New Mexican and Central American Ephemeroptera Records, with First Species Checklists for Mexican States Author(S): W

New Mexican and Central American Ephemeroptera Records, with First Species Checklists for Mexican States Author(s): W. P. McCafferty Source: Transactions of the American Entomological Society, 137(3 & 4):317-327. 2011. Published By: The American Entomological Society URL: http://www.bioone.org/doi/full/10.3157/061.137.0310

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BioOne sees sustainable scholarly publishing as an inherently collaborative enterprise connecting authors, nonprofit publishers, academic institutions, research libraries, and research funders in the common goal of maximizing access to critical research. TRANSACTIONS OF THE AMERICAN ENTOMOLOGICAL SOCIETY VOLUME 137, NUMBERS 3+4: 317-327, 2011

New Mexican and Central American Ephemeroptera records, with ﬁrst species checklists for Mexican states

W. P. McCafferty

Department of Entomology, Purdue University, West Lafayette, IN 47907

ABSTRACT

Twenty-three Ephemeroptera species are reported variously from four Central American countries (Costa Rica, Honduras, Nicaragua, Panama) and nine Mexican states (Chiapas, Durango, Guanajuato, Guerrero, Hidalgo, Oaxaca, Puebla, Queretaro, San Luis Potosi) for the first time. Campsurus decoloratus, Fallceon nikitai, and Tricorythodes explicatus are reported from Central America for the first time. Allenhyphes vescus is reported from southern Mexico for the first time, and second ever records are given for Choroterpes ungulus (Durango), and Leptohyphes priapus (Panama). Published record based checklists for the 28 Mexican states with reported mayflies are provided. The greatest diversity is found in Vera Cruz followed in order by Chihuahua, Chiapas, Tamaulipas, Neuvo Leon and Queretaro. Baetis magnus, Callibaetis pictus, Camelobaeitidus mexicanum, Fallceon quilleri, Neochoroterpes oklahoma, Thraulodes brunneus, Traverella albertana and Vacupernius packeri are among the most commonly encountered mayflies in Mexico.

INTRODUCTION

The following records are based on RECORD DATA identifications of material housed in the Purdue Entomological Research Collection and the Packer Allenhyphes vescus (Allen): MEXICO, Queretaro, Honduras Collection. The contribution continues Neblinas, Rio Verdito, 13-VII-2000, W. D. a series of records reports documenting mayfly Shepard. This species has been known from biodiversity and distribution in Central America Texas, Arizona, Chihuahua, Nuevo Leon and Mexico (McCafferty 1985, McCafferty and and Tamaulipas (see e.g., McCafferty 2011, Lugo-Ortiz 1992, Lugo-Ortiz and McCafferty McCafferty et al. 2012). Queretaro represents 1994, 1995a,b, 1996b, c, McCafferty and Lugo- the first record from as far south as southern Ortiz 1996, Randolph and McCafferty 2000, Mexico. 2001, 2002, McCafferty et al. 2004, Meyer et Asioplax isabelia Baumgardner, Meyer & al. 2008, McCafferty 2011) generated from McCafferty: HONDURAS, Dept. Cortes, sorting, identification and curation of acquisitions Chamelecon, Rio Chamelecon, 18-X-1964, associated with the entomological collections J. S Packer. This striking species has been program at Purdue University. Considerable known previously from Costa Rica and records are byproducts of systematic studies at Nicaragua (Baumgardner et al. 2006). Purdue (e.g., Waltz and McCafferty 1985, 1987, Callibaetis californicus Banks: MEXICO, Hidalgo, Lugo-Ortiz and McCafferty 1996a, McCafferty 10 mi N Monterey, Sabinus, 25-XII-1947, S. and Randolph 2000, McCafferty and Baumgardner Mulaik. This species is known in the USA 2003, Baumgardner and McCafferty 2010). All from California, Idaho and Texas, although materials reported here are in reference to larvae I have also seen material of it taken at a unless the data are appended with (A) indicating spring in Garfield County, Oklahoma (no that alate materials are referenced. other data). In Mexico, it has been known from Baja California, Baja California Sur, 318 MEXICAN AND CENTRAL AMERICAN EPHEMEROPTERA RECORDS

