Bibliography of Victorian (Insecta: )

I. Endersby

2006

Arthur Rylah Institute for Environmental Research

Technical Report Series No. 157

Arthur Rylah Institute for Environmental Research Technical Report Series No. 157

Bibliography of Victorian Dragonflies (Insecta: Odonata)

Ian Endersby

February 2006

Published by: Arthur Rylah Institute for Environmental Research Department of Sustainability and Environment PO Box 137 Heidelberg, Victoria 3094 Australia Telephone: (03) 9450 8600 www.dse.vic.gov.au/ari

This publication may be cited as: Endersby, I. (2006) Bibliography of Victorian Dragonflies (Insecta: Odonata). Arthur Rylah Institute for Environmental Research, Technical Report Series No.157. Department of Sustainability and Environment, Melbourne.

© The State of Victoria Department of Sustainability and Environment 2006 This publication is copyright. Apart from any fair dealing for private study, research, criticism or review allowed under the Copyright Act 1968, no part of this publication may be reproduced, stored in a retrieval system or transmitted in any forms or by any means, electronic, photocopying or other, without the prior permission of the copyright holder.

ISBN 1 74152 439 3 ISSN 1326 6446

Disclaimer This publication may be of assistance to you but the State of Victoria and its employees do not guarantee that the publication is without flaw of any kind or is wholly appropriate for your particular purposes and therefore disclaims all liability for any error, loss or other consequence which may arise from you relying on any information in this publication.

Front cover photos supplied by Reiner Richter (clockwise from top left): Orthetrum caledonicum (female), Adversaeschna brevistyla, Hemicordulia australiae, weyersii.

Bibliography of Victorian Dragonflies (Insecta: Odonata)

Contents

1 Overview...... 1 2 Format ...... 3 3 Species’ listing...... 4 Order Odonata...... 4 Suborder Zygoptera ...... 4 Family Hemiphlebiidae...... 4 Family ...... 4 Family ...... 4 Family ...... 5 Family ...... 6 Family ...... 6 Family ...... 6 Family Diphlebiidae...... 8 Suborder Epiproctophora ...... 8 Family ...... 8 Family ...... 8 Family ...... 10 Family ...... 10 Family ...... 12 Family ...... 13 Family ...... 13 Family Hemicorduliidae ...... 13 Family ...... 14 4 Acknowledgements...... 16 5 References...... 17 Appendix...... 29

i Bibliography of Victorian Dragonflies

1 Overview

One hundred and sixty-six scientific reports and papers have been located which refer to dragonflies in Victoria by species’ name. This literature can be divided into a number of eras:

Pre - 1900 Baron Edmund Sélys de Longchamps who lived in Belgium was a prolific worker in the late nineteenth century, decribing dragonflies provided by a world-wide network of collectors. His contribution to Victorian odonatology was in the naming of species where that State was the type locality.

1900- 1930 It is quite likely that R. Billinghurst was the source of some of Sélys’ specimens as he collected in Alexandra where the type of originated. He also sent specimens to R. Martin in France who described new species. Robin Tillyard, first Chief Entomologist of CSIR, specialised in the Odonata. While much of his research concentrated on collections from New South Wales and Queensland a number of his taxonomic and biological papers covered Victorian species.

1955 -1965 F.C. Fraser had a strong interest in “Far Eastern” dragonflies and contributed some taxonomic works including his Handbook which was the first comprehensive identification key to the Australian species.

Post 1973 Since the early 1970’s there has been an escalation in the literature within Australia so there is some value in dividing it into five-year periods. The number of papers classified by subject for Victoria has been:

biology distribution surveys excursions conservation theses Total 1971 - 1975 2 1 2 1 6 1976 - 1980 4 2 2 8 1981 - 1985 7 2 7 1 1 18 1986 - 1990 3 4 12 2 3 24 1991 - 1995 6 5 3 17 5 3 39 1996 - 2000 7 2 2 13 3 3 2 32 2001 - 2005 4 4 2 2 1 13

“Distribution” includes checklists and new distribution records for the State. “Surveys” have been carried out by statutory bodies or institutions, generally for environmental monitoring and river health analyses. “Excursions” were the results of amateur sampling or overseas visitors. “Theses” report research carried out at Monash or La Trobe Universities which included odonates amongst the taxa studied or sampled.

Considering each half-decade in turn:

1971 - 1975 Tony Watson’s paper on the distribution of Australian dragonflies was the stimulus for John Hutchinson to publish the first checklist for Victoria. Brian Timms produced the first higher degree that included Odonata in its species’ list.

1 Bibliography of Victorian Dragonflies

1976 - 1980 This is the start of the Watson era in Australian odonate studies with a number of taxonomic papers and new distribution records. John Hutchinson added additional species to the State’s list and the first major survey was conducted by the (then) National Museum of Victoria staff as part of the Dartmouth Dam project.

1981 - 1985 Taxonomy studies blossomed with contributions from Watson and Günther Theischinger, as well as Stewart and Maurits Lieftinck who, although based in the Netherlands, worked on specimens from the former Dutch East Indies colonies to Australia’s north. Survey work with an environmental health emphasis was started under the auspices of the Rural Water Commission, the Environmental Protection Authority and by Museum staff.

1986 - 1990 The Zoological Catalogue of Australia’s volume containing the Odonata was published as was John Hawking’s identification guide to larvae. This was a big half-decade for environmental survey reports and theses from Monash and La Trobe Universities included dragonfly taxa. Hemiphlebia mirabilis spawned what was probably the first conservation oriented analysis for the State and it was then used as case study material in broader contexts.

1991 - 1995 The big event was the publication of the Australian Dragonflies by Watson, Theischinger & Abbey and Theischinger continued with a number of additional taxonomic studies. Dragonflies as flagship species were introduced into the conservation literature and Hemiphlebia mirabilis was listed under the State Flora & Fauna Guarantee. Many environmental surveys included odonate taxa in their sampling. John Hawking’s thesis produced a number of ancillary papers on the biology of the species he studied.

