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Habitat Characterization and Assessment of the Northern Range Limit of the Regionally Rare Crepidomanes Intricatum in Northeast North America
Habitat characterization and assessment of the northern range limit of the regionally rare Crepidomanes intricatum in Northeast North America 2019 Les Mehrhoff Botanical Research Award Final Report Submitted by Matthew Charpentier & Laura Green November 30, 2019 Laura Green gesturing towards the location of a newly documented station of Crepidomanes intricatum. Introduction: Crepidomanes intricatum (Syn. Trichomanes intricatum; weft fern), is a diminutive member of the filmy fern family (Hymenophyllaceae) known only from its gametophyte form. Weft fern is known to grow as single-cell thick filaments in dark crevices of ledges, rockhouses, talus, and glacial erratics throughout the Eastern United States from Alabama to Vermont (Natureserve Explorer). Lacking the ability to produce spores, reproduction is achieved asexually through the production of gemmae. C. intricatum’s stature and cryptic nature requires detailed observation, under 20x magnification or greater, to document the presence of rhizoids, gemmae, and gemmifer cells, which distinguish it from co-occurring algae and bryophyte protonemata. Vectors of distribution are currently unknown, though, research has indicated gemma are too large to be wind dispersed (Don Farrar, Pers. Comm.). Throughout its northern range in the eastern United States, Crepidomanes intricatum is considered rare by State Heritage programs (S1, NJ; S1, NY; S3, CT; S1, MA; S1, VT; S1/Historic NH; Natureserve Explorer) and categorized as a Division II, Regionally Rare Taxon, in the Flora Conservanda New England 2012 by Brumback and Gerke (2013), with 16 known stations at the time of publication. The rarity of C. intricatum is attributed to the small number of stations known throughout the region; New York: 4 stations , Connecticut: 3 stations , Massachusetts: 11 stations , Vermont: 4 stations (Massachusetts Natural Heritage and Endangered Species Program Data Release Agreement; New York Natural Heritage Program, Vermont Department of Fish and Wildlife, Connecticut Department of Energy and Environmental Protection, Pers. -
A Taxonomic Revision of Hymenophyllaceae
BLUMEA 51: 221–280 Published on 27 July 2006 http://dx.doi.org/10.3767/000651906X622210 A TAXONOMIC REVISION OF HYMENOPHYLLACEAE ATSUSHI EBIHARA1, 2, JEAN-YVES DUBUISSON3, KUNIO IWATSUKI4, SABINE HENNEQUIN3 & MOTOMI ITO1 SUMMARY A new classification of Hymenophyllaceae, consisting of nine genera (Hymenophyllum, Didymoglos- sum, Crepidomanes, Polyphlebium, Vandenboschia, Abrodictyum, Trichomanes, Cephalomanes and Callistopteris) is proposed. Every genus, subgenus and section chiefly corresponds to the mono- phyletic group elucidated in molecular phylogenetic analyses based on chloroplast sequences. Brief descriptions and keys to the higher taxa are given, and their representative members are enumerated, including some new combinations. Key words: filmy ferns, Hymenophyllaceae, Hymenophyllum, Trichomanes. INTRODUCTION The Hymenophyllaceae, or ‘filmy ferns’, is the largest basal family of leptosporangiate ferns and comprises around 600 species (Iwatsuki, 1990). Members are easily distin- guished by their usually single-cell-thick laminae, and the monophyly of the family has not been questioned. The intrafamilial classification of the family, on the other hand, is highly controversial – several fundamentally different classifications are used by indi- vidual researchers and/or areas. Traditionally, only two genera – Hymenophyllum with bivalved involucres and Trichomanes with tubular involucres – have been recognized in this family. This scheme was expanded by Morton (1968) who hierarchically placed many subgenera, sections and subsections under -
