Atlas of Yorkshire Coleoptera (VCs 61-65) Part 9 – Derodontoidea, and Lymexyloidea

Introduction

This section of the atlas deals with the Superfamilies Derodontoidea, Bostrichoidea and Lymexyloidea, a total of 104 , of which there are 57 recorded in Yorkshire.

Each species in the database is considered and in each case a distribution map representing records on the database (at 1/10/2017) is presented. The number of records on the database for each species is given in the account in the form (a,b,c,d,e) where 'a' to 'e' are the number of records from VC61 to VC65 respectively. These figures include undated records (see comment on undated records in the paragraph below on mapping). As a recorder, I shall continue to use the vice-county recording system, as the county is thereby divided up into manageable, roughly equal, areas for recording purposes. For an explanation of the vice-county recording system, under a system devised in Watson (1883) and subsequently documented by Dandy (1969), Britain was divided into convenient recording areas ("vice-counties"). Thus Yorkshire was divided into vice-counties numbered 61 to 65 inclusive, and notwithstanding fairly recent county boundary reorganisations and changes, the vice-county system remains a constant and convenient one for recording purposes; in the text, reference to “Yorkshire” implies VC61 to VC65 ignoring modern boundary changes.

For some species there are many records, and for others only one or two. In cases where there are five records or less full details of the known records are given. Many common species have quite a high proportion of recent records. This is because the older county recorders tended not to record species they regarded as "common" quite as assiduously as we do today.

“Recent” records are those considered to be later than 1 January 2000 – this is also the break point used in the production of the distribution maps. (In previous parts of this Atlas, the break point for recent and otherwise was 1 January 1990). Habitat A brief statement on habitat is included where known and relevant. Taxon status Inclusion of a statement on the status of species presents problems in that those designations in Recorder 6 are often out of date. Recently Natural England has been publishing reports on various groups of Coleoptera which bring the national status of species in line with present knowledge and recording. Recorder 6 statuses are in many cases based largely on Hyman and Parsons (1992 and 1994) - see below for status category explanations. For the saproxylic species in this section I have consulted Alexander (2002).

Phenology Details are given for the range of months in the year for which there are records, with or without a peak month of occurrence, where the number and detail of records are sufficient to give a meaningful statement.

Maps Distribution maps have been produced using Dr Alan Morton’s DMAP software (version 7.0b). It may be worthwhile to note that as DMAP is asked to map pre- and post-2000 records in this application, any undated records will not be mapped. While efforts have been made to establish dates of records, some are difficult to ascertain, and I do not want to make guesses which may be wrong. In addition to this, records flagged in the database as unconfirmed will also not be mapped. The YNU database possesses a few peat sub-fossil records. These are not considered for inclusion in the atlas.

Nomenclature The YNU database is presently held in a copy of Recorder 6, which uses a Coleoptera checklist based on Duff (2008) and this new list incorporates a very large number of taxon name changes since the checklist of Pope (1977); there is now a second edition of the latest Coleoptera checklist (Duff, 2012) which has not yet been incorporated fully into Recorder 6. The nomenclature and arrangement of species used in this section of the atlas follows Duff (2012). Occasionally I receive records using “noddy names” (see Cooter, (1999) and Key (1999)) for an excellent treatment of this subject), e.g. “Snail Hunter”, “Crucifix ” and “Heath-glory ” (Cychrus caraboides, Panagaeus crux-major and nitens respectively). Such inventions were designed for use in political documents and seem to have

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originated somewhere within Natural England (previously English Nature) or JNCC. For the purposes of serious science these terms are, in my opinion, either vague, ridiculous, confusing or unhelpful. I do not accept records using such names only, unless accompanied by specimens or other convincing evidence.

Quality of the records A regrettable aspect of many records within the YNU database is the lack of detail accompanying the record, especially of accurate grid references, lack of pertinent comments as to the micro-habitat of the capture, its association with plants/, and sometimes, in the case of some modern records, a lack of care in determination, and the recording of the sex of the specimen. Old records, although mainly sound, tend not to give much or indeed any of this detail. Old records on the database so rarely give habitat details but sometimes interpretation of the site name can give clues to the habitat type. Grid references could not be applied before 1930 anyway, and grid references do not figure in the paper archive of the YNU Coleoptera database until the late 1970s. Apparently it was not considered necessary to give that level of accuracy. Entering records into Recorder 6 requires a grid reference, and with older records I have added a grid reference where I believe this has been possible. If the site name is a vague one I have used a 10km reference, and where a 1km reference is deemed accurate enough I have included that. There can be little excuse with modern records not to include some accurate comment with the record, and at least a six-figure grid reference. Taxonomic knowledge and expertise have increased greatly since the days of Walsh, Bayford and Stainforth. Much good literature now exists for the taxonomist in the form of dichotomous keys, and when identifying specimens, especially with critical species pairs or groups, too much reliance on the outcome of keys can be placed on a determination, without apparently considering the habitat in which the specimen was taken, or the host plant/. Atlases of the national distributions of various orders of are now becoming available and should always be consulted. These often give a guide to the likelihood or otherwise of a determination, especially in scarce taxa. Museum collections containing correctly determined and labelled specimens are invaluable in accurate recording; museum collections are there for our use and should be consulted. If a species is ‘new’ to a vice-county or is very significant in some other respect, a second opinion should always be sought or an acknowledged specialist in the group consulted. A voucher should always be retained by the collector if possible. The recorder can only deal with the information he is given, and often has to beware of identifications made on photographic evidence only, and identifications of ‘difficult’ species by non-specialists. It takes many years to become competent in many areas of entomology, and often the recorder has to rely on the expertise of co-workers, especially when records covering large Orders are being documented. I have adopted a policy in my county recording whereby any received record of a species afforded Notable B or above, especially if a member of a ‘difficult’ beetle group, should be confirmed by a specialist in that group before the record is entered into the database as a correct record.

