A Trek Through the Wilds of the US Botanic Garden Conservatory!
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RI Equisetopsida and Lycopodiopsida.Indd
IIntroductionntroduction byby FFrancisrancis UnderwoodUnderwood Rhode Island Equisetopsida, Lycopodiopsida and Isoetopsida Special Th anks to the following for giving permission for the use their images. Robbin Moran New York Botanical Garden George Yatskievych and Ann Larson Missouri Botanical Garden Jan De Laet, plantsystematics.org Th is pdf is a companion publication to Rhode Island Equisetopsida, Lycopodiopsida & Isoetopsida at among-ri-wildfl owers.org Th e Elfi n Press 2016 Introduction Formerly known as fern allies, Horsetails, Club-mosses, Fir-mosses, Spike-mosses and Quillworts are plants that have an alternate generation life-cycle similar to ferns, having both sporophyte and gametophyte stages. Equisetopsida Horsetails date from the Devonian period (416 to 359 million years ago) in earth’s history where they were trees up to 110 feet in height and helped to form the coal deposits of the Carboniferous period. Only one genus has survived to modern times (Equisetum). Horsetails Horsetails (Equisetum) have jointed stems with whorls of thin narrow leaves. In the sporophyte stage, they have a sterile and fertile form. Th ey produce only one type of spore. While the gametophytes produced from the spores appear to be plentiful, the successful reproduction of the sporophyte form is low with most Horsetails reproducing vegetatively. Lycopodiopsida Lycopodiopsida includes the clubmosses (Dendrolycopodium, Diphasiastrum, Lycopodiella, Lycopodium , Spinulum) and Fir-mosses (Huperzia) Clubmosses Clubmosses are evergreen plants that produce only microspores that develop into a gametophyte capable of producing both sperm and egg cells. Club-mosses can produce the spores either in leaf axils or at the top of their stems. Th e spore capsules form in a cone-like structures (strobili) at the top of the plants. -
Ozzie Da Ros Pond Completed
NEWSLETTER • WINTER 2019 A LOTUSLAND LEGEND Ozzie Da Ros JAPANESE GARDEN RENOVATION Pond Completed LETTER FROM THE CHIEF EXECUTIVE OFFICER Dear Members and Friends, IN THIS FIRST ISSUE OF YOUR 2019 newsletter we share stories of the past and plans for the future. 695 Ashley Road We dedicate the lead article to Ozzie Da Ros, whose recent Santa Barbara, California 93108 passing signaled the end of an era. Ozzie was the last of 805.969.3767 • www.lotusland.org many talented craftsmen and expert plantsmen who worked 2019 BOARD OF TRUSTEES directly with Ganna Walska over several decades to make Daniel Bifano, President her dream of a “most outstanding center of horticultural Geoff Crane significance and educational use” become a reality, a top Lesley Cunningham Dorothy H. Gardner ten garden of the world. Fortunately, we were able to gather Anthony Grumbine written or oral memoirs from most of Walska’s collaborators Belle Hahn so their accounts may guide us as we tend the Garden now David M. Jones and in perpetuity. Joseph Marek Suzanne Mathews Nowhere is this commitment better exemplified than in the renovation of the Japanese Mimi Michaelis Garden. Over six years we conducted intensive research, peeling away the multiple historic Alexandra Morse Connie Pearcy layers of the garden to understand and preserve the significant aspects of each. We honor Eileen Rasmussen the garden and its makers even as we conscientiously add a new layer that addresses Stephen P. Schaible the modern obligations of an historic estate turned public garden. With the garden’s George Schoellkopf completion this year, all guests will have safe, comfortable access to the garden and new, Mick Thomas Caroline R. -
<I>Equisetum Giganteum</I>
