Lillig 2015 Zoogeographie Westpaläarktischer Tenebrionidae.Pdf

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Lillig 2015 Zoogeographie Westpaläarktischer Tenebrionidae.Pdf Zur Zoogeographie westpaläarktischer Tenebrionidae (Insecta: Coleoptera) Inauguraldissertation zur Erlangung der Würde eines Doktors der Philosophie vorgelegt der Philosophisch-Naturwissenschaftlichen Fakultät der Universität Basel von Martin LILLIG aus Saarbrücken (Deutschland) Basel, 2015 Originaldokument gespeichert auf dem Dokumentenserver der Universität Basel edoc.unibas.ch Dieses Werk ist lizenziert unter einer Creative Commons Namensnennung-Nicht kommerziell 4.0 International Lizenz. Genehmigt von der Philosophisch-Naturwissenschaftlichen Fakultät auf Antrag von Prof. Dr. Peter Nagel (Fakultätsvertreter) und Prof. Dr. Thomas Wagner. Basel, den 08.12.2015 Prof. Dr. Jörg Schibler Dekan res severa verum gaudium Seneca Foto Umschlagseite: Helops rossii GERMAR, 1817 (Türkei) (Foto: M. Lillig) Inhalt Zusammenfassung ..............................................................................................................................6 Summary ............................................................................................................................................9 1. Einleitung .................................................................................................................................. 12 1.1 Systematik ......................................................................................................................... 13 1.2 Ökologie ............................................................................................................................ 16 1.3 Biodiversität ...................................................................................................................... 17 1.4 Faunistik und Zoogeographie ............................................................................................. 18 1.5 Nachweismethoden ........................................................................................................... 20 1.6 Offene Fragen und Ziel der Dissertation ............................................................................. 22 1.7 Literatur ............................................................................................................................ 24 2. Zur Geschichte der Erforschung der Tenebrionidae ................................................................... 29 2.1 Die Werke verstorbener Coleopterologen.......................................................................... 30 2.2 Frauen und Tenebrionidae ................................................................................................. 40 2.3 Nach 2000 aktive Tenebrionidologen ................................................................................. 44 2.4 Literatur ............................................................................................................................ 50 3. Einzelbeschreibungen ............................................................................................................... 69 3.1 Zwei neue Arten der Gattung Erodius ................................................................................ 73 3.2 Astorthocnemis becvarorum LILLIG & PAVLÍČEK, 2002 nov. gen. nov. spec. ............................ 81 3.3 Cryphaeus laticeps LILLIG, 2006 ........................................................................................... 90 3.4 Hedyphanes (Microhedyphanes) chikatunovi NABOZHENKO & LILLIG, 2013 ............................ 98 3.5 Zusammenfassung ........................................................................................................... 107 4. Faunistik ................................................................................................................................. 108 4.1 Schwarzkäfer des Saarlandes ........................................................................................... 114 4.2 Tenebrionidae von Malta ................................................................................................. 157 4.3 Zusammenfassung ........................................................................................................... 211 5. Revisionen .............................................................................................................................. 212 5.1 Nomenklatorische Änderungen bei Adesmia ................................................................... 214 5.2 Revision von Oxycara subgen. Symphoxycara .................................................................. 225 5.3 Zusammenfassung ........................................................................................................... 279 6. Biogeographie ......................................................................................................................... 