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Bryozoan Studies 2019
BRYOZOAN STUDIES 2019 Edited by Patrick Wyse Jackson & Kamil Zágoršek Czech Geological Survey 1 BRYOZOAN STUDIES 2019 2 Dedication This volume is dedicated with deep gratitude to Paul Taylor. Throughout his career Paul has worked at the Natural History Museum, London which he joined soon after completing post-doctoral studies in Swansea which in turn followed his completion of a PhD in Durham. Paul’s research interests are polymatic within the sphere of bryozoology – he has studied fossil bryozoans from all of the geological periods, and modern bryozoans from all oceanic basins. His interests include taxonomy, biodiversity, skeletal structure, ecology, evolution, history to name a few subject areas; in fact there are probably none in bryozoology that have not been the subject of his many publications. His office in the Natural History Museum quickly became a magnet for visiting bryozoological colleagues whom he always welcomed: he has always been highly encouraging of the research efforts of others, quick to collaborate, and generous with advice and information. A long-standing member of the International Bryozoology Association, Paul presided over the conference held in Boone in 2007. 3 BRYOZOAN STUDIES 2019 Contents Kamil Zágoršek and Patrick N. Wyse Jackson Foreword ...................................................................................................................................................... 6 Caroline J. Buttler and Paul D. Taylor Review of symbioses between bryozoans and primary and secondary occupants of gastropod -
Devonian Bryozoans of the Holy Cross Mountains, Poland
ACT A PAL A EON T 0 LOG ICA POLONICA Vol. XVIII 1973 No.4 MARIA KIEPURA DEVONIAN BRYOZOANS OF THE HOLY CROSS MOUNTAINS, POLAND. PART II. CYCLOSTOMATA AND CYSTOPORATA Abstract. - Descriptions are given of 33, species of Middle Devonian Bryozoa from the Holy Cross Mts., belonging to the orders Cyclostomata and Cystoporata. Cyclo stomata are represented by 26 species (4 new: Corynotrypa (Corynotrypa) skalensis, C. (C.) basiplata, Stomatopora varigemmata and Diversipora bitubulata n. gen.). Cystoporata are represented by 7 new species: Ceramoporella orbiculata, C. grandi cystica, Favositella integrimuralis, Fistulipora boardmani, F. emphantica, Cyclotrypa nekhoroshevi and Fistuliramus astrovae. The systematic assignment of the genus Hederella Hall, 1881 has been discussed and the "tabulate-like" microstructure of the zooecial walls of this genus examined. In addition, the epifauna and associated assemblages have been characterized as well as the geological and palaeoecological conditions in which the described Bryozoa occurred. INTRODUCTION Among the Middle Devonian Bryozoa of the Holy Cross Mts., the present author has, up to now, described Ctenostomata (Kiepura, 1965). The present paper is a continuation of her investigations on the Middle Devonian Bryozoa of this region and contains descriptions of 33 species belonging to 9 genera of the orders Cyclostomata and Cystoporata occur ring in the Grzegorzowice - Skaly profile. The bryozoan collection of the Palaeozoological Institute of the Polish Academy of Sciences also contains a rich material representing the orders Trepostomata and Cryptostomata from the Middle Devonian of the Holy Cross Mts., which will be described at a future date. The material described in the present paper was collected by the author during several years of fieldwork (1950-1954). -
Reduced El Niño–Southern Oscillation During the Last Glacial