Florida International University FIU Digital Commons FIU Electronic Theses and Dissertations University Graduate School 3-24-2009 Ecophysiology and Biomechanics of Equisetum Giganteum in South America Chad Eric Husby Florida International University, [email protected] DOI: 10.25148/etd.FI10022522 Follow this and additional works at: https://digitalcommons.fiu.edu/etd Recommended Citation Husby, Chad Eric, "Ecophysiology and Biomechanics of Equisetum Giganteum in South America" (2009). FIU Electronic Theses and Dissertations. 200. https://digitalcommons.fiu.edu/etd/200 This work is brought to you for free and open access by the University Graduate School at FIU Digital Commons. It has been accepted for inclusion in FIU Electronic Theses and Dissertations by an authorized administrator of FIU Digital Commons. For more information, please contact [email protected]. FLORIDA INTERNATIONAL UNIVERSITY Miami, Florida ECOPHYSIOLOGY AND BIOMECHANICS OF EQUISETUM GIGANTEUM IN SOUTH AMERICA A dissertation submitted in partial fulfillment of the requirements for the degree of DOCTOR OF PHILOSOPHY in BIOLOGY by Chad Eric Husby 2009 To: Dean Kenneth Furton choose the name of dean of your college/school College of Arts and Sciences choose the name of your college/school This dissertation, written by Chad Eric Husby, and entitled Ecophysiology and Biomechanics of Equisetum Giganteum in South America, having been approved in respect to style and intellectual content, is referred to you for judgment. We have read this dissertation and recommend that it be approved. _______________________________________ Bradley C. Bennett _______________________________________ Jack B. Fisher _______________________________________ David W. Lee _______________________________________ Leonel Da Silveira Lobo O'Reilly Sternberg _______________________________________ Steven F. Oberbauer, Major Professor Date of Defense: March 24, 2009 The dissertation of Chad Eric Husby is approved. -
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ISSN 2473-442X CONTENTS Message from Dr. Patrick Griffith, Co-chair, IUCN/SSC CSG 3 Official newsletter of IUCN/SSC Cycad Specialist Group Botanic Garden: In Focus Vol. IV I Issue 2 I December 2019 Montgomery Botanical Center’s Cycad Collection – Focus on research and conservation 5 Michael Calonje & Patrick Griffith Feature Articles Towards an approach for the conservation and illegal trade prevention of South Africa’s endangered Encephalartos spp. 10 James A. R. Clugston, Michelle Van Der Bankand Ronny M. Kobongo Fire is the most important threat for conservation of Dioon merolae (espadaña) in the hill Nambiyigua, municipality of Villaflores, Chiapas, Mexico 13 Miguel Angel Pérez-Farrera & Mauricio Martínez Martínez Ex-situ Cycad Conservation [1]: Public and Private Collections 16 Chip Jones & JS Khuraijam The Cycad Specialist Group (CSG) is a component of the IUCN Species Research and Conservation News Survival Commission (IUCN/SSC). It consists of a group of volunteer The Cycad Extinction Crisis in South Africa 19 experts addressing conservation Wynand van Eeden & Tim Gregory issues related to cycads, a highly What is Ceratozamia becerrae ? 21 threatened group of land plants. The Andrew P. Vovides, Miguel Angel Pérez-Farrera & José Said Gutiérrez-Ortega CSG exists to bring together the world’s cycad conservation expertise, Preliminary Finding: Seed longevity of Encephalartos in controlled storage 23 and to disseminate this expertise to Ngawethu Ngaka and Phakamani Xaba organizations and agencies which can use this guidance to advance cycad Meeting Reports conservation. 2nd Nong Nooch Cycad Horticulture Workshop 25 Official website of CSG: Anders Lindstrom http://www.cycadgroup.org/ Plant Conservation Genetics Workshop 26 Co-Chairs Caroline Iacuaniello, Stephanie Steele & Christy Powell John Donaldson Patrick Griffith CSG Members 28 Vice Chairs Michael Calonje Cristina Lopez-Gallego Red List Authority Coordinator De Wet Bosenberg CSG Newsletter Committee JS Khuraijam, Editor Irene Terry Andrew P. -