Guerrero, Mexico State, Morelos and Nuevo Fallceon nikitai McCafferty & Lugo-Ortiz, COSTA Leon (see e.g., Randolph and McCafferty RICA, Guanacaste Province, Rio Tenorio at 2000, McCafferty et al. 2012). Finca La Pacifica, E Pan Am Hwy, 2-8-II- Camelobaetidius arriaga (Traver & Edmunds): 1969, W. P. McCafferty. This species was COSTA RICA, Guanacaste Province, Rio previously known from Cuba (McCafferty Tenorio at Finca La Pacifica, E Pan Am and Lugo-Ortiz 1994). Hwy, 2-II-1969, W. P. McCafferty; Alajuela, Farrodes otiesa Lugo-Ortiz & McCafferty, Rio Pizote, ca 5 km S Brasilia, 12-III- MEXICO, Oaxaca, Oaxaca Comino, at 1986, Holzenthal, Fasth. This species has Guelatao, 9-III-1978, Zapién. This species been correctly reported only from Chiapas was previously known only from Costa Rica, and Vera Cruz in Mexico, and in Central Queretaro and Tamaulipas (Lugo-Ortiz and American from Belize and Guatemala (see McCafferty 1996a, Randolph and McCafferty McCafferty 2011). 2000, 2001). Camelobaetidius warreni (Traver & Edmunds): Farrodes texanus Davis, COSTA RICA, San PANAMA, Canal Zone, Rio Guanabano, 3 Jose, Rio Parrita Chiquito, 10-VI-1987, mi N Fort Clayton, and Rio Camaron, N edge Holzenthal, Hamilton, Heyn. This species Fort Clayton, Chiva-Chiva Rd, 9-IX-1963, is also known from Belize, Guatemala, and Rio Venado, Fort Kobbe, 10-IX-1964, Nicaragua, Tamaulipas and Texas (see W. L. Peters, C. M. Keenan. This relatively McCafferty et al. 2004, Meyer et al. 2008). common species is known from throughout Hexagenia albivitta (Walker): MEXICO, Chiapas, Mexico, Central America, and much of Agua Azul, 1-V-1978, H. Brailowsky (A). western North America north of Mexico (see In Mexico, this species has been known also review in McCafferty et al. 2012). Panama from the adjacent southern state of Vera Cruz represents the southernmost part of its known (Lugo-Ortiz and McCafferty 1996b). It has range. also been known from Guatemala and Costa Campsurus decoloratus (Hagen): NICARAGUA, Rica (see McCafferty et al. 2004), and is best La Flores, Finca Tullez, 20-XI-1987, J. M. known from much of tropical South America. Maes (A). This represents the first Central Leptohyphes apache Allen: COSTA RICA, American record of this species, which Guanacaste Province, Rio Tenorio, at Finca otherwise had been known only from Texas La Pacifica, E Pan Am Hwy, 2-II-1969, W. and Tamaulipas (see McCafferty 1975). P. McCafferty; Heredia Province, Quebrada Choroterpes ungulus Lugo-Ortiz & McCafferty: Salto in SW section of teaching area at La MEXICO, Durango, La Michilia, Arroyo Selva, 25-II-1969, W. P. McCafferty. This Nana Juana, Bosque seco de pino-encino,15- species is known from the southwestern USA IV-1988, R. Novelo. This species had (see McCafferty et al. 2012), in Mexico from previously been known only from its the states of Chiapas, Chihuahua, Morelos, type locality in Chiapas (Lugo-Ortiz and Oaxaca, Queretaro, Sonora, Tamaulipas McCafferty 1996a). and Vera Cruz, and in Central America it Epiphrades undatus (Lugo-Ortiz & McCafferty): is also known from Belize, Guatemala, HONDURAS, Dept. Francisco, Morazan, Honduras, Nicaragua and El Salvador (see 6.5 mi from Junction Hwys 3 and 5, 7-XI- also Baumgardner and McCafferty 2010, 1964, J. S. Packer. This Central American McCafferty 2011). species has also been taken from Costa Leptohyphes priapus Traver: PANAMA, Pueblo Rica (Lugo-Ortiz and McCafferty 1995b), Nuevo Cave, 16-VI-1999, W. Reeves (A). Guatemala (McCafferty et al. 2004), and This species is known only in the adult stage Nicaragua (Meyer et al. 2008). and was previously known only from its type locality in Costa Rica (see Baumgardner and McCafferty 2010). W. P. MCCAFFERTY 319