1996 - 2000 A number of phylogenetic studies by overseas workers had relevance to Australian species. John Hawking and Günther Theischinger incorporated some of those decisions in their revision of the Australian checklist which they incorporated into their identification guide to larvae. The guide covered all Australian families and to species level for New South Wales. All Victorian species are included either within monotypic familes or the generic and species accounts. Theischinger complemented the earlier NSW identification guide with additional guides for selected taxa examined Australia-wide. These were presented at annual taxonomy seminars hosted by the Murray Darling Freshwater Research Centre at Albury. Endersby adopted Hawking & Theischinger’s classification to produce a checklist for Victoria which documented taxonomic and distribution changes since Hutchinson’s list of 1975. Environmental and river health surveys continued and some higher degree theses were presented which contained reference to Odonates. John Hawking prepared a review of the conservation status of all Australian species

2001 - 2005 Theischinger continued to present identification guides to the larvae of particular families at the annual taxonomic seminars and in 2003 he and Hawking produced the field guide to Victorian adults and larvae for the World Dragonfly Association Symposium held in Beechworth that year. Some excursion reports also resulted from that Symposium.

2 Bibliography of Victorian Dragonflies

2 Format

The primary criterion for inclusion in this State bibliography is that the reference must cite, at identified species’ level, an occurrence within Victoria. The Species Listing follows the order of Endersby (2000) and categorises each reference by one or more of:

[td] type description [t] taxonomy and identification [d] distribution [b] biology (including behaviour) [c] conservation

Strictly all papers should have been designated as “distribution” as that is one of the criteria for inclusion. If a reference mentions the species as occurring Victoria, but has no additional information, the paper is not included in the distribution classification; that is reserved for reports of surveys which have an identifiable locality but little additional biological information. Odonata are rarely recorded in surveys and they play little part in statistical analyses or project discussion. Site specific reports would be useful in determining species’ ranges within the State.

Type descriptions are included only for those species where the type locality is Victoria or a site within the State. A list of those type localities is given after the reference section. Each species has been allocated a unique identification number. The Reference citations are followed by a list of identification numbers of the species which it includes. References which cover almost the entire fauna have not had their citation annotated with species number. They are:

Endersby 2000a, b Houston & Watson 1988 Watson 1974

Fraser 1960 Hutchinson 1975 Watson, Theischinger & Abbey 1991

Hawking 1986 Nuttall 1982a

Hawking & Theischinger, G. & Hawking, J.H., Theischinger 1999 2003

3 Bibliography of Victorian Dragonflies

3 Species’ listing

ORDER ODONATA SUBORDER ZYGOPTERA FAMILY HEMIPHLEBIIDAE 1. Hemiphlebia mirabilis Selys, 1869 Selys 1869a (td); Martin 1901; Billinghurst 1902 (d); Tillyard 1912 (b); Tillyard 1928a (b); Tillyard 1928b (t); Burns 1955 (d); Fraser 1955 (b); Fraser 1960 (t); Neboiss 1962 (d); Watson 1974; Donnelly 1974 (d); Hutchinson 1975; Nuttall 1982a (t); Watson 1982 (c); Davies 1985 (d,c); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (b,c); Yen et al. 1990 (c); SAC 1991 (c); Watson et al. 1991 (t); Endersby 1992 (d,b); Trueman et al. 1992 (d,c); Birkin, Quin & Jelinek 1993 (c); Endersby 1993(d); New 1993 (c); Watson 1995 (c); Hawking 1997 (c); Hawking 1999 (c); Trueman 1999 (t); Endersby 2000a; Paulson 2003 (d).

FAMILY SYNLESTIDAE 2. Selys, 1869 Selys 1869b (td); Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (as Synlestes tillyardi) (t); Watson 1974 (as Synlestes tillyardi & S. weyersi); Hutchinson 1975 (as S. tillyardi & S. weyersi); Nuttall 1982a(t); Hawking 1986 (as S. tillyardi & S. weyersi) (t,d); Houston & Watson 1988 (as S. tillyardi & S. weyersi) (t); Bennison et al. 1989 (as S. tillyardi) (d); Pettigrove 1989 (as S. tillyardi) (d); Watson et al. 1991 (as S. w. weyersii & S. w. tillyardi) (t); O’Connor 1993 (d); Theischinger, Watson & Rowe 1993 (d); Hawking 1994 (t,d,b); Marchant et al. 1997 (d); Hawking 1999 (c); Hawking & New 1999(d,b); Hawking & Theischinger 1999 (t); Pettigrove & Coleman 1999 (d); Endersby 2000a; Hawking & New 2002 (as S. tillyardi) (d), Endersby 2005 (d).

FAMILY LESTIDAE 3. analis (Rambur, 1842) Martin 1901 (as Lestes analis); Billinghurst 1902 (as L. analis); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Nuttall 1982a (t); Hawking 1986 (t); Houston & Watson 1988 (t); Sant & New 1988 (d); Lake et al. 1989 (d,b); Dobson 1990 (d); Watson et al. 1991 (t); Cameron 1992 (d); Hawking 1994 (t,d,b); Crowther 1995 (d); Dobrosak 1997 9d); Hawking 1999 9c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a.

4. Austrolestes annulosus (Selys, 1862) Martin 1901 (as Lestes annulosa); Billinghurst 1902 (as Lestes annulosa) (d); Tillyard 1913 (b); Fraser 1960 (t); Hutchinson 1973 (d); Timms 1973a (d,b); Watson 1974; Hutchinson 1975; Nuttall 1982a (t); Nuttall 1982b (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Robinson 1988 (d); Driver 1991 (d); Watson et al. 1991 (t); Cameron 1992 (d); Reed 1992 (d); Crowther 1995 (d); Dobson 1995 (d); Kefford & Robley 1996 (d); Endersby 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

5. Austrolestes aridus (Tillyard, 1908) Hutchinson 1975; Watson 1977; Nuttall 1982a (t); Hawking 1986 (t); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d).

4 Bibliography of Victorian Dragonflies

6. Austrolestes cingulatus (Burmeister, 1839) Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Nuttall 1982a (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Dobson 1990 (d); Watson et al. 1991 (t); Cameron 1992 (d); Crowther 1995 (d); Endersby 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

7. Austrolestes io (Selys, 1862) Martin 1901 (as Lestes io); Watson 1974; Hutchinson 1975; Nuttall 1982a (t); Hawking 1986 (t,d); Boulton 1988 (d); Houston & Watson 1988 (t); Boulton 1989 (d,b); Watson et al. 1991 (t); Boulton & Lake 1992a (d); Endersby 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

8. Austrolestes leda (Selys, 1862) Selys 1862a (as Lestes leda) (td); Martin 1901 (as Lestes leda); Billinghurst 1902 (as L. leda) (d); Tillyard 1913 (b); Chapman 1925 (as L. leda) (d,b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (d); Dobson 1990 (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Crowther 1995 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a, Endersby 2005 (d).