Ultimately, the responsibility for verification of records on the YNU database rests with me. With modern records received from reputable coleopterists there is little problem. Some older records requiring vouchers for confirmation have been retained in the database but flagged as requiring confirmation.

Species notes

There is much detailed information in Peacock (1993) regarding the , and in Alexander (2002) regarding , Anobiidae and Lymexylidae.

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Contributors (alphabetically by initials as appearing in the text, not alphabetically by surname)

ABD – Tony Drane JMa – John Martin AL – Alan Lazenby JMe – J Merryweather BC – Barry Constantine JRD – J R Dibb CH – Chris Hopkinson JWC – John William Carter CHo – Colin Howes KNAA – Keith Alexander DB – D Burgess LB – Leslie Barringer ECH – E C Horrell MC – Mike Clegg EFG – E F Gilmour MH – Martin Hammond EGB – E G Bayford MLD – Mike Denton EJS – Eric Smith PK – Peter Kendall EW – E Wood PS – Peter Skidmore EWA – Ted Aubrook RGB – Roger Booth GBW – G B Walsh RJM – Bob Marsh GM – Graham Maynard RL – R Lawson HB – Harry Britten TS – T Stainforth HBe – H Bean WAE – Bill Ely HK – Harry Kenward WDH – Walter Douglas Hincks JB – Joe Botting WWF – William Weekes Fowler

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Superfamily DERODONTOIDEA LeConte, 1861 4 433. Family DERODONTIDAE LeConte, 1861 43. Family DERODONTIDAE, LeConte, 1861

Subfamily LARICOBIINAE Mulsant & Rey, 1864

LARICOBIUS Rosenhauer, 1846 erichsonii Rosenhauer, 1846

Laricobius erichsonii Rosenhauer, 1846

3 WATSONIAN (0,2,0,0,0). Very rare in Yorkshire with only two known records, both from the same site: 2 YORKSHIRE Newburgh Priory Park House Pasture SE5675, 1 16/7/2003, ABD, on oak; Newburgh Priory 0 NY NZ NZ OV SD SE SE TA Pine Plantation SE5675, 8/8/2003, ABD, 9 beaten from oak. According to Peacock 8 (1993:67) and other literature this species is associated with woolly aphid Adelges piceae 7 on conifers, but has been reported on the 6 foliage of other trees probably fortuitously; in 5 Britain first discovered in Suffolk in 1971. Our 4 records presumably indicate establishment 3 but only further records will confirm this. 2 1

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Superfamily BOSTRICHOIDEA Latreille, 1802 44. Family DERMESTIDAE Latreille, 1804 530 44. Family DERMESTIDAE Latreille, 1804

Subfamily THYLODRIINAE Semenov, 1909

THYLODRIAS Motschulsky, 1839 contractus Motschulsky, 1839

NO RECORDS

Subfamily Agassiz, 1846

THORICTODES Reitter, 1875 heydeni Reitter, 1875

NO RECORDS

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Subfamily Latreille, 1804

DERMESTES Linnaeus, 1758

Subgenus DERMESTINUS Zhantiev, 1967 carnivorus Fabricius, 1775

NO RECORDS frischii Kugelann, 1792

Dermestes (Dermestinus) frischii Kugelann, 1792

3 WATSONIAN (0,2,1,0,0). GBES-Established. Rarely 2 YORKSHIRE reported with three records on the YNU database, two of these of uncertain date so 1 therefore unmapped: Langdale End SE9492, 0 NY NZ NZ OV SD SE SE TA date unknown, ECH; Scarborough TA08, date 9 unknown, WWF; Batley SE2325, 10/1967, 8 MC (Naturalist 1970:23). An imported 7 species of hides, bonemeal and animal feed, and also associated with carrion and dried 6 fish. There is a record from Scarborough 5 (1869) attributed to Wilkinson. This record 4 does not appear in the YNU records but is 3 quoted in Peacock (1993:47). 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 13/1/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4 maculatus De Geer, 1774

Dermestes (Dermestinus) maculatus De Geer, 1774

3 WATSONIAN (3,3,6,3,0). GBES-Indoors. Rarely reported with 15 known Yorkshire records. Reported 2 YORKSHIRE from an antelope skin in Scarborough 1 Museum, in a dead porpoise on the beach at 0 NY NZ NZ OV SD SE SE TA Spurn and in restaurant premises in 9 Huddersfield. The only recent (post-2000) 8 records are for Cadeby SE5101, 2005, RJM, in a dead fox by the roadside, and from Goole 7 (SE7223), 2005, PK, around a dead roe deer. 6 Nationally, sometimes reported as an import 5 in animal products such as hides, bones and 4 dried fish. 3 2 1

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murinus Linnaeus, 1758

Dermestes (Dermestinus) murinus Linnaeus, 1758

3 WATSONIAN (7,2,5,1,0). There are a known 15 records on the YNU database, only three of these recent. 2 YORKSHIRE Apparently an outdoor species in Britain in 1 carrion of mammals and birds, and in the 0 NY NZ NZ OV SD SE SE TA writer’s experience fairly common in 9 gamekeepers’ gibbets. A pest of stored 8 products in other parts of the world. 7 6 5 4 3 2 1

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undulatus Brahm, 1790

Dermestes (Dermestinus) undulatus Brahm, 1790

3 WATSONIAN (2,1,0,0,0) A rarity with only three known 2 YORKSHIRE records and not reported for many decades: Scarborough TA08, c.1910, RL; Breighton SE73, 1 1913, WJF; Holme on Spalding Moor SE8138, 0 NY NZ NZ OV SD SE SE TA 17/5/1924, TS. As an outdoor species now very 9 rare in Britain (Peacock (1993) in carrion of 8 mammals and birds. 7 6 5 4 3 2 1

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Subgenus DERMESTES Linnaeus, 1758 ater De Geer, 1774

