The Dorcatoma Herbst, 1792 (Coleoptera: Anobiidae) Species of Estonia

116© Entomologica Fennica. 3 July 2002 Süda & Nagirnyi • ENTOMOL. FENNICA Vol. 13

The Dorcatoma Herbst, 1792 (Coleoptera: Anobiidae) species of Estonia

Ilmar Süda & Vitali Nagirnyi

Süda, I. & Nagirnyi, V. 2002: The Dorcatoma Herbst, 1792 (Coleoptera: Anobiidae) species of Estonia. — Entomol. Fennica 13: 116–122. The collection specimens of the genus Dorcatoma Herbst, 1792 were revised for Estonia. A substantial part of the material was reared from fruiting bodies of bracket fungi. Seven species of the genus are found in the country. Five of them, D. ﬂavicornis F., D. substriata Hummel, D. punctulata Mulsnat & Rey, D. robusta Strand and D. lomnickii Rtt. are new for Estonia, the last also being new for the Northern Europe and Eastern Baltic regions. Data on the ecology and distribution of the species in Estonia are presented. Comments are given on the systematics and geographical distribution of D. lomnickii and D. obtrita Logv. Ilmar Süda, Forest Research Institute, Estonian Agricultural University, Kreutzwaldi 5, 51014 Tartu, Estonia; E-mail: [email protected] Vitali Nagirnyi, Narva mnt. 131-1, 51008 Tartu, Estonia; E-mail: vetal@ﬁ.tartu.ee Received 15 October 2001, accepted 24 January 2002

1. Introduction would suspect that widespread European species, such as D. robusta or D. punctulata, might sim- The genus Dorcatoma Herbst, 1792 comprises ply have been overlooked in Estonia. With this over 70 species worldwide (Borowski 1999). The concern in mind, we planned to study the mate- genus is poorly investigated even in Europe, rial of this genus available in Estonia. Since the mainly because of the morphological similarities material in collections was relatively scanty, we and secretive habits of adult beetles. Therefore, it also reared out beetles from fruiting bodies of is not surprising that new Dorcatoma species have bracket fungi collected in the field. This study recently been described from Europe (Baranowski resulted in the discovery of five species new for 1985, Zahradnik 1993a), and that entomologists Estonia, making the total number of Dorcatoma are still arguing about the synonymity of several species seven. Distribution maps are given for all species found in Estonia. species (e.g. Borowski 1999). Presently, only two species, D. chrysomelina Sturm, 1837 and D. dresdensis Herbst, 1792 have been reported from Estonia (Szeliga-Mierzeyewski 1942, Miländer 2. Material and methods 1993, 1999), while eight species in all are known We examined material from the collections of the Institute from Northern Europe and the Baltic countries of Zoology and Botany (Tartu), including the collection of (Silfverberg 1992, Jonsell 1998). Considering the J. Mieländer; the Estonian Museum of Natural History general distribution of Dorcatoma species, one (Tallinn), including the collection of V. Soo; the Zoologi- ENTOMOL. FENNICA Vol. 13 • The Dorcatoma species of Estonia 117 cal Museum of Tartu University (Tartu); the Institute of (Szeliga-Mierzeyewski 1942); Rakvere, Vinni, Plant Protection (Tartu) and the Forest Research Institute from dry oak rot, 4.VII.1965, 8, V. Soo; Laane- (Tartu), as well as the private collections of the authors and metsa 4 km SW, Laetiporus sulphureus on wil- those of G. Miländer, U. Roosileht and H. Õunap. The bulk of material studied was reared from dead fruit- low, 30.VII.1996, 1, I. Süda; Saaremaa Island, ing bodies of Fomes fomentarius, Phellinus igniarius, Kuressaare, Loode oak grove, Laetiporus sul- Phellinus nigricans, Phellinus cinereus, Phellinus tremulae, phureus on oak, 20.VII.1997, 1, I. Süda; Saare- Phellinus populicola, Fomitopsis pinicola, Inonotus maa Island, Kuressaare, Loode oak grove, Laeti- radiatus, Inonotus rheades and Ganoderma applanatum. porus sulphureus on oak, 19.VII.1998, 1, V. The fruiting bodies were collected in early spring or late Nagirnyi; Saaremaa Island, Viidumäe Nature autumn. They were placed into a light-tight box supplied Reserve, Audaku 3 km N, reared from oak rot, with a small transparent window in the upper part that can be used for hatched beetles attracted to light (photoeclector). 