9. Austrolestes psyche (Hagen in Selys, 1862) Martin 1901 (as L. psyche); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Watson & Moulds 1979 (d); Nuttall 1982a (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Sant & New 1989 (b); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Crowther 1995 (d); Endersby 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a.

FAMILY MEGAPODAGRIONIDAE 10. Austroargiolestes calcaris (Fraser, 1958) Watson 1974 (as Argiolestes calcaris); Hutchinson 1975 (as Argiolestes calcaris); Theischinger & O’Farrell 1986 (t); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Theischinger 1998a (t); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a, Endersby 2005 (d).

11. Austroargiolestes icteromelas (Selys, 1862) Selys 1862b (as Argiolestes icteromelas) (td); Martin 1901 (as Argiolestes icteromelas); Billinghurst 1902 (as Argiolestes icteromelas) (d); Tillyard 1913 (as Argiolestes icteromelas) (b); Watson 1974 (as Argiolestes icteromelas); Hutchinson 1975 (as Argiolestes icteromelas); Smith et al. 1977 (as Argiolestes icteromelas) (d); Smith et al. 1978 (as Argiolestes icteromelas) (d); Malipatil & Blyth 1982 (as Argiolestes ?icteromelas) (d,b); Hawking 1986 (as Argiolestes icteromelas) (t,d); Theischinger & O’Farrell 1986 (t); Hawking 1994 (t,d,b); Houston & Watson 1988 (t); Watson et al. 1991 (t); Theischinger 1998a (t); Pettigrove & Coleman 1998 (as Argiolestes ictermelas sic) (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Pettigrove & Coleman 1999 (as Argiolestes ictermelas sic) (d); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

12. Griseargiolestes eboracus (Tillyard, 1913) Martin 1901 (within Argiolestes grisea); Billinghurst 1902 (within Argiolestes grisea) (d); Tillyard 1913 (within Argiolestes griseus) (b); Watson 1974 (within Argiolestes griseus); Hutchinson 1975 (within Argiolestes griseus); Hawking 1986 (within Argiolestes griseus) (t,d); Houston & Watson 1988 (as Argiolestes griseus eboracus) (t); Watson et al. 1991 (as Argiolestes eboracus) (t); Dobrosak 1997 (as Argiolestes eboracus) (d); Theischinger 1998a (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

5 Bibliography of Victorian Dragonflies

13. Griseargiolestes intermedius (Tillyard, 1913) Martin 1901 (within Argiolestes grisea); Billinghurst 1902 (within Argiolestes grisea) (d); Tillyard 1912 (as Argiolestes griseus intermedius) (td); Fraser 1959 (as Argiolestes griseus intermedius) (d); Fraser 1960 (as Argiolestes griseus intermedius) (t); Watson 1974 (within Argiolestes griseus); Hutchinson 1975 (within Argiolestes griseus); Hawking 1986 (within Argiolestes griseus) (t,d); Houston & Watson 1988 (as Argiolestes griseus intermedius) (t); Watson et al. 1991 (as Argiolestes intermedius) (t); Theischinger 1998a (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

FAMILY PROTONEURIDAE 14. Nososticta solida (Hagen in Selys, 1860) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Watson 1974; Hutchinson 1975; Campbell et al. 1982 (as fam. Protoneuridae) (d); Watson & Theischinger 1984 (t,d); Hawking 1986 (t,d); Metzeling 1987 (d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson et al. 1991 (t); Metzeling et al. 1993 (d); Hawking 1994 (d,t,b); Kefford & Robley 1996 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

FAMILY ISOSTICTIDAE 15. Labidiosticta vallisi (Fraser, 1955) Watson 1974 (as Oristicta vallisi); Hutchinson 1975 (as O. vallisi); Houston & Watson 1988 (as O. vallisi) (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

16. Rhadinosticta simplex (Martin, 1901) Martin 1901 (as Isosticta simplex) (td); Billinghurst 1902 (as I. simplex) (d); Tillyard 1913 (as I. simplex) (b); Fraser 1960 (as I. simplex) (t); Watson 1974 (as I. simplex); Hutchinson 1975 (as I. simplex); Hawking 1986 (as I. simplex) (t,d); Houston & Watson 1988 (as I. simplex) (t); Bennison et al. 1989 (as I. simplex) (d); Pettigrove 1989 (as I. simplex) (d); Watson et al. 1991 (t); Metzeling et al. 1993 (as I. simplex) (d); Hawking 1994 (t,d,b); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Endersby 2001b (b), Endersby 2005 (d).

FAMILY COENAGRIONIDAE 17. Austroagrion watsoni Lieftinck, 1982 Martin 1901 (as Pseudagrion cyane); Billinghurst 1902 (as P. cyane) (d); Tillyard 1913 (as Austroagrion cyane) (b); Fraser 1960 (as A. cyane) (t); Watson 1974 (as A. cyane); Hutchinson 1975 (as A. cyane); Leiftinck 1982 (td); Nuttall 1982a ( as A. cyane) (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Robinson 1988 (d); Bennison et al. 1989 (as A. cyane) (d); Dobson, 1990 (as A. cyane) (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Cameron 1992 (also as A. cyane) (d); Hawking 1994 (t,d,b); Crowther 1995 (also as A. cyane) (d); Endersby 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a, Endersby 2005 (d).

18. Austrocnemis splendida (Martin, 1901) Martin 1901 (as Agriocnemis splendida) (td); Billinghurst 1902 (as Agriocnemis splendida) (d); Tillyard 1907 (as Agriocnemis splendida) (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Cameron 1992 (d); Reed 1992 (d); O’Connor 1993 (d); Ferdinands & Bennison 1994 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Deveraux et al. 2000 (d); Endersby 2000a.