Dermestes (Dermestes) ater De Geer, 1774

3 WATSONIAN (0,0,1,0,0) GBES-Indoors. There is a single record 2 YORKSHIRE on the YNU database: Sheffield SK3686, 1977, BC, imported in goods from South America 1 (records via Local Authority Environmental 0 NY NZ NZ OV SD SE SE TA Health). Nationally the most commonly 9 recorded Dermestes of imported cargoes, in a 8 wide range of animal products. 7 6 5 4 3 2 1

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haemorrhoidalis Küster, 1852

Dermestes (Dermestes) haemorrhoidalis Küster, 1852

3 WATSONIAN (0,0,2,0,0) There are two records of this species on the YNU database: Sprotbrough 2 YORKSHIRE SE5503, 10/12/2005, RJM, 1 specimen in 1 house, ex dried dog-feed; Leeds SE2833, 0 NY NZ NZ OV SD SE SE TA 5/8/2014, JMa det RJM, one specimen 9 collected with other dermestids in flat in 8 central Leeds. In Britain usually reported in 7 imported dried animal products, and very rare in the wild (Peacock, 1993). 6 5 4 3 2 1

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lardarius Linnaeus, 1758

Dermestes (Dermestes) lardarius Linnaeus, 1758

3 WATSONIAN (4,7,21,2,0) By far the most frequently 2 YORKSHIRE reported Dermestes, records being noted from houses generally, in carcases of rabbit, magpie 1 and crow in the wild, and from dried dog feed, 0 NY NZ NZ OV SD SE SE TA stored flour, and reported from within a 9 butcher’s shop. 8 7 6 5 4 3 2 1

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leechi Kalik, 1952

NO RECORDS

peruvianus Laporte, 1840

Dermestes peruv ianus Laporte de Castelnau, 1840

3 WATSONIAN (0,0,4,0,0) GBES-Established. There are no recent records of this rarely reported species 2 YORKSHIRE in the county but we have four on the 1 database: Huddersfield SE1416, 1/1954, DB, NY NZ NZ OV 0 SD SE SE TA in works canteen, see EMM 99:30 and Nat 9 1964:21; Rotherham SK4493, 1/10/1980, LB; 8 Sheffield SK38, 1986, EJS, see Sorby Record 24(1986):43; Sheffield SK3486, 20/11/1992, 7 AL. All records from indoor situations. 6 Nationally established as a pest of fish meal, 5 hides and other animal-related stored 4 products, and rarely observed in the wild. 3 2 1

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Subfamily Laporte, 1840

ATTAGENUS Latreille, 1802 brunneus Faldermann, 1835

NO RECORDS

cyphonoides Reitter, 1881

NO RECORDS

fasciatus (Thunberg, 1795)

NO RECORDS

pellio (Linnaeus, 1758)

Attagenus pellio (Linnaeus, 1758)

3 WATSONIAN (10,6,35,3,2) GBES-Established. Fairly 2 YORKSHIRE frequent and widespread in Yorkshire, in barns, warehouses, on sticky traps in 1 buildings, in a collection of birds’ eggs, and in 0 NY NZ NZ OV SD SE SE TA farmyard debris, but also on occasion by 9 sweeping hawthorn and hogweed flowers in 8 the wild. (ii-xii, peak v). 7 6 5 4 3 2 1

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smirnovi Zhantiev, 1973

NO RECORDS

trifasciatus (Fabricius, 1787)

NO RECORDS

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unicolor (Brahm, 1791)

Attagenus unicolor (Brahm, 1791)

3 WATSONIAN (0,0,0,1,0) GBES-Indoors. There is a single 2 YORKSHIRE record on our database: Leeds SE33, 1932, JRD, see EMM 1932:18 and Nat 1932:59. 1 Synanthropic in various imported 0 NY NZ NZ OV SD SE SE TA commodities (see Peacock, 1993 for more 9 detail). 8 7 6 5 4 3 2 1

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Subfamily Leach, 1815

TROGODERMA Dejean, 1821

angustum (Solier, 1849)

NO RECORDS

glabrum (Herbst, 1783)

NO RECORDS

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granarium Everts, 1898

Trogoderma granarium Ev erts, 1898

3 WATSONIAN (0,1,0,0,0) GBES-Not_Established. A single record is included in our database: 2 YORKSHIRE Scarborough TA08, 1921, GBW, imported in a 1 cargo of cereals. A native of southern Asia it is 0 NY NZ NZ OV SD SE SE TA synanthropic in Britain in imported vegetable- 9 based goods. Peacock (1993:56) mentions a 8 record from Leeds (“in barley”) but gives no other detail. This record does not appear in the 7 YNU records. 6 5 4 3 2 1

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inclusum LeConte, 1854

Trogoderma inclusum LeConte, 1854

3 WATSONIAN (0,0,1,0,0) GBES-Established. The single record on the YNU database gives us the following 2 YORKSHIRE data: Goole SE7223, 24/12/1984, PK, in a box 1 of Christmas nuts. Nationally this species is 0 NY NZ NZ OV SD SE SE TA reported in imports of a wide variety of 9 vegetable-based products, but may in warmer 8 climates breed in insect remains in beetle burrows and nests of various bees (see 7 Peacock, 1993:56) for more detail. 6 5 4 3 2 1

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variabile Ballion, 1878

NO RECORDS

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REESA Beal, 1967 vespulae (Milliron, 1939) Reesa v espulae (Milliron, 1939)

3 WATSONIAN (2,0,0,0,0) GBES-Established. We have two records on the YNU database: York University 2 YORKSHIRE SE6250, 1984, RGB, in laboratory premises; 1 York University SE6250, 25/2/1987, JMe, 1 ex 0 NY NZ NZ OV SD SE SE TA dead in herbarium collection. A rare, 9 synanthropic and parthenogenetic species in 8 Britain, it is reported internationally from plant material (living and dead), seeds, museum 7 collections of plants and fungi, bird skins and 6 dried museum-based animal specimens. 5 4 3 2 1

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GLOBICORNIS Latreille, 1829 rufitarsis (Creutzer in Panzer, 1796)

