrimp (Deca- Society of Washington 6:489-493. poda: Penaeidae).—Fishery Bulletin 86(3):595- 1986a. . Mud shrimps, Upogebia, from the 597. eastern Pacific (Thalassinoidea: Upogebi- 1988e. , & I. Dore. Lobsters of the world - An idae).—San Diego Society of Natural History, illustrated guide: Lobsters of the world in U.S. Memoir 14:1-60. trade. Osprey Books, Huntington. New York, 1986b. Bishop, G. A., & . The fossil lobster 186 pp. Unuparus canadensis. Carlile Shale (Creta- 1989a. . & C. A. Child. Comparison of some ceous), Black Hills.—National Geographic Re- genera and species of box crabs (Brachyura: search 2(3):372-387. Calappidae), southwestern North Atlantic, with 1986c. , & P. A. Rona. Two new caridean description of a new genus and species.—Fish- shrimps (Bresiliidae) from a hydrothermal field ery Bulletin 87(1):105-121. on the Mid-Atlantic Ridge.—^Journal of Crus- 1989b. , & R J. B. Scott. Upogebia corallifora. tacean Biology 6(3):446-462. a new species of coral-boring shrimp from the 1986d. , & R. D. "nimer. Squat lobsters (Gal- West Indies (Decapoda: Upogebiidae).—Pro- atheidae: Munidopsis) associated with mesh-en- ceedings of the Biological Society of Washing- closed wood panels submerged in the deep ton 102(2):405-410. sea.—Journal of Crustacean Biology 6(3):617- 1989c. , L. G. Abele, D. L. Felder, H. H. 624. Hobbs, Jr., R. B. Manning, P. A. McLaughlin, 1986e. , & D. L. Felder. Analysis of stone & I. P^rez Farfante. Common and scientific crabs: Menippe mercenaria (Say), restricted, names of aquatic invertebrates from the United and a previously unrecognized species de- States and Canada: decapod crustaceans.— scribed (Decapoda: Xanthidae).—Proceedings American Fisheries Society Special Publication of the Biological Society of Washington 99(3): No. 17:1-77. 517-543. 1990a. , & J. J. McDermott. An eastern United 1986f. . Foreword. In D. A. Wolfe, ed., Estu- States record for the western Indo-Pacific crab, arine variability. Academic Press, New York, Hemigrapsus sanguineus (Crustacea: Decapo- 509 pp. da: Grapsidae).—Proceedings of the Biological 1987a. . Lobsters—identification, world distri- Society of Washington 103(1): 108-109. bution, and U.S. trade.—Marine Fisheries Re- 1990b. , & K. Baba. New squat lobsters (Gal- view 48(2): 1-36. atheidae) from the Pacific : Mariana Back 1987b. . More records for shrimps of the genus Arc Basin, East Pacific Rise, and Cascadia Ba- Rimicaris (Decapoda: Caridea: Bresiliidae) sin.—Fishery Bulletin 87(4):899-910. from the Mid-Atlantic Rift.—^Journal of Crus- 1990c. Kensley, B., & . Axiopsis eximia, a new tacean Biology 7(1): 105. thalassinidian shrimp (Crustacea, Decapoda, 1987c. Wetherbcc, D. K., & . The late eigh- Axiidae) from the Middle Eocene of South Car- teenth century paintings by Rabie of Haitian olina.—Journal of Paleontology 64(5):798-802. moUusks and crustaceans. Further Contributions 1990d. , & E. E. Boschi. Panopeus margentus, on the History of Zoology in Hispaniola, Art. a new crab from the Argentine warm temperate 3:18-48. Privately published, Shelbume. Mas- subregion (Decapoda: Xanthidae).—Proceed- sachusetts. ings of the Biological Society of Washington 1987d. . Upogebia synagelas, new species, a 103(3):598-60l. commensal mud shrimp from in the 1990e. , & N. Ngoc-Ho. Pomatogebia. a new western central Atlantic (Decapoda: Upogebi- genus of thatassinidean shrimps from western idae).—Proceedings of the Biological Society hemisphere tropics (Crustacea: Upogebiidae).— of Washington 100(3):590-595. Proceedings of the Biological Society of Wash- 1988a. . Indo-Pacific spiny lobsters in the U.S. ington 103(3):614-616. National Museum of Natural History collected 1991a. , & R. W. Heard. Upogebia spinistipula, from 1963 to 1981 (Crustacea: Palinuridea).— a new burrowing shrimp from the Florida shelf, Crustaceana 55(3):313-316. northeastern Gulf of Mexico (Decapoda: Upo- VOLUME 113, NUMBER 1