16.VI.2001, 9, I. Süda; Saaremaa Island, Kures- A hole was made in the window and through this hole a saare, Loode oak grove, from oak rot, 5.VII.2001, transparent jar for collecting beetles was attached to the 3, I. Süda. box with the aid of a short plastic tube. Emerged beetles Distribution and ecology. In spite of the fact crawled up towards the light and into the jar. The rearing that D. chrysomelina is the first Dorcatoma spe- boxes were kept at room temperature or in an unheated room, cies reported from Estonia, it is one of the rarest where temperatures were several degrees higher than out- door temperatures. We did not, therefore, record the date of species in our fauna, found in five localities only hatching. The dates given in the material list for each spe- (Fig. 1b). It was found on oaks and once on a wil- cies refer to the dates of collecting of fungi. A few speci- low with red rot caused by the fungus Laetiporus mens were collected in the nature from bracket fungi, or sulphureus. were caught occasionally while sweeping the forest veg- etation with a net. A few specimens were reared from red wood rot taken from trees infested with Laetiporus sulphureus. *Dorcatoma substriata Hummel, 1829 The distribution maps were prepared using the UTM 10 ¥ 10-km grid system. = D. serra (Panzer, 1795 nec Fabricius, 1792) (Silfverberg 1992) Material. Soontaga, reared from Fomes fomen- 3. Results and discussion tarius on birch, 26.V.1995, 1 £, I. Süda; Soontaga 2 km NW, reared from Inonotus radiatus on al- An annotated list of Dorcatoma species found in der, 7.V.1999, 4 £, 1 ¥, I. Süda. Estonia is presented below. Species reported for Distribution and ecology. D. substriata is a the first time from the territory are denoted by an rare species in Estonia, known from two locali- asterisk (*). ties only (Fig. 1c). It was reared from Inonotus radiatus and Fomes fomentarius. To our knowl- edge, D. substriata has not been reared from *Dorcatoma flavicornis (Fabricius, 1792) Fomes fomentarius before. This is also true for Sweden, where 811 fruiting bodies of F. fomen- Material. Saaremaa Island, Kuressaare, Loode oak tarius have been analysed (Jonsell 1998). Up to grove, 19.VII.1998, 2 £, 1 ¥, V. Nagirnyi. now, D. substriata has been found on five fungal Distribution and ecology. D. flavicornis is rare species, all belonging to the genus Inonotus (Koch in Estonia, found in only one locality (Fig. 1a). 1989, Jonsell 1998, Nikitsky et al. 1998). On the The specimens were beaten from dead oak other hand, it is the species Dorcatoma minor branches lying on the ground under oak trees. Zahradník, 1993, habitually highly similar to D. substriata, which is known to breed in Fomes fomentarius (Jonsell 1998; Borowski 1999). Bear- Dorcatoma chrysomelina Sturm, 1837 ing all this in mind, we checked a preparation of the genitalia of the Estonian specimen reared from Literature. Szeliga-Mierzeyewski (1942). F. fomentarius. The result, however, confirmed Material. Ösel, Parrasmetz (= Saaremaa Is- that the specimen under discussion belongs to D. land, Parasmetsa), 26.VII.1933, 1, Mierzeyewski substriata. 118 Süda & Nagirnyi • ENTOMOL. FENNICA Vol. 13

Fig. 1. The distribution maps of Dorcatoma species in Estonia. — a. D. ﬂavicornis. — b. D. chrysomelina. — c. D. substriata. — d. D. punctulata. — e. D. dresdensis. — f. D. robusta. — g. D. lomnickii.