NO RECORDS

MEGATOMA Herbst, 1792 undata (Linnaeus, 1758) Megatoma undata (Linnaeus, 1758)

3 WATSONIAN (6,2,4,4,0) Nationally Scarce B. Very scarce. Usually found under bark of deciduous trees, 2 YORKSHIRE in fungi and spiders’ webs under loose bark. 1 Probably a scavenger on chitinous and 0 NY NZ NZ OV SD SE SE TA decaying fungal material. (iv-x). 9 8 7 6 5 4 3 2 1

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ANTHRENOCERUS Arrow, 1915 australis (Hope, 1843)

NO RECORDS

ORPHINUS Motschulsky, 1858 fulvipes (Guérin-Méneville, 1838)

NO RECORDS

CTESIAS Stephens, 1830 serra (Fabricius, 1792)

Ctesias serra (Fabricius, 1792)

3 WATSONIAN (0,9,6,4,0) Nationally Scarce B. Very scarce in 2 YORKSHIRE Yorkshire and recorded from beech, oak and horse chestnut, in insect galleries, old fungi 1 and in spiders’ webs where there are 0 NY NZ NZ OV SD SE SE TA remains, mainly in the increasingly 9 rare habitat of over-mature broad-leaved 8 trees in parkland and wood-pastures. (iv-viii). 7 6 5 4 3 2 1

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Subfamily ANTHRENINAE Gistel, 1848

ANTHRENUS Geoffroy, 1762

Subgenus ANTHRENUS Müller, O.F., 1764

flavipes LeConte, 1854

NO RECORDS

pimpinellae Fabricius, 1775

NO RECORDS

scrophulariae (Linnaeus, 1758)

NO RECORDS

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Subgenus ANTHRENODES Chobaut, 1898 sarnicus Mroczkowski, 1963

NO RECORDS

Subgenus ANTHRENOPS Reitter, 1881 coloratus Reitter, 1881

NO RECORDS

Subgenus FLORILINUS Mulsant & Rey, 1868 museorum (Linnaeus, 1761)

Anthrenus (Florilinus) museorum (Linnaeus, 1761)

3 WATSONIAN (1,3,6,0,0) Scarce: recorded from museum 2 YORKSHIRE collections and in houses. There is also a record from outdoors: a single specimen on 1 Heracleum (EWA). NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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olgae Kalik, 1946

NO RECORDS

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Subgenus HELOCERUS Mulsant & Rey, 1868 fuscus Olivier, 1789

Anthrenus (Helocerus) fuscus Oliv ier, 1789

3 WATSONIAN (8,10,52,5,0) Fairly common – mainly out of doors on flowers and vegetation but 2 YORKSHIRE occasionally in houses. (iii-x). 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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Subgenus NATHRENUS Casey, 1900 verbasci (Linnaeus, 1767)

Anthrenus (Nathrenus) v erbasci (Linnaeus, 1767)

3 WATSONIAN (7,15,53,5,0) Fairly common and apparently, 2 YORKSHIRE at least as far as our records indicate, more often indoors than out (where it is recorded 1 from hogweed flowers and low vegetation). 0 NY NZ NZ OV SD SE SE TA (iii-xi). 9 8 7 6 5 4 3 2 1

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Subfamily Casey, 1900

TRINODES Dejean, 1821 hirtus (Fabricius, 1781)

NO RECORDS 45. Family BOSTRICHIDAE Latreille, 1802

45. Family BOSTRICHIDAE Latreille, 1802

Subfamily Latreille, 1802

BOSTRICHUS Geoffroy, 1762 capucinus (Linnaeus, 1758)

Bostrichus capucinus (Linnaeus, 1758)

3 WATSONIAN (1,0,2,1,0). Extinct. Rare with no recent records on the YNU database. The species is 2 YORKSHIRE likely to be extinct in Britain (Alexander, 1 2002:43) but is occasionally imported. Our four 0 NY NZ NZ OV SD SE SE TA records are: Leeds SE33, 1867, YNU; Hull TA12, 9 1892, YNU, in imported oak from Odessa; 8 Doncaster SE50, 4/1992, MLD, several specimens found in a Doncaster garden centre; 7 Sheffield SE3581, 3/1992, GM, male and 6 female emerged from wooden supports on 5 imported air plants. 4 3 2 1

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Subfamily Thomson, C.G., 1863

RHYZOPERTHA Stephens, 1830 dominica (Fabricius, 1792)

Rhyzopertha dominica (Fabricius, 1792)

3 WATSONIAN (2,1,2,0,0) GBES-Indoors. We have five records for this stored-products species: Scarborough 2 YORKSHIRE TA08, 1900, RL, in flour mills; Hull TA12, 1 31/10/1930, EW, in stored flour; Hull TA12, 0 NY NZ NZ OV SD SE SE TA 1932, TS, in flour mills; Barnsley SE30, 1911, 9 EGB; Bradford SE13, 1919, HBe. 8 7 6 5 4 3 2 1

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STEPHANOPACHYS Waterhouse, C.O., 1888 substriatus (Paykull, 1800) NO RECORDS

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Subfamily LYCTINAE Billberg, 1820

LYCTUS Fabricius, 1792 brunneus (Stephens, 1830)

Lyctus brunneus (Stephens, 1830)

3 WATSONIAN (0,3,6,0,0). Nationally Scarce. We have nine records, four of which specify emergence 2 YORKSHIRE from wood products including flooring, 1 wooden shelves, furniture, and a window 0 NY NZ NZ OV SD SE SE TA frame derived from native hardwood. 9 8 7 6 5 4 3 2 1

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linearis (Goeze, 1777)

Lyctus linearis (Goeze, 1777)

3 WATSONIAN (0,3,1,0,0) Nationally Scarce B. Our four (very old) records are: Barnsley (SE30), 1922, YNU, 2 YORKSHIRE in the wood of a cabinet; Egton NZ8106, 1 1934, HB, in a house; Whitby NZ8911, 0 NY NZ NZ OV SD SE SE TA 22/7/1934, HB, in large numbers in a house; 9 Middlesbrough NZ52, 1935, YNU. 8 7 6 5 4 3 2 1