gebiidae).—Proceedings of the Biological So- (Brachyura: Majidae).—Crustacean Research ciety of Washington 104(l):49-54. 23:1-4. 1991b. , & R. B. Moffitt. Crabs from the Mar- 1995a. Rodriguez, G., & . Epilobocera weth- iana Archipelago: Bothromaia grijfini new ge- erheei. a new species of freshwater crab (De- nus and species (Brachyura: Majidae). and re- capoda: Brachyura: ) from marks on Poupinia hirsura Guinot (Homolo- Hispaniola.—Proceedings of the Biological So- idea, Poupiniidae).—Proceedings of the Biolog- ciety of Washington 108(l):76-83. ical Society of Washington 104(3):569-582. 1995b. Komfield, I., , & R. S. Steneck. As- 1991c. Bishop, G. A., & . Necrocarcinus ol- signment of Homarus capensis (Herbst, 1792), sonorum, new species, a crab (Decapoda: Ca- the of South Africa, to the new lappidae) from the Cretaceous Carlile Shale genus Homarinus (Decapoda: Nephropidae).— (TUronian), Western Interior United States.— Fi-shery Bulletin 93(1):97-I02. Journal of Crustacean Biology 11(3):451-459. 1995c. , & \\ C. Dobbs. A new genus and spe- 199Id. . Comments on the proposed conser- cies of caridean shrimp (Crustacea: Decapoda: vation of the specific name of Anemia francis- Bresiliidae) from hydrothermal vents on Loihi cana Kellogg, 1906 (Crustacea, Branchiopoda). Seamount, Hawaii.—Proceedings of the Biolog- (Case 2728; see BZN 47:178-183; 48:57).— ical Society of Washington 108(2):228-237. Bulletin of Zoological Nomenclature 48(3): I995d. . Chapter 2, Taxonomy and evolution. 246-247. Pp. 13-21 in i. R. Factor, ed.. Biology of the 1991e. Van Dover, C. L., & . Egg size in Homarus americanus. Academic Press, lobsters (): Constraint and free- New York, 529 pp. dom. Pp. 143-156 in A. Wenner & A. Kuris, 1995e. Pequegnat, L. H., & . TWo new species eds.. Crustacean Egg Production. Crustacean Is- of Munidopsis (Decapoda: Anomura: Galathei- sues, vol. 7, A. A. Balkema/Rotterdam, 401 pp. dae) from the Western Atlantic Ocean.—Journal 1992a. , & R. Wahle. Distinguishing juvenile of Crustacean Biology 15(4):786-792. stages of Jonah and Atlantic rock crabs. Cancer 1996a. . Comments on the proposed conser- borealis and C. irroralus (Crustacea: Decapoda: vation of the generic names Monstrilla Dana, Cancridae).—Journal of Crustacean Biology 1849 and Thaumaleus Kroyer, 1849 (Crustacea, l2(3):464-466. Copepoda) (Case 2894; see BZN 52(3):245- 1992b. . Review: Decapod Crustacea of the At- 249).—Bulletin of Zoological Nomenclature lantic Coast of Canada. By H. J. Squires, 1990, 53(2): 122-123. Canadian Bulletin of Fisheries and Aquatic Sci- 1996b. . Comments on the proposed conser- ences 221:532 pp.—Journal of Crustacean Bi- vation of the generic name Glomeris Latreille, ology I2(2):329. 1802 (Diplopoda) and the specific name of Ar- 1992c. . Review: Marine Lobsters of the World. madillo vulgaris Latreille, 1804 (Crustacea, Is- An Annotated and Illustrated Catalog of Species opoda), and the application for a ruling on the of Interest Known to Date. EAO Species Cata- status of the name Armadillo Latreille. 1802 log. By L. B. Hohhuis. 1991. FAO Fisheries (Crustacea, Isopoda). (Case 2909; see BZN Synopsis No. 125, vol. 13:292 pp. Rome.