6 Bibliography of Victorian Dragonflies

19 Caliagrion billinghursti (Martin, 1901) Martin 1901 (as Pseudagrion billinghursti) (td); Billinghurst 1902 (as P. billinghursti) (d); Fraser 1960 (t); Watson 1974; Hutchinson 1975 (as “Caliagrion lyelli”); Nuttall 1982a (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

20. Coenagrion lyelli (Tillyard, 1913) Tillyard 1912 (as Agrion lyelli) (t,b); Fraser 1960 (t); Watson 1974; Hutchinson 1975 (as “Caliagrion lyelli”); Nuttall 1982a (t); Houston & Watson 1988 (t); Sant & New 1988 (d); Crowther 1995 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

21. Ischnura aurora (Brauer, 1865) Martin 1901 (as I. delicata); Billinghurst 1902 (as I. delicata) (d); Tillyard 1913 (b); Fraser 1960 (t); Timms 1973a (d,b); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Williams 1981 (d,b); Nuttall 1982a (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Bennison et al. 1989 (d); Lake et al. 1989 (d,b); Dobson 1990 (d); Reed & Newall 1990 (d); Hart et al. 1991 (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Cameron 1992 (d); Hawking 1994 (d,t,b); Smith, Ferdinands & Bennison 1994 (d); Crowther 1995 (d); Marchant et al. 1997 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

22 Ischnura heterosticta (Burmeister, 1839) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Timms 1973a (d,b); Timms 1973b (d); Watson 1974; Hutchinson 1975; Watson 1976 (d); Timms 1981 (d); Nuttall 1982a (t); Nuttall 1982b (d); Blyth et al. 1984 (d); Marchant et al. 1984 (d); Metzeling et al. 1984 (d); Hawking 1986 (t,d); Chessman & Robinson 1987 (d,b); Robinson 1987 (d); Houston & Watson 1988 (t); Robinson 1988 (d); Sant & New 1988 (d); Bennison et al. 1989 (d); Dobson 1990 (b); Driver 1991 (d,b); Watson et al. 1991 (t); Williams et al. 1991 (d); Cameron 1992 (d); Reed 1992 (d); O’Connor 1993 (d); Ferdinands & Bennison 1994 (d); Hawking 1994 (t,d,b); Smith, Ferdinands & Bennison 1994 (d); Crowther 1995 (d); Kefford & Robley 1996 (d,b); Mitchell et al. 1996 (d); Marchant et al. 1997 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Marchant et al. 1999 (b); Deveraux et al. 2000 (d); Endersby 2000a; Quinn et al. 2000 (d,b); Hawking & New 2002 (d), Endersby 2005 (d).

23. Pseudagrion aureofrons Tillyard, 1906 Hawking 1986 (t,d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

24. Xanthagrion erythroneurum (Selys, 1876) Selys 1876 as (Agrion (Xanthagrion ) erythroneurum) (td); Martin 1901; Billinghurst 1902 (d); Timms 1973a (d,b); Watson 1974; Hutchinson 1975; Nuttall 1982a (t); Garrett 1984 (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Dobson 1990 (d); Driver 1991 (d); Watson et al. 1991 (t); Cameron 1992 (d); Crowther 1995 (d); Pettigrove & Coleman 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

7 Bibliography of Victorian Dragonflies

FAMILY DIPHLEBIIDAE 25. lestoides (Selys, 1853) Martin 1901; Billinghurst 1902 (d); Tillyard 1909; Tillyard 1913 (b); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Stewart 1980 t,d); Stewart 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Doeg 1995a (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

26. Diphlebia nymphoides Tillyard, 1911 Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Stewart 1980 (t,d); Stewart 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

SUBORDER EPIPROCTOPHORA FAMILY AUSTROPETALIIDAE 27. Austropetalia tonyana Theischinger 1995 Fraser 1960 (as A. patricia) (t); Neboiss 1962 (as A patricia) (d); Watson 1974 (as A. patricia); Hutchinson 1975 (as A. patricia); Hawking 1986 (as A. patricia) (t,d); Houston & Watson 1988 (as A. patricia) (t); Watson et al. 1991 (as A. patricia) (t); Hawking 1994 (as A. patricia) (t,d,b); Theischinger 1995a (td); Carle 1996 (as A. patricia and possibly as A. victoria) (t); Lohmann 1996 (t); Hawking 1997 (c); Hawking 1999 (c); Hawking & New 1999 (d); Hawking & Theischinger 1999 (t); Papas et al. 1999 (d); Endersby 2000a; Theischinger 2002 (t).

FAMILY TELEPHLEBIIDAE 28. Austroaeschna atrata Martin, 1909 Martin 1909 (td); Tillyard 1913 (b); Tillyard 1916 (t,d); Fraser 1960 (t); Watson 1974 (as Acanthaeschna atrata); Hutchinson 1975 (as Acanthaeschna atrata); Theischinger 1982 (t); Hawking 1986 (t,d); Houston & Watson 1988 (t); Hawking & Watson 1990 (b); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t), Endersby 2005 (d).

29. Austroaeschna flavomaculata Tillyard, 1916 Watson 1974 (as Acanthaeshna flavomaculata); Hutchinson 1975 (as Acanthaeshna flavomaculata); Theischinger 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Doeg 1996 (d); Endersby 1998 (d); Saddlier & Doeg 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2002 (t), Endersby 2005 (d).

30. Austroaeschna inermis Martin, 1901 Martin 1901 (td); Billinghurst 1902 (d); Tillyard 1916 (td); Fraser 1960 (t); Watson 1974 (as Acanthaeschna inermis); Hutchinson 1975 (as Acanthaeschna inermis); Theischinger 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Pettigrove 1989 (d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2002, Endersby 2005 (d).

8 Bibliography of Victorian Dragonflies

31. Austroaeschna multipunctata (Martin, 1901) Martin 1901 (as Planaeschna multipunctata) (td); Billinghurst 1902 (as P. multipunctata) (d); Tillyard 1913 (b); Tillyard 1916 (t,b); Fraser 1960 (t); Watson 1974 (as Acanthaeschna multipunctata); Hutchinson 1975 (as Acanthaeschna multipunctata); Theischinger 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Saddlier & Doeg 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t), Endersby 2005 (d).