A Review of the Fern Genus Polystichum (Pteropsida: Dryopteridaceae) in Madagascar and the Mascarene Region
A review of the fern genus Polystichum (Pteropsida: Dryopteridaceae) in Madagascar and the Mascarene region Jacobus P. ROUX National Botanical Institute, Compton Herbarium, Private Bag X7, Claremont 7735, South Africa. [email protected] ABSTRACT KEY WORDS The fern genus Polystichum Roth (Dryopteridaceae) in Madagascar and the Polystichum, Mascarene region is reviewed. Eight species are recorded, six being endemic Dryopteridaceae, Madagascar, to the region. Of these four are endemic to Madagascar. The Madagascan Mascarene region. species remain poorly known. RÉSUMÉ Révision du genre Polystichum (Pteropsida: Dryopteridaceae) de Madagascar et des Mascareignes. MOTS CLÉS Révision des fougères de Madagascar et des Mascareignes appartenant au Polystichum, genre Polystichum Roth (Dryopteridaceae). Huit espèces sont retenues, dont Dryopteridaceae, Madagascar, six endémiques de Madagascar. Les espèces malgaches demeurent peu Mascareignes. connues. INTRODUCTION Madagascar has been separated from Africa for at least 100 million years and is well known for its A review of the fern genus Polystichum in high level of endemism. RAVEN & AXELROD Madagascar and the Mascarene region (the (1974) ascribed the rich Madagascan flora to a now Madagascan and Comore archipelagos) is provid- submerged Madagascan plateau that connected it ed. Polystichum is a genus of between 160 (TRYON with India and Antarctica, thus allowing for migra- & TRYON 1982) and 200 species (DAIGOBO tion between these now distant continents until 1972), occurring throughout the temperate parts the late Cretaceous. LEROY (1978), on the other of the world as well as the montane tropics, but is hand, considered the rich Madagascan flora as an mostly absent from the lowland tropics. autochthonous flora that has differentiated princi- ADANSONIA, sér. -
Coordinadores) CONSEJO EDITORIAL INTERNACIONAL
Luis Fernando Plenge Tellechea Jorge Alberto Pérez León (Coordinadores) CONSEJO EDITORIAL INTERNACIONAL Álvaro Álvarez Parrilla Gaspar Ros Berruezo Fac. Ciencias, Matemáticas, UABC, Depto. de Bromatología e Inspección Ensenada, B. C. de Alimentos, Universidad de Murcia, Francisco Fernández Belda Murcia, España. Depto. de Bioquímica y Rocío Salceda Sacanelles Biología Molecular (A), Universidad Instituto de Fisiología Celular, Depto. Ciencia en la frontera: revista de ciencia y tecnología de Murcia, Murcia, España. Neuro ciencias, UNAM, México, D. F. de la Universidad Autónoma Alex Fragoso Sierra Fernando Soler de Ciudad Juárez Fac. de Química. Universidad Depto. de Bioquímica y Biología de La Habana, Cuba. Molecular (A), Universidad de DIRECTORIO Jorge Gardea Torresdey Murcia, Murcia, España. Javier Sánchez Carlos Chemistry, UTEP, El Paso, Texas. Marieta Tuena de Gómez Puyou Rector Armando Gómez Puyou Investigadora Emérita. Instituto de Fisiología David Ramírez Perea Investigador Emérito. Instituto de Ce lular, Depto. Bioquímica, UNAM. Secretario General Fisiología Celular, Depto. Bioquímica, México, D. F. UNAM. México, D. F. José Vázquez Tato Martha P. Barraza de Anda Coordinadora General de Gustavo González Fac. de Ciencias, Depto. de Investigación y Posgrado Tecnología de Alimentos de Química Física. Universidad de Origen Vegetal, CIAD Santiago de Compostela, Hugo Staines Orozco Hermosillo, Sonora, México. España. Director del ICB Louis Irwin Ricardo Tapia Ibargüengoytia Servando Pineda Jaimes Biological Science, UTEP, El Paso, Texas. Neurociencias Dirección General de Difusión José Luis Ochoa IFC-UNAM Cultural y Divulgación Científica CIBNOR, La Paz, B.C.S. Herminia Pasantes Esther Orozco Neurociencias CONSEJO EDITORIAL Emilio Álvarez Parrilla CINVESTAV, México, D. F. IFC-UNAM Leonel Barraza Pacheco Biomedicina Molecular. Thomas Kretzschmar Steinle Alejandro Donohue Cornejo María Jesús Periago Área de Geofísica Esaúl Jaramillo Depto. -