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planicollis LeConte

Lyctus planicollis LeConte, 1858

3 WATSONIAN (0,0,1,0,0). Naturalised. We have a single 2 YORKSHIRE record from Yorkshire in the database: Smarson Hills SK5282, 31/3/1984, WAE, in a 1 bracket fungus. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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PTINIDAE Latreille, 1802

Subfamily LeConte, 1861

HEDOBIA Dejean, 1821

Subgenus Mulsant & Rey, 1868 imperialis (Linnaeus, 1767)

Hedobia (Ptinomorphus) imperialis (Linnaeus, 1767)

3 WATSONIAN (8,7,16,6,0) Nationally Scarce B. A saproxylic 2 YORKSHIRE which is widespread but scarce. May be beaten from old hedges, particularly holly Ilex 1 aquifolium and hawthorn , with few 0 NY NZ NZ OV SD SE SE TA recent records. (v-vi). 9 8 7 6 5 4 3 2 1

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Subfamily PTININAE Latreille, 1802

Tribe GIBBIINI Jacquelin du Val, 1860

GIBBIUM Scopoli, 1777 aequinoctiale Boieldieu, 1854

Gibbium aequinoctiale Boieldieu, 1854

3 WATSONIAN (0,0,6,0,0). GBES-Unknown. Synanthropic and rarely recorded. We have six records, 2 YORKSHIRE once in a house (origin of specimen 1 unknown) and five records from latrines 0 NY NZ NZ OV SD SE SE TA underground in coal mines, all records 9 dating from 1968 to 1994. No recent 8 records. 7 6 5 4 3 2 1

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psylloides (de Czenpinski, 1778)

Gibbium psylloides (de Czenpinski, 1778)

3 WATSONIAN (1,1,7,0,0) Synanthropic. Our nine records 2 YORKSHIRE include specimens from underground latrines at coal mines, in sugar in a confectioner’s 1 shop, and once in the toilet of a restaurant. 0 NY NZ NZ OV SD SE SE TA Some of our records are old (pre-1920) and 9 may be erroneous owing to confusion with G. 8 aequinoctiale. 7 6 5 4 3 2 1

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STETHOMEZIUM Hinton, 1943 squamosum Hinton, 1943

Stethomezium squamosum Hinton, 1943

3 WATSONIAN (0,0,1,0,0) We have a single record of this 2 YORKSHIRE little-known synanthropic species: Huddersfield (in a restaurant) SE1416, 1 8/2/1986, MLD. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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MEZIUM Samouelle, 1819 affine Boieldieu, 1856

Mezium affine Boieldieu, 1856

3 WATSONIAN (0,1,1,1,0) Synanthropic in houses and farm 2 YORKSHIRE buildings. Our three records include the following scant detail: Leeds SE33, unknown 1 date, YNU (recorder noted as “Abbot”); York 0 NY NZ NZ OV SD SE SE TA area SE65, 1891, YNU; Wakefield SE32, 1879, 9 GBW. 8 7 6 5 4 3 2 1

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Tribe PTININI Latreille, 1802

TRIGONOGENIUS Solier, 1849 globulus Solier, 1849

Trigonogenius globulus Solier, 1849

3 WATSONIAN (1,2,1,1,0) GBES-Unknown. Synanthropic and associated with stored products. We have 2 YORKSHIRE five records: Leeds City Museum SE3033, 1 8/10/2009, JB, on sticky trap in museum 0 NY NZ NZ OV SD SE SE TA store; Scarborough TA0488, 1950, EFG, in 9 dried figs and soya flour; York SE6153, 8 12/4/1975, HK, in archaeological storage warehouse; Hull TA1030, unknown date, 7 YNU, in museum collections; Bradford SE13, 6 1924, JWC, in warehouse. 5 4 3 2 1

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SPHAERICUS Wollaston, 1854 gibboides (Boieldieu, 1854)

NO RECORDS

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NIPTUS Boieldieu, 1856 hololeucus (Faldermann, 1835)

Niptus hololeucus (Faldermann, 1835)

3 WATSONIAN (4,8,15,5,0) Widespread but infrequent. Synanthropic and found variously in houses, 2 YORKSHIRE on sticky traps in warehouses, in pigeon lofts, 1 underground in a colliery, in a church, in 0 NY NZ NZ OV SD SE SE TA museum collections and once in a sparrow’s 9 nest in an attic. 8 7 6 5 4 3 2 1

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TIPNUS Thomson, C.G., 1859

unicolor (Piller & Mitterpacher, 1783)

Tipnus unicolor (Piller & Mitterpacher, 1783)

3 WATSONIAN (4,3,6,2,0). Widespread, synanthropic and 2 YORKSHIRE scarce. Recorded in Yorkshire from pigeon droppings and in pigeon cotes, in henhouse 1 refuse, in old straw in stables, in oat rubbish 0 NY NZ NZ OV SD SE SE TA in barn, in nest of Formica lugubris and in 9 pellets of Barn Owl. 8 7 6 5 4 3 2 1

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PSEUDEUROSTUS von Heyden, L., 1906 hilleri (Reitter, 1877) hilleri (Reitter, 1877)

3 WATSONIAN (0,0,1,0,0). An established alien. We have a 2 YORKSHIRE single record on the database: Netherton SE1213, 25/11/1983, MLD, in discarded sacks 1 of rotting oats, barley and broad beans. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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PTINUS Linnaeus, 1767 clavipes Panzer, 1792

NO RECORDS – peat subfossil only dubius Sturm, 1837

NO RECORDS exulans Erichson, 1842

NO RECORDS

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fur (Linnaeus, 1758)

Ptinus fur (Linnaeus, 1758)