*Dorcatoma punctulata Mulsant & Rey, 1864 Tartu, Tiksoja, reared from F. pinicola, 17.IV.1999, 2, V. Nagirnyi; Soontaga, reared from F. pinicola Material. Lohu, mixed forest margin, from grass, on spruce stub, 7.V.1999, 13, I. Süda; Valmaotsa 4 19.VII.1960, 1, V. Soo; Kanaküla, mixed forest, km S, reared from F. pinicola on a pine stump, from grass, 17.VI.1964, 1, V. Soo; Krabi, Seveli, 25.V.1999, 3, I. Süda; Kaugjärv lake shore, reared clearing, rotten birch, 21.VI.1967, 1, V. Soo; from F. pinicola on birch, 12.VI.1999, 3, I. Süda; Viljandi, Aimla, clearing, from grass, 29.VI.1971, Jõemõisa lake shore, reared from F. pinicola on 1, V. Soo; Vormsi Island, Suuremõisa 1 km S, birch lime, 22.IV.2001, 8, I. Süda. stump, polypore fungus, 8.VI.1993, 1, G. Miländer Distribution and ecology. D. punctulata is a (Miländer 1999, as D. dresdensis); Kanaküla, relatively common species in Estonia, spread all 9.VII.1993, 1, U. Roosileht; Soontaga, reared from over the territory (Fig. 1d). Reared only from Fomitopsis pinicola on birch, 26.V.1995, 3, I. Süda; Fomitopsis pinicola on spruce, pine, birch and lime. ENTOMOL. FENNICA Vol. 13 • The Dorcatoma species of Estonia 119

Dorcatoma dresdensis Herbst, 1792 Koobassaare 4 km E, from Phellinus nigricans on birch, 13.VI.1999, 1, I. Süda; Koikküla, Fomes Literature. Miländer (1993), Miländer (1999). fomentarius on birch stub, 14.VI.1999, 1, I. Süda; Material. Võsu, Esku, VII.1931, 1, Vannary; Alam-Pedja Nature Reserve, Nõmmeotsa, Inonotus Aegviidu, Mustjõe, fungus on rotten birch, radiatus on alder, 20.VI.1999, 3, I. Süda; Luke, 27.VI.1947, 1, V. Soo; Tohkri, 6.VI.1949, 1, J. park, Ganoderma applanatum on willow, 24.VI.1999, Mieländer; Risti, Metsa, woodland, from grass, 2, I. Süda; Riimaru 2 km N, reared from Phellinus 6.VII.1952, 1, V. Soo; Krabi, Seveli, 3.VI.1960, tremulae on aspen, 7.V.2000, 7, I. Süda; Kaiu lake 5, J. Mieländer; Risti, Metsa, woodland, dry as- shore, reared from Phellinus nigricans on birch, pen, 30.VII.1961, 1, V. Soo; Hiiumaa Island, 22.IV.2001, 47, I. Süda; Pataste 4 km N, reared Jausa, 22.VI.1972, 1, J. Mieländer; Kanaküla, from Phellinus igniarius on willow, 22.IV.2001, Riimaru, clearing margin, from grass, 11.VII.1977, 4, V. Nagirnyi; Vaitka, Koiva woodland, Phellinus 1, V. Soo; Järvselja, under spruce bark, dead, igniarius on willow, 6.VI.2001, 1, I. Süda; Vaitka, 7.V.1984, 1, I. Süda; Risti, Metsa, 15.VI.1984, 2, Koiva woodland, reared from Phellinus igniarius U. Roosileht; Järvselja, 8.VI.1985, 1, U. Roosileht; on willow, 6.VI.2001, 2, I. Süda; Järvselja virgin Kanaküla, from grass, 21.VII.1992, 1, U. Roosileht; forest, reared from Phellinus nigricans on birch, Tartu, Kõrveküla, Phellinus igniarius on willow, 9.VI.2001, 12, B. Ehnström; Tudu 5 km NE, reared 2.VI.1993, 8, V. Nagirnyi; Tallinn, Tammetalu, from Phellinus tremulae on aspen, 10.VI.2001, 4, 5.VI.1993, 2, U. Roosileht; Vormsi Island, Hullo I. Süda; Tudu 5 km NE, reared from Phellinus 1.5 km E, fungus on birch, 5.VI.1993, 1, G. nigricans on birch, 10.VI.2001, 3, I. Süda; Miländer; Kanaküla, 25.VI.1993, 1, U. Roosileht; Saaremaa Island, Viidumäe Nature Reserve, Tartu, Kõrveküla, Phellinus igniarius on