—^The 52(3):236-244).—Bulletin of Zoological No- Lobster Newsletter 5:14. menclature 53(2): 121-122. [Resolved in Opin- 1993a. . Comments on the proposed conser- ion 1897, Bulletin of Zoological Nomenclature vation of the specific names of Gebia major ca- 55(2): 124-128, June 1998.] pensis Krauss, 1843 and G. africana Ortmann, 1997a. . Bylhograea ihermydron Williams, 1894 (currently Upogehia capensis and U. af- 1980. Arthropoda, Crustacea, Decapoda, Brach- ricana; Crustacea, Decapoda) by the designa- yura, Brachyrhyncha, Bythograeoidea, Bytho- tion of a replacement neotype for U. capensis. graeidac. p. 212 in D. Desbruy&res and M. Se- (Case 2827; BZN 49(3): 187-190.).—Bulletin gonzac, eds.. Handbook of deep-sea hydrother- of Zoological Nomenclature 50(2): 143-144. mal vent fauna. Editions IFREMER, Brest, 279 1993b. . Mud shrimps, Upogebiidae, from the PP- western Atlantic (Crustacea: Decapoda: Thal- 1997b. Curran. H. A., & — Ichnology of an in- assinidea).—Smithsonian Contributions to Zo- tertidal carbonate sand flat: Pigeon Creek, San ology 544:i-iii, 1-77. Salvador Island. Bahamas. Pp. 33-46 in J. L. 1993c. . Reflections on crab research in North Carew, ed.. Proceedings of the 8th Symposium America since 1758. Pp. 259-273 in F. Trues- on the Geology of the Bahamas: San Salvador, dale, cd.. History of Carcinology, Crustacean Is- Bahamian Field Station, June 9-13, 1995. sues, vol. 8., A. A. Balkema/Rotterdam, 445 pp. 1997c. . Occurrence of three species of mud 1994. , & L. G. Eldredge. A new species of shrimps in aquaculture ponds on Caribbean spider crab from Guam, Rochinia decipiata coasts of Venezuela and Colombia, with a re- 10 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

description of Upogebia omissago Williams, Paulo, Brazil; SDSM, Museum of Geology, 1993 (Oecapoda: Upogebiidae).—Proceedings South Dakota School of Mines, Rapid City; of the Biological Society of Washington 110(3): 412-416. SDSNH, San Diego Society of Natural His- 1997d. . Two new species and a range exten- tory, California; UKMNH, University of sion of mud shrimps. Upogebia, from Pacific Kansas Museum of Natural History, Costa Rica and Mexico (Decapoda: Thalassin- Lawrence; USNM, National Museum of idea: Upogebiidae).—Proceedings of the Bio- Natural History, Smithsonian Institution, logical Society of Washington 110(4):617-623. 1998a. Baba, K., & . New Galatheoidea (Crus- Washington, D.C. tacea, Decapoda, Anomura) from hydrothermal systems in the West Pacific Ocean: Bismarck Decapoda Archipelago and Okinawa Trough.—Zoosyste- ma 20(2): 143-156. Caridea 1998b. , & J. L. Hernandez-Aguilera. A new species of mud shrimp. Upogebia loralae, from Alvinocaris Williams & Chace, 1982b. Veracruz. Mexico (Decapoda: Thalassinidea: Alvinocaris lusca Williams & Chace, Upogebiidae).—Proceedings of the Biological 1982b. USNM 184534. Society of Washington 111(4):908-911. Alvinocaris markensis Williams, 1988c. 2(XX)a. , & R. Vargas. A new species of mud USNM 234286. shrimp, Upogebia cortesi, from Pacific Costa Rica (Decapoda: Thalassinidea: Upogebi- Alvinocaris muricola Williams, 1988c. idae).—Proceedings of the Biological Society USNM 234288. of Washington 113(1):13-16. Alvinocaris stactophila Williams, 1988c. 2(XX)b. Bishop, G. A., & . Fossil crabs from USNM 234291. Tepee Buttes, submarine seeps of the Late Cre- Leptalpheus Williams, 196Sb. taceous Pierre Shale, South Dakota and Colo- rado, U.S.A.