Neochoroterpes oklahoma (Traver): MEXICO, Traverella albertana (McDunnough): MEXICO, Puebla, Rio Lacan, Ha Km 22.5 carratera, Puebla, Rio Lacan, Ha Km 22.5 carretera, Agua Azul-Ocotzingo, 22-VII-1978, J. Agua Azul, Ocotzingo, 22-VII-1978, Bueno, J. Padilla. This species has been J. Bueno, J. Padilla. This is the most known throughout much of Mexico as well widespread Traverella species in North as prairie regions in the western USA (see America, and should be found in nearly all Review in McCafferty et al. 2012). of Mexico. It has also been found in Central Paracloeodes minutus (Daggy): MEXICO, San America in Nicaragua (Meyer et al. 2008). Luis Potosi, Ruinas de Tamuin, 19-V-1978, Traverella presidiana (Traver): MEXICO, Hidalgo, J. Bueno (A). Surprisingly, this species, Rio Venados, 6-X-1979, Zapién. This which is known as a transcontinental species species has been known from Texas and in the USA (e.g., see McCafferty et al. 2012) eastern Mexico, including the neighboring and is also known from several Central states of Coahuila, Neuvo Leon and American countries including Belize, Costa Tamaulipas (all of which border Texas) as Rica, Honduras and Nicaragua (e.g., see well as Chiapas to the south and Chihuahua Meyer et al. 2008), has been reported from to the north. Mexico only from Baja California Sur (Lugo- Tricorythodes explicatus (Eaton): HONDURAS, Ortiz and McCafferty 1994). It is expected Dept. El Paraiso, Escuela Agricola to occur throughout Mexico, but probably Panamericana 10-III-1967, J. S. Packer (A). because of its small size and difﬁculty in This is the ﬁrst report of this widespread sampling, it has been overlooked in much of western North American species from Mexico. south of Mexico, although it recently was Thraulodes brunneus Koss: MEXICO, also reported from southern Mexico by Guanajuato, San Miguel, VI-1984, Zapién; McCafferty (2011). San Luis Potosi, Rio Xilitla, 12-IV-1980, Tricorythodes sordidus Allen, 1967: HONDURAS, Zapién. This species was originally known Dept. Olancho, 1.6 mi W Campamento from the extreme southwestern USA, but Galera turn-off at Bridge on Hwy 3, 7-XI- is now known throughout much of Mexico 1964, J. S. Packer; Dept. El Paraiso, 6 mi (e.g., Randolph and McCafferty 2000, 2001) E Danli, 6-VIII-1966, J. S. Packer. These and is known to extend to Central America are the northernmost records of this Central in Costa Rica, Guatemala and Honduras (see American species. It has also been reported McCafferty et al. 2004). from Costa Rica (Allen 1967), Guatemala Thraulodes pacaya McCafferty, Baumgardner & (McCafferty et al. 2004), and Nicaragua Guenther: MEXICO, Puebla, Rio Lacan, Ha (Baumgardner and Ávila 2006, Meyer et al. Km 22.5 carretera, Agua Azul, Ocotzingo, 2008). 22-VII-1978, J. Bueno, J. Padilla. This species has been known from Chiapas, San Luis Potosi, Tabasco, Vera Cruz, Guatemala, Honduras, Nicaragua, and Panama (Allen and Brusca 1978, McCafferty et al. 2004, Meyer et al. 2008). Thraulodes tenulineus Lugo-Ortiz & McCafferty, MEXICO, Guerrero, Km 15 carretera Bejucos. Altamirano, 22-XI-1984. This species is known from Arizona and almost all northern states of Mexico as well as some southern Mexican states (Randolph and McCafferty 2000, McCafferty et al. 2012). 320 MEXICAN AND CENTRAL AMERICAN EPHEMEROPTERA RECORDS

CHECKLISTS Camelobaetidius musseri (Traver & Edmunds) Camelobaetidius warreni (Traver & Edmunds) The Mexican species Thraulodes humeralis Choroterpes inornata Eaton Navás, T. mexicanus (Eaton) and Traverella sallei Cloeodes excogitatus Waltz & McCafferty (Navás), while known from Mexico, are based on Cloeodes peninsulus Waltz & McCafferty records that did not include state designation nor Epeorus margarita Edmunds & Allen can such be ascertained with certainty, and thus Fallceon quilleri (Dodds) they do not appear in the following Mexican state Paracloeodes minutus (Daggy) checklists. No mayflies have been reported from Serratella micheneri (Traver) the Mexican states of Colima, Compeche, Quintana Thraulodes brunneus Koss Roo and Yucatan, the latter three of which generally Tricorythodes explicatus (Eaton) lack habitat to support mayflies. With respect to diversity and distribution of mayflies in Central Chiapas American countries, a comprehensive up-to-date Americabaetis lugoi Waltz & McCafferty accounting can be found at Mayfly Central at http:// Baetis magnus McCafferty & Waltz www.entm.purdue.edu/mayfly/ca-species-list.php. Baetodes caritus Cohen & Allen Baetodes pallidus Cohen & Allen Aguascalientes Baetodes tritus Cohen & Allen Callibaetis floridanus Banks Caenis bajaensis Allen & Murvosh Caenis latipennis Banks Baja California Callibaetis floridanus Banks Baetis adonis Traver Callibaetis pictus (Eaton) Baetis tricaudatus Dodds Camelobaetidius arriaga (Traver & Edmunds) Caenis bajaensis Allen & Murvosh Camelobaetidius mexicanus (Traver & Edmunds) Callibaetis californicus Banks Camelobaetidius musseri (Traver & Edmunds) Callibaetis pictus (Eaton) Camelobaetidius warreni (Traver & Edmunds) Choroterpes inornata Eaton Choroterpes ungulus Lugo-Ortiz & McCafferty Drunella coloradensis Dodds Cloeodes excogitatus Waltz & McCafferty Drunella flavilinea (McDunnough) Fallceon quilleri (Dodds) Epeorus margarita Edmunds & Allen Farrodes maya Domínguez Fallceon eatoni (Kimmins) Farrodes tulija Domínguez, Molineri & Peters Fallceon quilleri (Dodds) Hexagenia albivitta (Walker) Ironodes nitidus (Eaton) Homoeoneuria salviniae Eaton Paraleptophlebia memorialis (Eaton) Lachania iops Allen & Cohen Rhithrogena morrisoni (Banks) Leptohyphes apache Allen Serratella micheneri (Traver) Leptohyphes musseri Allen Thraulodes brunneus Koss Leptohyphes zalope Traver Tricorythodes explicatus (Eaton) Moribaetis macaferti Waltz Vacupernius packeri (Allen) Neochoroterpes oklahoma (Traver) Thraulodes cryptodrilus Nieto & Domínguez Baja California Sur Thraulodes eccentricus Logo-Ortiz & McCafferty Baetis adonis Traver Thraulodes ephippiatus Traver & Edmunds Baetodes longus Mayo Thraulodes pacaya McCafferty, Baumgardner & Caenis bajaensis Allen & Murvosh Guenther Callibaetis californicus Banks Thraulodes packeri Traver & Edmunds Callibaetis montanus Eaton Thraulodes spangleri Traver & Edmunds Callibaetis pictus (Eaton) Thraulodes tenulineus Lugo-Ortiz & McCafferty Camelobaetidius mexicanus (Traver & Edmunds) Thraulodes zonalis Traver & Edmunds W. P. MCCAFFERTY 321