willow, Mäebe, reared from Phellinus nigricans on birch, 28.VI.1993, 1, V. Nagirnyi; Kõmsi, Salevere, 14.VI.2001, 1, I. Süda; Saaremaa Island, Viidumäe reared from Fomes fomentarius on birch, 8.VI.1994, Nature Reserve, Kivesselja, reared from Inonotus 9, H. Õunap; Taagepera 6 km S, reared from radiatus on alder, 14.VI.2001, 2, I. Süda; Saaremaa Fomes fomentarius on birch, 26.IV.1995, 2, I. Island, Viidumäe Nature Reserve, Mäebe, reared Süda; Viisjaagu lake shore, reared from Phellinus from Inonotus radiatus on hazel, 15.VI.2001, 3, igniarius on willow, 1.V.1995, 1, I. Süda; I. Süda; Padakõrve 1 km N, reared from Phellinus Soontaga, reared from Fomes fomentarius on populicola on aspen, 29.VI.2001, 5, H. Õunap; birch, 26.V.1995, 2, I. Süda; Himma, Sika, reared Nõo 2 km SE, Phellinus igniarius on willow, from Ganoderma applanatum on lime, 29.V.1995, 12.VII.2001, 2, I. Süda. 154, I. Süda; Puhja, Saare, reared from Fomes Distribution and ecology. D. dresdensis is the fomentarius on birch, 23.VI.1996, 2, V. Nagirnyi; most common species in Estonia, spread all over Saaremaa Island, Kõljala, swept from bushes, the territory (Fig. 1e). It is a polyphagous species 15.VI.1998, 1, I. Süda; Elva, Peedu, 24.VI.1998, reared in Estonia from Fomes fomentarius on birch 1, V. Nagirnyi; Alam-Pedja Nature Reserve, Selli, and alder, from Phellinus igniarius on willow, reared from Inonotus rheades on aspen, from Phellinus tremulae and Phellinus populicola 22.IV.1999, 5, I. Süda; Tartu, Maramaa, reared on aspen, from Phellinus cinereus on birch, from from Phellinus tremulae on aspen, 24.IV.1999, 2, I. Inonotus radiatus on alder and hazel, from Süda; Tartu, Maramaa, reared from Phellinus Inonotus rheades on aspen and from Ganoderma cinereus on birch, 24.IV.1999, 4, I. Süda; Tartu, applanatum on lime. Maramaa, reared from Fomes fomentarius on alder, 24.IV.1999, 3, I. Süda; Tartu, Maramaa, reared from Inonotus radiatus on alder, 24.IV.1999, *Dorcatoma robusta Strand, 1938 3, I. Süda; Soontaga 2 km NW, reared from Inonotus radiatus on alder, 7.V.1999, 9, I. Süda; Material. Aegviidu, Mustjõe, fungus on rotten Alam-Pedja Nature Reserve, Utsali, reared from birch, 27.VI.1947, 1, V. Soo; Aegviidu, Mustjõe, Phellinus tremulae on aspen, 3.VI.1999, 5, I. Süda; fungous birch, 9.VI.1948, 1, V. Soo; Krabi, Seveli, Kaugjärv lake shore, reared from Phellinus 1.VI.1960, 1, J. Mieländer; Krabi, Seveli, 3.VI.1960, nigricans on birch, 12.VI.1999, 4, I. Süda; 1, J. Mieländer; Krabi, Seveli, 4.VI.1960, 1, J. 120 Süda & Nagirnyi • ENTOMOL. FENNICA Vol. 13

Mieländer; Krabi, Seveli, 8.VI.1960, 2, J. Mieländer; der, 14.V.1999, 25, I. Süda; Puhja, Saare, reared Hiiumaa Island, Männamaa, 13.VI.1969, 2, J. from Fomes fomentarius on birch, 10.III.2001, 5, Mieländer (Miländer 1993, as D. dresdensis); V. Nagirnyi; Kaiu lake shore, reared from Fomes Oidrema, 8.VII.1974, 2, J. Mieländer; Karinõmme, fomentarius on birch, 22.IV.2001, 1, I. Süda; Tarva, Fomes fomentarius on birch stub, Pangodi 2 km SE, reared from Fomes fomentarius 16.VII.1985, 1, I. Süda; Kanaküla, 9.VII.1993, 1, on birch, 19.V.2001, 8, I. Süda; Tudu, Oonurme U. Roosileht; Taagepera 6 km S, reared from 4.5 km E, reared