The Cochylid Fauna of the Southern Ural Mountains, with Description of Cochylimorpha Ignicolorana Junnilainen & K
Total Page：16
File Type：pdf, Size：1020Kb
94© Entomologica Fennica. 22 October 2001 Nupponen et al. • ENTOMOL. FENNICA Vol. 12 The cochylid fauna of the Southern Ural Mountains, with description of Cochylimorpha ignicolorana Junnilainen & K. Nupponen sp. n. (Lepidoptera: Tortricidae: Cochylini) Kari Nupponen, Jari Junnilainen, Timo Nupponen & Vladimir Olschwang Nupponen, K., Junnilainen, J., Nupponen, T. & Olschwang, V. 2001: The cochylid fauna of the Southern Ural Mountains, with description of Cochylimorpha ignicolorana Junnilainen & K. Nupponen sp. n. (Lepidoptera: Tortricidae: Cochylini). — Entomol. Fennica 12: 94–107. A list of 78 species of the tortricoid tribe Cochylini from the southern Ural Mountains is presented. The material was collected during 1996–2000 on nine different Finnish-Russian expeditions. Cochylimorpha ignicolorana Junnilainen & K. Nupponen sp. n. is described. The new taxon occurs on dry steppe slopes in the headland region of the southern Urals, and it is rather easy to separate from closely related taxa both externally and by the male genitalia. In addition, 7 species are reported as new for Europe and 4 species as new for Russia. The known distribution range of each species is given as well as further notes on some poorly known taxa. Kari Nupponen, Miniatontie 1 B 9, FIN-02360 Espoo, Finland Jari Junnilainen, Mahlapolku 3, FIN-01730 Vantaa, Finland Timo Nupponen, Riilahdentie 5 D 15, FIN-02360 Espoo, Finland Vladimir Olschwang, Nagornaja Street 11-32, RUS-620028 Ekaterinburg, Russia Received 2 January 2001, accepted 3 March 2001 1. Introduction has not been any serious collecting activities in the region for almost one hundred years and many There is a long tradition of lepidopterological stud- of the previously discovered species have been ies in the southern Ural region, the southeastern- considered great rarities to date. most corner of Europe. Prof. Eduard Eversmann The recently changed political situation in Rus- (1844) made thorough faunistic investigations in sia has made visits to the southern Urals possible the area in the 19th century and described numer- again. The present article is based on our own stud- ous new species, among them several cochylids. ies of the cochylid fauna in that area. The cochylid fauna is very rich in the headlands of the southern Ural Mountains and the adjacent lowland steppes, and many further cochylids were 2. The investigated area, material and described from there and adjacent regions at the methods end of the 19th century by several authors (Christoph, Kennel, Möschler, Staudinger). How- The investigated area is situated in Cheliabinsk and ever, since the beginning of the 20th century there Orenburg oblasts and Bashkiria in the southern Ural Moun- ENTOMOL. FENNICA Vol. 12 • The cochylid fauna of the Southern Ural Mountains 95 Fig. 1. Map of southern Ural region with collecting localities. — 1: Ajat river. — 2: Arkaim. — 3: Bajmak. — 4: Berlin. — 5: Chalk Hills. — 6: Iremel. — 7: Kidriasovo. — 8: Kizilskoye. — 9: Kuvandyk. — 10: Kuvandyk 2. — 11: Miass. — 12: Moskovo. — 13: Burannoe, Novoiletzk. — 14: Sakmara river. — 15: Verbljushka. — 16: Zirgan. — 17: Shkunovka. — 18: Bishtiryak. — 19: Sanarskii Bor. — 20: Uchaly. — 21: Ekaterinburg biol.st. — 22: Kosmokovo. — 23: Tavatui. tains, between 50°40´N–56°30´N and 54°26´E–62°06´E. — 7: 19.VI.2000; Sverdlovsk oblast (near Ekaterinburg); The majority of collecting places was located on the east- K. Nupponen, T. Nupponen. ern-southern foothill region and at low altitude. The habi- — 8: 25.VII.