PG and Research Deparment of Botany Pteridophytes, Gymnosperms and Paleobotany
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RI Equisetopsida and Lycopodiopsida.Indd
IIntroductionntroduction byby FFrancisrancis UnderwoodUnderwood Rhode Island Equisetopsida, Lycopodiopsida and Isoetopsida Special Th anks to the following for giving permission for the use their images. Robbin Moran New York Botanical Garden George Yatskievych and Ann Larson Missouri Botanical Garden Jan De Laet, plantsystematics.org Th is pdf is a companion publication to Rhode Island Equisetopsida, Lycopodiopsida & Isoetopsida at among-ri-wildfl owers.org Th e Elfi n Press 2016 Introduction Formerly known as fern allies, Horsetails, Club-mosses, Fir-mosses, Spike-mosses and Quillworts are plants that have an alternate generation life-cycle similar to ferns, having both sporophyte and gametophyte stages. Equisetopsida Horsetails date from the Devonian period (416 to 359 million years ago) in earth’s history where they were trees up to 110 feet in height and helped to form the coal deposits of the Carboniferous period. Only one genus has survived to modern times (Equisetum). Horsetails Horsetails (Equisetum) have jointed stems with whorls of thin narrow leaves. In the sporophyte stage, they have a sterile and fertile form. Th ey produce only one type of spore. While the gametophytes produced from the spores appear to be plentiful, the successful reproduction of the sporophyte form is low with most Horsetails reproducing vegetatively. Lycopodiopsida Lycopodiopsida includes the clubmosses (Dendrolycopodium, Diphasiastrum, Lycopodiella, Lycopodium , Spinulum) and Fir-mosses (Huperzia) Clubmosses Clubmosses are evergreen plants that produce only microspores that develop into a gametophyte capable of producing both sperm and egg cells. Club-mosses can produce the spores either in leaf axils or at the top of their stems. Th e spore capsules form in a cone-like structures (strobili) at the top of the plants. -
<I>Equisetum Giganteum</I>
Florida International University FIU Digital Commons FIU Electronic Theses and Dissertations University Graduate School 3-24-2009 Ecophysiology and Biomechanics of Equisetum Giganteum in South America Chad Eric Husby Florida International University, [email protected] DOI: 10.25148/etd.FI10022522 Follow this and additional works at: https://digitalcommons.fiu.edu/etd Recommended Citation Husby, Chad Eric, "Ecophysiology and Biomechanics of Equisetum Giganteum in South America" (2009). FIU Electronic Theses and Dissertations. 200. https://digitalcommons.fiu.edu/etd/200 This work is brought to you for free and open access by the University Graduate School at FIU Digital Commons. It has been accepted for inclusion in FIU Electronic Theses and Dissertations by an authorized administrator of FIU Digital Commons. For more information, please contact [email protected]. FLORIDA INTERNATIONAL UNIVERSITY Miami, Florida ECOPHYSIOLOGY AND BIOMECHANICS OF EQUISETUM GIGANTEUM IN SOUTH AMERICA A dissertation submitted in partial fulfillment of the requirements for the degree of DOCTOR OF PHILOSOPHY in BIOLOGY by Chad Eric Husby 2009 To: Dean Kenneth Furton choose the name of dean of your college/school College of Arts and Sciences choose the name of your college/school This dissertation, written by Chad Eric Husby, and entitled Ecophysiology and Biomechanics of Equisetum Giganteum in South America, having been approved in respect to style and intellectual content, is referred to you for judgment. We have read this dissertation and recommend that it be approved. _______________________________________ Bradley C. Bennett _______________________________________ Jack B. Fisher _______________________________________ David W. Lee _______________________________________ Leonel Da Silveira Lobo O'Reilly Sternberg _______________________________________ Steven F. Oberbauer, Major Professor Date of Defense: March 24, 2009 The dissertation of Chad Eric Husby is approved. -
Further Interpretation of Wodehouseia Spinata Stanley from the Late Maastrichtian of the Far East (China) M