3 WATSONIAN (12,7,13,1,0) Widespread and fairly frequent. Synanthropic. Recorded from a ’s nest, 2 YORKSHIRE in sticky traps in warehouses, in houses and 1 attics, in pigeon cotes, in farmyard debris, in 0 NY NZ NZ OV SD SE SE TA dry dog biscuits, and in the peat subfossil 9 record (Thorne Moor archaeological surveys). 8 (i-xii). 7 6 5 4 3 2 1

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latefasciatus Gorham, 1883

NO RECORDS

lichenum Marsham, 1802

Ptinus lichenum Marsham, 1802

3 WATSONIAN (0,0,2,0,0) RedList_pre94-NR. We have two records in the YNU database: Kings Wood 2 YORKSHIRE Elsecar SK39Z, 4/1998, CH; Netherton 1 SE1213, 20/11/1983, MLD, in discarded sacks 0 NY NZ NZ OV SD SE SE TA of rotting oats, barley and broad beans. 9 8 7 6 5 4 3 2 1

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palliatus Perris, 1847

NO RECORDS

pusillus Sturm, 1837

Ptinus pusillus Sturm, 1837

3 WATSONIAN (0,0,1,0,0) A single record exists in the YNU database: Austerfield SK6695, 1977, CH det 2 YORKSHIRE PS, in Barn Owl pellets. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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raptor Sturm, 1837

NO RECORDS

sexpunctatus Panzer, 1792

Ptinus sexpunctatus Panzer, 1792

3 WATSONIAN (0,3,5,1,0) Nationally Scarce B. Widespread and scarce. With nine records from five sites. 2 YORKSHIRE This species is associated with solitary bees 1 with nests in lawns and buildings. The 0 NY NZ NZ OV SD SE SE TA species is perhaps becoming more 9 commonly met with nationally – see Mendel 8 (2017). Recorded in Yorkshire in sticky traps in a warehouse, in a suburban garden, in the 7 upstairs room of a house and in an MV moth 6 trap. 5 4 3 2 1

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subpilosus Sturm, 1837

Ptinus subpilosus Sturm, 1837

3 WATSONIAN (0,0,2,0,0) With only two records known in Yorkshire, this species is a rarity: Barnsley 2 YORKSHIRE SE30, undated (therefore not mapped), GBW, 1 noted as ‘introduced’ but without further 0 NY NZ NZ OV SD SE SE TA details; Cadeby SE5201, 16/7/1966, PS. 9 8 7 6 5 4 3 2 1

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tectus Boieldieu, 1856

Ptinus tectus Boieldieu, 1856

3 WATSONIAN (5,13,20,3,0) GBES-Indoors. Widespread mainly in the southern half of the county 2 YORKSHIRE and a fairly frequent species. Recorded from 1 sticky traps, in wheat debris, in ground rice, 0 NY NZ NZ OV SD SE SE TA in a flour mill, among rolled oat rubbish in a 9 barn, and in domestic premises. 8 7 6 5 4 3 2 1

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villiger Reitter, 1884

NO RECORDS

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Subfamily DRYOPHILINAE Gistel, 1848

GRYNOBIUS Thomson, C.G., 1859 planus (Fabricius, 1787)

Grynobius planus (Fabricius, 1787)

3 WATSONIAN (4,18,9,6,1) Widespread and saproxylic, and 2 YORKSHIRE rather infrequent. Recorded from dead or moribund trees in parkland, neglected 1 churchyards and old hedgerows, trees 0 NY NZ NZ OV SD SE SE TA including lime, , oak, holly and apple. 9 (v-vii). 8 7 6 5 4 3 2 1

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DRYOPHILUS Chevrolat, 1832 anobioides Chevrolat, 1832

NO RECORDS pusillus (Gyllenhal, 1808)

Dryophilus pusillus (Gyllenhal, 1808)

3 WATSONIAN (5,22,17,3,5) Naturalised. Widespread and 2 YORKSHIRE fairly frequent. Reported from oaks and various conifers including larch and Scots 1 Pine by beating the foliage and also under 0 NY NZ NZ OV SD SE SE TA bark of oak, beaten from old hedgerows, in 9 a Malaise trap in a suburban garden and in a 8 MV moth trap. (vi-viii). 7 6 5 4 3 2 1

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Subfamily Pic, 1912

OCHINA Dejean, 1821 ptinoides (Marsham, 1802)

Ochina ptinoides (Marsham, 1802)

3 WATSONIAN (0,2,5,4,0) A widespread saproxylic species in a central north-south band in the county. 2 YORKSHIRE Scarce, with 11 known records. Recorded 1 from ivy on old trees and on 0 NY NZ NZ OV SD SE SE TA walls in neglected churchyards. The beetle 9 seeks out aged plants and bores and develops 8 in the thick stems. 7 6 5 4 3 2 1

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XESTOBIUM Motschulsky, 1845 rufovillosum (De Geer, 1774)

Xestobium rufov illosum (De Geer, 1774)

3 WATSONIAN (5,8,8,6,1) The . 2 YORKSHIRE Saproxylic. Centrally distributed in the county and possibly declining owing to the 1 increasing scarcity of its habitat – mainly old 0 NY NZ NZ OV SD SE SE TA oak dead heartwood where decay is present. 9 The species occurs in old buildings either 8 where the population has been imported in 7 the original wood, or is attracted to the finished timbers by the presence of attractive 6 fungi species (especially Donkiopora expansa 5 – see Alexander, 2002:44). The larval period 4 of the life cycle may last for up to 13 years. 3 2 1

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ERNOBIUS Thomson, C.G., 1859 abietis (Fabricius, 1792)

NO RECORDS

angusticollis (Ratzeburg, 1837)

NO RECORDS

gigas (Mulsant & Rey, 1863)

NO RECORDS

mollis (Linnaeus, 1758)

Ernobius mollis (Linnaeus, 1758)

3 WATSONIAN (5,8,13,3,2) Saproxylic. Very widespread but scarce. Mainly reported from Scots Pine in 2 YORKSHIRE Yorkshire but also recorded by beating a mixture 1 of oak, and hazel. The larvae feed in the 0 NY NZ NZ OV SD SE SE TA cambium layer beneath the bark. (iv-ix). 9 8 7 6 5 4 3 2 1