32. Austroaeschna parvistigma Selys, 1883 Martin 1901; Billinghurst 1902 (d); Watson 1974 (as Acanthaeschna parvistigma); Hutchinson 1975 (as Acanthaeschna parvistigma); Theischinger 1982 (t,d); Hawking 1986 (t,d); Boulton 1988 9b); Houston & Watson 1988 (t); Watson et al. 1991 (t); Williams et al. 1991 (d); Boulton & Lake 1992a (d); Doeg 1996 (d); Saddlier & Doeg 1998 (d); Vazquez 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t).

33. Austroaeschna pulchra Tillyard in Martin, 1909 Martin 1901 (as Acanthaeschna unicornis); Tillyard 1913 (as Austroaeschna unicornis) (b); Tillyard 1916 (as Austroaeschna unicornis) (t); Fraser 1960 (as Austroaeschna unicornis) (t); Watson 1974 (as Acanthaeschna unicornis); Hutchinson 1975 (as Acanthaeshna unicornis); Malipatil & Blyth 1982 (d,b); Theischinger 1982 (t,d); Barmuta 1986 (d); Hawking 1986 (t,d); Metzeling & Newall 1987 (d); Robinson 1987 (d); Houston & Watson 1988 (t); Metzeling 1990a, 1900b (d); Watson et al. 1991 (t); Doeg & Saddlier 1992, 1993, 1994 (d); Hawking 1994 (t,d,b); Doeg 1995a (d); Endersby 1998 (d); Pettigrove & Coleman 1998 (d); Saddlier & Doeg 1998 (d); Vazquez 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Marchant et al. 1999 (b); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t), Endersby 2005 (d).

34. Austroaeschna subapicalis Theischinger, 1982 Theischinger 1982 (td); Barmuta 1986 9d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Doeg 1996 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t).

35. Austroaeschna unicornis (Martin, 1901) Martin 1901 (as Planaeschna longissima) (td); Billinghurst 1902 (as P. longissima) (d); Tillyard 1913 (as Austroaeschna longissima) (b); Tillyard 1916 (as Austroaeschna longissima) (t,d,b); Fraser 1960 (as Austroaeshna longissima) (t); Watson 1974 (as Acanthaeschna longissima); Hutchinson 1975 (as Acanthaeshna longissima); Smith et al. 1977 (as Acanthaeschna longissima) (d); Smith et al. 1978 (as Acanthaeschna longissima) (d); Theischinger 1982 (t,d); Hawking 1986 (t,d); Metzeling & Newall 1987 ( as Anacantha [sic] unicornis) (d); Robinson 1987 (d); Houston & Watson 1988 (t); Bennison et al. 1989 (as Anacantha [sic] unicornis) (d); Metzeling 1990a, 1990b (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Doeg & Saddlier 1993 (d); O’Connor 1993 (d); Ferdinands & Bennison 1994 (d); Hawking 1994 (t,d,b); Smith, Ferdinands & Bennison 1994 (d); Marchant et al. 1997 (d); Hawking 1999 (c); Hawking & New 1995a (b); Hawking & New 1996 (b); Saddlier & Doeg 1997, 1998 (d); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Hawking & New 2002 (d); Theischinger 2002 (t), Endersby 2005 (d).

36. Dendroaeschna conspersa (Tillyard, 1907) Houston & Watson 1988 (t); Hawking 1991 (d); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2002 (t).

9 Bibliography of Victorian Dragonflies

37. Notoaeschna sagittata (Martin, 1901) Martin 1901 (as Planaeschna sagittata) (td); Billinghurst 1902 (as P. sagitta [sic]) (d); Tillyard 1913 (as Austroaeschna sagittata) (b); Tillyard 1916 (td-male,b); Watson 1974; Hutchinson 1975; Malipatil & Blyth 1982 (d,b); Theischinger 1982 (t,d); Metzeling et al. 1984 (d); Hawking 1986 (t,d); Robinson 1987 (d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Doeg 1995a (d); Saddlier & Doeg 1998 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Marchant et al. 1999 (b); Dean 2000 (d); Endersby 2000a; Theischinger 2002 (t); Hawking et al. 2004 (d,b), Endersby 2005 (d).

38. Spinaeschna tripunctata (Martin, 1901) Martin 1901 (as Planaeschna tripunctata) (td); Billinghurst 1902 (as P. tripunctata) (d); Tillyard 1913 (as Austroaeschna tripunctata) (b); Tillyard 1916 (as Austroaeschna tripunctata) (d); Watson 1974 (as Acanthaeshna tripunctata); Hutchinson 1975 (as Acanthaeshna tripuncta); Theischinger 1982 (t,d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Pettigrove 1989 (d); Watson et al. 1991 (t); Doeg 1995a (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2002 (t).

39. Telephlebia brevicauda Tillyard, 1916 Martin 1901 (as T. godefroyi); Billinghurst 1902 (as T. godefroyi) (d); Tillyard 1916 (as T. godeffroyi brevicauda) (td); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Theischinger 1985 (t,d); Hawking 1986 (t,d); Metzeling 1986 (d); Houston & Watson 1988 (t); Metzeling 1990b (d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Papas et al. 1999 (d); Endersby 2000a; Theischinger 2002 (t), Endersby 2005 (d).

FAMILY AESHNIDAE 40. brevistyla (Rambur, 1842) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Timms 1973b (d); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Malipatil & Blyth 1982 (d,b); Hawking 1986 (d,b); Houston & Watson 1988 (t); Robinson 1988 (d); Sant & New 1988 (d); Lake et al. 1989 (d,b); Dobson 1990 (d); Reed & Newall 1990 (d); Watson et al. 1991 (t); Williams et al. 1991(d); Cameron 1992 (d); Reed 1992 (d); Crowther 1995 (d); Mitchell et al. 1996 (d); Endersby 1998 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d); Theischinger 2002 (t), Endersby 2005 (d).

41. Hemianax papuensis (Burmeister, 1839) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (as Anax papuensis) (b); Tillyard 1916 (as Anax papuensis) (t,b); Fraser 1960 (t); Hutchinson 1973 (d); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Metzeling & Pettigrove 1988 (d); Bennison et al. 1989 (d); Lake et al. 1989 (d,b); Dobson 1990 (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Williams et al. 1991 (d); Cameron 1992 (d); O’Connor 1993 (d); Pouliet 1993 (d); Crowther 1995 (d); Hawking & New 1995a (b); Hawking & New 1996 (b); Endersby 1998 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d); Theischinger 2002 (t), Endersby 2005 (d).