280 6.1 Verbreitungstypen der Tenebrionidae in der Sahara ........................................................ 282 6.2 Mögliche Ausbreitungszentren in der Sahara ................................................................... 291 6.3 Verbreitungsbilder der Tenebrionidae der Sinai-Halbinsel ............................................... 302 6.4 Biodiversität von Insekten eines Mikrohabitats ................................................................ 307 6.5 Zusammenfassung ........................................................................................................... 322 4 7. Diskussion ............................................................................................................................... 323 7.1 Sind nahezu alle Arten der westlichen Paläarktis bekannt? .............................................. 323 7.2 Liegen für alle westpaäarktischen Regionen aktuelle Faunenlisten/kommentierte Artenverzeichnisse vor? .............................................................................................................. 326 7.3 Lassen sich alle Arten der Westpaläarktis mit Hilfe von existierenden Bestimmungstabellen eindeutig erkennen? ................................................................................................................... 329 7.4 Ist die Ausbreitungsgeschichte der westpaläarktischen Tenebrionidae bekannt? ............. 332 7.5 Schlußfolgerungen ........................................................................................................... 335 7.6 Literatur .......................................................................................................................... 336 8. Dank ....................................................................................................................................... 345 9. Anhang ................................................................................................................................... 346 9.1 Curriculum vitae .............................................................................................................. 346 9.2 Publikationen .................................................................................................................. 348 5 Zusammenfassung Die Tenebrionidae gehören mit mehr als 20.000 beschriebenen Arten zu den großen Käferfamilien. In den Bereichen Taxonomie, Faunistik, Biogeographie und Ökologie werden sie seit mehr als 250 Jahren erforscht. Die Geschichte der Erforschung der Tenebrionidae wird von Carl von LINNÉ, der als erster Naturforscher Vertreter dieser Familie beschrieben hat, bis zu den heutigen Bearbeitern dargestellt. Im 18. Jahrhundert beschränkten sich die Coleopterologen auf kurze, nach heutigem Maßstab in der Regel unzureichende Beschreibungen. Im 19. Jahrhundert wurden die Informationen zu den Arten meist umfangreicher, erste Monographien, Bestimmungstabellen und Kataloge wurden veröffentlicht. Die Spezialisierung der Coleopterologen, die sich mit der Taxonomie beschäftigten, auf eine oder auf wenige Familien begann zwar im 19. Jahrhundert, setzte sich aber verstärkt im 20. Jahrhundert fort. Jetzt traten auch die ersten Tenebrionidologinnen in Erscheinung. Neue Teilgebiete wurden erschlossen: Einige Forscher bearbeiteten Larven der Tenebrionidae, andere die Systematik. Es kam zu gravierenden Änderungen in der Zusammensetzung der Familie. Zu Beginn des 21. Jahrhunderts hielt die Molekulargenetik Einzug in die Bearbeitung der Tenebrionidae. Nach dieser langen Zeit könnte die Erforschung der Familie weitgehend abgeschlossen sein. Daher werden vier Hypothesen aufgestellt: Hypothese 1: Es sind inzwischen nahezu alle Arten bekannt. Dies sollte vor allem auf die westliche Paläarktis zutreffen, da aus diesem Gebiet die meisten Coleopterologen stammen. Hypothese 2: Die Faunistik der Tenebrionidae ist weit fortgeschritten. Aus den meisten Staaten und Regionen der westlichen Paläarktis liegen Faunenlisten mit zusätzlichen Informationen zu den Arten vor. Hypothese 3: Die beschriebenen Arten lassen sich mit Hilfe der Beschreibungen und/oder Bestimmungstabellen auch ohne Vergleich mit dem Holo-, Lecto- oder Neotypus voneinander eindeutig unterscheiden. Neue Revisionen sind nicht mehr notwendig. Dies gilt zumindest für die Westpaläarktis. 6 Hypothese 4: Die Ausbreitungsgeschichte der Tenebrionidae ist zumindest für die Arten der westlichen Paläarktis bekannt. Anhand eigener taxonomischer, faunistischer und biogeographischer Untersuchungen zu Tenebrionidae der Westpaläarktis, die vollständig oder in Ausschnitten in diese Arbeit eingebunden sind, und mit Hilfe von Publikationen von Kollegen werden die Hypothesen geprüft. Zunächst werden Einzelbeschreibungen von Arten, einer Untergattung und einer Gattung aus dem Nahen Osten vorgestellt: zwei Arten
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