–04.VIII.2000; Orenburg oblast, Cheliabinsk tats were mainly different kinds of steppes, but also taiga oblast, Bashkiria; T. Nupponen. forests, alpine meadows and mountain tundra. The lowest — 9: 26.VIII.–06.IX.2000; Orenburg oblast, Cheliabinsk locality was in the valley of the river Ilek, Novoiletzk oblast; K. Nupponen. (100 m a.s.l.) and the highest one the Iremel Mountain (1580 The material was collected both by artificial light at night m a.s.l.). Most localities were lying at an elevation of 200– and by sweeping and netting during daytime. Altogether 450 m. over 2000 specimens of cochylids were collected, and a The present article is based on the material collected large amount of additional specimens were observed and during 1996–2000 on 9 different expeditions. The dates, determined in the field. The majority of the material was areas visited and collectors on each of the trips are as fol- examined by Kari Nupponen. The following colleagues also lows: determined parts of the material and/or confirmed the de- — 1: 13.–29.VI.1996; Cheliabinsk oblast, Bashkiria; K. termination of some critical taxa: J. Itämies, J. Junnilainen, Nupponen, J.-P. Kaitila, J. Junnilainen, M. Ahola. J.-P. Kaitila, M. Mutanen and T. Nupponen. The collected — 2: 26.VI.–16.VII.1997; Cheliabinsk oblast; K. Nup- material is mostly deposited in the private collections of the observers. ponen, J.-P. Kaitila, J. Junnilainen, M. Ahola. The collecting localities are mentioned below. Brief — 3: 25.V.–22.VI.1998; Orenburg oblast, Cheliabinsk variants of locality names are given in uppercase letters oblast, Bashkiria; K. Nupponen, T. Nupponen, J. before each locality and used later in the species list. The Junnilainen. italicized dates indicate daytime collecting only in the lo- — 4: 11.–31.VII.1998; Orenburg oblast, Cheliabinsk oblast, cality. The number given to each of the localities is con- Bashkiria; K. Nupponen. nected with that on the map (Fig. 1). — 5: 11.–20.V.1999; Orenburg oblast, Cheliabinsk oblast; K. Nupponen. — 1: AJAT RIVER: Cheliabinsk oblast, 53°02´N 62°06´E, — 6: 13.–30.VI.1999; Orenburg oblast, Cheliabinsk oblast, 200 m, Ajat river near Nikolaevka village. A rocky hill Bashkiria; K. Nupponen, T. Nupponen. in a riverbank, surrounded by a moist place on a riverside 96 Nupponen et al. • ENTOMOL. FENNICA Vol. 12 and a large Artemisia steppe. 03.–05.VII.1997, 19.VI.2000. 24.25.–VII.1998, 04.05.IX.2000. — 9: KUVANDYK: Orenburg oblast, 51°26´N 57°26´E, — 2: ARKAIM: Cheliabinsk oblast, 52°39´N 59°34´E, 350 250 m, Kuvandyk 12 km SE. A foothill region with m, Arkaim reserve near Amurskii village. A large reserve different kinds of steppes and old Quercus – Populus – with different kinds of steppe habitats. 