FAMILY GOMPHIDAE 42. Antipodogomphus acolythus (Martin, 1901) Martin 1901 (as Austrogomphus acolythus) (td); Fraser 1951 (as Antipodogomphus acolytus) (t); Watson 1974; Hutchinson 1975 (as Antipodogomphys acolythus); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson 1991 (t,d); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t).

10 Bibliography of Victorian Dragonflies

43. Austrogomphus angelorum (Tillyard, 1913) Houston & Watson 1988 (as Austrogomphus angeli) (t); Watson 1991 ( as A. angeli) (t,d); Watson et al. 1991 (as A. angeli) (t); Campbell 1992 (as A. angeli) (b); Peterson 1993 (t,d); Hawking 1997 ( as A. angeli) (c); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t).

44. Austrogomphus australis Dale in Selys, 1854 Martin 1901; Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Metzeling 1986 (d); Metzeling 1987 (d); Metzeling & Newall 1987 (d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Pettigrove 1989 (d); Metzeling 1990b (d); Watson 1991 (t,d); Watson et al. 1991 (t); Metzeling et al. 1993 (d); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1999 (d); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000.

45. Austrogomphus cornutus Watson, 1991 Watson 1974 (as Austrogomphus sp. “c”); Hutchinson 1975 (as A. sp. “c”); Hawking 1986 (as A. sp. “c”) (t,d); Metzeling 1986 ( as A. Watson m/s) (d); Houston & Watson 1988 ( as Austrogomphus doddi) (t); Bennison et al. 1989 (d); Metzeling 1990a (d); Watson 1991 (td,d); Watson et al. 1991 (t); Campbell 1992 (t); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1995a (b); Hawking & New 1995b (d,b); Hawking & New 1996 (b); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t); Hawking & New 2002 (d).

46. Austrogomphus guerini (Rambur, 1842) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Blyth et al. 1984 (d); Marchant et al. 1984 (d); Metzeling et al. 1984 (d); Hawking 1986 (t,d); Houston & Watson 1988 9t); Watson 1991 (t,d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Endersby 1998 (d); Theischinger 1998b (t,d); Endersby 1999 (d); Hawking 1999 (c). Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t), Endersby 2005 (d).

47. Austrogomphus ochraceus (Selys, 1869) Selys 1869c (td); Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988(t); Bennison et al. 1989 (d); Pettigrove 1989 (d); Reed & Newall 1990 (d); Watson 1991 (t,d); Watson et al. 1991 (t); Williams et al. 1991 (d); O’Connor 1993 (d); Hawking 1994 (t,d,b); Doeg 1995a (d); Hawking 1999 (c); Hawking & New 1995a (b); Hawking & New 1995b (d,b); Hawking & New 1996 (b); Marchant et al. 1997 (d); Theischinger 1998b (t,d); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t); Hawking & New 2002 (d), Endersby 2005 (d).

48. Hemigomphus gouldii (Selys, 1854) Martin 1901 (as Austrogomphus. gouldi); Billinghurst 1902 (as A. gouldi) (d); Watson 1974; Hawking 1986 (t,d); Metzeling & Newall 1987 (d); Houston & Watson 1988 (t); Bennison et al. 1989 9d); Pettigrove 1989 (d); Metzeling 1990a (d); Watson 1991 (t,d); Watson et al. 1991 (t), Campbell 1992 (b); Doeg & Saddlier 1992 (d); Doeg & Saddlier 1993 (d); Doeg & Saddlier 1994 (d); Hawking 1994 (t,d,b); Marshall 1996 (d,b); Hawking 1999 (c); Doeg 1995b (d); Hawking & New 1995a (b); Hawking & New 1995b (d,b); Hawking & New 1996 (b); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2000 (t); Dean 2000 (d), Endersby 2005 (d).

11 Bibliography of Victorian Dragonflies

49. Hemigomphus heteroclytus Selys, 1854 Martin 1901 (as Austrogomphus heteroclytus); Tillyard 1913 (as A. heteroclitus) (b); Fraser 1960 (t); Watson 1974 (within Hemigomphus gouldii); Hawking 1986 (t); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson 1991 (t,d); Watson et al. 1991 (t); Campbell 1992 (b); Hawking 1994 (t,d,b); Doeg 1995a (d); Hawking 1999 (c); Hawking & New 1999 (d); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2000 (t); Hawking & New 2002 (d), Endersby 2005 (d).

FAMILY SYNTHEMISTIDAE 50. Archaeosynthemis orientalis (Hagen, ) Tillyard 1913 (as Synthemis macrostigma orientalis) (b); Fraser 1960 (as S. m. orientalis) (t); Watson 1974 (as S. m. orientalis); Hutchinson 1975 (as S. m. orientalis); Hawking 1986 (as S. m. orientalis) (t,d); Houston & Watson 1988 (as S. m. orientalis) (t); Watson et al. 1991 (as S. m. orientalis) (t); Hawking 1999 (as A. macrostigma) (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t); Endersby 2001a (t).

51. Eusynthemis brevistyla (Selys, 1871) Martin 1901 (as Synthemis brevistyla); Billinghurst 1902 (as S. brevistyla) (d); Tillyard 1913 (as Metathemis brevistyla) (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Pettigrove 1989 (d); Watson et al. 1991 (as Eusynthemis brevistyla brevistyla) (t); Doeg & Saddlier 1992 (d); Doeg & Saddlier 1993 (d); Hawking 1994 (t,d,b); Doeg 1995b (d); Saddlier & Doeg 1998 (d); Vazquez 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Dean 2000 (d); Endersby 2000a; Theischinger 2001 (t), Endersby 2005 (d).

52. Eusynthemis guttata (Selys, 1871) Martin 1901 (as Synthemis guttata); Billinghurst 1902 (as S. guttata) (d); Tillyard 1913 (as Metathemis guttata) (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (as Eusynthemis guttata guttata) (t); Hawking 1994 (t,d,b); Theischinger 1995b (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t); Kellar et al. n.d. (d), Endersby 2005 (d).

53. Eusynthemis tillyardi Theischinger 1995 Hawking 1986 (within E. guttata) (t,d); Hawking 1994 (within E. guttata) (t,d,b); Theischinger 1995b (td); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t); Kellar et al. n.d. (d).