ISSN 0031-0301, Paleontological Journal, 2019, Vol. 53, No. 2, pp. 203–213. © Pleiades Publishing, Ltd., 2019. Russian Text © M.V. Tekleva, S.V. Polevova, E.V. Bugdaeva, V.S. Markevich, Sun Ge, 2019, published in Paleontologicheskii Zhurnal, 2019, No. 2, pp. 94–105. Further Interpretation of Wodehouseia spinata Stanley from the Late Maastrichtian of the Far East (China) M. V. Teklevaa, *, S. V. Polevovab, E. V. Bugdaevac, V. S. Markevichc, and Sun Ged aBorissiak Paleontological Institute, Russian Academy of Sciences, Moscow, 117647 Russia bMoscow State University, Moscow, 119991 Russia cFederal Scientific Center of the East Asia Terrestrial Biodiversity, Vladivostok, 690022 Russia dCollege of Paleontology, Shenyang Normal University, Shenyang, Liaoning Province, China *e-mail: [email protected] Received May 25, 2018; revised September 30, 2018; accepted October 15, 2018 Abstract—Dispersed pollen grains Wodehouseia spinata Stanley of unknown botanical affinity from the Maastrichtian of the Amur River Region, Far East are studied using transmitted light, scanning and trans- mission electron microscopy. The pollen was probably produced by wetland or aquatic plants, adapted to a sudden change in the water regime during the vegetation season. The pattern of the exine sculpture and spo- roderm ultrastructure suggests that insects contributed to pollination. The flange and unevenly thickened endexine could facilitate harmomegathy. A tetragonal or rhomboidal tetrad type seems to be most logical for Wodehouseia pollen. The infratectum structure suggests that Wodehouseia should be placed within an advanced group of eudicots. Keywords: Wodehouseia, exine morphology, sporoderm ultrastructure, “oculata” group, Maastrichtian DOI: 10.1134/S0031030119020126 INTRODUCTION that has lost its flange (see a review Wiggins, 1976). -
Morphological Features of the Anther Development in Tomato Plants with Non-Specific Male Sterility
biology Article Morphological Features of the Anther Development in Tomato Plants with Non-Specific Male Sterility Inna A. Chaban 1, Neonila V. Kononenko 1 , Alexander A. Gulevich 1, Liliya R. Bogoutdinova 1, Marat R. Khaliluev 1,2,* and Ekaterina N. Baranova 1,* 1 All-Russia Research Institute of Agricultural Biotechnology, Timiryazevskaya 42, 127550 Moscow, Russia; [email protected] (I.A.C.); [email protected] (N.V.K.); [email protected] (A.A.G.); [email protected] (L.R.B.) 2 Moscow Timiryazev Agricultural Academy, Agronomy and Biotechnology Faculty, Russian State Agrarian University, Timiryazevskaya 49, 127550 Moscow, Russia * Correspondence: [email protected] (M.R.K.); [email protected] (E.N.B.) Received: 3 January 2020; Accepted: 12 February 2020; Published: 17 February 2020 Abstract: The study was devoted to morphological and cytoembryological analysis of disorders in the anther and pollen development of transgenic tomato plants with a normal and abnormal phenotype, which is characterized by the impaired development of generative organs. Various abnormalities in the structural organization of anthers and microspores were revealed. Such abnormalities in microspores lead to the blocking of asymmetric cell division and, accordingly, the male gametophyte formation. Some of the non-degenerated microspores accumulate a large number of storage inclusions, forming sterile mononuclear pseudo-pollen, which is similar in size and appearance to fertile pollen grain (looks like pollen grain). It was discussed that the growth of tapetal cells in abnormal anthers by increasing the size and ploidy level of nuclei contributes to this process. It has been shown that in transgenic plants with a normal phenotype, individual disturbances are also observed in the development of both male and female gametophytes. -
Lichens of Alaska's South Coast
United States Department of Agriculture Lichens of Alaska’s South Coast Forest Service R10-RG-190 Alaska Region Reprint April 2014 WHAT IS A LICHEN? Lichens are specialized fungi that “farm” algae as a food source. Unlike molds, mildews, and mushrooms that parasitize or scavenge food from other organisms, the fungus of a lichen cultivates tiny algae and / or blue-green bacteria (called cyanobacteria) within the fabric of interwoven fungal threads that form the body of the lichen (or thallus). The algae and cyanobacteria produce food for themselves and for the fungus by converting carbon dioxide and water into sugars using the sun’s energy (photosynthesis). Thus, a lichen is a combination of two or sometimes three organisms living together. Perhaps the most important contribution of the fungus is to provide a protective habitat for the algae or