from Fomes fomentarius on birch, Fomes fomentarius on birch, 26.IV.1995, 13, I. 10.VI.2001, 4, I. Süda. Süda; Soontaga, reared from Fomes fomentarius Distribution and ecology. In Estonia, D. on birch, 26.V.1995, 2, I. Süda; Vorbuse 1.5 km lomnickii is a relatively common species found in SW, reared from Fomes fomentarius on birch, the eastern part of the territory (Fig. 1g). Analysis 11.V.1996, 8, I. Süda; Koikküla, Fomes fomen- of known localities revealed that unlike the other tarius on birch stub, 2.VII.1997, 2, I. Süda; Laeva, species, D. lomnickii prefers moist forests. It is Kärevere, reared from Fomes fomentarius, reared mainly from Fomes fomentarius on birch 1.XI.1998, 100, V. Nagirnyi; Palupõhja, reared and alder, sometimes together with D. dresdensis from Fomes fomentarius on birch, 15.V.1999, 4, or D. robusta. On one single occasion it was also V. Nagirnyi; Kaiu lake shore, reared from Fomes reared from Ganoderma applanatum on lime. fomentarius on birch, 22.IV.2001, 11, V. Nagirnyi; Remarks on systematics. D. lomnickii has been Koikküla, reared from Fomes fomentarius on described by Reitter (1903) on the basis of three birch, 2.VI.2001, 2, I. Süda. specimens collected in the vicinity of Minsk Distribution and ecology. D. robusta was de- (present Byelorussia). Only a few specimens of scribed as late as in 1938 and remained unknown this species have been found since its description. to Estonian collectors for a relatively long time. It It has been included in the checklist of Slovakian is often confused with D. dresdensis, as the two Anobiidae (Zahradnik 1993b). Only three female species are morphologically similar. All the ma- specimens have been reported from the European terial of this species, preserved in older collec- part of the former Soviet Union (Logvinovskij tions, was identified as D. dresdensis. D. robusta 1980, 1985). Males were stated to be unknown. is quite a common species in Estonia, spread all D. lomnickii has been included in several keys to over the territory (Fig. 1f). All the reared speci- European beetles, based mainly on Reitter’s origi- mens originated invariably from Fomes fomen- nal description but without indication of new finds tarius on birch. This confirms that D. robusta is a (Dominik 1955, Lohse 1968). Baranowski (1985) monophagous species (Palm 1959, Jonsell 1999). studied Reitter’s type series of D. lomnickii in the Hungarian Natural History Museum, Budapest and published the first drawings of male genitalia *Dorcatoma lomnickii Reitter, 1903 after two syntype specimens had been identified as male (Fig. 2a). He also presented a very so- Material. Kanaküla, 25.VI.1993, 1, U. Roosileht; phisticated and detailed identification key to Cen- Taagepera 6 km S, reared from Fomes fomentarius tral and North-European species, including D. on birch, 26.IV.1995, 4, I. Süda; Soontaga, reared lomnickii. No additional specimens were indicated from Fomes fomentarius on birch, 26.V.1995, 6, besides a type series. Recently, D. lomnickii has I. Süda; Himma, Sika, reared from Ganoderma been reported as a new species for Poland on the applanatum on lime, 29.V.1995, 8, I. Süda; basis of finds in several