Fern Gazette
THE FERN GAZETTE Edited by BoAoThomas lAoCrabbe & Mo6ibby THE BRITISH PTERIDOLOGICAL SOCIETY Volume 14 Part 3 1992 The British Pteridological Society THE FERN GAZETTE VOLUME 14 PART 3 1992 CONTENTS Page MAIN ARTICLES A Revised List of The Pteridophytes of Nevis - B.M. Graham, M.H. Rickard 85 Chloroplast DNA and Morphological Variation in the Fern Genus Platycerium(Polypodiaceae: Pteridophyta) - Johannes M. Sandbrink, Roe/and C.H.J. Van Ham, Jan Van Brederode 97 Pteridophytes of the State of Veracruz, Medico: New Records - M6nica Pa/acios-Rios 119 SHORT NOTES Chromosome Counts for Two Species of Gleichenia subgenus Mertensiafrom Ecuador - Trevor G. Walker 123 REVIEWS Spores of The Pteridophyta - A. C. Jermy 96 Flora Malesiana - A. C. Jermy 123 The pteridophytes of France and their affinities: systematics. chorology, biology, ecology. - B. A. Thoinas 124 THE FERN GAZ ETTE Volume 14 Pa rt 2 wa s publis hed on lO Octobe r 1991 Published by THE BRITISH PTERIDOLOGICAL SOCIETY, c/o Department of Botany, The Natural History Museum, London SW7 580 ISSN 0308-0838 Metloc Printers Ltd .. Caxton House, Old Station Road, Loughton, Essex, IG10 4PE ---------------------- FERN GAZ. 14(3) 1992 85 A REVISED LIST OF THE PTERIDOPHYTES OF NEVIS BMGRAHAM Polpey, Par, Cornwall PL24 2T W MHRICKARD The Old Rectory, Leinthall Starkes, Ludlow, Shropshire SY8 2HP ABSTRACT A revised list of the pteridophytes of Nevis in the Lesser Antilles is given. This includes 14 species not previously recorded for the island. INTRODUCTION Nevis is a small volcanic island in the West Indian Leeward Islands. No specific li st of the ferns has ev er been pu blished, although Proctor (1977) does record each of the species known to occur on the island. -
Updated 2017 Edition of Our RHS Science Strategy
RHS Science Strategy Update 2017 The RHS Vision To enrich everyone’s life through plants, and make the UK a greener and more beautiful place. Our four key themes A global knowledge bank for 1 gardening and garden plants Plant health 2 in gardens Gardening in a 3 changing world Plant science for all: 4 people, plants, planet 2 Welcome 2017 has been a year to remember in RHS Science & Collections, and I’m delighted to bring you this update full of exciting developments. The team has delivered against so many of our key objectives that it would be impossible to present everything that has been done, so please enjoy these highlights and keep an eye on “In particular, our rhs.org.uk for more science news and information. work on climate We need to raise £13m from donors and RHS members change has to help build the new Centre for Horticultural Science and Learning and refurbish the historic Laboratory Professor Alistair demonstrated the Griffiths building at RHS Garden Wisley. We have received initial importance of RHS Director of support from the Heritage Lottery Fund (HLF), with the Science & Collections understanding the award of a first-round pass for a grant of £4.8m, and role that plants, are extremely grateful for the many further generous donations and pledges. However, we still need help to gardens and reach our target in order to inspire future generations gardening play.” of horticultural scientists and gardeners. If you would like to offer support, please see rhs.org.uk/donate. The plants we choose to grow in our gardens can have a positive impact on our health and environment, from providing food for pollinators to helping us manage stress. -