Traverella albertana (McDunnough) Vacupernius packeri (Allen) Traverella presidiana (Traver) Traverella promifrons Lugo-Ortiz & McCafferty Coahuila Vacupernius packeri (Allen) Callibaetis punctilusus McCafferty & Provonsha Varipes lasiobrachius Lugo-Ortiz & McCafferty Camelobaetidius kickapoo McCafferty Fallceon quilleri (Dodds) Chihuahua Neochoroterpes oklahoma (Traver) Acentrella insignificans (McDunnough) Rhithrogena plana Allen & Chao Acentrella turbida (McDunnough) Thraulodes gonzalesi Traver & Edmunds Allenhyphes vescus (Allen) Traverella presidiana (Traver) Baetis adonis Traver Tricorythodes explicatus (Eaton) Baetis flavistriga McDunnough Baetis magnus McCafferty & Waltz Distrito Federal Baetodes caritus Cohen & Allen Baetis magnus McCafferty & Waltz Baetodes pictus Cohen & Allen Baetis tricaudatus Dodds Caenis amica Hagen Callibaetis montanus Eaton Caenis latipennis Banks Callibaetis pictus (Eaton) Callibaetis floridanus Banks Callibaetis undatus (Pictet) Callibaetis pictus (Eaton) Rhithrogena notialis Allen & Cohen Callibaetis punctilusus McCafferty & Provonsha Thraulodes eccentricus Lugo-Ortiz & McCafferty Camelobaetidius kickapoo McCafferty Camelobaetidius warreni (Traver & Edmunds) Durango Choroterpes inornata Eaton Baetis magnus McCafferty & Waltz Cloeodes macrolamellus Waltz & McCafferty Caenis bajaensis Allen & Murvosh Epeorus margarita Edmunds & Allen Callibaetis floridanus Banks Euthyplocia hecuba (Hagen) Callibaetis pictus (Eaton) Fallceon quilleri (Dodds) Choroterpes ungulus Lugo-Ortiz & McCafferty Heptagenia elegantula (Eaton) Cloeodes macrolamellus Waltz & McCafferty Heterocloeon amplum (Traver) Epeorus metlacensis Traver Homoeoneuria alleni Pescador & Peters Neochoroterpes oklahoma (Traver) Homoleptohyphes dimorphus (Allen) Thraulodes brunneus Koss Homoleptohyphes mirus (Allen) Tricorythodes explicatus (Eaton) Isonychia intermedia (Eaton) Tricorythodes fictus Traver Isonychia sicca (Walsh) Labiobaetis ephippiatus (Traver) Guanajuato Leptohyphes apache Allen Caenis bajaensis Allen & Murvosh Neochoroterpes oklahoma (Traver) Homoleptohyphes dimorphus (Allen) Rhithrogena plana Allen & Chao Thraulodes brunneus Koss Siphlonurus occidentalis (Eaton) Tricoryhyphes mulaiki (Traver) Thraulodes brunneus Koss Tricoythodes explicatus (Eaton) Thraulodes gonzalesi Traver & Edmunds Thraulodes speciosus Traver Guerrero Thraulodes tenulineus Lugo-Ortiz & McCafferty Ableptemetes dicinctus (Allen & Brusca) Traverella albertana (McDunnough) Baetis magnus McCafferty & Waltz Traverella presidiana (Traver) Baetodes caritus Cohen & Allen Tricoryhyphes condylus (Allen) Baetodes deficiens Cohen & Allen Tricorythodes explicatus (Eaton) Baetodes inermis Cohen & Allen Tricorythodes fictus Traver Baetodes tritus Cohen & Allen 322 MEXICAN AND CENTRAL AMERICAN EPHEMEROPTERA RECORDS