—Journal of Crustacean Biology Leptalpheus forceps Williams, 1965b. (Special no. 2):(in press). USNM 111084. In press. . Comment on the proposed designa- Ogyrides hayi Williams, 1981a. USNM tion of Scottia pseudobrowniana Kempf, 1971 47958. as the type species of Scottia Brady & Norman, Ogyrides limicola Williams, 1955a. USNM 1889 (Crustacea: Ostracoda) (Case 2896: see BZN 51(4):304-305).—Bulletin of Zoological 96675. Nomenclature. Opaepele Williams & Dobbs, 1995c. In press. . Comment on the proposed conser- Opaepele loihi Williams & Dobbs, 1995c. vation of both the generic and specific names USNM 251447. Cubans murina Brandt, 1833 (Case 2910; see Rimicaris Williams & Rona, 1986c. BZN 52(2): 133-156).—Bulletin of Zoological Nomenclature. Rimicaris chacei Williams & Rona, 1986c. USNM 228452. Rimicaris exoculata Williams & Rona, List of taxa named by Austin B. >\^lliams 1986c. USNM 228443. Taxa are listed alphabetic£illy within each major decapod group. Holotype deposition Astacidea and number is indicated for all species and subspecies. Asterisk indicates fossil taxa. Homarinus Komheld, Williams & Steneck, Abbreviations for repositories are as fol- 1995b. lows: AHF, Allan Hancock Foundation. Orconectes eupunctus Williams, 1952a. University of Southern California (now UKMNH T4250. Natural History Museum of Los Angeles Orconectes meeki brevis >\^liams, 1952a. County); AMNH, American Museum of UKMNH T8140. Natural History, New York; MCZ, Museum Orconectes nana marcus \Mlliams, 1952a. of Comparative Zoology, Harvard Univer- UKMNH T4970. sity, Cambridge, Massachusetts; MZUSP, Orconectes nana Williams, 1952a. Museu de Zoologia, Universidade de S3o UKMNH T6640. VOLUME 113, NUMBER 1 11

Orconectes neglectus chaenodactylus Wil- Upogebia lepta Williams, 1986a. USNM liams, 1952a. UKMNH T4420. 213270. Orconectes ozarkae Williams, 1952a. Upogebia maccraryae Williams, 1986a. UKMNH T6150. USNM 213202. Upogebia macginitieorum Williams, 1986a. Thalassinidea USNM 213219. Upogebia molipollex Williams, 1993b. Aethogebia Williams, 1993b. AMNH 6820. Aethogebia gorei Williams, 1993b. USNM Upogebia omissago Williams, 1993b. 251425. USNM 222057. *Axiopsis eximia Kensley & Williams, Upogebia onychion Williams, 1986a. AHF 1990c. USNM 219431. 4133. Calocaris (Calastacus) jenneri Williams, Upogebia paraffinis Williams, 1993b. 1974b. USNM 150472. MZUSP 8049 Calocaris {Calastacus) oxypleura Williams, Upogebia pillsbury Williams, 1993b. 1974b. USNM 101651. USNM 251435. Pomatogebia Williams & Ngoc-Ho, 1990e. Upogebia ramphula Williams, 1986a. USNM 213446. Upogebia acanthops Williams, 1986a. Upogebia schmitti Williams, 1986a. AHF USNM 213194. 3933. Upogebia aestuari Williams, 1993b. Upogebia spinistipula Williams & Heard, USNM 251407. 1991a. USNM 239251. Upogebia aquilina Williams, 1993b. Upogebia synagelas Williams, 1987d. USNM 251426. USNM 233572. Upogebia baldwini Williams, 1997d. Upogebia tenuipollex Williams, 1986a. USNM 251486. USNM 213236. Upogebia bermudensis Williams, 1993b. Upogebia thistlei Williams, 1986a. USNM MCZ 12873. 213251. Upogebia burkenroadi Williams, 1986a. Upogebia toralae Williams & Hemdndez- SDSNH 3985. Aguilera, 1998b. USNM 285522. Upogebia careospina Williams, 1993b. Upogebia vargasae Williams, 1997d. USNM 138899. USNM 251484. Upogebia casis Williams, 1993b. USNM Upogebia veleronis Williams, 1986a. 251224. (USNM 213272) Upogebia cocosia Williams, 1986a. USNM 213268. Anomura Upogebia coralliflora Williams & Scott, 1989b. USNM 230075. Munidopsis alvisca Williams, 1988c. Upogebia cortesi 2000a. USNM 291186. USNM 234294. Upogebia dawsoni Williams, 1986a. Munidopsis glabra Pequegnat & ^lliams, AHF2566. 