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nigrinus (Sturm, 1837)

Ernobius nigrinus (Sturm, 1837)

3 WATSONIAN (1,2,0,0,0) Very rare in the county with only three known records: Cloughton TA19, 2 YORKSHIRE 11/8/1925, GBW, 2 exx on felled pines; 1 Allerthorpe SE74, 1931, WDH, on Scots 0 NY NZ NZ OV SD SE SE TA Pine; Wykeham Forest SE8794, 12/7/2000, 9 LA (no other details – this record requires 8 confirmation). 7 6 5 4 3 2 1

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pini (Sturm, 1837)

NO RECORDS

Subfamily Fleming, 1821

STEGOBIUM Motschulsky, 1860 paniceum (Linnaeus, 1758)

Stegobium paniceum (Linnaeus, 1758)

3 WATSONIAN (2,7,30,3,0) GBES-Indoors. Widespread and fairly commonly encountered. Reported 2 YORKSHIRE from inside domestic premises, in dried pet 1 food, in pigeon dung in lofts and in sticky 0 NY NZ NZ OV SD SE SE TA traps in warehouses. (i-xii). 9 8 7 6 5 4 3 2 1

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GASTRALLUS Jacquelin du Val, 1860 immarginatus (Müller, P.W.J., 1821)

NO RECORDS laevigatus (Olivier, 1790)

NO RECORDS

ANOBIUM Fabricius, 1775 fulvicorne (Sturm, 1837)

Anobium fulv icorne (Sturm, 1837)

3 WATSONIAN (8,5,34,8,0) Saproxylic. Widespread in the central part of the county and commonly 2 YORKSHIRE encountered. Adults recorded mainly by 1 general sweeping in suitable habitats where 0 NY NZ NZ OV SD SE SE TA the larvae develop in broadleaved trees, in 9 small dying or dead branches. Has been 8 collected from a Malaise trap in a suburban 7 garden. Records indicate an increase and frequency in range. (v-viii). 6 5 4 3 2 1

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inexspectatum Lohse, 1954

Anobium inexspectatum Lohse, 1954

3 WATSONIAN (1,0,3,1,1) Nationally Scarce B. Saproxylic. Rarely recorded, with five known instances: 2 YORKSHIRE Drax SE6627, 10/6/1997, PK det MLD; 1 Braithwaite Hall SE1186, 17/8/1987, KNAA; 0 NY NZ NZ OV SD SE SE TA Manor Farm SE771665, MH det MLD; Pot 9 Ridings SE5200, 9/2002, MLD; Pugney’s 8 SE333175, 5/6/2008, MLD. Develops in old 7 ivy Hedera stems. Described in 1954, specimens have been found in British 6 collections of A. punctatum dating from the 5 1930s (Allen, 1976). Any “Anobium 4 punctatum” from old ivy Hedera should be 3 checked carefully. 2 1

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nitidum Fabricius, 1792

NO RECORDS

punctatum (De Geer, 1774)

Anobium punctatum (De Geer, 1774)

3 WATSONIAN (22,12,43,14,1) The Furniture Beetle or Woodworm. Saproxylic. Widespread and 2 YORKSHIRE commonly encountered, in the dead 1 sapwood of living and dead softwood and 0 NY NZ NZ OV SD SE SE TA hardwood trees. Will attack finished timber 9 products especially if fungal decay is present. 8 7 6 5 4 3 2 1

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HADROBREGMUS Thomson, C.G., 1859 denticollis (Creutzer in Panzer, 1796)

NO RECORDS

PRIOBIUM Motschulsky, 1845 carpini (Herbst, 1793)

NO RECORDS

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Subfamily Shuckard, 1839

PTILINUS Geoffroy, 1762 pectinicornis (Linnaeus, 1758)

Ptilinus pectinicornis (Linnaeus, 1758)

3 WATSONIAN (19,32,90,22,3). Saproxylic. Widespread with 2 YORKSHIRE many records from a wide variety of old broadleaved tree species, often in beech but 1 also in ash, and oak, mainly in the dry 0 NY NZ NZ OV SD SE SE TA heartwood. In woods, parkland and 9 hedgerows. (iii-x, peak vii). 8 7 6 5 4 3 2 1

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Subfamily Gistel, 1848

XYLETINUS Latreille, 1809 longitarsis Jansson, 1942

NO RECORDS

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LASIODERMA Stephens, 1835 serricorne (Fabricius, 1792)

Lasioderma serricorne (Fabricius, 1792)

3 WATSONIAN (0,0,2,0,0) Rarely recorded with only two Yorkshire records: Doncaster SE50, 1974, PS; 2 YORKSHIRE Doncaster SE50, 1975, PS. Both records are 1 backed up by specimens in the Doncaster 0 NY NZ NZ OV SD SE SE TA Museum collections, and these specimens 9 were passed to the Museum by the Local 8 Authority Health Department. The 7 circumstances of the captures are unknown. The beetle is primarily a pest of tobacco and its 6 products, but may be noted from other stored 5 products such as cereals. 4 3 2 1

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Subfamily Thomson, C.G., 1859

MESOCOELOPUS Jacquelin du Val, 1860 collaris Mulsant & Rey, 1864

NO RECORDS

DORCATOMA Herbst, 1792 ambjoerni Baranowski, 1985

NO RECORDS chrysomelina Sturm, 1837

Dorcatoma chrysomelina (Sturm, 1837)

3 WATSONIAN 1,3,4,9,0) Scarce in Yorkshire, centrally 2 YORKSHIRE distributed and with few recent records. This saproxylic develops in oak Quercus and 1 ash Fraxinus boughs. The fungus Laetiporus 0 NY NZ NZ OV SD SE SE TA sulphureus seems to be a common factor in 9 some records. In old orchards, and ancient 8 parkland sites such as Temple Newsam, 7 Studley Royal and Beningbrough Hall. (iii- viii). 6 5 4 3 2 1