Callibaetis californicus Banks Fallceon quilleri (Dodds) Callibaetis floridanus Banks Hexagenia limbata (Serville) Callibaetis pictus (Eaton) Isonychia sicca (Walsh) Camelobaetidius mexicanus (Traver & Edmunds) Leptohyphes musseri Allen Camelobetidius musseri (Traver & Edmunds) Thraulodes brunneus Koss Camelobaetidius warreni (Traver & Edmunds) Traverella albertana (McDunnough) Fallceon quilleri (Dodds) Lachlania saskatchewanensis Ide Mexico Leptohyphes lestes Allen & Brusca Americabaetis intermedius (Lugo-Ortiz & Leptohyphes musseri Allen McCafferty) Leptohyphes nigripunctum Traver Baetis magnus McCafferty & Waltz Leptohyphes peterseni Ulmer Baetis notos Allen & Murvosh Leptohyphes zalope Traver Baetis tricaudatus Dodds Moribaetis macaferti Waltz Baetodes pallidus Cohen & Allen Paracloeodes lugoi Randolph & McCafferty Callibaetis californicus Banks Thraulodes brunneus Koss Callibaetis pictus (Eaton) Thraulodes eccentricus Lugo-Ortiz & McCafferty Epeorus metlacensis Traver Thraulodes speciosus Traver Lachlania fusca (Navás) Thraulodes tenulineus Lugo-Ortiz & McCafferty Moribaetis macaferti Waltz Traverella albertana (McDunnough) Thraulodes brunneus Koss Tricoryhyphes mulaiki (Traver) Tricorythodes comus Traver Michoacan Baetis magnus McCafferty & Waltz Hidalgo Callibaetis pictus (Eaton) Baetis magnus McCafferty & Waltz Ecdyonurus bellus (Allen & Cohen) Callibaetis floridanus Banks Epeorus metlacensis Traver Callibaetis pictus (Eaton) Fallceon quilleri (Dodds) Epeorus metlacensis Traver Rhithrogena notialis Allen & Cohen Fallceon sageae McCafferty Thraulodes brunneus Koss Siphlonurus occidentalis (Eaton) Thraulodes eccentricus Lugo-Ortiz & McCafferty Morelos Thraulodes lunatus Traver & Edmunds Baetis magnus McCafferty & Waltz Traverella albertana (McDunnough) Baetis tricaudatus Dodds Traverella presidiana (Traver) Baetodes caritus Cohen & Allen Baetodes deficiens Cohen & Allen Baetodes fuscipes Cohen & Allen Jalisco Baetodes inermis Cohen & Allen Baetis magnus McCafferty & Waltz Baetodes tritus Cohen & Allen Baetis tricaudatus Dodds Callibaetis californicus Banks Baetodes deficiens Cohen & Allen Callibaetis floridanus Banks Baetodes inermis Cohen & Allen Callibaetis pictus (Eaton) Baetodes pallidus Cohen & Allen Camelobaetidius mexicanus (Traver & Edmunds) Baetodes tritus Cohen & Allen Camelobaetidius musseri (Traver & Edmunds) Callibaetis floridanus Banks Epeorus metlacensis Traver Callibaetis montanus Eaton Fallceon quilleri (Dodds) Callibaetis pictus (Eaton) Leptohyphes apache Allen Camelobaetidius mexicanus (Traver & Edmunds) Leptphyphes musseri Allen Camelobaetidius musseri (Traver & Edmunds) Paracloeodes lugoi Randolph & McCafferty W. P. MCCAFFERTY 323

