Proceedings of the Desert Council

VOLUME XXVI

1994 ANNUAL SYMPOSIUM 17 to 20 November Death Valley National Park Furnace Creek California U.S.A.

Edited by

Dean A. Hendrickson Texas Natural History Collection / R4000 University of Texas Austin, TX 78712-1100

Publication date: September 30, 1995 ISSN 1068-0381 P.O. Box 337 • Bishop, California 93515-0337 • (619) 872-8751 Voice and Fax MISSION

The mission of the Desert Fishes Council is to preserve the biological integrity of North America's desert aquatic ecosystems and their associated life forms, to hold symposia to report related research and management endeavors, and to effect rapid dissemination of information concerning activities of the Council and its members. OFFICERS

Chairman: Dean A. Hendrickson, Texas Natural History Collection/R4000, University of Texas, Austin, Texas Chairman-elect: vacant Executive Secretary: E. P. Pister, Bishop, CA, U.S.A. COMMITTEES

Awards: Gary K. Meffe Membership: Paul C. Marsh Proceedings: Dean A. Hendrickson (Chair/Editor) Proceedings Translation sub-committee (Dr. Samuel Ocaña, Director, Centro Ecológico de Sonora, sponsor): Biol. Guadalupe Morales-Abril (Chair), Biol. Mar. Andrés Villarreal Lizárraga, Ecól. Elvira Rojero Díaz, Biol. Mar. Alejandro Varela Romero, Biol. Eduardo López Saavedra, Ecól. José Martín E. Haro Rodríguez, Biol. Iván Eduardo Parra Salazar, Biol. Florentino Garza Salazar, Biol. Cristina Meléndez Torres, Ecól. María Isabel Granillo, Dr. Dean A. Hendrickson Program: Michael E. Douglas (Chair), Dean A. Hendrickson, Nadine Kanim Resolutions: Walter R. Courtenay, Jr. Local Committees: 1995 (Reno, Nevada, U.S.A.) Glenn Clemmer, Randy McNatt; 1996 (La Paz, Baja California, México) Francisco Reynoso-Mendoza; 1997 (Death Valley National Park, Furnace Creek, California, U.S.A.) E. Phil Pister. MEMBERSHIP

Membership in the Desert Fishes Council is open to any person or organization interested in or engaged in the management, protection, or scientific study of desert fishes, or some related phase of desert conservation. Membership includes subscription to the Proceedings of the Desert Fishes Council. Annual dues are $25 (regular: domestic or foreign), $15 (student), $35 (family: 1 Proceedings), >$35 (sustaining) and $1000 (patron: single payment). Send dues payments and general contributions along with address information (including affiliation, voice, fax, and e-mail) and indication of permission to include this information in a published directory of the Desert Fishes Council to: Paul C. Marsh, Membership Chair, Center for Environmental Studies, Arizona State University, Tempe, AZ 85287-3211 USA (602/965-2977 FAX 602/965-8087/ Internet [email protected] ABOUT PROCEEDINGS OF THE DESERT FISHES COUNCIL

It is the policy of the Council to publish in the annual Proceedings of the Desert Fishes Council papers, abstracts, discussion summaries, business items, resolutions, and other material submitted for presentation, whether actually presented at the Annual Symposium or not. The Proceedings are published and delivered to all members of the Desert Fishes Council and subscribing libraries in the year following the Annual Symposium. All contributions are published as received following automated electronic processing designed to standardize format only. Authors are responsible for their own technical editing and for any errors caused by failure to follow Instructions to Authors (published in each volume). Proofs are not provided to authors for review prior to publication, and neither abstracts nor full papers are subjected to peer review. Resolutions are published exactly as passed by the membership in the business meeting of the annual symposium. The Translation Subcommittee of the Proceedings Committee accepts responsibility for errors in translations to Spanish for those abstracts they translate. This committee provides original translations of all abstract