Agriculture I*l Canada

A catalog of Nearctic

A catalog of Catalogue des Nearctic Chironomidae Chironomidae delardgion ndarctique

D.R. Oliver and M.E. Dillon D.R. Oliver et M.E. Dillon Biosystematics Research Centre Centre de recherches biosyst6matiques Ottawa, Ontario Ottawa (Ontario) K1A 0C6 K1A 0C6 and et P.S. Cranston P.S. Cranston Commonwealth Scientific and Organisation de la recherche Industrial Research scientifique et industrielle du Organization, Entomology Commonwealth, Entomologie Canberra ACT 2601 Canberra ACT 2601 Australia Australie

Research Branch Direction g6n6rale de la recherche Agriculture Canada Agriculture Canada

Publication 185718 Publication 185718 1 990 1 990 @Minister of Supply and Services Canada 1990 oMinistre des Approvisionnement et Services Canada 1990 Available in Canada through En vente au Canada par I'entremise de nos Authorized Bookstore Agents agents libraires agr66s et autres and other bmkstores libraires. or by mail from ou par la poste au

Canadian Govemnent Publishing Centre Centre d'6dition du gouvemement du Supply and Servies Canada Canada Oltawa, Canada K1A 0S9 Approvisionnements et Seryies Canada Ottawa (Canada) K1A 0S9

Cat No. A43-I85'7ll99O N" de cat A43-785117990 ISBN 0-660-55839-4 ISBN 0-660-55839-4

Price subject to change without notic€ Prix sujet i changemenl sans pr6avis

Canadian Cataloguing in Publication Data Donn6ee de catalogage avant publication (Canada)

Oliver, D.R. Oliver, D.R.

A mtalog of Nearctic Chironomidae A atalog of Nearctic Chironomidae

(Publication ; 1857/8) (Publiation ; 18578)

Text in English and French- Texle en anglais et en frangais. Includes bibliographiel referenes. Comprend des r6f6rences bibliogr. Issued by Research Branch, Agriculture Canada. Publi6 par Direction g6n6rale de la recherche, Cat No. A43-1857/19908 Agriculture Canada ISBN 0-660-55839-4 N' de cal A43-1857/1990F ISBN 0-660-55839-4

1. Chironomidae--Ameria. 1. Chironomidae--Am6rique. 2. Chironomidae-- 2. Chironomidae--Nomenclature. I. Dillon, M.E Nonenclature. I. Dillon, M.E. II. Cranston, II. Cranston, P.S. III. Canada. Agriculture P.S. III. Canada. Agriculture Canada. IV. Canada. IV. Title. V. Title: Catalogue des Titre. V. Titre: C:talogue des Chironomidae de Chironomidae de la r6gion n6arctique. VI. la r6gion n6arctique. VI. Titre: Chironomidae Title: Nearctic Chironomidae. VII. Series: de la rEgion n6arctique. VII. Clllection: PubUcation (Canada. Agriculture Canada). Publiation (Canada. Agriculture Canada). English ; 1857,18. Franqais ; 1857,8.

QL537.C45604 1990 595.77'7 C90-099ttt-98 QL537.C45604 1990 595.77',1 C90-099111-9F

Cover Couverture Adult male of Chirnmw pfumm (Linnaeus) M6le adulte du Chironomw plumm (Linnaeus)

Produced by Research Program Senrice Production du Seruie aux programmes de recherches TABLE OF CONTENTS TABLE DES MATIERES

INTRODUCTION...... 1 INTRODUCTION ...... 4

FORMAT ,,...1 MODEDEPRESENTATION ...... 4 ABBREVIATIONS .....2 ABREVIATIONS ....,.5 ACKNOWLEDGMENTS ...... 3 REMERCIEMENTS ....6

FamilyCHIRONOMIDAE ...... 7 Famille des CHIRONOMIDAE ...... 7

Subfamily ...... 7 Sous-famille des TELMATOGETONINAE . . . 7

SubfamilyPODONOMINAE ...... 7 Sous-famille des PODONOMINAE ...... 7

TTibeBOREOCHLINI ...... 7 Tribu des BOREOCHLINI ...... 7

TTibePODONOMINI ...... 8 Tribu desPODONOMINI ...... 8

SubfamilyTANYPODINAE ...... 8 Sous-famille des ...... 8

Tribe COELOTANYPODINI . . .. 8 Tribu des COELOTANYPODINI ...... 8

TTibeMACROPELOPIINI ...... 8 Tribu des MACROPELOPIINI ...... 8

TTibeNATARSIINI ...10 TTibUdesNATARSIINI ..... 10

TTibePENTANEURINI ...... 10 Tribu des ...... 10

TTibePROCLADIINI ...... 74 Tribu des ...... 14

Tribe TANYPODINI ...... 16 Tribu desTANYPODINI ..... 16

SubfamilyDIAMESINAE ...16 Sous-famille des DIAMESINAE ...... 16

Tribe BOREOHEPTAGYIINL ...... '16 Tribu des BOREOHEPTAGYIINI . . . . . 16

TTibeDIAMESINI ...... 16 Tribu desDIAMESINI ...... 16

TTibePROTANYPINI ...... 19 Tribu des PROTANYPINI ...... 19

SubfamiIyPRODLAMESINAE ..... 19 Sous-famille des PRODIAMESINAE ...... 19

SubfamilyORTIIOCLADIINAE . ... 20 Sous-famille des ORTHOCLADIINAE . . . . . 20

SubfamilyCHIRONOMINAE ...... 40 Sous-famille des ...... 40

TTibeCHIRONOMINI ...... 40 Tribu des CHIRONOMINI ...... 40

Tribe PSEUDOCHIRONOMINI ...... 55 Tribu des PSEUDOCHIRONOMINL . . 55

TTibeTANYIARSINI ...... 55 Tribu des TANYTARSINI ...... 55

REFERENCES ,.., ... 60 BIBLIOGRAPHIE .. ... 60 rNDEX .... 76 INDEX ... 76

INTRODUCTION Adults of the family Chironomidae are Subtette 0[ iver, Di [ [on commonly called chironomids, , and and nonbiting midges, or chironomid midges, and Subtette Cranston 't965 1990 the red larvae of some groups are called bloodworms. Chironomid larvae occur in all G ene Fa 205 types of aquatic and semiaquatic habitats and to a lesser extent in brackish and marine Spec i es 601 1 051 waters, and semiterrestrial and terrestrial habitats. Aquatic ecosystems are usually dominated by larval chironomids, in terms of The need for an updated catalog of Nearctic number of species, biomass, or both. They Chironomidae is evident. are important in the breakdown of organic Accordingly we have prepared a catalog of material and recycling nutrients, as food for the taxa of Chironomidae recorded from the fish, birds, and both terrestrial and aquatic Nearctic region, with information on their invertebrates, and as predators of aquatic distribution and revelent literature. ThiS invertebrates. Some species can tolerate the catalog is based only on published data to the most polluted conditions, whereas others have end of 1989; it does not include unpublished very special requirements and quickly data in collections, such as the Canadian disappear when their habitat is stressed. National Collection of . We have Thus the Chironomidae have exceptional attempted to list all published names, valid or potential to act as biological indicators of not, that have been used in the literature aquatic environmental health. pertaining to the Nearctic region. A few The family is distributioned around the older faunal lists (Leonard 1928; Johnson world, occurring in all zoogeographical 1925; Brimley 1938; Petch and Maltais 1932) regions, including Antarctica. Currently 10 are not included because they contain many subfamilies and 24 tribes (Ashe 1983) are obviously incorrect determinations. As the recognized, of which 7 subfamilies and 14 taxa are presented uncritically in their most tribes occur in the Nearctic region. A recently published taxonomic position, it minimum of 5000 species are found around should not be construed that we are in the world and at least 2000 of these in the agreement with all the taxonomic placements. Nearctic region. This catalog will be continuously updated The nonspecialist was faced, until recently, with periodic publication of additions and with two generic classifications. This dual corrections. To assist us in keeping an system arose during the first half of this updated list, we request any errors or century, because one group of workers omissions, and copies of publications with principally emphasized the immature stages information on and distribution of and the other emphasized adults. Pinder Nearctic Chironomidae. be sent to either of (1983) gives a brief history of the the first two authors. development of the two systems and their reconciliation. Reconciliation of the two FORMAT systems has culminated in the classification The arrangement of names is alphabetical, used in the three volumes of the except for the subfamily-group names which Chironomidae of the Holarctic region edited follow the phylogenetic arrangement of the by Wiederholm (1983, 1986, 1989) and that Holarctic Chironomidae volumes. Species- classification is used herein. group names follow the subfamily, tribe, The last catalog of Nearctic Chironomidae , and subgenus headings (not all groups (Sublette and Sublette 1965) was published have been assigned tribes or subgenera). The about 25 years ago. The intervening years first occurrence of all valid names are printed has been a period of intense chironomid in boldface type along the left margin of each research, including much taxonomic and column. Valid taxa that have been ecological study. The following table shows incorrectly reported from the Nearctic region the increase in the number of valid generic- are also listed along the left margin but they and species-level names between 1965 and are printed in italics as are all other listing of 1989. names, valid or invalid. Citations for genus-group names are arranged as in the following example. NEARCTIC CHIRONOMIDAE

Genus NANOCLADIUS Kieffer original binomial citation is next, followed by Nanocladius Kieffer, 1913d: 31. subspecies or varieties and then synonyms (if Type-species, vitellinus Kieffer any), which are cited in their original form (orig. des.). with author, date, page, and type-locatity. Microcric otopas Thienemann & Type-localities for Canada and the United Ilarnisch, 1.932: 1.37. States are listed by province, territory or Type-species, Cricotopus state; all others are listed by country or al b icornis Goetghebuer larger geographical regions where country is (Goetghebuer in Goetghebuer & not known. Lenz, 1944: 114) = bicolor Zetterstedt. ABBREVIATIONS The abbreviations used are as follows: Subgenus Nanocladius Kieffer General &- and The original citation is given in the second o., E.- east, eastern line, indented, followed by synonyms (if any), emend.- emended also indented. The genus name is followed I., Is.- Island, Islands by the original author, date, page, name of ICZN- International Commission type-species, and nature of type fixation. The on Zoological Nomenclature originat type-species combination is cited, but misident.- misidentification the generic name is omitted if it is the same mon.- monotypy as the genus for which it is the type. If the nam.- name original type-species is now a junior synonym, n. nam.- new name it is given at the end following an equal nom. nud.- nomen nudum symbol (=). Only the authors of subsequent orig. des.- original designation type fixation are cited. Much of the preocc.- preoccupied information about type fixation and synonyms s., S.- south, southern has been obtained from Ashe (1983). This se., Se.- southeast, southeastern publication has detailed information on sub. mon.- subsequent monotypy generic synonymy, type-species, and subg.- subgenus nomenclatorial problems. Subgeneric-name syn.- synonym, synonymy citation format follows that of the generic var.- variety citations. Citations for nominate subgenera w.- west, western are omitted when they are the same as the generic citations; otherwise they follow the Geographical Names format of generic citations. AF.- Afrotropical Region Species-group names are arranged as Ala.- Alabama follows: Alas.- Alaska Alta.- Alberta triannulatus. Sask., Ont. to N.Y., s. to Amer.- America Ky. and Del.; PA. Ariz.- Arizona Chironomus triannulatus Macquart, Ark.- Arkansas 7826: 202.-Germany. AU.- Australasian Region Cricotopus exjlis Johannsen, 1905: Baja Calif.- Baja California 25.5.-N.Y. B.C.- British Columbia Calif.- California The species name is followed by a synoptic Can.- Canada account of the Nearctic region distribution Colo.- Colorado stated by territory, province, or state in a Conn.- Connecticut northwest to northeast and southwest to D.C.- District of Columbia southeast pattern, except for Greenland. Del.- Delaware Occurrence in other zoogeographical regions Fla.- Florida are listed next, separated by a semicolon. Ga.- Georgia When a Mexican locality record occurs near Grld.- Greenland the uncertain division between Nearctic and Ill.- Illinois Neotropical regions, it is listed as occurring in Ind.- Indiana both Mexico and the Neotropical region. The Kans.- Kansas Ky.- Kentucky CHIRONOMIDAE DE LA REGION NEARCNQUE

Ky.- Kentucky ACKNOWLEDGMENTS La.- Louisiana We gratefully acknowledge Mr. B. Labr.- Labrador Caldwell for checking several drafts. The late Man.- Manitoba Dr. Sam Roback checked an early draft of Mass.- Massachusetts the subfamily Tanypodinae. We thank Mr. L. Md.- Maryland Speers for formatting the manuscript for Mex.- Mexico printing, and Ms. L. Robillard for typing Mich.- Michigan some of the references. Minn.- Minnesota Miss.- Mississippi Mo.- Missouri Mont.- Montana N.B.- New Brunswick N.C.- North Carolina N. Dak.- North Dakota NE.- Nearctic Region Nebr.- Nebraska Nev.- Nevada Nfld.- Newfoundland N.H.- New Hampshire N.J.- New Jersey N. Mex.- New Mexico N.S.- Nova Scotia NT.- Neotropical Region N.W.T.- Northwest Territories N.Y.- New York Okla.- Oklahoma Ont.- Ontario OR.- Oriental Region Oreg.- Oregon PA.- Palaearctic Region Pa.- Pennsylvania P.E.L- Prince Edward Island Qu€.- Qudbec R.I.- Rhode Island Sask.- Saskatchewan S.C.- South Carolina S. Dak.- South Dakota Tenn.- Tennessee Tex.- Texas U.S.A.- United States of America U.S.S.R.- Union of Soviet Socialist Republics Va.- Virginia Vt.- Vermont Wash.- Washington Wis.- Wisconsin W. Va.- West Virginia Wyo.- Wyoming Y.T.- Yukon Territory NEARCTIC CHIRONOMIDAE