Thraulodes brunneus Koss Oaxaca Thraulodes eccentricus Lugo-Ortiz & McCafferty Americabaetis intermedius (Lugo-Ortiz & Thraulodes tenulineus Lugo-Ortiz & McCafferty McCafferty) Traverella albertana (McDunnough) Baetis magnus McCafferty & Waltz Tricorythodes notatus Allen & Brusca Baetis tricaudatus Dodds Tricorythodes ulmeri Allen & Brusca Baetodes inermis Cohen & Allen Vacupernius packeri (Allen) Callibaetis floridanus Banks Callibaetis pictus (Eaton) Nayarit Camelobaetidius maidu Jacobus & McCafferty Caenis bajaensis Allen & Murvosh Camelobaetidius mexicanus (Traver & Edmunds) Thraulodes tenulineus Lugo-Ortiz & McCafferty Camelobaetidius musseri (Traver & Edmunds) Vacupernius packeri (Allen) Camelobaetidius warreni (Traver & Edmunds) Choroterpes inornata Eaton Nuevo Leon Epeorus metlacensis Traver Allenhyphes vescus (Allen) Fallceon quilleri (Dodds) Americabaetis pleturus (Lugo-Ortiz & McCafferty) Farrodes otiesa Lugo-Ortiz & McCafferty Baetis magnus McCafferty & Waltz Hexagenia mexicana Eaton Baetodes bibranchius McCafferty & Provonsha Leptohyphes alleni Brusca Baetodes longus Mayo Leptoyphes apache Allen Baetodes tritus Cohen & Allen Leptohyphes musseri Allen Baetodes velmae Cohen & Allen Leptohyphes sabinas Traver Caenis latipennis Banks Leptohyphes zalope Traver Caenis punctata McDunnough Moribaetis macaferti Waltz Callibaetis californicus Banks Rhithrogena notialis Allen & Cohen Callibaetis floridanus Banks Thraulodes brunneus Koss Callibaetis pictus (Eaton) Thraulodes eccentricus Lugo-Ortiz & McCafferty Callibaetis punctilusus McCafferty & Provonsha Thraulodes tenulineus Lugo-Ortiz & McCafferty Camelobaetidius mexicanus (Traver & Edmunds) Tricoythodes notatus Allen & Brusca Camelobaetidius musseri (Traver & Edmunds) Vacupernius packeri (Allen) Camelobaetidius variabilis Wiersema Cloeodes excogitatus Waltz & McCafferty Puebla Fallceon eatoni (Kimmins) Baetis magnus McCafferty & Waltz Fallceon quilleri (Dodds) Baetodes inermis Cohen & Allen Heptagenia flavescens (Walsh) Callibaetis pictus (Eaton) Hexagenia limbata (Serville) Fallceon quilleri (Dodds) Isonychia sicca (Walsh) Moribaetis macaferti Waltz Leptohyphes sabinus Traver Neochoroterpes oklahoma (Traver) Leptohyphes zalope Traver Neochoroterpes orientalis Henry Neochoroterpes oklahoma (Traver) Thraulodes pacaya McCafferty, Baumgardner & Thraulodes eccentricus Lugo-Ortiz & McCafferty Guenther Thraulodes gonzalesi Traver & Edmunds Traverella albertana (McDunnough) Thraulodes lunatus Traver & Edmunds Thraulodes packeri Traver & Edmunds Queretaro Traverella presidiana (Traver) Acentrella insignificans (McDunnough) Traverella promifrons Lugo-Ortiz & McCafferty Allenhyphes vescus (Allen) Tricorythodes sordidus Allen Americabaetis pleturus (Lugo-Ortiz & McCafferty) Vacupernius packeri (Allen) Asioplax numinuh Wiersema, McCafferty & Baumgardner 324 MEXICAN AND CENTRAL AMERICAN EPHEMEROPTERA RECORDS

Baetis magnus McCafferty & Waltz Thraulodes tenulineus Lugo-Ortiz & McCafferty Baetodes deficiens Cohen & Allen Traverella longifrons Lugo-Ortiz & McCafferty Baetodes fuscipes Cohen & Allen Tricorythodes sordidus Allen Baetodes inermis Cohen & Allen Vacupernius packeri (Allen) Baetodes longus Mayo Baetodes tritus Cohen & Allen Sinaloa Baetodes velmae Cohen & Allen Baetodes edmundsi Koss Callibaetis pictus (Eaton) Baetodes fuscipes Cohen & Allen Camelobaetidius kickapoo McCafferty Callibaetis pictus (Eaton) Camelobaetidius mexicanus (Traver & Edmunds) Camelobaetidius maidu Jacobus & McCafferty Camelobaetidius shepardi Randolph & McCafferty Camelobaetidius mexicanus (Traver & Edmunds) Camelobaetidius variabilis Wiersema Camelobaetidius musseri (Traver & Edmunds) Camelobaetidius warreni (Traver & Edmunds) Camelobaetidius warreni (Traver & Edmunds) Choroterpes inornata Eaton Choroterpes inornata Eaton Cloeodes excogitatus Waltz & McCafferty Cloeodes excogitatus Waltz & McCafferty Ecdyonurus bellus (Allen & Cohen) Fallceon quilleri (Dodds) Epeorus metlacensis Traver Homoleptohyphes mirus (Allen) Fallceon quilleri Dodds) Leptohyphes lestes Allen & Brusca Farrodes otiesa Lugo-Ortiz & McCafferty Thraulodes brunneus Koss Leptohyphes apache Allen Thraulodes gonzalesi Traver & Edmunds Leptohyphes zalope Traver Thraulodes tenulineus Lugo-Ortiz & McCafferty Moribaetis macaferti Waltz Tortopsis primus (McDunnough) Neochoroterpes orientalis Henry Traverella albertana (McDunnough) Thraulodes brunneus Koss Tricorythodes explicatus (Eaton) Thraulodes eccentricus Lugo-Ortiz & McCafferty Vacupernius packeri (Allen) Thraulodes speciosus Traver Thraulodes zonalis Traver & Edmunds Sonora Tricorythodes fictus Traver Acentrella insignificans (McDunnough) Tricorythodes sordidus Allen Baetodes deficiens Cohen & Allen Baetodes edmundsi Koss San Luis Potosi Baetodes fuscipes Cohen & Allen Americabaetis pleturus (Lugo-Ortiz & McCafferty) Caenis bajaensis Allen & Murvosh Baetis magnus McCafferty & Waltz Callibaetis distinctus Lugo-Ortiz & McCafferty Caenis latipennis Banks Callibaetis pictus (Eaton) Camelobaetidius musseri (Traver & Edmunds) Callibaetis punctilusus McCafferty & Provonsha Campsurus cuspidatus Eaton Camelobaetidius mexicanus (Traver & Edmunds) Fallceon quilleri (Dodds) Camelobaetidius warreni (Traver & Edmunds) Farrodes reevesi Randolph & McCafferty Choroterpes inornata Eaton Hexagenia bilineata (Say) Ecdyonurus criddlei (McDunnough) Hexagenia limbata (Serville) Fallceon eatoni (Kimmins) Isonychia sicca (Walsh) Fallceon quilleri (Dodds) Lachlania iops Allen & Cohen Fallceon sonora (Allen & Murvosh) Leptohyphes zalope Traver Homoleptohyphes mirus (Allen) Neochoroterpes oklahoma (Traver) Isonychia sicca (Walsh) Paracloeodes minutus (Daggy) Leptohyphes apache Allen Thraulodes brunneus Koss Leptohyphes lestes Allen & Brusca Thraulodes pacaya McCafferty, Baumgardner & Leptohyphes musseri Allen Guenther Leptohyphes sabinus Traver W. P. MCCAFFERTY 325