cyanobacteria. The green or blue-green photosynthetic layer is often visible between two white fungal layers if a piece of lichen thallus is torn off. Most lichen-forming fungi cannot exist without the photosynthetic partner because they have become dependent on them for survival. But in all cases, a fungus looks quite different in the lichenized form compared to its free-living form. HOW DO LICHENS REPRODUCE? Lichens sexually reproduce with fruiting bodies of various shapes and colors that can often look like miniature mushrooms. These are called apothecia (Fig. 1) and contain spores that germinate and Figure 1. Apothecia, fruiting grow into the fungus. Each bodies fungus must find the right photosynthetic partner in order to become a lichen. Lichens reproduce asexually in several ways. -
Ordovician Land Plants and Fungi from Douglas Dam, Tennessee
PROOF The Palaeobotanist 68(2019): 1–33 The Palaeobotanist 68(2019): xxx–xxx 0031–0174/2019 0031–0174/2019 Ordovician land plants and fungi from Douglas Dam, Tennessee GREGORY J. RETALLACK Department of Earth Sciences, University of Oregon, Eugene, OR 97403, USA. *Email: gregr@uoregon. edu (Received 09 September, 2019; revised version accepted 15 December, 2019) ABSTRACT The Palaeobotanist 68(1–2): Retallack GJ 2019. Ordovician land plants and fungi from Douglas Dam, Tennessee. The Palaeobotanist 68(1–2): xxx–xxx. 1–33. Ordovician land plants have long been suspected from indirect evidence of fossil spores, plant fragments, carbon isotopic studies, and paleosols, but now can be visualized from plant compressions in a Middle Ordovician (Darriwilian or 460 Ma) sinkhole at Douglas Dam, Tennessee, U. S. A. Five bryophyte clades and two fungal clades are represented: hornwort (Casterlorum crispum, new form genus and species), liverwort (Cestites mirabilis Caster & Brooks), balloonwort (Janegraya sibylla, new form genus and species), peat moss (Dollyphyton boucotii, new form genus and species), harsh moss (Edwardsiphyton ovatum, new form genus and species), endomycorrhiza (Palaeoglomus strotheri, new species) and lichen (Prototaxites honeggeri, new species). The Douglas Dam Lagerstätte is a benchmark assemblage of early plants and fungi on land. Ordovician plant diversity now supports the idea that life on land had increased terrestrial weathering to induce the Great Ordovician Biodiversification Event in the sea and latest Ordovician (Hirnantian) -
Scouring-Rush Horsetail Scientific Name: Equisetum Hyemale Order
Common Name: Scouring-rush Horsetail Scientific Name: Equisetum hyemale Order: Equisetales Family: Equisetaceae Wetland Plant Status: Facultative Ecology & Description Scouring-rush horsetail is an evergreen, perennial plant that completes a growing season in two years. At maturity, scouring-rush horsetail usually averages 3 feet in height but can be range anywhere from 2 to 5 feet. It can survive in a variety of environments. One single plant can spread 6 feet in diameter. It has cylindrical stems that averages a third of an inch in diameter. Noticeably spotted are the jointed unions that are located down the plant. The stems are hollow and don’t branch off into additional stems. Also, scouring- rush horsetail has rough ridges that run longitudinal along the stem. Although not covered in leaves, tiny leaves are joined together around the stem which then forms a black or green band, or sheath at each individual joint on the stem. This plant has an enormous root system that can reach 6 feet deep and propagates in two ways: rhizomes and spores. Incredibly, due to the fact that this plant is not full of leaves, it is forced to photosynthesize through the stem rather than leaves. Habitat Scouring-rush horsetail is highly tolerant of tough conditions. It can survive and thrive in full sun or part shade and can successfully grow in a variety of soil types. It can also grow in moderate to wet soils, and can survive in up to 4 inches of water. Distribution Scouring-rush horsetail can be found throughout the United States, Eurasia, and Canada. -
Seed Plant Phylogeny: Demise of the Anthophyte Hypothesis? Michael J