INTRODUCTION suivant montre la multiplication des noms Les adultes de la famille des chironomidds valide au niveau des genres et des espCces sont communement appeles chironomes ou entre 1965 et 1989. moucherons et les larves d'un rouge sang de certains groupes sont les vers de vase utilisds par les pOcheurs. On trouve des larves de Subtette 0[iver, Ditton chironomidds dans tous les types d'habitats et et Cranston aquatiques et semi-aquatiques et, dans une Subtette 1965 1990 moindre mesure, dans les eaux saumAtres ou marines de m€me que dans les habitats semr- Genres 205 terrestres et terrestres. Les 6cosystdmes aquatiques sont habituellement domin6s par EspCces 601 1 051 les formes larvaires, en ce qui touche le nombre d'espdces, la biomasse ou les deux. Ces insectes jouent un r0le important dans la La n€cessitd d'un inventaire i jour des d6gradation de la matidre organique et le chironomid6s est dvidente. recyclage des 6l6ments nutritifs ou comme C'est pourquoi nous avons dressd un nourriture pour les poissons, les oiseaux ainsi rdpertoire des taxons des chironomid6s dont que les invertdbr6s tant terrestres la prdsence a 6t6 signal6e dans la rdgion qu'aquatiques ou encore comme pr€dateurs ndarctique, assorti d'informations sur leur des invertdbrds aquatiques. Certaines espdces repartition et la documentation pertinente. peuvent toldrer des conditions de pollution Ce catalogue n'est fond6 que sur des donndes extreme, alors que d'autres, trds sp6cialisdes, publides jusqu'd la fin de 1989 et ne disparaissent rapidement quand leur habitat comprend pas les donndes non publides que est menac6. Les chironomidds possddent l'on trouve dans les collections comme donc un potentiel exceptionnel comme l'Insectarium national canadien. Nous avons indicateurs biologiques de la santd du milieu essayd d'€num6rer tous les noms publies, aquatlque. valides ou pas, qui ont ete mentionn6s dans Cette famille, de rdpartition mondiale, est la documentation relative d la rdgion prdsente dans toutes les rdgions ndarctique. Quelques vieilles listes fauniques zoog6ographiques, y compris l'Antarctique. (Leonard 1928; Johnson 1925; Brimely 1938; On reconnait actuellement 10 sous-familles et Petch et Maltais 1932) n'ont pas 6td prises en 24 tribus (Ashe 1983), dont 7 sous-familles et consideration parce qu'elles contiennent de 14 tribus dans la rdgion nearctique. La nombreuses appellations qui sont hors de tout planete compte un minimum de doute incorrectes. Comme les taxons sans 5 000 espOces dont au moins 2 000 dans cette commentaires dans leur dernidre position rdgion-ld. taxinomique publi6e, il ne faudrait pas en Jusqu'A recemment, le profane se conclure que nous y souscrivons dans tous les retrouvait devant deux classifications cas. gdndriques. Ce double systdme a pris forme Le pr6sent inventaire sera continuellement au cours de la premidre moitid du sidcle, un tenu d jour par la publication pdriodique groupe de chercheurs ayant mis I'accent sur d'ajouts et de corrections. Pour nous aider les stades immatures et I'autre sur les dans cette teche, nous vous saurions grd de adultes. Pinder (1983) trace un bref faire part d l'un ou I'autre des deux premiers historique du ddveloppement des deux auteurs nomm6s des erreurs ou des omissions systemes et de leur rapprochement. La qui auraient pu s'y glisser et de leur envoyer reconciliation des deux systemes a atteint un des exemplaires de publications contenant de point culminant dans la classification utilis€e I'information sur la taxinomie et la 16oartition dans les trois volumes de Chironomidae of des chironomidds. the Holarctic region 6,dit€ par Wiederholm (1983, 1986, 1989), et c'est cette classification MODE DE PRESENTATION que I'on utilise dans le pr6sent ouvrage. Les noms apparaissent en ordre Le dernier inventaire des chironomid6s alphab€tique, sauf que ceux des groupes de nearctiques (Sublette et Sublette 1965) a 6t6 sous-familles suivent I'organisation publid il y environ 25 ans. Les anndes qui phylog6nique des volumes sur les suivirent furent une pdriode de recherches chironomid€s holarctiques. Les noms des intenses sur les chironomid6s, surtout leur groupes d'espdces suivent les titres des sous- taxinomie et leur €coloeie. Le tableau familles, tribus, genres et sous-genres (a CHIRONOMIDAE DE I-A REGION NEARCNQUE

noter que ce ne sont pas tous les groupes qui Chironomus ffiannulatus Macquart, se sont fait assigner des tribus et des sous- 1826: 202.-Germany. genres). La premidre fois qu'un nom valide Cricotopus exilis Johannsen, 1905: apparait, il est imprim6 en caractires gras 255.-N.Y. prds de la marge de gauche de chaque colonne. Les taxons valides qui ont 6t€ Le nom de I'espdce est suivi d'un compte signalds a tort comme de la rdgion n6arctique rendu synoptique de la rdpartition dans la sont aussi €numdr€s le long de la marge de rdgion ndarctique indiqude par le territoire, la gauche, mais en italique, comme le sont province ou I'Etat, dans le sens nord-ouest a d'ailleurs toutes les autres listes de noms, nord-est et sud-ouest d sud-est, sauf pour le valides ou non. Groenland. L'apparition de l'espdce dans Les citations pour les noms de groupes de d'autres rdgions zoog€,ographiques est genres se pr€sentent comme suit : 6num€r6e ensuite et sdpar6e par un point- virgule. Quand l'apparition d'une espdce est Genus NANOCLADIUS Kieffer indiqude dans une r6gion du Mexique situde Nanocladius Kieffer. 1913d: 31. prds de la division impr6cise entre les rdgions Type-species, vitellinus Kieffer n€arctique et n€otropicale, elle est indiqude (orig. des.). comme observable tant au Mexique que dans Microcricotopas Thienemann & la rdgion n6otropicale. La citation binomiale Harnisch, 7932: 737. originale apparait ensuite, suivie des sous- Type-species, Cricotopus especes ou varidtds, puis des synonymes (s'il y albicornis Goetghebuer a lieu), qui sont mentionnds sous leur forme (Goetghebuer in Goetghebuer & originale accompagn€s du nom de I'auteur, de Lenz,1944; 114) = Chironomus la date, de la page et de l'emplacement type. bicolor Zetterstedt. Les emplacements types pour le Canada et les Etats-Unis sont indiquds selon les Subgenus Nanocladius Kieffer frontidres des provinces, des territoires ou des Etats; tous les autres suivent les La citation originale se trouve sur la frontidres des pays ou encore des grandes deuxi0me ligne, en retrait, suivie des 169ions g6ographiques si celui-ci n'est pas synonymes (s'il y a lieu), eux aussi en retrait. connu. Le nom du genre est suivi du nom de I'auteur original, de la date, de la page, du ABREVIATIONS nom de I'espdce type et de la nature de la Les abrdviations utilis€es sont les fixation du type. L'esp0ce type originale est suivantes : mentionnee, mais le genre est omis s'il est G6n6ralitds identique d celui dont elle est le type. Si le &- et nom de l'espOce type originale est par contre e., E.- est, de l'est, orientale un synonyme mineur, il apparait i la fin emend.- corrigd aprds un symbole d'6galit6 (=). Seuls les I., Is.- ile, iles auteurs de fixation de type ultdrieure sont ICZN.- Commission internationale de mentionnds. Une grande partie des nomenclature zoologique informations sur la fixation des types et les misident.- erreur d'identification synonymes sont tirdes de Ashe (1983). Cette mon.- monotypie publication contient des renseignements nam.- nom d6taill6s sur la synonymie g6n6rique, les n. nam.- nouveau nom espoces types et les probldmes de nom. nud.- nom seul nomenclature. Les citations des noms des orig. des.- d€signation originale sous-genres suivent celles des genres, sauf s'il preocc.- d6jd appliqud s'agit de rdpdtition (dans ce cas, elles sont s., S.- sud, du sud, mdridional alors omises). se., Se.- sud-est, du sud-est Les noms des groupes d'espdces se sub. mon.- monotypie ulterieure presentent comme suit : subg.- sous-genre syn.- synonyme, synonymie triannulatus. Sask., Ont. to N.Y., s. to var.- varidtd Ky. and Del.; PA. w.- ouest, de I'ouest, occidental NEARCTIC CHIRONOMIDAE

Noms gdographiques R.L- Rhode Island AF.- r€gion afrotropicale Sask.- Saskatchewan Ala.- Alabama S.C.- Caroline du Sud Alas.- Alaska S. Dak.- Dakota du Sud Alta.- Alberta Tenn.- Tennessee Amer.- Amdrique Tex.- Texas Ariz.- Arizona U.S.A.- Etats-Unis d'Am6rique Ark.- Arkansas U.S.S.R.- Union des Rdpubliques AU.- r6gion australasienne Socialistes Sovi6tiques Baja Calif.- Basse-Californie Va.- Virginie B.C.- Colombie-Britannique Vt.- Vermont Calif.- Californie Wash.- Washington Can.- Canada Wis.- Wisconsin Colo.- Colorado W. Va.- Virginie de I'Ouest Conn.- Connecticut Wyo.- Wyoming D.C.- District de Columbia Y.T.- Territoire du Yukon Del.- Delaware Fla.- Floride Ga.- Gdorgie REMERCIEMENTS Grld.- Groenland Nous tenons d remercier M. B. Caldwell Ill.- Illinois d'avoir vdrifid plusieurs 6bauches. Feu D' Ind.- Indiana Sam Roback a eu la bont6 de vdrifier une Kans.- Kansas 6bauche antdrieure de la sous-famille des Ky.- Kentucky tanypodin6s. Nous remercions 6galement La.- Louisiane M. L. Speers d'avoir prdpard le manuscrit Labr.- Labrador pour I'impression et Mme L. Robitaille Man.- Manitoba d'avoir dactylographi6 une partie de la Mass.- Massachusetts bibliographie. Md.- Maryland Mex.- Mexique Mich.- Michigan Minn.- Minnesota Miss.- Mississippi Mo.- Missouri Mont.- Montana N.B.- Nouveau-Brunswick N.C.- Caroline du Nord N. Dak.- Dakota du Nord NE.- r€gion ndarctique Nebr.- Nebraska Nev.- Nevada Nfld.- Terre-Neuve N.H.- New Hampshire N.J.- New Jersey N. Mex.- Nouveau-Mexique N.S.- Nouvelle-Ecosse NT.- rdgion n€otropicale N.W.T.- Territoires du Nord- Ouest N.Y.- New York Okla.- Oklahoma Ont.- Ontario OR.- r6gion orientale Oreg.- Oregon PA.- rdgion paldarctique Pa.- Pennsylvanie P.E.I.- lle-du-Prince-Edouard Qu6.- Qudbec CHIRONOMIDAE DE LA REGION NEARCTIQUE

FAMILY CIIIRONOMIDAE sinuaticornis. Wash., Mont., Calif. Boreochlus sinuaticornis Brundin, 1966: Subfamily TELMATOGETONINAE 30-5.-Calif.

Genus Schiner Telmatogeton Schiner, 1866: 931. Genus LASIODIAMESA Kieffer Type-species, sanctipauli Schiner (orig. Lasiodiamesa Kieffer, I924b: 48 (as subg. des.). of Syndiamesa Kieffer, 1918a). Paraclunio Kieffer, 1911b: 103. Type-species, Syndiamesa gracilis Type-species, trilobatus Kieffer (orig. Kieffer (orig. des.). des.). arietina. Ont., Qu6., Wis., N.Y. alaskensis. Alas. s. to Calif.; Mex. Tanypus arietinus Coquillett, 1908: Telmatogeton alaskensis Coquillett, 1900: 144.-N.Y. 395.-Alas. brusti. N.W.T., Wyo., Minn. japonicus. Coastal, N.Y., s. to Miss.; PA.; Lasiodiamesa brusti Sather. 1969: NT.; OR.; AU. 15.-N.W.T. Tokunaga, 1933: rawsoni. Sask. 95.-Japan. Lasiodiamesa rawsoni Brundin, 1966: macswaini. Calif. 325.-Sask. Telmatogeton macswaini Wirth, 1949: tenebrosa. Alas., N.W.T., B.C., Minn., N.H. 170.-Calif. Tanypus tenebrosus Coquillett, 1905: spinosus. Calif. 66.-N.I{. Paraclunio spinosus Hashimoto, 1.973: Linacerus piloala Garrett, 1925a: 9.-B.C. 266.-Calit. Lasiodiamesa nearctica Brundin, 1966: trilobatus. Oreg., Calif. 326.-N.W.T. Paraclunio trilobatus Kieffer, 191 1b: sphagnicola. Ont.; PA. 105.-Calif. Isoplastus sphagnicola Kieffer, 7925c: 16.-U.S.S.R.