Siphlonurus occidentalis (Eaton) Isonychia sicca (Walsh) Thraulodes brunneus Koss Leptohyphes apache Allen Thraulodes gonzalesi Traver & Edmunds Leptohyphes sabinas Traver Thraulodes speciosus Traver Leptohyphes zalope Traver Thraulodes tenulineus Lugo-Ortiz & McCafferty Maccaffertium mexicanum (Ulmer) Traverella albertana (McDunnough) Neochoroterpes oklahoma (Traver) Tricoryhyphes condylus (Allen) Plauditus virilis (McDunnough) Tricoryhyphes mulaiki (Traver) Thraulodes gonzalesi Traver & Edmunds Tricorythodes explicatus (Eaton) Thraulodes lunatus Traver & Edmunds Tricorythodes fictus Traver Thraulodes tenulineus Lugo-Ortiz & McCafferty Vacupernius packeri (Allen) Traverella presidiana (Traver) Tricorythodes sordidus Allen Tabasco Tricorythodes ulmeri Allen & Brusca Hydrosmilodon primanus (Eaton) Vacupernius packeri (Allen) Leptohyphes zalope Traver Maccaffertium mexicanum (Ulmer) Tlaxcala Thraulodes hilaris (Eaton) Callibaetis floridanus Banks Thraulodes pacaya McCafferty, Baumgardner & Guenther Vera Cruz Tortopsis primus (McDunnough) Americabaetis intermedius (Lugo-Ortiz & Traverella albertana (McDunnough) McCafferty) Americabaetis pleturus (Lugo-Ortiz & McCafferty Tamaulipas Baetis magnus McCafferty & Waltz Allenhyphes vescus (Allen) Baetis notos Alen & Murvosh Americabaetis pleturus (Lugo-Ortiz & McCafferty) Baetis tricaudatus Dodds Apobaetis etowah (Traver) Baetodes adustus Cohen & Allen Asioplax numinuh Wiersema, McCafferty & Baetodes caritus Cohen & Allen Baumgardner Baetodes deficiens Cohen & Allen Asioplax texana (Traver) Baetodes fortinensis Mayo Baetodes bibranchius McCafferty & Provonsha Baetodes fuscipes Cohen & Allen Baetodes inermis Cohen & Allen Baetodes inermis Cohen & Allen Baetodes longus Mayo Baetodes obesus Mayo Baetodes tritus Cohen & Allen Baetodes pictus Cohen & Allen Caenis latipennis Banks Baetodes tritus Cohen & Allen Callibaetis floridanus Banks Caenis anceps Traver Callibaetis pictus (Eaton) Callibaetis floridanus Banks Camelobaetidius mexicanus (Traver & Edmunds) Callibaetis pictus (Eaton) Camelobaetidius variabilis Wiersema Camelobaetidius arriaga (Traver & Edmunds) Campsurus decoloratus (Hagen) Camelobaetidius mexicanus (Traver & Edmunds) Choroterpes inornata Eaton Camelobaetidius musseri (Traver & Edmunds) Cloeodes excogitatus Waltz & McCafferty Camelobaetidius warreni (Traver & Edmunds) Cloeodes waltzi Wiersema & Baumgardner Campsurus cuspidatus Eaton Fallceon eatoni (Kimmins) Cloeodes excogitatus Waltz & McCafferty Fallceon fortipalpus Lugo-Ortiz & McCafferty Ecdyonurus bellus (Allen & Cohen) Fallceon quilleri (Dodds) Epeorus metlacensis Traver Farrodes otiesa Lugo-Ortiz & McCafferty Euthyplocia hecuba (Hagen) Farrodes texanus Davis Fallceon quilleri (Dodds) Hexagenia limbata (Serville) Farrodes flavipennis Domínguez, Molineri & Peters 326 MEXICAN AND CENTRAL AMERICAN EPHEMEROPTERA RECORDS