RESEARCH | REPORTS PALEOCEANOGRAPHY vergent results and our newly generated data by considering geographic location, choice of fora- minifera species, and changes in thermocline – depth (see supplementary materials). Reduced El Niño Southern Oscillation ENSO variability is asymmetric (the El Niño warm phase is more extreme than the La Niña during the Last Glacial Maximum cold phase) (14), so temperature variations in the equatorial Pacific are not normally distrib- Heather L. Ford,1,2* A. Christina Ravelo,1 Pratigya J. Polissar2 uted (7, 15), and statistical tests that assume normality (e.g., standard deviation) can lead to El Niño–Southern Oscillation (ENSO) is a major source of global interannual variability, but erroneous conclusions with respect to changes its response to climate change is uncertain. Paleoclimate records from the Last Glacial in variance. Therefore, we use quantile-quantile Maximum (LGM) provide insight into ENSO behavior when global boundary conditions (Q-Q) plots—a simple, yet powerful way to vi- — (ice sheet extent, atmospheric partial pressure of CO2) were different from those today. sualize distribution data to compare the tem- In this work, we reconstruct LGM temperature variability at equatorial Pacific sites perature range and distribution recorded by two using measurements of individual planktonic foraminifera shells. A deep equatorial populations of individual foraminifera shells to thermocline altered the dynamics in the eastern equatorial cold tongue, resulting in interpret possible climate forcing mechanisms. reduced ENSO variability during the LGM compared to the Late Holocene. These results Sensitivity studies using modern hydrographic suggest that ENSO was not tied directly to the east-west temperature gradient, as data show how changes in ENSO and seasonality previously suggested. -
Exhibit Specimen List FLORIDA SUBMERGED the Cretaceous, Paleocene, and Eocene (145 to 34 Million Years Ago) PARADISE ISLAND
Exhibit Specimen List FLORIDA SUBMERGED The Cretaceous, Paleocene, and Eocene (145 to 34 million years ago) FLORIDA FORMATIONS Avon Park Formation, Dolostone from Eocene time; Citrus County, Florida; with echinoid sand dollar fossil (Periarchus lyelli); specimen from Florida Geological Survey Avon Park Formation, Limestone from Eocene time; Citrus County, Florida; with organic layers containing seagrass remains from formation in shallow marine environment; specimen from Florida Geological Survey Ocala Limestone (Upper), Limestone from Eocene time; Jackson County, Florida; with foraminifera; specimen from Florida Geological Survey Ocala Limestone (Lower), Limestone from Eocene time; Citrus County, Florida; specimens from Tanner Collection OTHER Anhydrite, Evaporite from early Cenozoic time; Unknown location, Florida; from subsurface core, showing evaporite sequence, older than Avon Park Formation; specimen from Florida Geological Survey FOSSILS Tethyan Gastropod Fossil, (Velates floridanus); In Ocala Limestone from Eocene time; Barge Canal spoil island, Levy County, Florida; specimen from Tanner Collection Echinoid Sea Biscuit Fossils, (Eupatagus antillarum); In Ocala Limestone from Eocene time; Barge Canal spoil island, Levy County, Florida; specimens from Tanner Collection Echinoid Sea Biscuit Fossils, (Eupatagus antillarum); In Ocala Limestone from Eocene time; Mouth of Withlacoochee River, Levy County, Florida; specimens from John Sacha Collection PARADISE ISLAND The Oligocene (34 to 23 million years ago) FLORIDA FORMATIONS Suwannee -
Uncorking the Bottle: What Triggered the Paleocene/Eocene Thermal Maximum Methane Release? Miriame
PALEOCEANOGRAPHY, VOL. 16, NO. 6, PAGES 549-562, DECEMBER 2001 Uncorking the bottle: What triggered the Paleocene/Eocene thermal maximum methane release? MiriamE. Katz,• BenjaminS. Cramer,Gregory S. Mountain,2 Samuel Katz, 3 and KennethG. Miller,1,2 Abstract. The Paleocene/Eocenethermal maximum (PETM) was a time of rapid global warming in both marine and continentalrealms that has been attributed to a massivemethane (CH4) releasefrom marine gas hydrate reservoirs. Previously proposedmechanisms for thismethane release rely on a changein deepwatersource region(s) to increasewater temperatures rapidly enoughto trigger the massivethermal dissociationof gas hydratereservoirs beneath the seafloor.To establish constraintson thermaldissociation, we modelheat flow throughthe sedimentcolumn and showthe effectof the temperature changeon the gashydrate stability zone throughtime. In addition,we provideseismic evidence tied to boreholedata for methanerelease along portions of the U.S. continentalslope; the releasesites are proximalto a buriedMesozoic reef front. Our modelresults, release site locations, published isotopic records, and oceancirculation models neither confirm nor refute thermaldissociation as the triggerfor the PETM methanerelease. In the absenceof definitiveevidence to confirmthermal dissociation,we investigatean altemativehypothesis in which continentalslope failure resulted in a catastrophicmethane release.Seismic and isotopic evidence indicates that Antarctic source deepwater circulation and seafloor erosion caused slope retreatalong -