1995e. USNM 251455. Upogebia felderi Williams, 1993b. USNM Munidopsis granosicorium Williams & 251430. Baba, 1990b. USNM 240205. Upogebia galapagensis Williams, 1986a. Munidopsis lentigo Williams & Van Etover, USNM 213223. 1983b. USNM 191160. Upogebia inomissa Williams, 1993b. Munidopsis lignaria Williams & Baba, USNM 251396. 1990b. USNM 240202. Upogebia jonesi Williams, 1986a. USNM Munidopsis marianica Williams & Baba, 213195. 1990b. USNM 240198. 12 PROCEEDINGS OF THE BIOLOGICAL SOCIETY OF WASHINGTON

Shinkaia Baba & Williams, 1998a. Ovalipes stephensoni Williams, 1976a. Shinkaia crosnieri Baba & Williams, USNM 155110. 1998a. USNM 251480. Panopeus austrobesus Williams, 1984a. Uroptychus edisonicus Baba & Williams, USNM 59462. 1998a. USNM 251479. Panopeus margentus Williams & Boschi, 1990d. USNM 239191. Brachyura Panopeus meridionalis Williams, 1984a. USNM 99846. Allactaea Williams, 1974a. *Plagiophthalmus bjorki Bishop & Wil- Allactaea lithorostrata A^^Uiams, 1974a. liams, 2000b. SDSM 11021. USNM 143770. *Raninella manningi Bishop & Williams, Bothromaia Williams & Moffit, 1991b. 2000b. SDSM 11018. Bothromaia griffini WUliams & Moffit, Rochinia decipiata Williams & Eldredge, 1991b. USNM 250884. 1994. USNM 251434. Bythograeoidea Williams, 1980. (Super- Stilbomastax Williams, Shaw & Hopkins, family). 1977a. Bythograeidae Williams, 1980. (Family). Bythograea Williams, 1980. Acknowledgments Bythograea mesatlantica Williams, 1988c. We would like to thank David McN. Wil- USNM 234300. liams for providing valuable family facts, Bythograea thermydron Williams, 1980. and also several of Austin's colleagues and USNM 172830. friends for contributing photographs and Callinectes similis Williams, 1966b. USNM other information. In particular, we would 113341. like to mention Norma Samuels who kindly Cyclozodion Williams & Child, 1989a. provided photographs and insight on as- Cyclozodion tuberatum Williams & Child, pects of Austin private life; Gale A. Bishop, 1989a. USNM 234462. for providing valuable and often emotional Epilobocera wetherbeei Rodriguez & Wil- testimony on his experiences in the field; liams, 1995a. USNM 268832. Raymond B. Manning and Brian Kensley Eplumula Williams, 1982a. for reminiscing on their interactions with *Heus Bishop & Williams, 2000b. Austin; and Keiji Baba for information and *Heus foersteri Bishop & Williams, 2000b. photographs. The invaluable help of Mi- SDSM 11016. chael Vecchione and the dedicated staff of Hypsophrys noar Williams, 1974c. USNM the Systematics Laboratory, National Ma- 150816. rine Fisheries Service, at the Smithsonian, Latreillia manningi Williams, 1982a. in locating and compiling files and data on USNM 57071. Austin's professional life, is gratefully ac- Latreillia metanesa Williams, 1982a. knowledged; these include Lara Cooper, USNM 74570. Ruth Gibbons, David Hardy, Keiko Hira- Menippe adina Williams & Felder, 1986e. tsuka Moore, Roosevelt McMillan, Mary USNM 228862. Mickevich, Tom Munroe, and Martha Ni- Mimilambridae Williams, 1979c. (Family). zinski. Thanks also to Rose A. Gulledge for Mimilambrus Williams, 1979c. processing electronic images of the photo- Mimilambrus wileyi Williams, 1979c. graphs, and compiling bibliographic infor- USNM 172222. mation; Chad Walter for proof-reading the *Necrocarcinus olsonorum Bishop & Wil- bibliography; and Molly K. Ryan for put- liams, 1991c. SDSM 11000. ting together the figure.