Scouring-Rush Horsetail Scientific Name: Equisetum Hyemale Order
Common Name: Scouring-rush Horsetail Scientific Name: Equisetum hyemale Order: Equisetales Family: Equisetaceae Wetland Plant Status: Facultative Ecology & Description Scouring-rush horsetail is an evergreen, perennial plant that completes a growing season in two years. At maturity, scouring-rush horsetail usually averages 3 feet in height but can be range anywhere from 2 to 5 feet. It can survive in a variety of environments. One single plant can spread 6 feet in diameter. It has cylindrical stems that averages a third of an inch in diameter. Noticeably spotted are the jointed unions that are located down the plant. The stems are hollow and don’t branch off into additional stems. Also, scouring- rush horsetail has rough ridges that run longitudinal along the stem. Although not covered in leaves, tiny leaves are joined together around the stem which then forms a black or green band, or sheath at each individual joint on the stem. This plant has an enormous root system that can reach 6 feet deep and propagates in two ways: rhizomes and spores. Incredibly, due to the fact that this plant is not full of leaves, it is forced to photosynthesize through the stem rather than leaves. Habitat Scouring-rush horsetail is highly tolerant of tough conditions. It can survive and thrive in full sun or part shade and can successfully grow in a variety of soil types. It can also grow in moderate to wet soils, and can survive in up to 4 inches of water. Distribution Scouring-rush horsetail can be found throughout the United States, Eurasia, and Canada. -
35 Ideal Landscape Cycads
3535 IdealIdeal LandscapeLandscape CycadsCycads Conserve Cycads by Growing Them -- Preservation Through Propagation Select Your Plant Based on these Features: Exposure: SunSun ShadeShade ☻☻ ColdCold☻☻ Filtered/CoastalFiltered/Coastal SunSun ▲▲ Leaf Length and Spread: Compact, Medium or Large? Growth Rate and Ultimate Plant Size Climate: Subtropical, Mediterranean, Temperate? Dry or Moist? Leaves -- Straight or Arching? Ocean-Loving, Salt-Tolerant, Wind-Tolerant CeratozamiaCeratozamiaCeratozamiaCeratozamia SpeciesSpeciesSpeciesSpecies ☻Shade Loving ☻Cold TolerTolerantant ▲Filtered/Coastal Sun 16 named + several undescribed species Native to Mexico, Guatemala & Belize Name originates from Greek ceratos (horned), and azaniae, (pine cone) Pinnate (feather-shaped) leaves, lacking a midrib, and horned, spiny cones Shiny, darker green leaves arching or upright, often emerging red or brown Less “formal” looking than other cycads Prefer Shade ½ - ¾ day, or afternoon shade Generally cold-tolerant CeratozamiaCeratozamia ---- SuggestedSuggested SpeciesSpecies ☻Shade Loving ☻Cold TolerTolerantant ▲Filtered/Coastal Sun Ceratozamia mexicana Tropical looking but cold-tolerant, native to dry mountainous areas in the Sierra Madre Mountains (Mexican Rockies). Landscape specimen works well with water features, due to arching habit. Prefers shade, modest height, with a spread of up to 10 feet. Trunk grows to 2 feet tall. Leaflets can be narrow or wider (0.75-2 inches). CeratozamiaCeratozamia ---- SuggestedSuggested SpeciesSpecies ☻Shade Loving ☻Cold TolerTolerantant ▲Filtered/Coastal Sun Ceratozamia latifolia Rare Ceratozamia named for its broad leaflets. Native to cloud forests of the Sierra Madre mountains of Mexico, underneath oak trees. Emergent trunk grows to 1 foot tall, 8 inches in diameter. New leaves emerge bronze, red or chocolate brown, hardening off to bright green, semiglossy, and grow to 6 feet long. They are flat lance-shaped, asymmetric, and are broadest above middle, growing to 10 inches long and 2 inches wide. -