Crepidomanes Shenzhenense (Subg
Phytotaxa 440 (2): 101–107 ISSN 1179-3155 (print edition) https://www.mapress.com/j/pt/ PHYTOTAXA Copyright © 2020 Magnolia Press Article ISSN 1179-3163 (online edition) https://doi.org/10.11646/phytotaxa.440.2.2 Crepidomanes shenzhenense (subg. Crepidomanes; Hymenophyllaceae), a new filmy fern species from Guangdong, southern China HUI WANG 1,2,4, GUO-HUA ZHAO 2,5, BASKARAN XAVIER RAVI2,3,6 & SHU-GANG LU 1,7* 1 Institute of Ecology and Geobotany, Yunnan University, Kunming 650091, Yunnan, China 2 Shenzhen Key Laboratory of Southern Subtropical Plant Diversity, Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Sciences, Shenzhen 518004, Guangdong, China 3 School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, Guangdong, China 4 �[email protected]; https://orcid.org/0000-0002-1102-4644 5 �[email protected]; https://orcid.org/0000-0003-2703-3139 6 �[email protected]; https://orcid.org/0000-0003-4748-4818 7 �[email protected]; https://orcid.org/0000-0001-7220-115X *Corresponding author Abstract A new filmy fern, Crepidomanes shenzhenense (Hymenophyllaceae) is described from Shenzhen, Guangdong, southern China. Crepidomanes shenzhenense is a tiny fern characterized by having a broadly obovate lamina only with internal false veinlets and crenate to dissected margins of involucre lips. The new species is presently known from two localities in Shenzhen. The distinctiveness between the new species and three relevatives, C. parvifolium, C. megistostomum and C. latealatum complex, are presented in detail. Keywords: Endangered species, filmy ferns, southern China Introduction Filmy fern family, Hymenophyllaceae, is the most diverse basal lineage of polypodiidae ferns which comprises ca. -
Molecular Systematics of the Filmy Ferns (Hymenophyllaceae) of South India P
Indian Fern J. 31 : 65-83 (2014) ISSN 0970-2741 MOLECULAR SYSTEMATICS OF THE FILMY FERNS (HYMENOPHYLLACEAE) OF SOUTH INDIA P. K. VIDYA VARMA1 AND P. V. MADUSOODANAN2* 1Govt. College, Madappally, Vatakara-673102, Kerala 2Malabar Botanical Garden, Calicut-673014, Kerala (Received April 22, 2014; Revised Accepted May 16, 2014) ABSTRACT Molecular systematics of the filmy ferns (Hymenophyllaceae) of South India is done based on rbcL nucleotide sequences. Live materials were collected from Western Ghats of South India. Fresh and silica dried fronds were used for the extraction of DNA. The rbcL gene was amplified and sequenced. For phylogenetic analysis 19 sequences (eight newly sequenced and 11 sequences from GenBank) were considered. Multiple sequence alignment was done with ClustalW2 software. The phylogenetic analysis was done with Neighbor Joining method, Maximum Likelihood method and Bayesian inference. As a result a conspicuous clustering pattern was obtained showing the Trichomanoid lineage of South India comprising of three strongly supported clades which correspond to the three genera recognized by Ebihara et al. (2006) viz., Abrodictyum, Crepidomanes and Didymoglossum. Key Words : Molecular systematics, Trichomanoid lineage, South India, rbcL gene. I NTRODUCTION The filmy ferns are unique group of leptosporangiate monilophytes belonging to the family of Hymenophyllaceae Link comprising around 600 species (Iwatsuki, 1990). They grow in the interior of dense humid forests either as epipetric forms or as epiphytes (except certain terrestrial species such as Abrodictyum obscurum) and are easily mistaken for thalloid bryophytes. The molecular phylogenetic study of the family done by Pryer et al. (2001) showed that there exist two lineages in the family viz., Trichomanoid lineage and Hymenophylloid lineage. -