54. Eusynthemis virgula (Selys, 1874) Martin 1901 (as Synthemis virgula); Billinghurst 1902 (as S. virgula) (d); Tillyard 1913 (as Metathemis virgula) (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson et al. 1991 (t); O’Connor 1993 9d); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t); Kellar et al. n.d. (d).

55. Parasynthemis regina (Selys, 1874) Fraser 1960 (as Synthemis regina) (t); Watson 1974 (as S. regina); Hutchinson 1975 (as S. regina); Hawking 1986 (as S. regina) (t); Houston & Watson 1988 (as S. regina) (t); Watson et al. 1991 (as S. regina) (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t).

12 Bibliography of Victorian Dragonflies

56. Synthemis eustalacta (Burmeister, 1839) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Malipatil & Blyth 1982 (d,b); Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Endersby 1998 (d); Saddlier & Doeg 1998 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t), Endersby 2005 (d).

FAMILY AUSTROCORDULIIDAE 57. Apocordulia macrops Watson, 1980 Watson 1980 (td); Theischinger & Watson 1984 (t,d); Hawking 1986 (t,d); Robinson 1987 (d); Houston & Watson 1988 (t); Bennison et al. 1989 (t); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Hawking 1999 (c); Hawking & New 1995a (b); Hawking & New 1996 (b); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Theischinger 2001 (t).

58. refracta Tillyard, 1909 Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Endersby 2001a (t); Theischinger 2001 (t).

FAMILY CORDULEPHYIDAE 59. pygmaea Selys, 1870 Selys 1870 (td); Billinghurst 1902 (d); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Endersby 1995 (d); Hawking 1999 (c); Hawking & New 1999 (d); Hawking & Theischinger 1999 (t); Endersby 2000a.

FAMILY HEMICORDULIIDAE 60. Hemicordulia australiae (Rambur, 1842) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Timms 1973b (d); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Dobson 1990 (d); Watson et al. 1991 (t); Crowther 1995 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a, Endersby 2005 (d).

61. Hemicordulia tau (Selys, 1871) Selys 1871 (as Cordulia (Hemicordulia) tau) (td); Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Hutchinson 1973 (d); Timms 1973a (d,b); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Timms 1981 (d); Garrett 1984 (d); Hawking 1986 (t,d); Metzeling 1986 (d); Metzeling 1987 (d); Metzeling & Newall 1987 (d); Boulton 1988 (d); Houston & Watson 1988 (t); Robinson 1988 (d); Bennison et al. 1989 (d); Boulton 1989 (d); Dobson 1990 (d); Metzeling 1990a (d); Reed & Newall 1990 (d); Hart et al. 1991 (d,b); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Boulton & Lake 1992a (d); Boulton & Lake 1992b (d); Cameron 1992 (d); Reed 1992 (d); Metzeling et al. 1993 (d); O’Connor 1993 (d); Pouliet 1993 (d); Ferdinands & Bennison 1994 (d); Hawking 1994 (t,d,b); Crowther 1995 (d); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

62. Procordulia jacksoniensis (Rambur, 1842) Martin 1901 (as Somatochlora jacksoniensis); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Mitchell & Clark 1991 (d); Watson et al. 1991 (t); Reed 1992 (d); O’Connor 1993 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

13 Bibliography of Victorian Dragonflies

FAMILY LIBELLULIDAE 63. Austrothemis nigrescens (Martin, 1901) Martin 1901 (as Diplax nigrescens) (td); Billinghurst 1902 (as D. nigrescens) (d); Tillyard 1913 (b); Watson 1974; Hutchinson 1975; Metzeling et al. 1984 (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

64. Crocothemis nigrifrons (Kirby, 1894) Hawking 1986 (t,d); Houston & Watson 1988 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

65. Diplacodes bipunctata (Brauer, 1865) Martin 1901 (as Diplax rubra); Billinghurst 1902 (as Diplax bipunctata) (d); Tillyard 1913 (b); Fraser 1960 (t); Hutchinson 1973 (d); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Metzeling & Pettigrove 1988 (d); Dobson 1990 (d); Watson et al. 1991 (t); Cameron 1992 (d); Crowther 1995 (d); Endersby 1998 (d): Hawking 1999 (c); Hawking & Theischinger 1999 (d); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

66. Diplacodes haematodes (Burmeister, 1839) Martin 1901 (as Diplax rubra); Billinghurst 1902 (as Diplax rubra) (d); Tillyard 1913 (b); Hutchinson 1973 (d); Watson 1974; Hutchinson 1975; Garrett 1984 (d); Hawking 1986 (t); Houston & Watson 1988 (t,d); Watson et al. 1991 (t); Williams et al. 1991 (d); Hawking & New 1995a (b); Hawking & New 1995b (d,b); Hawking & New 1996 (b); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

67. Diplacodes melanopsis (Martin, 1901) Martin 1901 (as Diplax melanopsis) (td); Billinghurst 1902 (as Diplax melanopsis) (d); Tillyard 1913 (b); Fraser 1960; Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Dobson 1990 (d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Crowther 1995 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a.

68. Nannophlebia risi Tillyard, 1913 Hawking 1986 (t,d); Houston & Watson 1988 (t); Bennison et al. 1989 (d); Watson et al. 1991 (t); Hawking 1994 (t,d,b); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a.

69. Nannophya australis Brauer, 1865 Martin 1901 (as Nannodythemis australis); Houston & Watson 1988 (t); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

70. Nannophya dalei (Tillyard 1908) Tillyard 1908 (as Nannodythemis dalei) (td); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Smith et al. 1977 (d); Smith et al. 1978 (d); Hawking 1986 (t,d); Houston & Watson 1988 (t); Sant & New 1988 (d); Watson et al. 1991 (t); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a.