Genus Schiner Thalassomya Schiner, 1856: 219. Genus PARABOREOCHLUS Thienemann Type-species, frauenfeld i Schiner Paraboreochlas Thienemann. 1939: 166. (mon.). Type-species, Tanypus minutissimus Strobl (orig. des.). bureni. Ky., N.C., s. to Miss. & Fla.; NT. Thalassomya bureni Wirth, 1949: stahli. Pa., Kans. 167.-Fla. Paraboreochlus stahli Coffman in Coffman, Ferrington & Seward, 1.988: 192.-Pa.

Subfamily PODONOMINAE Genus TRICIIOTANYPUS Kieffer Tribe BOREOCHLINI Trichotanypus Kieffer, 1906a:. 42. Type-species, Tanypus posticalis Genus BOREOCHLUS Edwards Lundbeck (mon.). Boreochlus Edwards in Edwards & Thienemann, t938: I52. Type-species, alaskensis. Alas. thienemanni Edwards (orig. des.). Trichotanypus alaskensis Brundin, 1966: 315.-Alas. gracilistylus. Alas., Alta., Wash., Wyo. foliaceus. Alas. Boreochlus gracilistylus Brundin, 1966: Trichotanypus foliaceus Wirth & Sublette, 304.-Alta. 1970: 343.-Alas. persimilis. Alas. to Qu€., Calif. to Va. hanseni. Wyo. Trichotanypus persimilis Johannsen, 1926: Trichotanypus hanseni Wirth & Sublette, 99.-N.Y. 1970: 347.-Wyo. NEARCTIC CHIRONOMIDAE mariae. Alas. Procladius caliginosus Johannsen, 190-5: Trichotanypus mariae Wirth & Sublette, 131.-N.Y. 1970: 345.-Alas. planus. Fla. posticalis. N.W.T., Grld.; PA. Clinotanypus planus Roback, 1971: Tanypus posticalis Lundbeck, 1898: 24.-Fla. 295.-Grld. wirthi. Fla. Clinotanypus wirthi Roback, 7977: Tribe PODONOMINI 24.-Fla.

Genus PAROCI{LUS Enderlein Parochlus Enderlein, l9I2: 109. Genus COELOTANYPUS Kieffer Type-species, fuegianus Enderlein (orig. Coelotanypus Kieffer, l9L3c: t54. des.). Type-species, Tanypus humeralis Loew Paratanypus Garrett, 1925a: 8. (orig. des.). Type-species kiefferi carrett (orig. des.). atus. Kans., La., Tex., Mex.; NT. Coelotanypus a/us Roback, 1977: 37.-Tex. kiefferi. Alas. to N.8., s. to Calif. & N. Mex concinnus. Ilt., Neb. to Pa., s. to N. Mex. & Grld.; PA. Fla. Paratanypus kiefferi Garrett, 7925a: Tanypus concinnus Coquillett, 1895: 8.-B.C. 308.-Fla. Tanypus tibialis Staeger, 1845: 354 Coelotanypus flavus Kieffer, 1923a: (preocc. Say, 1823).-Grld. 296.-Tex. Podonomus peregrinus Edwards, 1929: naelis. Tex.; NT. 296.-Scotland. Coelotanypus naelis Roback, 1.963b: 170.-Surinam. scapularis. Ont., Ohio, e. U.S.A., s. to N. Mex., Kans., Tex., Mex.; NT. Subfamily TANYPODINAE Tanypus scapularis Loew, 1866: 2.-D.C. tricolor. Kans., Ill., N. Mex. to N.C., s. to Tribe COELOTANYPODINI Fla. Tanypus tricolor Loew, 1861a: 309.-N.Y. Genus CLINOTANYPUS Kieffer Clinotanypus Kieffer, 1973c: 157. Type-species, Procladius fuscosignatus Tribe MACROPELOPIINI Kieffer (Sublette & Sublette, 1965: 14-5). Genus ALOTANYPUS Roback Alotanypus Roback, l97I: 95. Subgenus APONTEUS Roback Type-species, Tanypus venustus Aponteus Roback, I97I: 25. Type-species, Coquillett (orig. des.). Clinotanypus sabensis Roback (orig. des.). aris. N.H., W. Va., N.C., S.C., Ala., Ga. Alotanypus aris Roback, 7971: 97.-N.C. sabensis. Tex. venusta. Wash., Wyo., Utah, Calif. to N. Clinotanypus sabensis Roback, 1971: Mex. 25.-Tex. Tanypus venustus Coquillett, 1902b: 91.-N. Mex. Subgenus CLINOTANYPUS Kieffer aureus. Kans., La., Fla. Genus APSECTROTANYPUS Fittkau Clinotanypus aureus Roback, 1971: Apsectrotanypr.rs Fittkau, 7962: l4l. 25.-La. Type-species, Tanypus trifascipennis pinguis. Man., Ont., widespread U.S.A. Zetterstedt (orig. des.). Tanypus pinguis Loew, 1861a: 308.-N.Y. Tanypus flavicinctus Loew, 1861a: algens. Alas., Alta., Wash., Oreg. 309.-Pa. Tanypus algens Coqttillett, 1902b: Tanypus thoracicus Loew, 1866: 4.-D.C. 90.-Alas. CHIRONOMIDAE DE LA REGION NEARCNQUE

florens. Alta., Man., Wash., Wyo., Calif., Genus FITTKAUIMYIA Karunakaran Nev., Colo., Ariz. Fittkauimyia Karunakaran, 1969: 75. Tanypus florens Johannsen, 1908: Type-species, disparipes (orig. des.). 272.-Colo., Wash., N.Y. Parapelopia Roback, l97l: 91. johnsoni. Sask., Qu6., N.H., N.J., N.C., S.C., Type-species, serta Roback (mon). Fla. Tanypus johnsoni Coquillett, 1901b: serta. Fla. 609.-N.J. Parapelopia serta Roback, t97l: 91.-Fla.

Genus BETHBILBECKIA Fittkau & Murray Genus MACROPELOPIA Thienemann Bethbilbeckia Fittkau & Murray, 1988: Macropelopia Thienemann in Thienemann 253. Type-species, floridensis (orig. & Kieffer, 1916: 497. Type-species, des.). Isoplastus bimaculatus Kieffer (orig. des.) = Tanypus nebulosus Meigen. floridensis. Fla. Bethbilbeckia floridensis Fittkau & decedens. B.C. to Qu6. & R.I., Utah, Wyo., Murray, 1988: 2-54.-Fla. Colo., Ky., N.C., s. to Miss. & Fla. Tanypus decedens Walker, 7848: 22.-Ont. Tanypus hirtipennis Loew, 1866: Genus BRUNDINIELLA Roback -5.-Maine. Brundiniella Roback, 1978b: 141 (n. nam. Anatopynia dena Roback in Sublette & for Brundinia Roback). Sublette, 1965: 145 (n. nam. for Brundinia Roback, 1978a: 168 (preocc. Anatopynia brunnea Roback, 1955: 2, Tottenham, 1949). Type-species, preocc. Edwards, 1931).-Mass. Anatopynia eumorpha Sublette (orig. des.). Genus PSECTROTANYPUS Kieffer eumorpha. Ont., Wash. to Pa., s. to Calif. & Psectrotanyplrs Kieffer, 1,909: 42. W. Va. Type-species, brevicalcar Kieffer Anatopynia eumorpha Sublette, I964a: (Fittkau, 1962: 129) = Tipula varici 107.-Calif. Fabricius.

discolor. Wash., Wis., N.H., S.C. Genus DEROTANYPUS Roback Tanypus discolor Coquillett, 1902b: Derotanypus Roback, l97l:- 91. 89.-N.H. Type-species,Anatopynia aclines dyari. B.C. to Ont., s. to Calif. & FIa. Sublette (orig. des.). Tanypus dyari Coquillett,1902a: Merotanypus Roback, 1971: 94" 85.-Mich. Type-species, Tanypus alaskensis Tanypus guttularis Coquillett, I902b: Malloch (orig. des.). 92.-Wash. Tanypus garretti Walley, 1925: 275.-B-C. aclines. Alas., Wash., Oreg., Idaho, Utah, pictipennis. Grld.; PA. Wyo., Calif., Nev. Tanypus pictipennis Zetterstedt, 1838: Anatopynia aclines Sublette, 1,964a: 818.-Grld. 104.-Calif. Derotanypus aclines var. aclines Roback, L971,: 93.-Utah, Wyo. Genus RADOTANYPUS Fittkau & Murray alaskensis. AIas. to N.W.T., B.C. to Man. Radotanypus Fittkau & Murray, 1985: 209. Tanypus alaskensis Malloch, 1919: Type-species, Anatopynia submarginella 35.-Alas. Sublette (orig. des.). Anatopynia centralis Malloch, 1934: 13.-N.W.T. submarginella. Oreg., Calif., Colo., Wyo. Anatopynia submarginelld Sublette, 7964a: 103.-Calif. 10 NEARCTIC CHIRONOMIDAE

Tribe monilis var. nigra Roback, l97l;378.-Ariz. Genus FittKau Tanypus basalis Walley, 1925:. 273--Ont- Natarsia Fittkau, 1962: l5l. Type-species, Ablabesmyia americana Fittkau, 1.962: 430 Chironomus punctatus Fabricius (orig. (n. nam. for Tipula monilis, Johannsen, des.). 1905: 142, not Linnaeus 1758, misident.). miripes. Calif. parajanta. Ont., N.Y., Neb. to Pa., s. to Tanypus miripes Coquillett, 1905: Miss. & Fla. 6-5.-Calif. Ablabesmyia parajanta Roback, 1971: baltimoreus. N.W.T., e. of Rockies to Kans. 373.-S.C. & Fla. rasha. N.H. Tanypus baltimoreus Macquart, 1855: Ablabesmyia rasha Roback, I97I:. 35.-Md. 371.-N.H. Ablabesmyia fastuosa Johannsen, 190-5: rhamphe. Minn. to Ohio, s. to N. Mex. & 1.53.-N.Y. Fla. Ablabesmyia rhamphe Sublette, 1964b:. ll4.-La. Tribe PENTANEURINI simpsoni. N.Y., Pa., S.C. Ablabesmyia sintpsoni Roback 1985: Genus ABLABESMYIA Johannsen 188.-N.Y. Ablabesmyia Johannsen, 1905: 135. Type-species, Tipula monilis Linnaeus Subgenus ASAYIA Roback (Johannsen, 1,907a: 400). Asayia Roback, 7985: 177- Type-species Tanypus annulatus (orig. des.). Subgenus ABLABESMYIA Johannsen annulata. Sask. to Qu6., s. to Calif. & Fla. aspera. B.C. to Maine, s. to Calif. to Fla. Tanypus annulatus Say, 1823: 15.-Pa. Pentaneura aspera Roback, 1959a: 'i.24.-Maine. Subgenus KARELIA Roback Pentaneura gera Roback, L959a: Karelia l{oback, l97l: 357. Type-species, 126.-N.H. Tanypus illinoensis Malloch (orig. des.). hauberi. Man., Ont., N.Y., Pa., N.J., Ky., s. to Miss. & Fla. alaskensis. Alas. Ablabesmyia hauberi Beck & Beck, 1966: Ablabesmyia alaskensis Roback, 1971: 329.-Fla. 360.-Alas. janta. N.W.T., Md., Pa., Tenn., S.C., Okla., cinctipes. Calif., Nev., Ariz., N. Mex., Ky., s. Tex., Ga., Fla. to Miss. & Fla. Pentaneura janta Roback, 19-59a: Pentaneura cinctipes Johannsen, 1946: 131.-Md. 271..-Fla. johannseni. Ill. idei. Ont., N.Y., Pa., S.C. Pentaneura j ohannseni Roback, 19-59a: Tanypus idei tNalley, 7925: 272.-Ont. 131.-Ill. illinoensis. N.W.T., Alta. to Qu6., N.Y., mallochi. Widespread NE. Maine, Colo., N. Mex., Kans., W. Va. Tanypus mallochi Walley, 1925: 273.-Ont. Tanypus illinoensis Malloch, 191-5b: Pentaneura auriensis Roback, 7957 a: 376.-\t. 39.-Pa. peleensis. Alas., widespread Can. & U.S.A. Pentaneura aequifasciata Dendy & Tanypus peleensis Walley, 1926a: 64.-Ont. Sublette, 1959: 507.-Ala. philosphagnos. B.C., Ont., N.B., Pa. to Miss. Ablabesmyia ornata Beck & Beck, 1966: & FIa. 328.-Fla. Ablabesmyia philosphagnos Beck & Beck, Ablabesmyia tarella Roback, 1971: 1966: 330.-Fla. 368.-Fla. pulchripennis. N.W.T., B.C. to Qu6., Wash., monilis. Widespread NE.; PA.; OR. Wyo., S. Dak., D.C., Col., Kans., Grld. Tipula monilis Linnaeus, 1758: Tanypus pulchripennis Lundbeck, L898: 587.-'Europe'. 293.-Grld. Tanypus prudens Walley, L925: 275.-Man. CHIRONOMIDAE DE LA REGION NEARCTIQUE 11