The Timescale of Early Land Plant Evolution PNAS PLUS
The timescale of early land plant evolution PNAS PLUS Jennifer L. Morrisa,1, Mark N. Putticka,b,1, James W. Clarka, Dianne Edwardsc, Paul Kenrickb, Silvia Presseld, Charles H. Wellmane, Ziheng Yangf,g, Harald Schneidera,d,h,2, and Philip C. J. Donoghuea,2 aSchool of Earth Sciences, University of Bristol, Bristol BS8 1TQ, United Kingdom; bDepartment of Earth Sciences, Natural History Museum, London SW7 5BD, United Kingdom; cSchool of Earth and Ocean Sciences, Cardiff University, Cardiff CF10, United Kingdom; dDepartment of Life Sciences, Natural History Museum, London SW7 5BD, United Kingdom; eDepartment of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, United Kingdom; fDepartment of Genetics, Evolution and Environment, University College London, London WC1E 6BT, United Kingdom; gRadclie Institute for Advanced Studies, Harvard University, Cambridge, MA 02138; and hCenter of Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yunnan 666303, China Edited by Peter R. Crane, Oak Spring Garden Foundation, Upperville, VA, and approved January 17, 2018 (received for review November 10, 2017) Establishing the timescale of early land plant evolution is essential recourse but to molecular clock methodology, employing the for testing hypotheses on the coevolution of land plants and known fossil record to calibrate and constrain molecular evolu- Earth’s System. The sparseness of early land plant megafossils and tion to time. Unfortunately, the relationships among the four stratigraphic controls on their distribution make the fossil record principal lineages of land plants, namely, hornworts, liverworts, an unreliable guide, leaving only the molecular clock. However, mosses, and tracheophytes, are unresolved, with almost every the application of molecular clock methodology is challenged by possible solution currently considered viable (14). -
Subclase Equisetidae ¿Tienes Alguna Duda, Sugerencia O Corrección Acerca De Este Taxón? Envíanosla Y Con Gusto La Atenderemos
subclase Equisetidae ¿Tienes alguna duda, sugerencia o corrección acerca de este taxón? Envíanosla y con gusto la atenderemos. Ver todas las fotos etiquetadas con Equisetidae en Banco de Imagénes » Descripción de WIKIPEDIAES Ver en Wikipedia (español) → Ver Pteridophyta para una introducción a las plantas Equisetos vasculares sin semilla Rango temporal: Devónico-Holoceno PreЄ Є O S D C P T J K Pg N Los equisetos , llamados Equisetidae en la moderna clasificación de Christenhusz et al. 2011,[1] [2] [3] o también Equisetopsida o Equisetophyta, y en paleobotánica es más común Sphenopsida, son plantas vasculares afines a los helechos que aparecieron en el Devónico, pero que actualmente sobrevive únicamente el género Equisetum, si bien hay representantes de órdenes extintos que se verán en este artículo. Este grupo es monofilético, aun con sus representantes extintos, debido a su morfología distintiva. Son plantas pequeñas, aunque en el pasado una variedad de calamitácea alcanzó los 15 metros durante el pérmico.[4] Índice 1 Filogenia 1.1 Ecología y evolución 2 Taxonomía 2.1 Sinonimia Variedades de Equisetum 2.2 Sistema de Christenhusz et al. 2011 Taxonomía 2.3 Clasificación sensu Smith et al. 2006 2.4 Otras clasificaciones Reino: Plantae 3 Caracteres Viridiplantae 4 Véase también Streptophyta 5 Referencias Streptophytina 6 Bibliografía Embryophyta (sin rango) 7 Enlaces externos Tracheophyta Euphyllophyta Monilophyta Filogenia[editar] Equisetopsida o Sphenopsida Introducción teórica en Filogenia Clase: C.Agardh 1825 / Engler 1924 Equisetidae Los análisis moleculares y genéticos de filogenia solo Subclase: se pueden hacer sobre representantes vivientes, Warm. 1883 como circunscripto según Smith et al. (2006) (ver la Órdenes ficha), al menos Equisetales es monofilético (Pryer et Equisetales (DC. -
