Kenai National Wildlife Refuge Species List, Version 2016-11-18
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Kenai National Wildlife Refuge Species List, version 2016-11-18 Kenai National Wildlife Refuge biology staff November 18, 2016 2 Cover images represent changes to the checklist. Top left: Impatiens noli- tangere at Jims' Landing on August 15, 2016 (http://www.inaturalist. org/observations/3893699), Image CC BY Matt Bowser. Top right: Oryctolagus cuniculus observed at the Refuge's shop area on Ski Hill Road on August 26, 2016 (http://www.inaturalist.org/observations/ 3967164), Image CC BY Matt Bowser. Bottom left: Lycia rachelae ob- served at Headquarters Lake on April 7, 2016 (http://arctos.database. museum/guid/KNWR:Ento:10862), Image CC0 Matt Bowser. Bottom right: Onnia tomentosa observed near the Refuge's headquarters, August 10, 2016 (http://www.inaturalist.org/observations/3855184). Image CC BY Matt Bowser. Contents Contents 3 Introduction 5 Purpose............................................................ 5 About the list......................................................... 5 Acknowledgments....................................................... 5 Refuge checklist 7 Vertebrates .......................................................... 7 Phylum Chordata.................................................... 7 Invertebrates ......................................................... 12 Phylum Annelida.................................................... 12 Phylum Arthropoda .................................................. 13 Phylum Cnidaria.................................................... 31 Phylum Mollusca.................................................... 32 Phylum Platyhelminthes................................................ 32 Vascular Plants........................................................ 32 Division Tracheophyta................................................. 32 Bryophytes .......................................................... 45 Division Bryophyta................................................... 45 Fungi ............................................................. 49 Division Ascomycota.................................................. 49 Division Basidiomycota ................................................ 56 Division Chytridiomycota ............................................... 57 Division Myxomycota ................................................. 57 Unicellular Organisms.................................................... 57 Phylum Cercozoa.................................................... 57 Phylum Heterokonta.................................................. 57 Phylum Protozoa.................................................... 57 Extirpated species 59 Vertebrates .......................................................... 59 Phylum Chordata.................................................... 59 Vascular Plants........................................................ 59 Division Tracheophyta................................................. 59 Change log 61 References 63 Index 65 3 Introduction Purpose nasingham and Hebert, 2013). We have chosen not to fol- low the usual convention of italicizing genus and species A primary purpose for which the Kenai National Wildlife names because our current system does not enable us to Refuge was established in the Alaska National Interest handle accepted and provisional names separately. We want Lands Conservation Act of 1980 is, \to conserve fish and to avoid implying that any of the provisional names we wildlife populations and habitats in their natural diver- use are accepted by formatting them as proper scientific sity. ," where the term “fish and wildlife" is defined as names. Also, note that our database enforces lower case \any member of the animal kingdom, including without lim- letters in our specific epithets so that, for example, \Rab- itation any mammal, fish, bird. , amphibian, reptile, mol- dophaga sp. BOLD:ADA9342" becomes \Rabdophaga sp. lusk, crustacean, arthropod or other invertebrate." An obvi- bold:ada9342." ous first step toward accomplishing this purpose is to know In this draft we added handling of selected synonyms, what fish and wildlife, habitats, and natural diversity are to which are now included in the index. These synonymies do be conserved. This checklist is intended to be a frequently- not necessarily reflect our opinions and are not intended as updated document reflecting our current knowledge of which statements on correct taxonomy; we generally follow ITIS. living things call the Kenai National Wildlife Refuge home. Rather, synonymns are included in the index as an aid to the reader to help find taxa for which names may have changed. This list is dynamic and far from perfect with About the list many changes and corrections yet to be made. Please send