Reconstructions of Late Ordovician Crinoids and Bryozoans from the Decorah Shale, Upper Mississippi Valley Sibo Wang Senior Inte
Reconstructions of Late Ordovician crinoids and bryozoans from the Decorah Shale, Upper Mississippi Valley Sibo Wang Senior Integrative Exercise March 10, 2010 Submitted in partial fulfillment of the requirements for a Bachelor of Arts degree from Carleton College, Northfield, Minnesota TABLE OF CONTENTS ABSTRACT INTRODUCTION ........................................................................................................ 01 GEOLOGIC SETTING ................................................................................................ 03 Late Ordovician world ................................................................................. 03 Southern Minnesota and the Decorah Shale ............................................... 03 Benthic community ....................................................................................... 05 Marine conditions ........................................................................................ 05 CRINOIDS ................................................................................................................. 06 General background and fossil record ........................................................ 06 Anatomy ....................................................................................................... 07 Decorah Shale crinoids ................................................................................10 BRYOZOANS ............................................................................................................. 10 General background and -
Clay Minerals at the Paleocene–Eocene Thermal Maximum: Interpretations, Limits, and Perspectives
minerals Review Clay Minerals at the Paleocene–Eocene Thermal Maximum: Interpretations, Limits, and Perspectives Fabio Tateo Istituto di Geoscienze e Georisorse, Consiglio Nazionale delle Ricerche (IGG-CNR) Padova, c/o Dipartimento di Geoscienze, Università di Padova, Via Gradenigo 6, I-35131 Padova, Italy; [email protected] Received: 20 October 2020; Accepted: 26 November 2020; Published: 30 November 2020 Abstract: The Paleocene–Eocene Thermal Maximum (PETM) was an “extreme” episode of environmental stress that affected the Earth in the past, and it has numerous affinities concerning the rapid increase in the greenhouse effect. It has left several biological, compositional, and sedimentary facies footprints in sedimentary records. Clay minerals are frequently used to decipher environmental effects because they represent their source areas, essentially in terms of climatic conditions and of transport mechanisms (a more or less fast travel, from the bedrocks to the final site of recovery). Clay mineral variations at the PETM have been studied by several authors in terms of climatic and provenance indicators, but also as tracers of more complicated interplay among different factors requiring integrated interpretation (facies sorting, marine circulation, wind transport, early diagenesis, etc.). Clay minerals were also believed to play a role in the recovery of pre-episode climatic conditions after the PETM exordium, by becoming a sink of atmospheric CO2 that is considered a necessary step to switch off the greenhouse hyperthermal effect. This review aims to consider the use of clay minerals made by different authors to study the effects of the PETM and their possible role as effective (simple) proxy tools for environmental reconstructions. -
GEOLOGIC TIME SCALE V