Genus FittKau mera. Wash., Ariz. Arctopelopia Fittkau, 1962: 194. mera Roback, 1971: Type-species, Tanypus barbitarsis 246.-Wash. Zetterstedt (orig. des.). paramelanops. N.Y., Va. Conchapelopia paramelanops Roback, cana. N.W.T. 1971: 249.-Ya. Arctopelopia cana Roback, 1,977: rurika. Man., Qu6., s. to Kans. & Ga. 253.-N.W.T. Pentaneura rurika Roback. 1957a: 34--Pa. melanosoma. Grld.; PA. telema. N.W.T. to Ont., Mont., N.Y., Ill., Ablabesmyia melanosoma Goetghebuer, Kans.. N.C. 1933: 20.-Grld. Conchapelopia telema Roback, 1971: 247.-Alta. Yarna. W. Va. Genus CANTOPELOPIA Roback Conchapelopia varna Roback, 1981: Cantopelopia Roback, l97l: 270. 93.-W. Va. Type-species, Cantopelopia gesta Roback (orig. des.). Genus DENOPELOPIA Roback & Rutter gesta. Qu6., Ill., N.Y., Fla. Denopelopia Roback & Rutter, 1988: 118. Cantopelopia gesta Roback, 197I: Type-species, atria (orig. des.). 270.-Qu6.. atria. Fla. Denopelopia atria Roback & Rutter, 1988: Genus CONCHAPELOPIA Fittkau 120.-Fla. Conchapelopia Fittkau, L957: 317- Type-species, Tanypus pallidus Meigen (orig. des.). Genus Fittkau Mesopelopia Roback, 1977: 252. Type Guttipelopia Fittkau, 1962: 251. species, aleta (mon). Type-species, Tanypus guttipennis Wulp (orig. des.). aleta. Widespread in e. Can. & U.S.A. Conchapelopia aleta Roback, 1971: guttipennis. N.W.T., Man., Ont., s. to Ala. & 252.-N.Y. Fla.; PA. bruna. Alas. to N.W.T. Tanypus guttipennis Wulp, 1861: Conchapelopia bruna Roback, 1971: 16.-Netherlands. 249.-N.W.T. Tanypus multipunctatas Curran, 1930: currani. Man. to Qu€., Wyo., Iowa, N.Y., 29.-N.Y. Ohio. s. to Miss. & Fla.. Guttipelopia curuani Beck & Beck, 1966: Tanypus currani Walley, 1925: 276.-Qu€. 332.-Fla. fasciata. Alta. to Ont., s. to Kans. & Fla. rosenbergi. Man., Ont. Conchapelopia fasciata Beck & Beck, Guttipelopia rose nbergi Bilyj, 1984: 1966: 346.-Fla. 6-59.-Man. Conchapelopia dusena Roback, 1971: 247.-N.Y. melanops. Not NE. Genus IIAYESOMYIA Murray & Fittkau Tanypus melanops Meigen, 1818: Murray & Fittkau, 1985: 196. 65.- Not stated. Type-species Tanypus lripunctatus pallens. Alas. to Qu6., s. to Calif. & Fla. Goetghebuer (orig. des.). Tanypus pallens Coquillett, I902b: 91.-N. Mex. senata. Alta. to N.S., s. to Calif., Kans. & Pentaneura goniodes Sublette, 1964a: Fla. 89.-Calif. Tanypus senatus Walley, L925: 276: -Ont. Conchapelopia goniodes var. trifida ' Pentaneura chrysos Sublette, 1964a: Roback. I97I: 246.-Calif. 90.-Calif. ?Chironomus unicolor Walker. 1848: 19.-N.S. t2 NEARCTIC CIIIRONOMIDAE

Genus Roback neopilosella. Neb., Ky., N.Y., N.C., s. to Helopelopia Roback, l97l: 250 (as subg. Miss. & Fla. of Conchapelopia Fittkau, 1957). neopilosella Beck & Beck, Type-species, Tanypus cornuticaudatus 1966: 341.-Fla. Walley (orig. des.). pilosella. Alas. to Qu6., s. to Ariz. & Fla.; NT. cornuticaudata. Alta. to Qu6., N.Y., Ohio, Tanypus pilosellus Loew, 1866: 5.-D.C. Pa., Kans., N.C., S.C., Fla. Labrundinia floridana Beck & Beck, 1966: Tanypus cornuticaudatus Walley, 1,925: 339.-Fla. 277--Qu6. virescens. Pa. s. to Tex. & Fla. pilicaudata. N.W.T. to N.B., N.Y., N.C. Labrundinia virescens Beck & Beck, 1966: Tanypus pilicaudatus Walley, 1925: 341.-Fla. 277.-Ont.

Genus Fittkau Genus Roback. Larsia Fittkat, 1962.339. Type-species, Hudsonimyia Roback, 1979: I. Ablabesmyia atrocincta Goetghebuer Type-species karelena Roback (orig. (orig. des.). des. ). berneri. Minn., Fla. karelena. N.Y., N.C., S.C., Ga. Larsia berneri Beck & Beck, 1966: Hudsonimyia karelena Roback, 1979: 3-50.-Fla. 3.-S.C. canadensis. Can. e. of Rockies. parrishi. Ga. Larsia canadensis Bilyj, 1984: 663.-Man. Hudsonimyia parrishi Caldwell & Soponis, decolorata. Wis., Mich., N.Y., Kans., Ill., 1982: 507.-Ga. Ohio. La., Fla. Tanypus decoloratus Malloch, 191-5b: 370.-Ill. Genus Fittkau Larsia lurida Beck & Beck, 1966: Krenopelopia Fittkau, 1962: 262. 3-51.-Fla. Type-species, Chironomus binotatus Larsia indistincta Beck & Beck, 1966: Wiedemann (orig. des.). 3-52.-Fla. Iyra. Calif. hudsoni. S.C. Pentaneura /yra Sublette, 1964a. Krenopelopia hudsoni Roback, 1983: 96.-Calif. 254.-S.C. marginella. Qu6., Ill., Kans. narda. Alas., ?Oreg. Tanypus marginellus Malloch, 1915b: Krenopelopia narda Roback, l97l: 374--Ill. 274.-Alas. planensis. Ont., Qu€., Calif., Tex., Ohio, N.Y., Va., Mex. Pentaneura planensis Johannsen, 1946: Genus LABRUNDINIA Fittkau 284.-Tex. Labrundinia Fittkau, 1962: 372. sequoiaensis. Calif. Type-species, Tanypus longipalpis Pentaneura sequoiaensis Sublette, 1964a: Goetghebuer (orig. des.). 99.-Calif.

becki. Okla., Ky., N.C., s. to Miss. & Fla. Labrundinia becki Roback, I97I: Genus MEROPELOPIA Roback 279.-Fla- Meropelopia Roback, l97l: 253 (as subg. johannseni. Ky., N.C., s. to Miss. & Fla. of Arctopelopia Fittkau, 7962). Labrundinia johannseni Beck & Beck, Type-species, Ablabesmyia flavifrons 1966: 342.-Fla. Johannsen (orig. des.). maculata. Calif., Kans., N. Mex., Tex., N.C., Mex.; NT. Labrundinia maculata Roback. 1971: 278.-Calif. CHIRONOMIDAE DE I-A REGION NEARCTIQUE l-t

americana. N.Y., Conn., N.J., Ohio, N.C., Genus Fittkau S.C., Kans. Paramerina Fittkau, 1962: 317. Conchapelopia americana Fittkau, 19-57: Type-species, Chironomus cingulatus 320 (n. nam. for Pentaneura vitellina Walker (orig. des.). Johannsen, t946: 281, not Kieffer, 1916, misident.).-Conn. anomala. Ky., N.C., s. to Miss. & Fla. flavifrons. Alas., Alta. to Qu6., s. to Calif., Paramerina anomala Beck & Beck, 1966: Fla. 344.-Fla. Ablabesmyia flavifrons Johannsen, 1905: fragilis. N.W.T., B.C. to Qud., Oreg., Va. 150.-N.Y. Tanypus fragilis Walley, 1926b. 205.-Ont. Pentaneura gesra Roback in Sublette & hanseni, Wash. Sublette, 1965: I47 (n. nam. for Paramerina hanseni Roback, 1971: Pentaneura alba Roback, 1957a: 32, 274.-Wash. preocc. Tokunaga, 1937).-Pa. smithae. Wash., N. Dak., S. Dak., Calif., Arctopelopia fittkaui Beck & Beck, 1966: IItah, Wyo., N. Mex., Kans. 348.-Fla. Pentaneura smithae Sublette, 1964a: 100.-Calif. testa. Okla., Tex. Genus Fittkau Paramerina testa Roback. l97 l: 272.-Tex. Monopelopia Fittkau, 1962: 394. Type-species, Tanypus tenuic al c ar Kieffer (orig. des.). Genus PENTANEURA Philippi Pentaneura Philippi, 18651. 629. s. to Miss. & Type-species, grisea Philippi (mon.). Fla. Monopelopia boliekae Beck & Beck, 1966: inconspicua. Ont., Wash., s. to Ariz., Kans., 335.-Fla. Okla., Ill. to Pa., s. to Fla. tenuicalcar. 8.C., Ont., N.8., Nfld., Calif., Tanypus inconspicuus Malloch, 191-5b: Fla.: PA. 371.-Ill. Tanypus tenuicalcar Kieffer, 1918a: Pentaneura comosa Sublette, 1,964a: 182.-Czechoslovakia. 93.-Calif. tillandsia. Ky., N.C., s. to Miss. & Fla. Pentaneura inculta Beck & Beck, 1966: Monopelopia tillandsia Beck & Beck, 334.-Fla. t966: 336.-Fla. inyoensis. Calif., N.Y., Ill., Ark. Pentaneura inyoensis Sublette, 1964a: 98.-Calif. Genus Kieffer Nilotanypus Kieffer, 1923b: 191. Type-species, r emot is s imus Kieffer Genus REOMYIA Roback (mon.). Reomyia Roback, 1986b: 283. Type- species, wartinbei Roback americanus. Ga., N.C., S.C., Fla. (orig. des.). Nilotanypus americanus Beck & Beck, 1966: 345.-Fla. wartinbei. Alas., Oreg. du,bius. Not NE. Zavrelimyia wartinbei Roback, 1984: Tanypus dubius Meigen, 1804: 25.-not 17.-Alas. stated. fimbriatus. Widespread NE. Chironomus fimbriatus Walker, 1848: Genus Fittkau 20.-Ont. Rheopelopia Fittkau, 1962: 209. ka.nsensis. Kans. Type-species, Tanypus ornatus Meigen Nilotanypus kansensis Roback, 1986a: (orig. des.). 458.-Kans. acra. Qu6., Maine, Mich., s. to W. Va., Ga. acra Roback, 1971: 243.- Mich. L4 NEARCTIC CIIIRONOMIDAE

paramaculipennis. Qu6., Ill., Va. tincta. Calif. Thie n emannimy i a p ar amacul ipe nnis tincta Roback, l97I: Roback, I97l: 243.-Ya. 256.-Calif . perda. Alta., Idaho, Ont., Minn., N.Y., Pa. Thienemannimyia perda Roback, 1971: 244.-N.Y. Genus ZAVRELIMYIA Fittkau Zavrelimyia Fittkau, 1962: 285- Type-species, Tanypus melanurus Genus TELOPELOPIA Roback Meigen (orig. des.). Telopelopia Roback, l97l: 255. Type-species, Tanypus okoboji Walley bifasciata. Man. to Nfld., U.S.A. s. to S.C. (orig. des.). Tanypus bifasciatus Coquillett, 1901b: 609.-N.J. okoboji. Man., Minn., Iowa, Ohio, Md., Va., sinuosa. Alta. to Qu6., s. to Kans. & Fla., Kans.. N. Mex. Calif. Tanypus okoboji Walley, I928b: Tanypus sinuosus Coquillett, 1905: 582.-Iowa. 6-5.-N.H. Zavrelimyia carneosa Fittkau, 1,962: 375 (n. nam. for Ablabesmyia carnea Genus THIENEMANNIMYIA Fittkau Johannsen, 190-5: 140, not Fabricius, Thienemannimyia Fittkau, 1.9 57 : 375. 180-5, misident.).-N.Y. Type-species, Ablabesmyia geij skesi Pentaneura fluminalis Sublette, 1964a: Goetghebuer (orig. des.). 88.-Calif. thryptica. Alas., Alta., Oreg., Ont., N.Y., s. barberi. Wash. s. to Calif., Nev., Ariz., Kans., to Kans. & S.C., Calif.; ?PA. N. Mex. Pentaneura thryptica Sublette, L964a: Tanypus barberi Coquillett, 1902b: 90.-N. 9-5.-Calif. Mex. fusciceps. Alas., N.W.T., B.C., Man., Calif., Incerta sedis in Pentaneurini Wyo.; PA. Tanypus futilis Wulp, 1867: Pentaneura fusciceps Edwards, 1929: 130.-Wis. 290.-England. norena. Man., Ont., Mich., Maine s. to Pa., Nominen dubium in Pentaneurini Oreg., Kans. Ablabesmyia aurea J ohannsen, 1907b: Pentaneura norena Roback, 7957a: 110.-Kans. 38.-Pa. Thie nemannimyia norena var. quadrata Roback, 197L: Tribe PROCLADIINI 242.-Ill. Genus Fittkau Dj almab atis ta Fittkau, I968a: 328. Genus Kieffer Type-species, director Fittkau (orig. Trissopelopia Kieffer, 1923d: L78. des.). Type-species, flavida Kieffer (orig. Calotanypus Roback, 7977: I52 (as subg. des.). of Procladius Skuse, 1889). Type-species, Protenthes pulcher ogemawi. Alta., Ont., Qud., Mich., N.Y., Pa., Johannsen (orig. des.). Ga. Trissopelopia ogemawi Roback, 1971: pulcher. Ont., Qu6., N.Y., Mich., Ohio, Pa., 268.-Mich. N.C., S.C., Fla., Ga. Protenthes pulcher Johannsen, 1908: 273.-N.Y. Genus XENOPELOPIA Fittkau Tanypus apicalis Walley, L926a: 64.-Qu6. Xenopelopia Fittkau, 1962: 381. Protenthes fasciger Curran, 1930: Type-species, Pelopia falcigera Kieffer 29.-N.Y. (orig. des.). Procladius maculatus Roback. 1971: 152.-Fla. CHIRONOMIDAE DE LA REGION NEARCTIQUE 1-5