bb10c06.qxd 02/29/2000 04:18 Page R106 R106 Dispatch Seed plant phylogeny: Demise of the anthophyte hypothesis? Michael J. Donoghue* and James A. Doyle† Recent molecular phylogenetic studies indicate, The first suggestions that Gnetales are related to surprisingly, that Gnetales are related to conifers, or angiosperms were based on several obvious morphological even derived from them, and that no other extant seed similarities — vessels in the wood, net-veined leaves in plants are closely related to angiosperms. Are these Gnetum, and reproductive organs made up of simple, results believable? Is this a clash between molecules unisexual, flower-like structures, which some considered and morphology? evolutionary precursors of the flowers of wind-pollinated Amentiferae, but others viewed as being reduced from Addresses: *Harvard University Herbaria, 22 Divinity Avenue, Cambridge, Massachusetts 02138, USA. †Section of Evolution and more complex flowers in the common ancestor of Ecology, University of California, Davis, California 95616, USA. angiosperms, Gnetales and Mesozoic Bennettitales [1]. E-mail: [email protected] These ideas went into eclipse with evidence that simple [email protected] flowers really are a derived, rather than primitive, feature Current Biology 2000, 10:R106–R109 of the Amentiferae, and that vessels arose independently in angiosperms and Gnetales. Vessels in angiosperms 0960-9822/00/$ – see front matter seem derived from tracheids with scalariform pits, whereas © 2000 Elsevier Science Ltd. All rights reserved. in Gnetales they resemble tracheids with circular bor- dered pits, as in conifers. Gnetales are also like conifers in These are exciting times for those interested in plant lacking scalariform pitting in the primary xylem, and in evolution. -
The Timescale of Early Land Plant Evolution PNAS PLUS
The timescale of early land plant evolution PNAS PLUS Jennifer L. Morrisa,1, Mark N. Putticka,b,1, James W. Clarka, Dianne Edwardsc, Paul Kenrickb, Silvia Presseld, Charles H. Wellmane, Ziheng Yangf,g, Harald Schneidera,d,h,2, and Philip C. J. Donoghuea,2 aSchool of Earth Sciences, University of Bristol, Bristol BS8 1TQ, United Kingdom; bDepartment of Earth Sciences, Natural History Museum, London SW7 5BD, United Kingdom; cSchool of Earth and Ocean Sciences, Cardiff University, Cardiff CF10, United Kingdom; dDepartment of Life Sciences, Natural History Museum, London SW7 5BD, United Kingdom; eDepartment of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, United Kingdom; fDepartment of Genetics, Evolution and Environment, University College London, London WC1E 6BT, United Kingdom; gRadclie Institute for Advanced Studies, Harvard University, Cambridge, MA 02138; and hCenter of Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Yunnan 666303, China Edited by Peter R. Crane, Oak Spring Garden Foundation, Upperville, VA, and approved January 17, 2018 (received for review November 10, 2017) Establishing the timescale of early land plant evolution is essential recourse but to molecular clock methodology, employing the for testing hypotheses on the coevolution of land plants and known fossil record to calibrate and constrain molecular evolu- Earth’s System. The sparseness of early land plant megafossils and tion to time. Unfortunately, the relationships among the four stratigraphic controls on their distribution make the fossil record principal lineages of land plants, namely, hornworts, liverworts, an unreliable guide, leaving only the molecular clock. However, mosses, and tracheophytes, are unresolved, with almost every the application of molecular clock methodology is challenged by possible solution currently considered viable (14). -
On the Evolution of Angiosperms in the Himalayan Region: a Summary
The Palaeobotanist 57(2008) : 453-457 0031-0174/2008 $2.00 On the evolution of Angiosperms in the Himalayan region: A summary J.S. GULERIA Birbal Sahni Institute of Palaeobotany, 53 University Road, Lucknow 226007, India. [email protected] (Received 10 June, 2007; revised version accepted 15 September, 2008) ABSTRACT Guleria JS 2008. On the evolution of Angiosperms in the Himalayan region: A summary. The Palaeobotanist 57(3): 453-457. The paper summarises the evolution of angiosperms in different zones of Himalaya. The Himalayan Cenozoic flora has been divided age-wise as Palaeogene and Neogene flora. The Himalayan Palaeogene flora is largely a continuation of tropical peninsular flora of India. The early Miocene flora of Lesser Himalaya is also moist tropical. However, temperate plants started appearing during Miocene in the Higher Himalaya and their occurrence in