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dresdensis Herbst, 1792

Dorcatoma dresdensis Herbst, 1792

3 WATSONIAN (4,3,2,0,0) Nationally Scarce A. Rare with 2 YORKSHIRE only nine records for the county. Several records mention Fomes on beech Fagus 1 sylvatica. NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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flavicornis (Fabricius, 1792)

Dorcatoma flav icornis (Fabricius, 1792)

3 WATSONIAN (1,4,5,5,0) Nationally Scarce B. Scarce in 2 YORKSHIRE Yorkshire with 15 known records. Recorded from Corylus sp., Malus sp., Salix fragilis, in old 1 Quercus robur. Fungal associations include 0 NY NZ NZ OV SD SE SE TA Piptoporus betulinus and Laetiporus 9 sulphureus. 8 7 6 5 4 3 2 1

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substriata Hummel, 1829

Dorcatoma substriata Hummel, 1829

3 WATSONIAN (0,4,0,0,0) Nationally Scarce A. Rare in 2 YORKSHIRE Yorkshire with only four certain records: Duncombe Park and Plockwoods Bank Wood 1 areas (SE6082, SE6083, SE6182), 2005-2006, 0 NY NZ NZ OV SD SE SE TA KNAA, in Inonotus dryadeus on Quercus sp.. [A 9 fifth record from Fenwick Churchyard SE5815 8 requires confirmation, beaten from Herera 7 helix on Fraxinus so may be erroneous.] 6 5 4 3 2 1

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CAENOCARA Thomson, C.G., 1859

affinis (Sturm, 1837)

NO RECORDS

bovistae (Hoffmann, J., 1803)

NO RECORDS

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ANITYS Thomson, C.G., 1863 rubens (Hoffmann, J., 1803)

Anitys rubens (Hoffmann, J., 1803)

3 WATSONIAN (0,1,3,3,0) Nationally Scarce B. Rare in the county with seven records. In red-rotted 2 YORKSHIRE heartwood of old oaks Quercus sp.. 1

NY NZ NZ OV 0 SD SE SE TA 9 8 7 6 5 4 3 2 1

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MIROSTERNOMORPHUS Español, 1977 heali Bercedo & Arnáiz, 2010

NO RECORDS

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Superfamily LYMEXYLOIDEA Fleming, 1821 47. Family LYMEXYLIDAE Fleming, 1821

47. Family LYMEXYLIDAE Fleming, 1821

Subfamily HYLECOETINAE Germar, 1818

HYLECOETUS Latreille, 1806 dermestoides (Linnaeus, 1761)

Hylecoetus dermestoides (Linnaeus, 1761)

3 WATSONIAN (8,17,37,12,2) Nationally Scarce B. 2 YORKSHIRE Widespread and fairly frequent in a central north-south band in Yorkshire. In a wide 1 range of softwood and hardwood species. 0 NY NZ NZ OV SD SE SE TA Develops in the heartwood. A mainly 9 northern species in Britain and . (iv-ix). 8 7 6 5 4 3 2 1

SD SE SE TA 0 SJ SK SK TF 9 8 post 31/12/1999 Map produced for Yorkshire Naturalists' Union by DMAP 13/1/2018 pre 31/12/1999 6 7 8 9 0 1 2 3 4 5 6 7 8 9 0 1 2 3 4

Subfamily LYMEXYLINAE Fleming, 1821

LYMEXYLON Fabricius, 1775 navale (Linnaeus, 1758) NO RECORDS

Atlas of Yorkshire Coleoptera - part 9 – Derodontidae to Lymexylidae 40 | P a g e

Acknowledgements I am grateful to Mike Denton for proof reading the manuscript and identifying errors.

References

Alexander, K.N.A., 2002. English Nature Reports 467. The invertebrates of living & decaying timber in Britain and Ireland – a provisional annotated checklist.

Allen, A.A., 1976. Notes on some British Serricorn Coleoptera, with adjustments to the list. 3. A new British Anobium with notes on three others of the family. EMM 112 (1976):151-154.

Cooter, J., 1999. On ‘English’ names for British Coleoptera. Coleopterist 8 (1): 43-45.

Duff, A.G., (ed.), 2008 Checklist of the of the British Isles, 2008 edition. A.G. Duff publishing, Wells, Somerset.

Duff, A.G., (ed.), 2012 Checklist of Beetles of the British Isles, 2nd edition. Pemberley Books.

Hyman, P.S. and Parsons, M.S., 1994. A review of the scarce and threatened Coleoptera of Great Britain, part 2. JNCC, Peterborough.

Hyman, P.S., and Parsons, M.S., 1992. A review of the scarce and threatened Coleoptera of Great Britain, part 1. JNCC, Peterborough.

Key, R.S., 1999. More on ‘noddy names’ for British beetles. Coleopterist 8 (2): 94.

Mendel, H., 2017. Is Ptinus sexpuntatus Panzer () becoming more common as a result of ‘bee hotels’ in gardens? Coleopterist 26(3): 156-157.

Peacock, E.R., 1993. Adults and Larvae of Hide, Larder and Carpet Beetles and their relatives (Coleoptera: Dermestidae) and of Derodontid Beetles (Coleoptera: Derodontidae). Handbooks Ident. Brit. volume 5(3), pp. 144. Royal Entomological Society, London.

Pope, R.D., 1977. Kloet & Hincks. A Check List of British Insects. Part 3: Coleoptera and Strepsiptera. Second revised edition. Handbooks Ident. Brit. Insects volume 11(3), pp. xiv+105. Royal Entomological Society, London.

Errors and omissions

The author would be pleased to receive feedback regarding any errors or omissions noted by users of this Atlas.

R J Marsh FRES YNU Coleoptera recorder 1/5/2018

Atlas of Yorkshire Coleoptera - part 9 – Derodontidae to Lymexylidae 41 | P a g e