GSA GEOLOGIC TIME SCALE v. 4.0 CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST HIST. ANOM. (Ma) ANOM. CHRON. CHRO HOLOCENE 1 C1 QUATER- 0.01 30 C30 66.0 541 CALABRIAN NARY PLEISTOCENE* 1.8 31 C31 MAASTRICHTIAN 252 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 Lopin- 254 32 C32 72.1 635 2A C2A PIACENZIAN WUCHIAPINGIAN PLIOCENE 3.6 gian 33 260 260 3 ZANCLEAN CAPITANIAN NEOPRO- 5 C3 CAMPANIAN Guada- 265 750 CRYOGENIAN 5.3 80 C33 WORDIAN TEROZOIC 3A MESSINIAN LATE lupian 269 C3A 83.6 ROADIAN 272 850 7.2 SANTONIAN 4 KUNGURIAN C4 86.3 279 TONIAN CONIACIAN 280 4A Cisura- C4A TORTONIAN 90 89.8 1000 1000 PERMIAN ARTINSKIAN 10 5 TURONIAN lian C5 93.9 290 SAKMARIAN STENIAN 11.6 CENOMANIAN 296 SERRAVALLIAN 34 C34 ASSELIAN 299 5A 100 100 300 GZHELIAN 1200 C5A 13.8 LATE 304 KASIMOVIAN 307 1250 MESOPRO- 15 LANGHIAN ECTASIAN 5B C5B ALBIAN MIDDLE MOSCOVIAN 16.0 TEROZOIC 5C C5C 110 VANIAN 315 PENNSYL- 1400 EARLY 5D C5D MIOCENE 113 320 BASHKIRIAN 323 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 1500 CALYMMIAN 6 C6 APTIAN LATE 20 120 331 6A C6A 20.4 EARLY 1600 M0r 126 6B C6B AQUITANIAN M1 340 MIDDLE VISEAN MISSIS- M3 BARREMIAN SIPPIAN STATHERIAN C6C 23.0 6C 130 M5 CRETACEOUS 131 347 1750 HAUTERIVIAN 7 C7 CARBONIFEROUS EARLY TOURNAISIAN 1800 M10 134 25 7A C7A 359 8 C8 CHATTIAN VALANGINIAN M12 360 140 M14 139 FAMENNIAN OROSIRIAN 9 C9 M16 28.1 M18 BERRIASIAN 2000 PROTEROZOIC 10 C10 LATE -
A Middle Eocene Lowland Humid Subtropical “Shangri-La” Ecosystem in Central Tibet
A Middle Eocene lowland humid subtropical “Shangri-La” ecosystem in central Tibet Tao Sua,b,c,1, Robert A. Spicera,d, Fei-Xiang Wue,f, Alexander Farnsworthg, Jian Huanga,b, Cédric Del Rioa, Tao Dengc,e,f, Lin Dingh,i, Wei-Yu-Dong Denga,c, Yong-Jiang Huangj, Alice Hughesk, Lin-Bo Jiaj, Jian-Hua Jinl, Shu-Feng Lia,b, Shui-Qing Liangm, Jia Liua,b, Xiao-Yan Liun, Sarah Sherlockd, Teresa Spicera, Gaurav Srivastavao, He Tanga,c, Paul Valdesg, Teng-Xiang Wanga,c, Mike Widdowsonp, Meng-Xiao Wua,c, Yao-Wu Xinga,b, Cong-Li Xua, Jian Yangq, Cong Zhangr, Shi-Tao Zhangs, Xin-Wen Zhanga,c, Fan Zhaoa, and Zhe-Kun Zhoua,b,j,1 aCAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, China; bCenter of Plant Ecology, Core Botanical Gardens, Chinese Academy of Sciences, Mengla 666303, China; cUniversity of Chinese Academy of Sciences, 100049 Beijing, China; dSchool of Environment, Earth and Ecosystem Sciences, The Open University, Milton Keynes, MK7 6AA, United Kingdom; eKey Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 100044 Beijing, China; fCenter for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 100101 Beijing, China; gSchool of Geographical Sciences and Cabot Institute, University of Bristol, Bristol, BS8 1TH, United Kingdom; hCAS Center for Excellence in Tibetan Plateau Earth Sciences, Chinese Academy of Sciences, 100101 Beijing, China; iKey Laboratory of -
The Palaeocene – Eocene Thermal Maximum Super Greenhouse
The Palaeocene–Eocene Thermal Maximum super greenhouse: biotic and geochemical signatures, age models and mechanisms of global change A. SLUIJS1, G. J. BOWEN2, H. BRINKHUIS1, L. J. LOURENS3 & E. THOMAS4 1Palaeoecology, Institute of Environmental Biology, Utrecht University, Laboratory of Palaeobotany and Palynology, Budapestlaan 4, 3584 CD Utrecht, The Netherlands (e-mail: [email protected]) 2Earth and Atmospheric Sciences, Purdue University, 550 Stadium Mall Drive, West Lafayette, IN 47907, USA 3Faculty of Geosciences, Department of Earth Sciences, Utrecht University, Budapestlaan 4, 3584 CD Utrecht, The Netherlands 4Center for the Study of Global Change, Department of Geology and Geophysics, Yale University, New Haven CT 06520-8109, USA; also at Department of Earth & Environmental Sciences, Wesleyan University, Middletown, CT, USA Abstract: The Palaeocene–Eocene Thermal Maximum (PETM), a geologically brief episode of global warming associated with the Palaeocene–Eocene boundary, has been studied extensively since its discovery in 1991. The PETM is characterized by a globally quasi-uniform 5–8 8C warming and large changes in ocean chemistry and biotic response. The warming is associated with a negative carbon isotope excursion (CIE), reflecting geologically rapid input of large amounts of isotopically light CO2 and/or CH4 into the exogenic (ocean–atmosphere) carbon pool. The biotic response on land and in the oceans was heterogeneous in nature and severity, including radiations, extinctions and migrations. Recently, several events that appear -