Plio-Pleistocene flora of Kashmir reflect uplift of the Himalaya. The sub-Himalayan flora indicates existence of warm humid conditions in this belt which became drier by the end of Pliocene. The northern floral elements appeared to have invaded India all along the Himalayan belt. Since its birth the Himalaya has played a significant role in the immigration of plants from the adjoining regions, i.e. east, west and north, thereby enriching the Indian flora. The development of the Cenozoic flora of the Himalayan region is an expression of changing patterns of geography, topography and climate. Key-words—Cenozoic, Angiosperms, Himalayan flora, Migration, Climate (India). fgeky;h -
Subclase Equisetidae ¿Tienes Alguna Duda, Sugerencia O Corrección Acerca De Este Taxón? Envíanosla Y Con Gusto La Atenderemos
subclase Equisetidae ¿Tienes alguna duda, sugerencia o corrección acerca de este taxón? Envíanosla y con gusto la atenderemos. Ver todas las fotos etiquetadas con Equisetidae en Banco de Imagénes » Descripción de WIKIPEDIAES Ver en Wikipedia (español) → Ver Pteridophyta para una introducción a las plantas Equisetos vasculares sin semilla Rango temporal: Devónico-Holoceno PreЄ Є O S D C P T J K Pg N Los equisetos , llamados Equisetidae en la moderna clasificación de Christenhusz et al. 2011,[1] [2] [3] o también Equisetopsida o Equisetophyta, y en paleobotánica es más común Sphenopsida, son plantas vasculares afines a los helechos que aparecieron en el Devónico, pero que actualmente sobrevive únicamente el género Equisetum, si bien hay representantes de órdenes extintos que se verán en este artículo. Este grupo es monofilético, aun con sus representantes extintos, debido a su morfología distintiva. Son plantas pequeñas, aunque en el pasado una variedad de calamitácea alcanzó los 15 metros durante el pérmico.[4] Índice 1 Filogenia 1.1 Ecología y evolución 2 Taxonomía 2.1 Sinonimia Variedades de Equisetum 2.2 Sistema de Christenhusz et al. 2011 Taxonomía 2.3 Clasificación sensu Smith et al. 2006 2.4 Otras clasificaciones Reino: Plantae 3 Caracteres Viridiplantae 4 Véase también Streptophyta 5 Referencias Streptophytina 6 Bibliografía Embryophyta (sin rango) 7 Enlaces externos Tracheophyta Euphyllophyta Monilophyta Filogenia[editar] Equisetopsida o Sphenopsida Introducción teórica en Filogenia Clase: C.Agardh 1825 / Engler 1924 Equisetidae Los análisis moleculares y genéticos de filogenia solo Subclase: se pueden hacer sobre representantes vivientes, Warm. 1883 como circunscripto según Smith et al. (2006) (ver la Órdenes ficha), al menos Equisetales es monofilético (Pryer et Equisetales (DC. -
Characteristics of Fungi 1. Thallus
CHARACTERISTICS OF FUNGI 1. THALLUS ORGANIZATION Except some unicellular forms (e.g. yeasts, Synchytrium). The fungal body is a thallus called mycelium. The mycelium is an interwoven mass of thread-like hyphae (Sing., hypha). Hyphae may be septate (with cross wall) and aseptate (without cross wall). Some fungi are dimorphic that found as both unicellular and mycelial forms e.g. Candida albicans. 1 2. DIFFERENT FORMS OF MYCELIUM (a) Plectenchyma (fungal tissue): In a fungal mycelium, hyphae organized loosely or compactly woven to form a tissue called plectenchyma. It is two types: i. Prosenchyma or Prosoplectenchyma: In these fungal tissue hyphae are loosely interwoven lying more or less parallel to each other. ii. Pseudoparenchyma or paraplectenchyma: In these fungal tissue hyphae are compactly interwoven looking like a parenchyma in cross-section. (b) Sclerotia (Gr. Skleros=haid): These are hard dormant bodies consist of compact hyphae protected by external thickened hyphae. Each Sclerotium germinates into a mycelium, on return of favourable condition, e.g., Penicillium. 2 (c) Rhizomorphs: They are root-like compactly interwoven hyphae with distinct growing tip. They help in absorption and perennation (to tide over the unfavourable periods), e.g., Armillaria mellea. 3. NUTRITION The fungi lack chlorophyll. Therefore, they cannot synthesize their own food. Depending on from where and how they get nutrition, fungi are of following types: (a) Saprotrophs (= saprobes): They obtain food from dead and decaying organic matter. They secrete digesting enzymes to outside which digest the substratum and then absorb nutrients, e.g., Mucor, Rhizopus (bread mould) etc. (b) Parasitic: They obtain food from living.