Genus PROCLADIUS Skuse johnsoni. Ont., N.Y. Procladius Skuse. 1889: 283. Procladius johnsoni Roback, 1980a: Type-species, paludic ola Skuse 43.-Ont. (coquillett, 1910: 594). paragretis. ?Alas., Grld. Procladius paragretis Roback, 1971: Subgenus HOLOTANYPUS Roback 185.-Grld. Holotanypus Roback, 1982: 99 (n. nam. prolongatus. Alas., N.W.T. for subg. Procladius sensu Roback Procladius prolongatus Roback, 1971: 1971, not Skuse 1.889, misident.). 188.-N.W.T. Type-species, Tipul a culicifor mis ruris. N.W.T., B.C. to Labr., Minn. Linnaeus (orig. des.). Procladius ruris Roback, 1971.: 182.-Alta. Procladius ruris var. grandis Roback, abetus. Alta., Ont. 1977: 183.-N.w.T., 8.C., Nfld. Procladius abetus Roback. 1971: sublettei. Widespread NE. 185.-Alta. Procladius sublettei Roback, 1971: barbatulus. Calif. 191.-Calif. Procladius b arb atulus Sublette. 1964a; Procladius sublettei var. grandis Roback, l2l.-Calif. l9'71: 793.-N.W.T., Sask., Ont. clavus. N.W.T., B.C. Procladius sublettei var. minuta Roback, Procladius clavus Roback, 1971: 1971.: 194.-N.C., S.C., Fla. 187.-N.W.T. vesus. Alas., N.W.T. culiciformis. N.W.T., B.C. to Ont., s. to Procladius vesas Roback, 1971: Kans. & Fla., Calif., Nev., Ariz.; PA. 181.-N.W.T. Tanypus culiciformis Linnaeus, L767: wilhmi. Tenn. 978.-. Procladius wilhmi Roback, 1966: Tanypus crassinervis Zetterstedt, 1,838: 177.-Tenn. 817. Protenthes claripennis Malloch, 1915b: Subgenus PSILOTANYPUS Kieffer 387.-Mich. Psilotanypus Kieffer, 1906a: 39. curtus. Fla. Type-species, Tanypus bellus Loew Procladius curlas Roback, l97l: 181.-Fla. (Coquillett, 1910: -597). denticulatus. Widespread NE. Procladius denticulatus Sublette. 1964a; bellus. N.W.T., B.C., Sask. to N.B., s. to I24.-Calif. Calif. & Fla. dentus. Alas., N.W.T., B.C. to Labr. Tanypus bellus Loew, 1866;4.-D.C. Procladius dentus Roback. 1971: Procladius adumbratus Johannsen, 190-5: 186.-Man. 132.-N.Y. desis. Alas.. N.W.T. Tanypus pusillus Loew, 1866: 5.-D.C. Procladius desis Roback, l97l: 178.-Alas. Procladius flavidus Kieffer, 1923a: freemani. Alas., N.W.T., B.C. to Qu€., s. to 297.-Tex. Calif. & Ga. Procladius malifero Garrett, 1925: Procladius freemani Sublette, 7964a: 10.-8.C. 122.-Calif. deltaensis. Man. Procladius freemani var. asetus Roback, Procladius deltaensis Roback in Wrubleski l97l: 172.-Ga., N.C., S.C. & Roback. 1987: 208.-Man. Procladius freemani var. brunneus Roback, macrotrichus. Minn. 1971: 17l.-Nfld. Procladius macrotrichus Roback, 1971: Procladius freemani var. cretis Roback, 168.-Minn. l97l: 172.-8.C. nietus. B.C., Alta., Sask., Man. Procladius freemani var. elongatus Roback, Procladius nietus Roback. 1,971:. l97t: L7Z.-Fla., N.C., S.C. 168.-B.C. gretis. N.W.T. riparius. Kans., Ill. Procladius Sretis Roback, 1971: Protenthes riparius Malloch, 1915b: 184.-N.W.T. 389.-Ill. jeris. Alas.. N.W.T. Procladius jeris Roback, 197 l: 187.-Alas. t6 NEARCTIC CHIRONOMIDAE

Incerta sedis in Procladius concavus. Iowa, Kans., Ark., Mich., Ohio, Procladius lundstromi Goetghebuer in N.Y., Va., N.C., S.C., Tex. Goetghebuer & Lenz, 1936: 11.-not Tanypus concavus Roback, 1971: stated. 63.-Mich. Tanypus abrupta Garrett, I925a: 8.-B.C. parastellatus. Calif., Nev., Ariz., Tex. Tanypus arcuata Garrett, 7925a: 7.-I|.C. Tanypus parastellatus Sublette, 1964a: Tanypus bifida Garrett, 1925a: 8.-B.C. 113.-Calif. Tanypus trifolia Garrett, 1925a: 7.-B.C. punctipennis. B.C., Sask. to Qu6., Calif. to Trichotanypus vestitipennis Kieffer, 1917: Md., Ala., S.C., Fla.; PA.; OR. 339.-N.Y. Tanypus punctipennis Meigen, 1818: 61.-Not stated. stellatus. Alta., Ont., Utah to Pa., s. to Tex. Tribe TANYPODINI & Fla. Tanypus stellatus Coquillett, 1902b: Genus TANYPUS Meigen 89.-Tex. Tanypus Meigen, 1,803: 261. Type-species, telus. Fla. Tipula cincta Fabricius (Latreille, 1810: Tanypus relus Roback, l97l. 68.-Fla. 442; misident.). = Tanypus vilipennis. Man., Wis., Mich.; PA. punctipennis Meigen. Protenthes vilipennis Kieffer, 1918a: Pelopia Meigen, 1800: 18. Type-species, 164.-Hungary. Tipula cincta Fabricius (Coquillett, 1910: -586; misident.) = Tanypus Nomen dubium in Tanypodinae punctipennis Meigen. Supressed by Tanypus pubitarsis Zetterstedt, 1850: ICZN, 1963: 339. 3603.-Grld.; Sweden. Protenthes Johannsen, 7907a: 400. Type-species, Tanypus punc tipennis Meigen (orig. des.). Subfamily DIAMESINAE Subgenus APELOPIA Roback Apelopia Roback, l97L: 55. Type-species, Tribe B OREOHEPTAGYIINI Tanypus neopunctipennls Sublette (orig. des). Genus BOREOHEPTAGYIA Brundin Boreoheptagyla Brundin, L966: 420. clavatus. Ky. to N.C., s. to Tex. & Fla. Type-species, Heptagia rugosa Saunders Tanypus clavatus Beck, 1962: 92.-Fla. (orig. des.). grodhausi. Calif. Tanypus grodhausi Sublette, I964a: lurida. Alas., B.C., N.Y., Wash., Wyo. 116.-Calif. Diamesa lurida Garrett. t925a: 6.-8.C. imperialis. Calif., N. Mex. Tanypus imperialis Sublette, 1964a: II2.-Calif. Tribe DIAMESINI neopunctipennis. Widespread U.S.A. e. of Rockies, Mex.; NT. Genus ARCTODIAMESA Makarchenko Tanypus neopunctipennis Sublette, 1964a: Arctodiamesa Makarchenko, 1983; 264. 118.-Ill. Type-species, Diamesa appendiculata nubifer. Man., Calif., Nev., Utah, N. Mex., LundstrOm (orig. des.). Neb., Kans. appendiculata. Alas.; PA. Tanypus nubifer Coquillett, 190-5: Diamesa appendiculata LundstrOm, 1915: 66.-Utah. 23.-Siberia.

Subgenus TANYPUS Meigen Genus DIAMESA Meigen carinatus. Calif., Miss., Pa., S.C. Diamesa Meigen in Gistl, 1835: 66. Tanypus carinatus Sublette, 1964a: Type-species, cinerella Meigen in Gistl 110.-Calif. (orig. des.). CHIRONOMIDAE DE LA REGION NEARCNQUE l7

Adiamesa Kieffer, 1906a: 36. geminata. N.W.T., Grld.; PA. Type-species, Chironomus tonsus Diamesa geminata Kieffer, 1926: Walker, 19-56a: 195 (mon.). 79.-Grld. Eutanypus Coquillett, 1899: 341.. Diamesa biappendiculata Goetghebuer in Type-species, borealis Coquillett (orig. Remy, 1928: 52.-Gtld- des.). Diamesa furcata Edwards, 1933: Psilodiamesa Kieffer, 1918a: 104. 616.-N.W.T. Type-species, Psilodiamesa gregsoni. N.W.T., N.B.; PA. spitzbergensis Kieffer (orig. des.). Diamesa gregsoni Edwards, 1.933: 618.-N.W.T. aberrata. Alas., Wyo., Grld.; PA.; OR. haydaki. Alas., Wyo., Minn., Col., Ariz. Diamesa abetata Lundbeck, 1898: Diamesa haydaki Hansen in Hansen & 289.-Grld. Cook, 1976: 93.-WYo. alpina. Alas.; PA. heteropus. Alas. s. to N. Mex., to Neb. & Diamesa alpina Tokunaga, 1936: Minn. 539.-Japan. 'Tanypus heteropus Coquillett, 1905: amplexivirilia. B.C., Alta., Wash., Mont.; PA. 66.-?N.H., N. Mex., Wash. Diamesa amplexivirilia Ilansen in I{ansen Adiamesa confusa Garrett, 192-5: 5.-B.C. & Cook, 19?6: -53.-Wash. Diamesa banana Garrett, 1925: 6.-8.C. ancysta. Alas., Wash., Idaho, Mont., Nev., Diamesa onteona Roback, 1957b: Utah. Colo. 6.-Utah. Diamesa ancysta Roback, 1959b: hyperborea. Not NE. l.-Mont. Diamesa hyperborea Holmgren, 1869: arctica. Alas., N.W.T.; PA. 48.-Bear Island. Chironomus arcticus Boheman, 1865: Adiamesa ursus Kieffer, 1918a: 104.-Bear 574.-Spitzbergen. Island. bertrami. Grld.; PA. incallida. Alas., Y.T., Man., Ont., Wash., Diamesa bertrami Edwards, 1935: Utah, Wyo.; PA. 470.-Grld. Chironomus incallidus Walker, L956a: bohemani. N.W.T., Grld.; PA. 183.-England. Diamesa bohemani Goetghebuer, 1932a: Diamesa fonticola Sether, 1969: 181.-Spitzbergen. 24.-Man. cheimatophila. N.Y. insignipes. Wyo., Utah; PA. Diamesa cheimatophila Hansen in Hansen Diamesa insignipes Kieffer in Kieffer & & Cook, 1976: 68.-N.Y. Thienemann, 1908: 3.-Germany. chiobates. Minn., Wis. japonica. Wash., Mont., Wyo., Utah, Calif.; Diamesa chiobates Hansen in Hansen & PA. Cook, 1976: 70.-Wis. Diamesa japonica Tokunaga, 1936: chorea. Alas., Calif., Wyo., Grld. 542.-Japan. Diamesa chorea Lundbeck. 1898: Diamesa leoniella Hansen in Hansen & 291.-Grld. Cook, 1976: 111.-Wyo. clavata. N.W.T. leona. Wash. to N. Mex., to Minn., Colo., Diamesa clavata Edwards, 1933: Qu6., N.B.; PA. 61-5.-N.W.T. Diamesa leona Roback, 1957b: 7.-Utah. colenae. Alas., Y.T., Wyo. Diamesa caena Roback, 1957b: 9.-Utah. Diarnesa colenae Hansen in Hansen & Diamesa pieta Roback, 1957b: 8.-Utah. Cook, 1976: 76.-Wyo. lindrothi. Grld.; PA. davisi. N.W.T., Utah, Wyo., N.H., Grld. Diamesa lindrothi Goetghebuer in Diamesa davisi Edwards, 1933: Goetghebuer & Lindroth, 1931: 614.-N.W.T. 281.-Iceland. garretti. B.C., Wash., Idaho, Mont., Wyo. lupus. Alas., Alta. Diamesa garretti Sublette & Sublette, Diamesa /apas Willassen, 1985: 1965: 151 (n. nam. for Diamesa 127.-Alas. borealis Garrett, 1925: 6, preocc. Kieffer, 1915b).-B.C. mendotae. S. Dak., Minn., Wis. Diamesa mendotae Muttkowski. 1915: 116.-Wis. 18 NEARCTIC CHIRONOMIDAE