Decoding the Fossil Record of Early Lophophorates
Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 1284 Decoding the fossil record of early lophophorates Systematics and phylogeny of problematic Cambrian Lophotrochozoa AODHÁN D. BUTLER ACTA UNIVERSITATIS UPSALIENSIS ISSN 1651-6214 ISBN 978-91-554-9327-1 UPPSALA urn:nbn:se:uu:diva-261907 2015 Dissertation presented at Uppsala University to be publicly examined in Hambergsalen, Geocentrum, Villavägen 16, Uppsala, Friday, 23 October 2015 at 13:15 for the degree of Doctor of Philosophy. The examination will be conducted in English. Faculty examiner: Professor Maggie Cusack (School of Geographical and Earth Sciences, University of Glasgow). Abstract Butler, A. D. 2015. Decoding the fossil record of early lophophorates. Systematics and phylogeny of problematic Cambrian Lophotrochozoa. (De tidigaste fossila lofoforaterna. Problematiska kambriska lofotrochozoers systematik och fylogeni). Digital Comprehensive Summaries of Uppsala Dissertations from the Faculty of Science and Technology 1284. 65 pp. Uppsala: Acta Universitatis Upsaliensis. ISBN 978-91-554-9327-1. The evolutionary origins of animal phyla are intimately linked with the Cambrian explosion, a period of radical ecological and evolutionary innovation that begins approximately 540 Mya and continues for some 20 million years, during which most major animal groups appear. Lophotrochozoa, a major group of protostome animals that includes molluscs, annelids and brachiopods, represent a significant component of the oldest known fossil records of biomineralised animals, as disclosed by the enigmatic ‘small shelly fossil’ faunas of the early Cambrian. Determining the affinities of these scleritome taxa is highly informative for examining Cambrian evolutionary patterns, since many are supposed stem- group Lophotrochozoa. The main focus of this thesis pertained to the stem-group of the Brachiopoda, a highly diverse and important clade of suspension feeding animals in the Palaeozoic era, which are still extant but with only with a fraction of past diversity. -
Eocene, Oligocene, and Miocene Rocks and \Fertebrate Fossils at the Emerald Lake Locality, 3 Miles South of Ifellowstone National Park,Wyoming
Eocene, Oligocene, and Miocene Rocks and \fertebrate Fossils at the Emerald Lake Locality, 3 Miles South of Ifellowstone National Park,Wyoming GEOLOGICAL SURVEY PROFESSIONAL PAPER 932-A Prepared in cooperation with the Geological Survey of Wyoming, the Department of Geology of the University of Wyoming, the American Museum of Natural History, and the Carnegie Museum Eocene, Oligocene, and Miocene Rocks and \fertebrate Fossils at the Emerald Lake Locality 3 Miles South of Ifellowstone National Park,Wyoming By J. D. LOVE, MALCOLM C. McKENNA, and MARY R. DAWSON GEOLOGY OF THE TETON-JACKSON HOLE REGION, NORTHWESTERN WYOMING GEOLOGICAL SURVEY PROFESSIONAL PAPER 932-A Prepared in cooperation with the Geological Survey of Wyoming, the Department of Geology of the University of Wyoming, the American Museum of Natural History, and the Carnegie Museum UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1976 UNITED STATES DEPARTMENT OF THE INTERIOR THOMAS S. KLEPPE, Secretary GEOLOGICAL SURVEY V. E. McKelvey, Director Library of Congress Cataloging in Publication Data Love, John David, 1913- Eocene, Oligocene, and Miocene rocks and vertebrate fossils at the Emerald Lake locality, 3 miles south of Yellowstone National Park, Wyoming. (Geology of the Teton-Jackson Hole region, northwestern Wyoming) (Geological Survey Professional Paper 932-A) Bibliography: p. Includes index. 1. Geology, Stratigraphic Tertiary. 2. Vertebrates, Fossil. 3. Geology Wyoming Teton Co. I. McKenna, Malcolm C., joint author. II. Dawson, Mary R., joint author. III. Wyoming Geological Survey. IV. Title: Eocene, Oligocene, and Miocene rocks and vertebrate fossils at the Emerald Lake locality ... V. Series. VI. Series: United States Geological Survey Professional Paper 932-A. QE691.L79 551.?'8 76-8159 For sale by the Superintendent of Documents, U.S.