PTERIDOLOGIST 2012 Contents: Volume 5 Part 5, 2012 Scale Insect Pests of Ornamental Ferns Grown Indoors in Britain
PTERIDOLOGIST 2012 Contents: Volume 5 Part 5, 2012 Scale insect pests of ornamental ferns grown indoors in Britain. Dr. Chris Malumphy 306 Familiar Ferns in a Far Flung Paradise. Georgina A.Snelling 313 Book Review: A Field Guide to the Flora of South Georgia. Graham Ackers 318 Survivors. Neill Timm 320 The Dead of Winter? Keeping Tree Ferns Alive in the U.K. Mike Fletcher 322 Samuel Salt. Snapshots of a Victorian Fern Enthusiast. Nigel Gilligan 327 New faces at the Spore Exchange. Brian and Sue Dockerill 331 Footnote: Musotima nitidalis - a fern-feeding moth new to Britain. Chris Malumphy 331 Leaf-mining moths in Britain. Roger Golding 332 Book Review: Ferns of Southern Africa. A Comprehensive Guide. Tim Pyner 335 Stem dichotomy in Cyathea australis. Peter Bostock and Laurence Knight 336 Mrs Puffer’s Marsh Fern. Graham Ackers 340 Young Ponga Frond. Guenther K. Machol 343 Polypodium Species and Hybrids in the Yorkshire Dales. Ken Trewren 344 A Challenge to all Fern Lovers! Jennifer M. Ide 348 Lycopodiums: Trials in Pot Cultivation. Jerry Copeland 349 Book Review: Fern Fever. Alec Greening 359 Fern hunting in China, 2010. Yvonne Golding 360 Stamp collecting. Martin Rickard 365 Dreaming of Ferns. Tim Penrose 366 Variation in Asplenium scolopendrium. John Fielding 368 The Case for Filmy Ferns. Kylie Stocks 370 Polystichum setiferum ‘Cristato-gracile’. Julian Reed 372 Why is Chris Page’s “Ferns” So Expensive? Graham Ackers 374 A Magificent Housefern - Goniophlebium Subauriculatum. Bryan Smith 377 A Bolton Collection. Jack Bouckley 378 360 Snails, Slugs, Grasshoppers and Caterpillars. Steve Lamont 379 Sphenomeris chinensis. -
Scientific Name Common Name Taxon Group Aceria Pseudoplatani
Scientific Name Common Name Taxon Group Aceria pseudoplatani Aceria pseudoplatani acarine (Acari) Alabidocarpus acarine (Acari) Phytoptus avellanae Phytoptus avellanae acarine (Acari) Tetranychidae Tetranychidae acarine (Acari) Tetranychus urticae Tetranychus urticae acarine (Acari) Trombidiidae Trombidiidae acarine (Acari) Bufo bufo Common Toad amphibian Lissotriton helveticus Palmate Newt amphibian Lissotriton vulgaris Smooth Newt amphibian Rana Frog amphibian Rana temporaria Common Frog amphibian Triturus Newt amphibian Triturus cristatus Great Crested Newt amphibian Triturus helveticus Palmate Newt amphibian Annelida Worms annelid Eisenia fetida Manure Worm annelid Erpobdella testacea Erpobdella testacea annelid Lumbricus terrestris Common Earthworm annelid Acanthis flammea/cabaret agg. Redpoll (Common\Lesser) agg. bird Accipiter nisus Sparrowhawk bird Acrocephalus schoenobaenus Sedge Warbler bird Aegithalos caudatus Long-tailed Tit bird Alauda arvensis Skylark bird Anas platyrhynchos Mallard bird Anthus pratensis Meadow Pipit bird Apodidae Swifts bird Apus apus Swift bird Scientific Name Common Name Taxon Group Ardea cinerea Grey Heron bird Aythya fuligula Tufted Duck bird Branta canadensis Canada Goose bird Buteo buteo Buzzard bird Caprimulgus europaeus Nightjar bird Carduelis carduelis Goldfinch bird Certhia familiaris Treecreeper bird Chloris chloris Greenfinch bird Chroicocephalus ridibundus Black-headed Gull bird Cinclus cinclus Dipper bird Coloeus monedula Jackdaw bird Columba Pigeon bird Columba livia Feral Pigeon/Rock Dove -
