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Review of adventive of Coleoptera (Insecta) recorded from eastern Canada

Article · June 2010

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Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada

by

Jan Klimaszewski, David Langor, Christopher G. Majka, Patrice Bouchard, Yves Bousquet, Laurent LeSage, Aleš Smetana, Patricia Sylvestre, Georges Pelletier, Anthony Davies, Pierre DesRochers, Henri Goulet, Reginald Webster & Jon Sweeney

Sofia–Moscow 2010 Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada by Jan Klimaszewski 1, David Langor 2, Christopher G. Majka 4, Patrice Bouchard 5, Yves Bousquet 5, Laurent LeSage 5, Aleš Smetana 5, Patricia Sylvestre 1, Georges Pelletier 1, Anthony Davies 5, Pierre DesRochers 1, Henri Goulet 5, Reginald Webster 3 & Jon Sweeney 3

1) Natural Resources Canada, Canadian Forest Service, Laurentian Forestry Centre, 1055 du P.E.P.S., P.O. Box 10380, Stn. Sainte-Foy, Quebec, Quebec, Canada G1V 4C7 2) Natural Resources Canada, Canadian Forest Service, Northern Forestry Centre, 5320-122 Street, Edmonton, Alberta, Canada T6H 3S5 3) Natural Resources Canada, Canadian Forest Service, Atlantic Forestry Centre, 1350 Regent Street, P.O. Box 4000, Fredericton, New Brunswick, Canada E3B 5P7 4) Natural History Resources, 6252 Jubilee Road, Halifax, Nova Scotia Canada B3H 2G5 5) Agriculture and Agri-Food Canada, Canadian National Collection of , Arachnids and Nematodes, Ottawa, Ontario, Canada K1A 0C6

First published 2010 ISBN 978-954-642-552-2 (hardback)

Pensoft Publishers Geo Milev Str. 13a, Sofi a 1111, Bulgaria Fax: +359-2-870-42-82 [email protected] www.pensoft.net

Printed in Bulgaria, April 2010 This book is dedicated to the 50th anniversary of research conducted at the Laurentian Forestry Centre, Quebec City, Quebec, Canada

Contents

Introduction 9

Material and methods 15

Results 17

Checklist of taxa with adventive species in eastern Canada (in taxonomic order) 21

Key to families of Coleoptera with adventive species recorded from eastern Canada 25

Review of families, adventive species and literature 37

Acknowledgements 192

References 193

Index of families, genera and species listed in the text 229

Plates 241

Review of adventive species of Coleoptera (Insecta) recorded from eastern Canada 9

Introduction

For as long as entomologists have turned their attention to Coleoptera in eastern Can- ada, the subject of adventive species has been a topic of considerable interest. In the fi rst paper about Coleoptera from eastern Canada, Kirby (1837) listed eleven adven- tive species including Pterostichus melanarius (Illiger), Philonthus politus (Linnaeus), Philonthus ventralis (Gravenhorst), pellio L