comments and corrections to Matt Bowser at The present list includes a total of 1,952 species: 206 matt [email protected]. vertebrates, 681 invertebrates, 494 vascular plants, 181 bryophytes, 361 fungi, and 29 unicellular organisms. Of these, 1,861 are considered to be native, 87 species are con- Acknowledgments sidered to be non-native, and four species are represented by both native and non-native populations. In addition, 17 Many acknowledgments are due to many people who have non-native species are believed to have been extirpated from reviewed this list, provided identifications, and helped in the Refuge. other ways. The list below is by no means exhaustive. Since the last version of this list (Kenai National Wildlife Refuge biology staff, 2016) where 1,925 species were listed, James Bergdahl (Conservation Biology Center, Spokane, there has been a net change of 27 species added including Washington) reviewed and made helpful comments on the non-native species Oryctolagus cuniculus and Leontodon the Refuge's list of Carabidae. autumnalis. Two invertebrate phyla (Cnidaria and Platy- helminthes) have been added. See the change log (page 61) Peter Hovingh provided a list of leeches (Hirudinea) col- for details. lected on the Refuge. The list was produced from an in-house database using John Hudson (USFWS, Juneau, Alaska) provided lists of ITIS (http://www.itis.gov) for its taxonomic backbone. Odonata observed on the Refuge. Accordingly, names and hierarchies generally follow ITIS ex- cept where taxa are not represented in ITIS. Richard Payne (University of York, Heslington, UK) pro- Although accepted scientific names are used wherever vided a list of protists from Jigsaw Lake from Payne these are known, we have included provisional identifications et al.(2006). (for example, \Typhula sp.") where these represent the best information available. Species recognized only as molecu- David Wartinbee (retired from Kenai Peninsula College, lar operational taxonomic units are also included, mainly Soldotna, Alaska) shared a list of Chironomidae he had entities recognized by Barcode Index Numbers (BINs, Rat- collected on the Refuge. 5 Checklist of species occurring on the Kenai National Wildlife Refuge Vertebrates Common names: Dolly varden iNaturalist observations: 811997 Phylum Chordata Salvelinus namaycush Class Actinopterygii Common names: Lake trout Order Cypriniformes yThymallus arcticus Common names: Arctic grayling Family Catostomidae Order Scorpaeniformes Catostomus catostomus Common names: Longnose sucker Family Cottidae Literature records: Dean and Rickabough(2005), Friedersdorff Cottus aleuticus (1986), Friedersdorff and Jakubas(1984a), Friedersdorff and Jakubas Common names: Coastrange sculpin (1984b) Cottus cognatus Order Gasterosteiformes Common names: Slimy sculpin Family Gasterosteidae Class Amphibia Gasterosteus aculeatus Order Anura Common names: Threespine stickleback iNaturalist observations: 2138344 Family Ranidae Pungitius pungitius Lithobates sylvaticus : Tenspined stickleback Common names Synonyms: Rana sylvatica Common names: Wood frog Order Salmoniformes iNaturalist observations: 115158, 115166, 117220, 1816454 Family Salmonidae Class Aves Oncorhynchus gorbuscha Common names: Humpbacked salmon Order Accipitriformes Oncorhynchus keta Family Accipitridae Common names: Chum salmon Accipiter gentilis Oncorhynchus kisutch Common names: Northern goshawk Common names: Coho salmon or silver salmon iNaturalist observations: 114273 Oncorhynchus mykiss Accipiter striatus Common names: Steelhead Common names: Sharp-shinned hawk iNaturalist observations: 811982, 1814201 Aquila chrysaetos Oncorhynchus nerka Common names: Golden eagle Common names: Sockeye salmon or kokanee Buteo jamaicensis iNaturalist observations: 1021584 Common names: Red-tailed hawk Oncorhynchus tshawytscha Buteo lagopus Common names: Chinook salmon or king salmon Common names: Rough-legged hawk Prosopium cylindraceum Circus cyaneus Common names: Round whitefish Common names: Northern harrier Salvelinus alpinus Haliaeetus leucocephalus Common names: Arctic char Common names: Bald eagle Salvelinus malma iNaturalist observations: 353902, 2318743 7 8 REFUGE CHECKLIST Family Pandionidae Melanitta fusca Pandion haliaetus Common names: White-winged scoter Common names: Osprey Melanitta nigra iNaturalist observations: 17348 Common names: Black scoter Melanitta perspicillata Order Anseriformes Common names: Surf scoter Family Anatidae Mergus merganser Anas acuta Common names: Common merganser Common names: Northern pintail Mergus serrator Anas americana Common names: Red-breasted merganser iNaturalist observations: 1741201 Common names: American wigeon iNaturalist observations: 38971 Order Apodiformes Anas clypeata Common names: Northern shoveler Family Trochilidae Anas crecca Selasphorus rufus Common names: Green-winged