nivicavernicola. Alas.. Wash. Genus POTTHASTIA Kieffer Diamesa nivicavernicola Hansen in Hansen Potthastia Kieffer. 7922c: 362. & Cook, 1976: 123.-Wash. Type-species,longimana Kieffer (orig. nivoriunda. Minn. to Nfld., Ky., N.C., s. to des.). Miss. & Fla. Chironomus nivoriundus Fitch, 1847: gaedii. E. U.S.A.; PA. 282.-N.Y. Diamesa gaedii Meigen, 1838: simplex. Alas., N.W.T., B.C., Qu6., Wyo., 13.-Belgium. Grld. iberica. Idaho, Wyo., Ga.; PA. Diamesa simplex Kieffer, 1926: Potthastia iberica Serra-Tosio, 1971: 81.-N.W.T. 160.--Spain. sommermani. Alas. longimana. Ont., Qu6., N.Y., N.C., S.C.; PA. Diamesa sommermani Hansen in Hansen Potthastia longimanus Kieffer, 1922c: & Cook, 7976: 134.-Alas. 362.-Germany. sonorae. Calif. montium. Se. U.S.A.; PA. Diamesa sonorae Willassen, 1985: Diamesa montium Edwards, 1929: 125.-Calif. 307.-Great Britain. spinacies. Alas., Alta., Mont., Idaho, Wyo., utah, col., calif. Diamesa spinacies Sether, 1969: Genus PSEUDODIAMESA Goetgbebuer 21.-Alta. Pseudodiamesa Goetghebuer in subletti. Alta. Goetghebuer & Lenz, 1939a: 9 (as Diamesa subletti Makarchenko, 1986: subg. of Syndiamesa Kieffer, 1918a). 155.-Alta. Type-species, Syndiamesa pilosa Kieffer vockerothi. Ont., Qu6. (orig. des.) = Diamesa branickii Diamesa vockerothi Hansen in Hansen & Nowicki. Cook, 1976: 139.-Ont. Subgenus PACIIYDIAMESA Oliver Pachydiamesa Oliver, 1959: 54. Genus LAPPODIAMESA Serra-Tosio Type-species, Diamesa arctica Malloch Lappodiamesa Serra-Tosio, 1968: 140. (orig. des.). Type-species, b rundini Serra-Tosio (orig. des.). arctica. N.W.T. Diamesa arctica Malloch, 1919: boltoni. Ohio. 37.-N.W.T. Lappodiamesa boltoni Sether & Willassen, 1988: 78.-Ohio. Subgenus PSEUDODIAMESA Goetghebuer

Genus PAGASTIA Oliver branickii. B.C., Alta., Calif., N. Mex., Grld.; Pagastia Oliver, 1959: 49. Type-species, PA.; OR. orthogonia Oliver (orig. des.). Diamesa branickii Nowicki, 1873: Hesperodiamesa Sublette, I967a: 305. 3.-Poland. Type-species, Prodiamesa sequax pertinax. N.W.T., B.C., Utah, Wyo., Colo. Garrett (orig. des.). Prodiamesa pertinax Garrett, 1925: 6.-8.C. orthogonia. Alas., Mich. Pagastia orthogonia Oliver, 1959: 51.-Alas. Genus PSEUDOKIEFFERIELLA Zavrel partica. Alas., Y.T., Utah, Wyo. Pseudokiefferiella Zavrel, 1947: 1.1. Syndiamesa partica Roback, I95'7b: Type-species, Diplomesa 4.-Utah. lapponica Pagast = Diamesa Syndiamesa artisia Roback, 19-57b: parva Edwards (orig. des.). (see 5.-Utah. Cranston, 1975: 88; Ashe, 1983: sequax. B.C., Alta. 46- ICZN ruling required). Prodiamesa sequax Garrett, 1925: 7.-8.C. Diplomesa Pagast, 1947: 539. Type-species, lapponica Pagast (mon.). CHIRONOMIDAE DE LA REGION NEARCTIQUE 19

parva. N.W.T.; PA. ramosus. N.W.T., Ont. Diamesa parva Edwards, 1.932a: Protanypus ramosus Sether, 1975c: 4-5.-Scotland. 368.-Ont. saetheri, Alas. Protanypus saetheri Wiederholm, 797 5: Genus SYMPOTTHASTIA Pagast 224.-Alas. Sympotthastia Pagast, 1947: 457. Type-species, zavreli Pagast (mon.).

diastena. Oreg., Colo., Calif. Subfamily PRODIAMESINAE Pseudodiamesa diastena Sublette. I964a: 128.-Calif. Genus COMPTEROMESA Sether fulva. N.Y., N.C., S.C. Compteromesa Sather, 1987c: 794. Diamesa fulva Johannsen, 192L: Type-species, oconeensis Srether (orig. 229.-N.Y. des.). zavreli. N.C. Sympotthastia zavreli Pagast, 1947: oconeensis. S.C. 458.-Germany. Compteromesa oconeensis Saether, 1981c: 195.-S.C.

Genus SYNDIAMESA Kieffer Syndiamesa Kieffer, 1918a: 103. Genus MONODIAMESA Kieffer Type-species, Diamesa hygropetrica Monodiamesa Kieffer, 1922a: 287. Kieffer (Edwards, 1929: 303). Type-species, Prodiamesa bathyphila Kieffer, Goetghebuer in Goetghebuer Unnamed species.-N.W.T., Ont. & Lenz, 7939a: 3).

Nomina dubia in Diamesinae bathyphila. Alas., N.W.T., Minn., Ill. to Ohio, Diamesa waltlii Meigen, 1838: Grld.; PA. l3.-Germany. Prodiamesa bathyphila Kieffer, 1918a: Syndiamesa polaris Kieffer, 1926: 102.-Germany. 81.-N.W.T. depectinata. Man., Ont., Pa. Monodiamesa depectinata Srther, I973b: 666.-Man. Incerta sedis in Diamesinae prolilobata, B.C. Diamesa polaris Kieffer, 1926: Monodiamesa prolilobata Sether, 1,973b: 79.-N.W.T. 673.-B.C. tuberculata. N.W.T., 8.C., Ont., Mich., Ind. Monodiamesa tuberculata Srther. 1,973b:. Tribe PROTANYPINI 671.-Mich.

Genus PROTANYPUS Kieffer Protanypus Kieffer, I9O6a: 40. Genus ODONTOMESA Pagast Type-species, Tanypus morio Odontomesa Pagast, 1947: 502. Zetterstedt (Edwards, 1924a: 719). Type-species, Prodiamesa fulva Kietfer (orig. des.). caudatus. N.W.T.: PA. Protanypus caudatus Edwards, 1,924a: ferringtoni. Colo. 122.-. Odontomesa ferringtoni S&ther, 1985e: hamiltoni. N.W.T., B.C. 25.-Colo. Protanypus hamiltoni Saether, 197-5c: fulva. Mont. s. to N. Mex., Wis., II[., Pa. s. to 375.-8.C. Miss. & Fla.: PA. pseudomorio. Alas.; PA. Prodiamesa fulva Kief.fer, 1919a: Protanypus pseudomorio Makarchenko, 132.-Hungary. 1982: L33.-U.S.S.R. Odontomesa fulva nearctica Srther, 1985e: 20.-Ga. 20 NEARCTIC CHIRONOMIDAE

lutosopra. B.C. Chironomus lucens lucens Prodiamesa lutosopra Garrett, 7925: Zetterstedt.-Not NE. 7.-B.C. Chironomus lucens var. baueri Goetghebuer, I955. 167. Unrecognized. Genus PRODIAMESA Kieffer nitidellus. B.C. to Man., S. Dak., Ill., N.8., Prodiamesa Kieffer, 1906a: 37. N.Y., Pa. Type-species, Diamesa praecox Kieffer Orthocladius nitidellus Malloch, 1915b: (orig. des.) = Chironomus olivaceus 515.-Ill. Meigen. Spaniotoma senex Johannsen, 1937a: 63.-N.Y. cubita. B.C. Prodiamesa cubita Garrett, 1925: 7.-8.C. olivacea. Widespread NE.; PA. Genus ANTILLOCLADIUS Sether Chironomus olivaceus Meigen, 1818: Antillocladius Srether, 1981b: 4. 29.-Not stated, ?Germany. Type-species, antecalvus Sether (orig. Chironomus turpis Zetterstedt, 1838: des.). 81 1.-Sweden. Tanypus occidentalis Coquillett, 1902b: arcuatus. Kans., S.C. 92.-Colo. Antillocladius arcuatus Srether, 1982: 474.-S.C. pluspilalus. Kans., S.C. Antillocladius pluspilalus Sether, 1982: Subfamily ORTHOCLADIINAE 474.-S.C.

Genus ABISKOMYIA Edwards Abiskomyia Edwards, 1937: I4O. Genus APOMETRIOCNEMUS Sether Type-species, virgo Edwards (orig. Apometriocnemus Snther, 1985c: 536. des.). Type-species, fontinalis Sether (orig. des.). virgo. Alas., N.W.T.; PA. Abiskomyia virgo Edwards, 1937: beringensis. Y.T. l41.-Sweden. Apometriocnemus beringensis Cranston & Oliver. 1988a: 428.-Y.T. fontinalis. Tenn. Genus ACAMPTOCLADIUS Brundin Apometiocnemus fontinalis Srether, 1985c: Acamptocladlas Brundin, 1956: 162. 537.-Tenn. Type-species, Spaniotoma submontana Edwards (orig. des.). Phycoidella Srether, 1971: 1810. Genus BAEOCTENUS Srether Type-species, dentolatens Srether (orig. Baeoctenus Srether, \976:. 4I. des.). Type-species, bicolor Sether (orig. des.). dentolatens. Ont. Phycoidella dentolatens Sether, 1971: bicolor. Man., N.B. 1814.-Ont. Baeoctenus bicolor Sether, 1976: 43.-Man.

Genus ACRICOTOPUS Kieffer Acricotopus Kieffer, l92la: 90. Genus BRILLIA Kieffer Type-species, grandis Kieffer (orig. Brillia Kieffer, 1913b: 34. Type-species, des.) - Chironomus lucens Zetterstedt. Metriocnemus bifidus Kieffer (orig. des.) = Chironomus modestus Meigen. Iucens. N.W.T., Alta.; PA. Chironomus lucens Zetterstedt. 1850: 3574.-Denmark. CHIRONOMIDAE DE LA REGION NEARCTIQUE 2l

flavifrons. B.C. to N.B., s. to Oreg., N. Mex. laticaudus. S. Dak. to La., N.C., S.C., Ga. Bryophaenocladius I atic audus Sether, Metriocnemus flavifrons Johannsen, L905: 1973a: 55.-S. Dak. 301.-N.Y. pleuralis. Ill. laculata. Wash., Oreg., Wyo. Orthocladius pleuralis Malloch, 1915b: Brillia laculata Oliver & Roussel, 1983: 527.-rll. 268.-Oreg. psilacrus. S.C. longifurca. Not NE. Bryophaenocladius psilacrus Sether, 1982: Brillia longifurca Kieffer, I92la:. 500.-s.c. 86.-Germany. subparallelus. S. Dak., Ill., Ohio. parva. Ont. to N.B., s. to S.C. Orthocladius subparallelus Malloch, 1915b: Brillia parva Johannsen, 1934: 3-51.-N.Y. 522.-ilL. retifinis. Alas., B.C. to Man., s. to Calif. & Colo. Brillia retifinis Srether, t969: 37.-8.C. Genus CAMPTOCLADIUS WulP sera. N.If., Pa., S.C., Ga. Camptocladirzs WulP, 1874: 133. Brillia sera Roback. 1951a:. 64.-Pa. Type-species, Tipula byssina Schrank (Coquillett, 1910: 518) = Tiqula stercorarius De Geer. Genus BRYOPHAENOCLADIUS Thiennemann stercorarius. Widespread NE.; PA.; AU. Bryophaenocladius Thienemann, 1934: 36. Tipula stercorarius De Geet, I776: Type-species, Orthocladius mus cic ola 388.-Sweden. Kieffer (orig. des.). Tipula byssina Schrank, 1803: Clinocladius Sublette. 1970: -50 76.-Germany. Type-species, Orthocl adius subparallelus Malloch (orig. des.). Platycladius Sublette, t97O: 82. Genus CARDIOCLADIUS Kieffer Type-species, Orthocladius pleuralis Cardiocladius Kieffer, L9l2a: 22. Malloch (orig. des.). Type-species, ceylanicus Kieffer (orig. des.). aestivus. Alas.; PA. Eudacrylocladius aestivus Brundin, 1947: albiplumus. E|.C., Ont., Qu6., s. to N.C- 30.-Sweden. Cardiocladius albiplumus Saether, 1969: astis. 8.C., Utah. 34.-Ont. Hydrobaenus astis Roback, 1957b: fulvus. Mich., N.Y., Ill., Miss' IZ.-Utah. Thalassomyia fulva Johannsen, 1908: Hydrobaenus traenis Roback, 1957b; 275.-N.Y. 12.-Utah. obscurus. Ont., Qu6., Wy., Utah to N.Y-' s. digitatus. S. Dak., N.C., S.C. to Miss. & Fla. Bryophaenocladius digitatrrs Sether, 1973a: Thalassomyia obscura Johannsen, 55.-S. Dak. 1903: 437.-N.Y. flavoscutellatus. Ont., N.8., Ill. Thalassomyia fusca Johannsen, \903:. 437 Orthocladius flavoscutellalas Malloch, (nom. nud.).-N.Y. 1915b: 523.-lll. platypus. Calif., Nev., Ariz., N. Mex., ?Qu6. fumosinus. N.Y. Orthocladius platypus Coquillett, L902b'- Camptocladius fumosinus Curran, 1930: 93.-Ariz. 33.-N.Y. furcatus. N.Y.; PA. Dactylocladius furcatus Kieffer in Genus CHAETOCLADIUS Kieffer Thienemann & Kieffer. 1916: Chaetocladius Kieffer, 191lc: 182 (as 535.-Sweden. subg. of Dacrylocladias Kieffer, 1906a). impectinus. N.C., S.C. Type-species, Dacrylocladius setiger Bryophaenocladius impeuinus Sather, Kieffer (Goetghebuer in Goetghebuer 1976: 273.-5.C. & Lenz, 1942: 57) = Chironomus perennis Meigen. zz NEARCTTC CHIRONOMIDAE