The Potential Ecological Impact of Ash Dieback in the UK
JNCC Report No. 483 The potential ecological impact of ash dieback in the UK Mitchell, R.J., Bailey, S., Beaton, J.K., Bellamy, P.E., Brooker, R.W., Broome, A., Chetcuti, J., Eaton, S., Ellis, C.J., Farren, J., Gimona, A., Goldberg, E., Hall, J., Harmer, R., Hester, A.J., Hewison, R.L., Hodgetts, N.G., Hooper, R.J., Howe, L., Iason, G.R., Kerr, G., Littlewood, N.A., Morgan, V., Newey, S., Potts, J.M., Pozsgai, G., Ray, D., Sim, D.A., Stockan, J.A., Taylor, A.F.S. & Woodward, S. January 2014 © JNCC, Peterborough 2014 ISSN 0963 8091 For further information please contact: Joint Nature Conservation Committee Monkstone House City Road Peterborough PE1 1JY www.jncc.defra.gov.uk This report should be cited as: Mitchell, R.J., Bailey, S., Beaton, J.K., Bellamy, P.E., Brooker, R.W., Broome, A., Chetcuti, J., Eaton, S., Ellis, C.J., Farren, J., Gimona, A., Goldberg, E., Hall, J., Harmer, R., Hester, A.J., Hewison, R.L., Hodgetts, N.G., Hooper, R.J., Howe, L., Iason, G.R., Kerr, G., Littlewood, N.A., Morgan, V., Newey, S., Potts, J.M., Pozsgai, G., Ray, D., Sim, D.A., Stockan, J.A., Taylor, A.F.S. & Woodward, S. 2014. The potential ecological impact of ash dieback in the UK. JNCC Report No. 483 Acknowledgements: We thank Keith Kirby for his valuable comments on vegetation change associated with ash dieback. For assistance, advice and comments on the invertebrate species involved in this review we would like to thank Richard Askew, John Badmin, Tristan Bantock, Joseph Botting, Sally Lucker, Chris Malumphy, Bernard Nau, Colin Plant, Mark Shaw, Alan Stewart and Alan Stubbs. -
Cheilanthes (Cheilanthoideae, Pteridaceae), with Emphasis on South American Species
Organisms Diversity & Evolution (2018) 18:175–186 https://doi.org/10.1007/s13127-018-0366-6 ORIGINAL ARTICLE Further progress towards the delimitation of Cheilanthes (Cheilanthoideae, Pteridaceae), with emphasis on South American species M. Mónica Ponce1 & M. Amalia Scataglini1 Received: 20 July 2017 /Accepted: 22 April 2018 /Published online: 5 May 2018 # Gesellschaft für Biologische Systematik 2018 Abstract Cheilanthoid ferns (Cheilanthoideae sensu PPG 1 2016) constitute an important group within the Pteridaceae and are cosmopolitan in distribution. In South America, there are 155 species distributed in 13 genera, among which the largest are Adiantopsis (35), Cheilanthes (27), and Doryopteris (22). Most of the cheilanthoid species are morphologically adapted to grow in arid to semi-arid conditions and show convergent evolution, which has implied difficulties in defining the genera throughout their taxonomic history (Copeland 1947,Tryon&Tryon1973,Gastony&Rollo 1995, 1998,KirkpatrickSystematic Botany, 32:504–518, 2007, Rothfels et al. Taxon, 57: 712–724, 2008). Here, we sequenced two plastid markers (rbcL + trnL-F) of 33 South American cheilanthoid species, most of which have not been included in phylogenetic analyses previously. The South American species were analyzed together with South African and Australasian Cheilanthes and representatives of related cheilanthoid genera. The phylogenetic analysis showed that most Cheilanthes species are related to the genus Hemionitis, constituting different groups according to their distribu- tion; moreover, three species—C. hassleri, C. pantanalensis,andC. obducta—appear as the sister clade of Hemionitis. Cheilanthes micropteris, the type species, is strongly supported in a clade with Australasian Cheilanthes plus five South American Cheilanthes species, all of which show a reduction in the number of spores per sporangium; this feature would be a synapomorphy for core Cheilanthes s.s.