localities (Borowski Vorbuse 1.5 km SW, reared from Fomes fomen- 1999). Borowski produced a subgeneric division tarius on birch, 11.V.1996, 8, I. Süda; Laeva, as well as compiled an interesting identification Kärevere, reared from Fomes fomentarius, key based on new characteristics for Central Eu- 1.XI.1998, 4, V. Nagirnyi; Ontika, reared from ropean Dorcatoma species. In the same paper, a Fomes fomentarius on birch, 17.IV.1999, 13, I. drawing of the male genitalia (Fig. 2b) differs Süda; Soontaga, reared from Fomes fomentarius slightly from the one drawn by Baranowski on alder, 7.V.1999, 11, I. Süda; Puurmani, (1985). For comparison, we have produced a pho- Jüriküla, reared from Fomes fomentarius on al- tographic image of typical male genitalia taken ENTOMOL. FENNICA Vol. 13 • The Dorcatoma species of Estonia 121

Fig. 2. Male genitalia of D. lomnickii (a–c) and D. obtrita (d). — a. Type series after Baranowski (1985). — b. Poland, after Borowski (1999). — c. Estonia. — d. After Log- vinovskij (1980). from Estonian specimens (Fig. 2c). Both undoubt- material of D. obtrita, as well as the distribution edly confirm that the beetles collected in Estonia of both species, are yet to be examined. belong to the species D. lomnickii. The finds in Estonia are the northernmost ones The distribution of D. lomnickii is not yet clear. for D. lomnickii. The species is new not only for Borowski (1999) treated D. obtrita Logvinovskij, Estonia but also for the Baltic countries and North- 1980, a species described from the Russian Far ern Europe. Taking into account the finds of D. East (Logvinovskij 1980), as a synonym of D. lomnickii in Byelorussia, Poland and Estonia, the lomnickii without indicating any plausible reasons occurrence of this species in Lithuania and Latvia for synonymizing. In the same study, D. lomnickii seems to be highly probable. is named a Siberian species with its distribution extending as far west as Poland. This seems doubt- Acknowledgements. The present investigation was sup- ful for several reasons. The known material of D. ported by the Estonian Science Foundation (grant no. 2684). obtrita is confined to the area of the Russian Far We are much indebted to Prof. E. Parmasto for the identification of some polypores and to G. Miländer, U. Roosileht East. There are no indications of the occurrence and H. Õunap for allowing the use of material from their of this species over the huge territory from the private collections for study. We are grateful to Roger de Khabarovsk region in the Far East to the Central Keyzer and George Hangay for valuable comments on the European part of Russia. On the other hand, reli- paper and to Ester Jäigma for language correction. able easternmost finds of D. lomnickii have been reported from the Bryansk and Kirov districts in European Russia (Logvinovskij 1985). References The original description indicates several ex- plicit characteristics for distinguishing D. obtrita Baranowski, R. 1985: Central and Northern European from D. lomnickii; however, it appears to be too Dorcatoma (Coleoptera: Anobiidae), with a key and de- brief and schematic. These characteristics are a scription of a new species. — Ent. Scand. 