Vascular Plants of Williamson County Equisetum Hyemale L. Subsp. Affine
Vascular Plants of Williamson County Equisetum hyemale subsp. affine − COMMON SCOURING RUSH [Equisetaceae] Equisetum hyemale L. subsp. affine (Engelm.) Calder & Roy L. Taylor, COMMON SCOURING RUSH. Perennial herb, clonal, evergreen with photosynthetic stems, rhizomatous, fibrous-rooted, many-stemmed at base, stems jointed, unbranched above base (year 1), sometimes with 1−5 ascending branches at particular nodes (older stems), erect to ascending, in range 100–275 cm tall; shoots without foliage, each green stem persisting for 2 or more years and potentially with a terminal, spore-producing cone (in range most shoots sterile), glabrous; rhizomes deep-seated, horizontal and ascending, ± 7 mm diameter, dull black, in ×-section oblong and hollow (internodes) with a ring of 20+ canals. Stems: with 22−30(−44+) low ridges, 6−8(−12) mm diameter at base, straight, tough, slightly constricted at nodes, each segment green to blue-green with a broad black band on lower internode underneath sheath of older stem segment, internodes typically 15−28 per shoot and 25−160 mm long, scabrous due to silica projections (transverse ridges) in cell walls, with 2 longitudinal lines of sunken stomates within each stem valley; internode hollow, mature wall 0.5–2 mm thick, in ×-section with a canal beneath each ridge; lateral branches whorled but not symmetrically so, ± 16-ridged, ca. 3 mm diameter. Leaves: as many as stem ridges, whorled and fused forming sheath around node with scalelike free portions; sheath cylindric, (7−)10−13(−17) mm long, diameter < length, initially pale yellow green above node, with a black band formed by upper sheath and lobes, in age forming a second black band near midpoint between base and tip separated by an ash-gray band; free portion of leaf flat needle-shaped, 0.5−1 mm wide, soon dead and brownish, 1- veined, mostly abscised; free portions on microphylls of lateral branches 0.5 mm wide, black base to tip. -
PG and Research Deparment of Botany Pteridophytes, Gymnosperms and Paleobotany
PG and Research Deparment of Botany Pteridophytes, Gymnosperms and Paleobotany Part-A 1. The vascular tissue is confined to the central region of the stem forming: (a) Bundles (b) stele (c) Cortex (d) Pericycle 2. The leaves which bear the sporangia are called: (a) sporophylIs (b) Bract (c) Cone (d) Strobilus 3. One or two peripheral layers persist for the nourishment of the developing spores. These nourishing cells form: (a) Elators (b) Sopores (c) Jacket (d) tapetum. 4. Match gametophyte with one of the followings: (a) Prothallus (b) Thallus (c) Cone (d) Strobilus 5. Selaginella belongs to division (a) Lycopsida (b) Pteropsida (c) Psilopsida (d) Sphenopsida 6. Hone tails are: (a) Lycopsida (b) Pteropsida (c ) Psilopsida (d) Sphenopsida 7. Marsilea belongs: (a) Lycopsida (b) Pteropsida (c) Psilopsida (d) Sphenopsida 8. Which of the followings is fern? (a) Psilopsida (b) Pteropsida (c) Lycopsida (d) Sphenopsida 9. Which of the followings is most primitive division? (a) Lycopsida (b) Pteropsida (c) Psilopsida (d) Sphenopsida 10. Club mosses are: (a) Lycopsida (b) Pteropsida (c) Psilopsida (d) Sphenopsida 11. The protostele in which xylem core is Smooth and rounded is: (a) Haplostele (b) Actinostlele (c) Plectostele (d) Siphonostele 12. The protostele in which xylem core is star like is called: (a) Haplostele (b) Actinostlele (c) Plectostele (d) Siphonostele 13. The siphonostele in–which two cylinders of vascular tissue are present in the stele is: (a) Haplostele (b) Actinostlele (c) Plectostele (d) Polycyclic 14. In Xylem in which protoxylem is lying in the middle of Metaxylem is: (a) Exarch (b) Mesarch (c) Endarch (d) Diarch 15. -