Farrodes mexicanus Domínguez Baumgardner, D. E. and W. P. McCafferty. 2010. Hexagenia albivitta (Walker) Revision of the genus Leptohyphes Eaton Hexagenia mexicana Eaton (Ephemeroptera: Leptohyphidae) in North and Hydrosmilodon primanus (Eaton) Central America. Zootaxa 2360: 1-33. Isonychia sicca (Walsh) Baumgardner, D. E., M. D. Meyer, and W. P. Leptohyphes apache Allen McCafferty. 2006. A new species of Asioplax Leptohyphes berneri Traver (Ephemeroptera: Leptohyphidae) from Costa Leptohyphes ferruginus Allen & Brusca Rica and Nicaragua. Pan-Pacific Entomologist Leptohyphes pilosus Allen & Brusca 82: 346-350. Leptohyphes sabinas Traver Lugo-Ortiz, C. R. and W. P. McCafferty. 1994. Leptohyphes zalope Traver New records of Ephemeroptera from Mexico. Moribaetis macaferti Waltz Entomological News 105: 17-26. Moribaetis salvini (Eaton) Lugo-Ortiz, C. R. and W. P. McCafferty. 1995a. Neochoroterpes oklahoma (Traver) New species, stage description, and records Rhithrogena notialis Allen & Cohen of Baetodes (Ephemeroptera: Baetidae) from Thraulodes brunneus Koss Mexico and Central America. Entomological Thraulodes eccentricus Lugo-Ortiz & McCafferty News 106: 81-86. Thraulodes pacaya McCafferty, Baumgardner & Lugo-Ortiz, C. R. and W. P. McCafferty Guenther 1995b. Contribution to the taxonomy of the Thraulodes spangleri Traver & Edmunds Leptohyphidae (Insecta: Ephemeroptera) of Thraulodes tenulineus Lugo-Ortiz& McCafferty Central America. Studies on Neotropical Fauna Traverella albertana (McDunnough) and Environment 30: 165-176. Vacupernius packeri (Allen) Lugo-Ortiz, C. R. and W. P. McCafferty. 1996a. New species of Leptophlebiidae Zacatecas (Ephemeroptera) from Mexico and Central Callibaetis pictus (Eaton) America. Annales de Limnologie 32: 3-18. Camelobaetidius mexicanus (Traver & Edmunds) Lugo-Ortiz, C. R. and W. P. McCafferty 1996b. Fallceon quilleri (Dodds) New Central American and Mexican records of Isonychia sicca (Walsh) Ephemeroptera species. Entomological News Neochoroterpes oklahoma (Traver) 107: 303-310. Thraulodes eccentricus Lugo-Ortiz& McCafferty Lugo-Ortiz, C. R. and W. P. McCafferty. 1996c. Thaulodes lunatus Traver & Edmunds Central American Tortopus (Ephemeroptera: Traverella albertana (McDunnough) Polymitarcyidae): a unique new species and new country records. Entomological News 107: 23- LITERATURE CITED 27. Allen, R. K. 1967. New species of New World McCafferty, W. P. 1975. The burrowing mayflies Leptohyphinae (Ephemeroptera: Tricorythidae). (Ephemeroptera: Ephemeroidea) of the Canadian Entomologist 99: 350-375. United States. Transactions of the American Allen, R. K. and R. C. Brusca. 1978. Generic Entomological Society 101: 447-504. revisions of mayfly nymphs. II. Thraulodes in McCafferty, W. P. 1985. New records of North and Central America (Leptophlebiidae). Ephemeroptera from Middle America. Canadian Entomologist 110: 413-433. International Quarterly of Entomology 1: 9-11. Baumgardner, D. E. and S. A. Ávila. 2006. McCafferty, W. P. 2011. Notable new North and Cabecar serratus, a new genus and species of Central American records of Ephemeroptera leptohyphid mayfly from Central America, and species. Transactions of the American description of the imagal stage of Tricorythodes Entomological Society 137: 1-10. sordidus Allen (Ephemeroptera: Leptohyphidae). McCafferty, W. P. and D. E. Baumgardner. 2003. Zootaxa 1187: 47-59. Lugoiops maya, a new genus and species W. P. MCCAFFERTY 327

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