14 Bibliography of Victorian Dragonflies

71. Orthetrum caledonicum (Brauer, 1865) Martin 1901; Billinghurst 1902 (d); Tillyard 1913 (b); Fraser 1960 (t); Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Dobson 1990 (d); Watson et al. 1991 (t); Williams et al. 1991 (d); Hawking 1994 (t,d,b); Crowther 1995 (d); Hawking & New 1995b (d,b); Hawking & New 1996 (d,b); Hawking & New 1996 (b); Endersby 1999 (d); Hawking 1999 (c); Hawking & New 1999 (d,b); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

72. Orthetrum villosovittatum (Brauer, 1868) Watson 1974; Hutchinson 1975; Hawking 1986 (t,d); Houston & Watson 1988 (t); Watson et al. 1991 (t); Crowther 1995 (d); Hawking & New 1995b (d,b); Hawking 1999 (c); Hawking & Theischinger 1999 (t); Endersby 2000a; Hawking & New 2002 (d), Endersby 2005 (d).

73. Pantala flavescens (Fabricius, 1798) Hawking & Ingram 1994 (d); Hawking 1999 (c); Hawking & Theischinger 1999 (t); van Damme & Dumont 1999 (b); Endersby 2000a.

74. Trapezostigma loewii (Kaup in Brauer, 1866) Hutchinson 1977 (d); Watson 1977; Hawking 1986 (t,d); Houston & Watson 1988 (as T. loewii tillyardi Lieftinck 1942) (t); Watson et al. 1991 (t); Hawking 1999 (c): Hawking & Theischinger 1999 (t); Endersby 2000a.

15 Bibliography of Victorian Dragonflies

4 Acknowledgements

Thanks are due to: John Hawking for access to his reprint collection, Phil Papas and Carol Harris and for access to the Arthur Rylah Library, Alena Glaister and Carol Logan for access to Monash University theses. Tim New for access to La Trobe University theses, John Dean for access to Environmental Protection Authority references. Tim Doeg and Ben Kefford for details from their reports. Phil Papas for formatting the report and Reiner Richter for supplying the photographs used on the front cover.

16 Bibliography of Victorian Dragonflies

5 References

Barmuta, L.A., 1986. Community structure of the macroinvertebrate benthos in an Australian upland stream. Ph.D. Thesis, Monash University, Melbourne. [33,34]

Bennison, G.L., Hillman, T.J. & Suter, P.J., 1989. Macroinvertebrates of the River Murray. Survey and Monitoring: 1980-1985. Water Quality Report No. 3. Murray Darling Basin Commission, Canberra. [2, 14, 16, 17, 21-24, 35, 41, 44, 45, 47-49, 54, 57, 59, 61, 68]

Billinghurst, F.L., 1902. Some notes on the Dragon-flies of the Alexandra district. Victorian Naturalist 19: 24-28. [1-4, 8, 11-14, 16-19, 21, 22, 24, 25, 30-32, 35, 37-41, 46-48, 51, 52, 54, 56, 59-61, 63, 65-67, 71]

Birkin, E., Quin, B & Jelinek, A., 1993. Hemiphlebia . Hemiphlebia mirabilis. Flora & Fauna Guarantee Action Statement No. 46. Department of Conservation & Natural Resources - Victoria. 6 pp. [1]

Blyth, J.D., Doeg, T.J. & St.Clair, R.M., 1984. Response of the macroinvertebrate fauna of the Mitta Mitta River, Victoria, to the construction and operation of Dartmouth Dam. 1: Construction and initial filling period. Occasional Papers from the Museum of Victoria 1: 83-100. [22,46]

Boulton, A.J., 1988. Composition and dynamics of macroinvertebrate communities in two intermittent streams. Ph.D. Thesis, Monash University, Melbourne. [7, 32, 61]

Boulton, A.J., 1989. Over-summering refuges of aquatic macroinvertebrates in two intermittent streams in Central Victoria. Transactions of the Royal Society of South Australia 113: 23- 34. [7, 61]

Boulton, A.J. & Lake, P.S., 1992a. The macroinvertebrate assemblages in pools and riffles in two intermittent streams (Werribee and Lerderderg Rivers, south central Victoria). Occasional Papers from the Museum of Victoria 5: 55-71. [7, 32, 61]

Boulton, A.J. & Lake, P.S., 1992b. The ecology of two intermittent streams in Victoria, Australia. II. Comparisons of faunal composition between habitats, rivers and years. Freshwater Biology 27: 99-121. [61]

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26 Bibliography of Victorian Dragonflies

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Watson, J.A.L., 1976. The synonymy of Ischnura heterosticta (Burmeister) and Ischnura torresiana Tillyard (Odonata: Coenagrionidae). Journal of the Australian Entomological Society 15: 71-78. [22]

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Watson, J.A.L., 1995. The conservation status of the enigmatic Australian Dragonfly Hemiphlebia mirabilis Selys. In Proceedings of the International Symposium on the Conservation of Dragonflies and their habitats. P.S. Corbet, S.W. Dunkle & H. Ubukata (eds), pp. viii and 16-18. Japanese Society for Preservation of Birds, Kushiro. xii + 70 pp. January 1995. [1]

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28 Bibliography of Victorian Dragonflies

Appendix

Type Localities within Victoria.

Hemiphlebia mirabilis Selys, 1869 Port Denison Qld (error) Synlestes weyersii Selys, 1869 Port Denison Qld. (error) Austrolestes leda (Selys, 1862) Melbourne Austroargiolestes icteromelas (Selys, 1862) Melbourne Griseargiolestes intermedius (Tillyard, 1913) Alexandria Vic Austrocnemis splendida (Martin, 1901) Victoria Caliagrion billinghursti (Martin, 1901) Victoria Coenagrion lyelli (Tillyard, 1913) Gisborne, Victoria Xanthagrion erythroneurum (Selys, 1876) Melbourne, Victoria Austroaeschna atrata Martin, 1909 Victoria Austroaeschna inermis Martin, 1901 Alexandra, Vic Austroaeschna multipunctata (Martin, 1901) Victoria Notoaeschna sagittata (Martin, 1901) Alexandra Vic Spinaeschna tripunctata (Martin, 1901) Alexandra Vic Telephlebia brevicauda Tillyard, 1916 Mount Macedon Vic Austrogomphus ochraceus (Selys, 1869) Melbourne Vic Apocordulia macrops Watson, 1980 Kiewa R., Killara, Vic Cordulephya pygmaea Selys, 1870 Melbourne Hemicordulia tau (Selys, 1871) Melbourne, Vic Austrothemis nigrescens (Martin, 1901) Vic & NSW Diplacodes melanopsis (Martin, 1901) Vic & NSW

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