adsimilis. Grld. Genus CLUNIO Haliday Orthocladius adsimilis Goetghebuer, 1933: Clunio Haliday, 1855: 62. Type-species, 25.-Grld. marinus Haliday (mon.). grandilobus. N.W.T.; PA. Chaetocladius grandilobus Brundin, 1956: cal iforniensis. Calif. 126.-Sweden. Clunio californiensis Hashimoto, 1.97 4: holmgreni. N.W.T.; PA. 108.-Calif. Chironomus holmgreni Jacobson, 1898: marshalli. Ky., N.C., s. to Fla. 204 (n. nam. for Chironomus festivus Clunio marshalli Stone & Wirth. 1947: Holmgren, 1869: 43, preocc. Say, 214.-Fla. 1823).-Spitsbergen & Bear I. ligni. Oreg. Chaetocladius ligni Cranston & Oliver, Genus COMPTEROSMITTIA Srether 1988b: 144.-Oreg. Compterosmittia Se,ther, 1981b: 20. perennis. Alas., N.W.T.; PA. Type-species, dentispina Sather (orig. Chironomus perennis Meigen, 1830: des.). 249.-Europe. stamfordi. Qu6., Ont., S. Dak., Ohio, Conn. nerius. N.Y., S.C. Hydrob aenus stamfordi Johannsen, 1947: Camptocladius nerius Curran, 1930: 171.-Conn. 34.-N.Y. Chaetocladitts oliveri Sether, 1969: Compterosmittia clavigera Sather, 1982: 93'-Que' 491.-S.C.

Genus CHASMATONOTUS Loew Genus CORYNONEURA Winnertz Chasmatonotas Loew, 1864: -51. Type- Corynoneura Winnertz, 1846: 12. species, unimaculatus Loew (mon.). Type-species, s cutell ata Winnertz (Coquillett, 1910: 528). atripes. Ont. to N.B., s. to Ill., Ohio, Md. Chasmatonotus atripes Rempel, 1937 a: arctica. N.W.T.; PA. 2_53.-N.Y. Corynoneura arctica Kieffer, 1923c: bicolor. N.Y., Pa., Tenn. 4.-U.S.S.R. Chasmatonotus bicolor Rempel, 1937 a: celeripes, Ont., Maine, N.Y., Ill., Ind., Ohio, 254.-Tenn. Grld.: PA. bimaculatus. Qu€. s. to Pa., Tenn. Corynoneura celeripes Winnertz, 18-52: Chasmatonotus bimaculalas Osten Sacken, -50.-Germany. 1877: 19l.-N.Y. Corynoneura atra Winnertz, 1852: fascipennis. B.C., Idaho, Mont., Wyo. 50.-Germany. Chas matonotus fasc ipe nnis Coqu illett, diara. Utah. 1905: 66.-8.C. Corynoneura diara Roback, 1957b: hyalinus. Calif. 10.-Utah. Chasmatonotus hyalinus Coquillett, 190-5: fittkaui. N.C., S.C.; PA. 67.-Calit. Corynoneura littkaui Schlee, 1968: maculipennis. B.C. s. to Calif. 19.-Germany. Chasmatonotus maculipennls Rempel, lacustris, Man., Ont.; PA. 1937a:. 254.-Wash. Corynoneura lacustris Edwards, unimaculatus. Ont. to N.S., s. to Mich., N.C., 7924c: I87.-Great Britain. S.C. lobata. Alta., Ont., N.B., N.C., S.C.; Chasmatonotus unimaculatus Loew, 1864: PA. 50.-N.H. Corynoneura lobata Edwards, I924c: univittatus. Alas. s. to Wash., Calif. 186.-England. Chasmatonotus univittatus Coquillett, minuscula. Ont.; PA. 1900: 395.-Alas. Corynoneura minuscula Brundin, 1949: 833.-Sweden. oxfordana. Ohio. Corynoneura oxfordana Boesel & Winner 1980: -50.5.-0hio. CHIRONOMIDAE DE I-A REGION NEARCTIQUE z-)

scutellata. N.W.T. s. to Fla., Grld.; PA.; NT; festivellus. Man., Ont.; PA. AU. Chironomus festivellus Kieffer, 1906a: 18 Corynoneura scutellata Winnertz, 1846: (n. nam. for Chironomus festivus 13.-Germany. Meigen, 1830:. 252, Preocc. SaY, taris. Ohio, N.Y., s. to Miss. & Fla. 1823).-Europe. Corynoneura /aris Roback, 1957a: flavocinctus. Man.; PA. 61.-N.J. Tricho cl adius fl av o c inc tus Kieffer, 1924b: 92.-Germany. fugax. Ont., N.Y., Ky., N.C., s. to Miss. & Genus CRICOTOPUS Wulp Fla. Cricotopus Wulp 1874: 132. Type-species, Orthocladius fugax Johannsen, 1905: Chironomus tibialis Meigen (Coquillett 279.-N.Y. 1910b: 528). fuscus. Ont.; PA. Trichocladius fuscus Kieffer, 1909: Subgenus CRICOTOPUS Wulp 46.-Germany. gelidus. N.W.T., Grld.; PA. abanus. B.C., Man. Trichocladius gelidus Kieffer, 1922b: Cricotopus abana Curran, 1929: 1.-Man. 18.-Novaya Zemlya. absurdus. N.Y., Ohio, N. Mex. infuscatus. Ont., S. Dak., Ill., Ohio, Pa., N. Orthocladius absurdus Johannsen. 190-5: Mex. 277.-N.v. Orthocladius infuscatus Malloch, 1915b: annulator. Ont., Calif., N. Mex.; PA. 517.-Ill. Cricotopus annulator Goetghebuer, 1927:. Cricotopus aratus Roback, 1957a: 73.-Pa. 52.-Belgium. Cricotopus ceris Roback, 7957a: 72.-Pa. Cricotopus irwini Sublette & Sublette, lestralis. Alas., N.W.T., Wash., Grld.; PA. l97l: 97.-Calif. Trichocladius lestralis Edwards, I924b: baptistensis. Ont. 172.-Spitsbergen. Cricotopus baptistensis Boesel, 1983: luciae. Ont. 87.-Ont. Cricotopus luciae LeSage in LeSage & beringensis. Alas. Harrison, 1980: 91.-Ont. Cricotopus beringensis Oliver & Dillon, mackenziensis. N.W.T. 1988: 475.-Alas. Cricotopus mackenziensis Oliver, 1977 : bicinctus. Widespread NE.; PA.; NT.; AF. 99.-N.W.T. Chironomus bicinctus Meigen, 1818: macraei. Man. 41.-Austria. Cricotopus macraei Srether, 1971: coronatus. N.W.T., Man., Qu6.; PA. 1799.-Man. Cricotopus coronatus Hirvenoja, 1973: Cricotopus flannagani Srether, 1971: 22Z.-. 1809.-Man. cumulatus. N.W.T., Grld.; PA. patens. Y.T., N.W.T.; PA. Cricotopus cumulatus Hirvenoja, 1973: Cricotoptts patens Hirvenoja, 1973: 146.-Grld. 218.-Finland. curtus. Sask., N.Y.; PA. pedatus. Mont., N. Mex. Cric otopus curtus Hirvenoja, 1973: Cricotopus pedatus Sublette, 1967b:. 199.-Germany. 558.-Mont. cylindraceus. Man., N.Y.; PA. pilosellus. Y.T., N.W.T., Grld.; PA. Trichocladius cylindraceus Kieffer in Cricotopus pilosellus Brundin, 1956: Kieffer & Thienemann. 1908: 114.-Sweden. 8.-Germany. politus. Sask., Colo:, Ill., Ohio, N.Y., D.C., elegans. Mich., Pa., Ohio. Md., N. Mex. to Fla. Cricotopus elegans Johannsen, 1943: Orthocladius politus Coquillett, 1902b: 78.-Mich. 93.-D.C. ephippium. Alas.; PA. pulchripes. N.Y.; PA. Chironomus ephippium Zetterstedt, 1838: Cric otopus pulchripe s Y errall, 19 12: 814.-Sweden. 22--Wales. Ceratopogon humeralis Zetterstedt, 1838: 820.-Sweden. aA NEARCTIC CHIRONOMIDAE

slossonae. Sask. to Ont., s. to Ill., N.Y., glacialis. Not NE. Ohio, Ga. Cricotopus glacialis Edwards, 7922: Cricotopus slossonae Malloch, 1915b: 209.-Spitsbergen. 506.-Iil. intersectus. Sask., Man., Ohio; PA. tibialis. Y.T., N.W.T., Man., N.B., Grld.; PA. Chironomus intersectus Staeger, 1839: Chironomus tibialis Meigen, 1804: 574.-Denmark. 16.-Europe. laetus. Ont.: PA. Chironomus basalis Staeger, 184-5: Cricotopus laetus Hirvenoja, 1,973: 351.-Grld. 264.-Finland. Chironomus pavidus Holmgren, 1869: laricomalis. Alas., N.W.T., Qu6., Grld.; PA. 42.-Spitsbergen. Cricotopus laricomalis Edwards, 1932a: Cricotopus polaris Kieffer, 1926: 46.-Scotland. 84.-Grld. myriophylli. B.C., Ont. Trichocladius b acilliger Kieffer, 1926: Cricotopus myriophylli OIiver, 1984: 86.-Grld. L287.-8.C. tremulus. Alta. to N.S., Oreg., Ohio, N.Y., ornatus. Alas. to N.W.T., Sask., Man., Ont., N'J'; Pa' N.Y.; PA. Tipula tremula Linnaeus, 17-58: Chironomus ornatus Meigen, 1818: 587.-Sweden. 43.-Germany. triannulatus. Sask., Ont. to N.Y., s. to Miss. pilitarsis. Sask., N.Y.; PA. & Fla.; PA. Chironomus pilitarsis Zetterstedt, 18-50: Chironomus lriannulatus Macquart, 1826: 3-56-5.-Sweden. 202.-Germany. sylvestris. Widespread NE.; PA. Cricotopus exjlis Johannsen, 1905: Tipula sylvestris Fabricius, 1794: 2.5-5.-N.Y. 252.-Etrope. trifascia. Sask., Ont., Ohio, Pa., N.Y., N.C.; tricinctus. Utah to N.Y., s. to Tex.; PA. PA. Chironomus tricinctus Meigen, 1818: Cricotopus trifascia Edwards, 1929: 41.-Europe. 322.-England. Cricotopus lebetis Sublette, t964b: Cricotopus ithacanensis Sublette, 1967b: 118.-La. 555.-N.Y. trifasciatus. 8.C., Ont., Idaho to N.Y., s. to trilobus. Alas. N. Mex. & Fla.; PA. Cricotopus trilobus Oliver & Dillon, 1988: Chironomus trifasciatus Meigen in Panzer, 485.-Alas. 1813: l8.-Germany. tristis. Y.T., N.W.T.; PA. Cricotopus remus Sublette, 1,964b: Cricotopus /rislis Flirvenoja, 1973; 11-5.-La. 196.-Finland. varipes. N.W.T., Ont., Wash., Ohio, N.Y., Subgenus NOSTOCOCLADIUS Ashe & Conn., Ga., Fla. Murray Cricotopus varipes Coquillett, l9l2b: Nostococladias Ashe & Murray, 1980: 93.-Md. 105. Type-species, lygropis Edwards vierriensis. Man., Ohio, Pa., N.Y., N. Mex.; (orig. des.). PA. Cricotopus vierriensis Goetghebuer, 1935: nostocicola. Alta., Mont., N.Y., Ohio s. to 417.-Belgium. Ga., Oreg., Calif. Cricotopus nostocicola Wirth, 1957: Subgenus ISOCLADIUS Kieffer 722.-Calif. Isocladius Kieffer, I9O9: 44 (as genus). Type-species, albipes Kieffer (mon.) = Subgenus PSEUDOCRICOTOPUS Nishida Tipula silvestris Fabricius. Pseudocricoropus Nishida, 1987'. 460 (as subg. of Cricotopus Wulp). flavibasis. B.C., Alta., S. Dak., Ill. Type-species, montanus Tokunaga Cricotopus flavibasis Malloch, 1915b: (orig. des.). 502.-Iil. flavipes. Ohio, Mich., Conn. bifurcatus, 8.C., Wash. Cricotopus flavipes Johannsen, 1942: Cricotopus bifurcatus Cranston & Oliver, 73.-Mich. 1988a: 430.-B.C. CHIRONOMIDAE DE LA REGION NEARCTIQUE 25