16: 203–206. gold-coloured and shiny pubescence and a rela- Borowski, J. 1999: A contribution to the Central European species of the genus Dorcatoma Herbst, 1792 (Cole- tively sparse punctuation of the elytra. Moreover, optera, Anobiidae, Dorcatominae). — Annals of War- the male genitalia of D. lomnickii display a dis- saw Agricultural University, SGGW, Forestry and tinctive feature (arrow in Fig. 2b) which is very Wood Technology 49: 127–136. stable and can hardly be overlooked. On the draw- Dominik, J. 1955: Klucze do oznaczania owadów Polski. ing of the genitalia of D. obtrita (Fig. 2d, repro- 19, Chrzaszcze — Coleoptera, 41, Kolatki — Anobiidae. duced from Logvinovskij 1980, 1985), this fea- — PWN, Warszawa. 69 pp. Jonsell, M. 1998: En ny tickgnagare i Sverige, Dorcatoma ture is lacking altogether. In our opinion, making minor Zahradnik (Coleoptera: Anobiidae), och dess a judgement on the synonymity of D. obtrita and värdval. — Ent. Tidskr. 119 (2): 105–109. D. lomnickii requires further research. The type Jonsell, M. 1999: Insects on wood decaying polypores — 122 Süda & Nagirnyi • ENTOMOL. FENNICA Vol. 13

conservation aspects. — Ph. D. thesis, SLU, Uppsala. M.) 1998: The beetles of the Prioksko-Terrasny Bio- Koch, K. 1989: Die Käfer Mitteleuropas. Ökologie. 2. — sphere Reserve — xylobiontes, mycetobiontes and Goecke & Evers, Krefeld. 382 pp. Scarabaeidae (with the review of the Moscow region Logvinovskij, V. D. (Logvinovskij, V. D.) 1980: A re- fauna of these groups). — Arch. Zool. Mus. Moscow view of the genus Dorcatoma Herbst (Coleoptera, Univ. 36, Suppl. 1. 61 pp. [In Russian]. Anobiidae) in fauna of the USSR. — Revue d’Entomo- Palm, T. 1959: Die Holz- und Rinden Käfer der Süd- und logie de l’URSS 59 (1): 148–153. [In Russian]. Mittelschwedishe Laubbäume. — Opusc. Entomol. Logvinovskij, V. D. (Logvinovskij, V. D.) 1985: [Fauna Suppl. 16. of USSR. Coleopterous insects. 14 (2). Deathwatch Bee- Reitter, E. 1903: Dorcatoma Lomnickii n. sp. — Wiener tles — family Anobiidae.] — Nauka, Leningrad. 175 Entomologische Zeitung 22 (9): 239. pp. [In Russian]. Silfverberg, H. 1992: Enumeratio Coleopterorum Fenno- Lohse, G.A. 1968: Familie Anobiidae. — In: Freude, H., scandiae, Daniae et Baltiae. — Helsingin Hyönteis- Harde, K. W., Lohse, G. A. (eds.), Die Käfer Mittel- vaihtoyhdistys, Helsinki. 94 pp. europas 8: 26–59. Goecke and Evers, Krefeld. 388 pp. Szeliga-Mierzeyewski, W. v. 1942: Verzeichnis der Käfer Miländer, G. 1993: Hiiumaa mardikalised (Coleoptera). — Ösels in meiner Insektensammlung. — Korrespondenzbl. Pirrujaak 2. Biosfääri Kaitseala Hiiumaa Keskus, Naturt. (Riga) 64: 180–230. Kärdla. 120 pp. Zahradnik, P. 1993a: New species of the genus Dorcatoma Miländer, G. 1999: Huvitavamaid leide Eesti mardikalistest from Central Europe (Coleoptera, Anobiidae). — Fo- (Coleoptera). — Lepidopteroloogiline Informatsioon lia Heyrovskyana 1(8): 80–83. 11: 52–58. Zahradnik, P. 1993b: Anobiidae. — In: Jelinek, Y. (ed.), Nikitsky, N. B., Semenov, V. B. & Dolgin, M. M. Check-list of Czechoslovak Insects 4 (Coleoptera): 86– (Nikitskij, N. B., Semenov, V. B. & Dolgin, M. 88. Folia Heyrovskyana, Suppl. 1, Prague.