14.–19.VI.1996, Betula forests on the top of the hills.13.–16.VI.1998, 06.–10.VII.1997, 22.–23.VII.1998, 17.V.1999, 15.– 19.–21.VII.1998, 15.V.1998, 02.VIII.2000, 16.VI.1999. 02.IX.2000. — 3: BAJMAK: Bashkiria, 52°40´N 58°34´E, 450 m, — 10: KUVANDYK 2: Orenburg oblast, 51°37´N 57°34´E, Bajmak 15 km E. Open foothill steppe locality. 17.– 300 m, Kuvandyk 30 km NE. Rocky hills and meadows, 18.VI.1998. at the slopes some blackish coloured, hot, gravelly spots — 4: BERLIN: Cheliabinsk oblast, 53°59´N 61°12´E, with sparse vegetation. 16.–17.VI.1998, 03.VIII.2000. 250 m, Troizkii reserve near Berlin village. A small, — 11: MIASS: Cheliabinsk oblast, 55°01´N 60°06´E, 350 mainly grassland steppe surrounded by a bog and young m, Miass, Ilmen State reserve. Forest steppes and old forest. 30.VI.–02.VII.1997. conifer forests. The records from Miasovo lake be- —18: BISHTIRYAK: Bashkiria, 51°48´N 57°05´E, 500 longing to the same reserve (appr. 10 km NE) are m, Kasmarka river near Bishtiryak village. Steep rocky included in the list of Miass. 13.VI.1996, 28.–29.VI.1996, slopes, dry meadows and deciduous forests. 13.– 26.–29.VI.1997, 15.–16.VII.1997, 25.–26.V.1998, 14.VII.1998. 19.–20.VI.1998, 11.V.1999, 18.–20.V.1999, 29.– — 13: BURANNOE: Orenburg oblast, 50°58´N 54°25´E, 30.VI.1999, 26.VIII.2000, 06.IX.2000; In addition, light 100 m, near Burannoe village, Ilek river valley. Lowland trap collecting during 15.VI.–24.VIII.1999 and 25.VII.– Artemisia steppes, wet meadows and wetlands. 20.– 05.IX.2000. 21.VI.1999, 30.VII.2000, 29.VIII.2000. — 12: MOSKOVO: Cheliabinsk oblast, 53°57´N 59°03´E, — 5: CHALK HILLS: Orenburg oblast, 50°40–45´N 650 m, near Moskovo village. Open, rocky foothill 54°26–28´E, 170–230 m, Pokrovka village 20 km S, region with different kinds of steppes and wet meadows Schibendy valley. A dry, open, lowland Artemisia along the riverside. 22.–23.VI.1996, 10.–11.VII.1997, steppe with wet meadows along the small riverside. 26.V.1998, 18.VI.1998, 11.–13.VII.1998, 04.VIII.2000, Whitish limestone rocks surround the flat valley, the 26.VIII.2000. vegetation being luxurious in northern slopes and very — 13: NOVOILETZK: Orenburg oblast, 50°59´N 54°17– sparse in southern slopes. 03.–07.VI.1998, 17.– 22´E, 100 m, Novoiletzk 8 km E, Ilek river valley. Sand 18.VII.1998, 21.–24.VI.1999, 31.VII.–01.VIII.2000, dune region with few Artemisia steppe spots, wet 30.–31.VIII.2000. meadows and wetlands. 08.–09.VI.1998. — EKATERINBURG: Sverdlovsk oblast, Ekaterinburg — 14: SAKMARA RIVER: Bashkiria, 51°54´N 57°43´E, city. 27.VII.1998. 450 m, Sakmara river near Jantyshevo village. Forest — 21: EKATERINBURG BIOL.ST.: Sverdlovsk oblast, steppes, meadows and mixed forests. 20.–21.VI.1996. Ekaterinburg 50 km S, near Dvurechensk village, bio- — 19: SANARSKII BOR: Cheliabinsk oblast, 54°06´N logical station of Ural’s university. Mixed forests, bogs 60°30´E, 400 m, Sanarskii bor near Sanarka village. and meadows. 20.–22.VI.1998. Old conifer forest. 26.–27.VII.1998. — 6: IREMEL: Cheliabinsk oblast, 54°31–35´N 58°49– — 17: SHKUNOVKA: Orenburg oblast, 50°48´N 55°18´E, 54´E, 900–1580 m, Iremel Mountain reserve. Taiga 200 m, Malaja Hobda river near Shkunovka village.