Diversity and Evolution of the Megaphyll in Euphyllophytes
G Model PALEVO-665; No. of Pages 16 ARTICLE IN PRESS C. R. Palevol xxx (2012) xxx–xxx Contents lists available at SciVerse ScienceDirect Comptes Rendus Palevol w ww.sciencedirect.com General palaeontology, systematics and evolution (Palaeobotany) Diversity and evolution of the megaphyll in Euphyllophytes: Phylogenetic hypotheses and the problem of foliar organ definition Diversité et évolution de la mégaphylle chez les Euphyllophytes : hypothèses phylogénétiques et le problème de la définition de l’organe foliaire ∗ Adèle Corvez , Véronique Barriel , Jean-Yves Dubuisson UMR 7207 CNRS-MNHN-UPMC, centre de recherches en paléobiodiversité et paléoenvironnements, 57, rue Cuvier, CP 48, 75005 Paris, France a r t i c l e i n f o a b s t r a c t Article history: Recent paleobotanical studies suggest that megaphylls evolved several times in land plant st Received 1 February 2012 evolution, implying that behind the single word “megaphyll” are hidden very differ- Accepted after revision 23 May 2012 ent notions and concepts. We therefore review current knowledge about diverse foliar Available online xxx organs and related characters observed in fossil and living plants, using one phylogenetic hypothesis to infer their origins and evolution. Four foliar organs and one lateral axis are Presented by Philippe Taquet described in detail and differ by the different combination of four main characters: lateral organ symmetry, abdaxity, planation and webbing. Phylogenetic analyses show that the Keywords: “true” megaphyll appeared at least twice in Euphyllophytes, and that the history of the Euphyllophytes Megaphyll four main characters is different in each case. The current definition of the megaphyll is questioned; we propose a clear and accurate terminology in order to remove ambiguities Bilateral symmetry Abdaxity of the current vocabulary. -
Horsetail-Scouring Rush (Equisetum Spp.) Description: Horsetail Is in a Family of Plants Containing About 35 Species Within the Genus Equisetum
INTEGRATED PEST MANAGEMENT PRESCRIPTION Horsetail-Scouring rush (Equisetum spp.) Description: Horsetail is in a family of plants containing about 35 species within the genus Equisetum. More than 230 million years ago, the horsetail family was one of the most dominant plants in the world, with some species reaching 100 feet tall. The remains of these giants contributed greatly to the formation of coal beds around most of the world. Horsetail species are natives nearly everywhere, the few exceptions being Australia, New Zealand, and Antarctica. They are found in great abundance in the Pacific Northwest, which is home to nearly half the world’s species. Three common Equisetum species in Thurston County are: ▪ Field horsetail (Equisetum arvense) is a perennial with a spreading rhizome system that produces numerous shoots and tubers. It has two types of stems; the leafless, fertile, cone bearing stems that grow 6 to 12 inches tall, and the sterile, hollow, vegetative stems that grow to 2 feet tall with whorls of branches growing form banded joints. Fertile stems are few in number, tan colored, and die back shortly after the pale green spores are shed. The vegetative stems all die back in the fall. ▪ Giant horsetail (Equisetum telmateia) is very similar to field horsetail only taller, more robust, and produces an abundance of fertile stems. ▪ Scouring rush (Equisetum hyemale) is evergreen, with leafless, hollow, segmented stems with ash-colored bands. Stems grow to about 1/2 inch in diameter and reach up to five feet tall. They grow from aggressively spreading rhizomes and each stem is fertile, having small rounded cones containing reproductive spores at the tips.