Cricotopus unassigned to subgenus Genus DIPLOSMITTIA Sether Diplosmittia Sather, I98lb: 29- belkini. N. Mex. to Ala. Type-species, hatisoni Srether (orig. Cricotopus belkini Dendy & Sublette, des.). 1959:51O.-Ala. brunnicans. Sask. carinata. Mich. Cricotopus brunnicans Walley, L928a: Diplosmittia carinata Sether, 1985h: 21.-Sask. 55.-Mich. diversus. Ont., Ohio., Mich., N.Y., Del. Cricotopus diversus Boesel, 1983: 85.-Ohio. Genus DOITHRIX Saether & Sublette eilurus. Calif. Doithrix Srether & Sublette, 1983: 6. Cricotopus edurus Sublette & Sublette, Type-species, villosa Sether & Sublette 1971: 85.-Calif. (orig. des.). fuscatus. Calif. Cricotopus fuscatus Wirth, 1957: barberi. Calif. t24.-Calif. Doithrix barberi Sether & Sublette, 1983: furtivus. Calif. 2I.-Calif. Cricotopus furtivus Sublette & Sublette, dillonae. N.S. L97L: 99.-Calif. Doithrix dillonae Cranston & Oliver, geminatus. Pa. 1988a: 432.-N.S. Chironomus geminatus Say, 1823: 14.-Pa ensifer. Calif. globistylus. Utah, N. Mex. Doithrix ensifer Sether & Sublette, 1983: Cricotopus globistylus Roback, I957b: 15.-Calif. 10.-Utah. hamiltoni. B.C. junus. Pa. Doithrk hamiltoni Sather & Sublette, Cricotopus jznr.rs Roback, 1957a; 72.-Pa. 1983: 17.-B.C. obscurifuscus. Calif. parcivillosa. Tenn. Cricotopus obscurifuscus Sublette & Doithrix parcivillosa Sether & Sublette, Sublette, I97 l: 96.-Calif. 1983: 12.-Tenn. oceanicus. Mass. vitlosa. Man., N.8., N.S., S.C., Tenn. Chironomus oceanicus Packard, 1869: Doithrit villosa Sather & Sublette, 1983: 42.-Mass. 9.-Tenn. olivetus. Ohio. Cricotopus olivetus Boesel, 1983: 88.-Ohio. Genus DONCRICOTOPUS Sather parafuscatus. Calif. Doncricotopas Srether, l98la:. 223. Cricotopus parafuscatus Sublette & Type-species, bicaudatus Saether (orig. Sublette, 1971: 9O.-Calif. des.). subfuscus. Calif. Cricotopus subfuscus Sublette & Sublette, bicaudatus. N.W.T. 1971: 98.-Calif. Doncricotopus bicaudatus Srether, 1981a: 225.-N.W.T.

Genus DIPLOCLADIUS Kieffer Diplocladius Kieffer in Kieffer & Genus EPOICOCLADIUS Zavrel Thienemann, 1908: 6. Type-species, Epoicocladius Zavrel in Sulc & Zavrel, cultriger Kieffer (mon.). 1,924: 368 (as subg. of Camptocladius Wulp, 1874). Type-species, cultriger. Widespread Can. to Nebr., Ohio, Camptocladius ephemerae Kieffer (orig. N.C., S.C., Grld.; PA. des.) - Camptocladius flavens Malloch. Diplocladius cultiger Kieffer in Kieffer & Thienemann, 1908: 6.-Germany flavens. Ont., Mont., N. Dak., s. to Tex., Diplocladius bilobatus Brundin, L956: Mich., Ill., s. to Fla.; PA. 71.-Sweden. Camptocladius f-lavens Malloch, 191-5b: 511.-Ill. 26 NEARCTIC CHIRONOMIDAE

Camptocladius ephemerae Kieffer in Sulc illoba. Calif. & Zavrel, 1924: 385.-Czechoslovakia. Euryhapsis illoba Oliver, 1981: 720.-Calif.

Genus ERETMOPTERA Kellogg Genus GEORTHOCLADIUS Strenzke Eretmoptera Kellogg, 1900: 82. Georthocladias Strenzke, 1941: 185. Type-species, browni Kellogg (orig. Type-species, Orthocladius luteicornis des.). Goetghebuer (orig. des.).

browni. Calif. Subgenus ATELOPODELLA Sether Eretmoptera browni Kellogg, 1900: Atelopodella Sather, 7982: 488. 82.-Calif. Type-species, curlicornus Sether (orig. des.).

Genus EUKIEFFERIELLA Thienemann curticornus. S.C. Eukiefferiella Thienemann, 1926: 325. Georthocladius curticornrzs Sether, 1982: Type-species, Dactylocladius longicalcar 488.-5.C. Kieffer, 1.9llc, sensu Thienemann (misident. = Spaniotoma gracei Subgenus GEORTIIOCLADIUS Strenzke Edwards, 1929), (orig. des.) (ICZN ruling required on the true type - fimbriosus. Tenn. species (ICZN 70a)). Georthocladius fimbriosus Sether & Adactylocladizrs Sether, 1,968: 464. Sublette, 1983: 32.-Tenn. Type-species,finsensis Sether (mon.) platystylus. Calif. = Spaniotoma minor Edwards. Georthocladius platystylus Sether & Sublette, 1983: 28.-Calif. brevinervis. Ill., Ohio, Pa., N. Mex. triquetrus. S.C. Orthocladius brevinervis Malloch, 1915b: Georthocladiu.s tiquetus Seether & 526.-Ill. Sublette, 1983: 25.-S.C. claripennis. N.W.T., B.C., Alta., N. Mex., wirthi. Calif. Grld.; PA.; Hawaii. Georthocladius wirthi Sather & Sublette, Chironomus claripennis Lundbeck, 1898: 1983: 30.-Calif. 281.-Grld. Spaniotoma hospita Edwards, 1929: 351.-England. Genus GYMNOMETRIOCNEMUS cyanea. Alas., ?Oreg.; PA. Goetghebuer Eukiefferiella cyanea Thienemann, 1936b: Gymnometriocnemus Goetghebuer, 1932a: 61.-Germany. 23 (as subg. of Metriocnemus Wulp, devonica. Widespread Can., U.S.A.; PA. 1874). Type-species, Metriocnemus Spaniotoma devonica Edwards, 1929: subnudus Edwards (Edwards, 7932b: 349.-England. 141). (Technically invalid as no unicalcar. Alta. type-species originally designated Adactylocladius unicalcar Sather, 1969: (ICZN 13b), but treated as above by 59.-Alta. usage).

. Subgenus GYMNOMETRIOCNEMUS Genus EURYHAPSIS Oliver Goetghebuer Euryhapsis Oliver, 7981: 7ll. Type-species, cilium Oliver (orig. des.). subnudus. B.C., Ont., N.S., Tenn., N.C.; PA. Metiocnemus subnudus Edwards, 1929: annuliventris. Calif. 316.-England. Metriocnemeus annulivenrrls Malloch, Gymnometriocnemus marionensis Sather, 191-5a: 46.-Calif. 1969: 729.-8.C. cilium. Y.T., N.W.T., Alta.; PA. Euryhapsis cilium Oliver, 1981: 714.-N.W.T. CHIRONOMIDAE DE I-A REGION NEARCNQUE 27

Subgenus RAPIIIDOCLADIUS Sether Genus HETEROTRISSOCLADIUS Spdrck Raphidocladizs Srether, 1983b: 211. Heterotris socl adius SpAr ck,'1.923: 94. Type-species, acigus Sether (orig. des.) Type-species, Metriocnemus cubitalis = Metriocnemus brumalis Edwards. Kieffer (Goetghebuer in Goetghebuer & Lenz,1940: 6) ?= Chironomus brumalis. 8.C., Man. to N.S., Tenn., S.C.; marcidus Walker. PA. Metriocnemus brumalis Edwards, changi. N.W.T., Alta. to Ont., N.Y., s. to Ill., 1929: 316.-England. Ind. Gymnometriocnemus acigus Snther, 1983b: Heterotrissocladius changi Srether, 197 5a: 214.-S.C. 47.-Man. cooki. Minn. Heterotrissocladius cooki Sether, I97 5a:. Genus HALOCLADIUS Hirvenoja 41.-Minn. Halocladius Hirvenoja, 1973: 106. hirtapex. Ont. Type-species, Chironomus varians Heterotrissocladius hirtape-x Sather, 1975a; Staeger (orig. des.). 39.-Ont. latilaminus. 8.C., Mont. Subgenus HALOCLADIUS Hirvenoja Heterotrissocladius latilaminas Sather, 1975a: 36.-8.C. variabilis. Man., Grld.; PA. maeaeri. Alas.: PA. Chironomus variabilis Staeger, 1839: Heterotrissocladius mriiiri Brundin, L949: 571.-Denmark. 812.-Sweden. marcidus, B.C., Ont., Qu6., N.Y., Mich., Nomen dubium in Ilalocladius N.C., S.C., Fla.; PA. Chironomus vitripennis Meigen, 1818: Chironomus marcidus Walker, 1'956a; 32.-Europe. 177.-England. oliveri. N.W.T., B.C. to Ont., s. to Calif., N.y.: pA. Genus HELENIELLA Gowin Heterotrissocladius oliveri Sether, I975a.: Heleniella Gowin. 1943: 116. 19.-N.W.T. Type-species, thienemanni Gowin (orig. subpilosus. N.W.T.; PA. des.). = Spaniotoma ornaticollis Dactylocladius subpilosus Kieffer in E,dwards. Kieffer & Lundbeck. 1911: 273.-Bear Island. curtistila. B.C., Alta., Ont. Heleniella curtistila Sether, 1969: 130.-8.C. Genus HYDROBAENUS Fries hirta. Ont., Qud., N.S. Hydrobaenus Fries, 1830: 177. Heleniella hirta Snther. 1969: 134.-Qu6. Type-species, lugubris Fries (orig. des.). parva. S.C., Tenn. Heleniella parva Sather, 1985b: -532.-5.C. calvescens. S. Dak. Hydrob aenus c alvescens Saether, 1976: 158.-5. Dak. Genus HETEROTANYTARSUS Splirck conformis. N.W.T., Qu6., Labr., Grld.; PA. Heterotanytarsas Spdrck, t923: 92. Chironomus conformis Holmgren, 1869: Type-species, Metiocnemus apicalis 42.-Spitsbergen. Kieffer (mon.). Hydrobaenus conformis conformis Holmgren (Srether, 1976: 83). nudalus. Ont. Hydrobaenus conformis Heterotanytarsus nudalus Sether, 1975b: labradorensis Sether. 1976: 263.-Ont. 89.-Qu6., Labr. perennis. B.C. Orthocladius natvigi Goetghebuer, 1933: Heterotanytarsus perennis Srether, I975b: 25.-Franz Joseoh Land. 260.-B.C. 28 NEARCTIC CHIRONOMIDAE

fusistylus. Alas., N.W.T., Alta., Mont., Colo., camptophleps Edwards, 1,929) (orig. Grld. des.). Orthocladius fusistylus Goetghebuer, 1933: 26.-Grld. ?boreoalpina. B.C., Alta.; PA. Hydrobaenus fusistylus var. fusistylus Smittia boreoalpina Goetghebuer, 1944: (Goetghebuer), Sather, 7976:- 94. 43.-Austria. Hydrobaenus fusistylus var. octomerus ?camptophleps. B.C.; PA. Srether, 7976: 99. Spaniotoma camptophleps Edwards, 1929: hudsoni. S. Dak. 353.-Great Britain. Hydrobaenus hudsoni Sether, 1976: halvorseni. N.W.T., N.B.; PA. 131.-S. Dak. Rheosmittia halvorseni Cranston & Sather. johannseni. Man., Ont., S. Dak., Minn., Ohio, 1986: 45.-Norwav. N.Y., Md., Ark., N.C., S.C. Orthocladius johannseni Sublette, 1967b: 504 (n. nam. for Orthocladius Genus LIMNOPHYES Eaton nivoriundus Johannsen, 1934: 348, Limnophyes Eaton, 1875: 60. preocc. Fitch, 1847).-N.Y. Type-species, pusillus Eaton (mon.). Orthocladius domus Sublette, 1966: 594.-Md. asquamatus. N.Y., Grld.; PA. Trissocladius hamiltoni Sether, 1969: Limnophyes eltoni var. asquamatus 42.-Man. Andersen, 1937 : 7 Z.-Grld. laticaudus. Alas. atomarius. N.Y., Grld.; PA. Hydrobaenus laticaudus Sether, 1976: Chironomus atomarius Zetterstedt, 1850: 80.-Alas. 3522.-Sweden. martini. N.W.T., N.B.; PA. borealis. Y.T. to Grld., N.8., s. to Minn., Hydrobaenus martini Sether, 1976: Tenn., N.C., S.C.; PA. 134.-N.W.T. Limnophyes borealis Goetghebuer, 1933: pilipes. Ont., Qu€., S. Dak., Ohio, Ill., Calif., 29.-Grld. N. Mex., N.C., S.C., Fla. Limnophyes spatulosus Srether, t975d: Orthocladius pilipes Malloch, 1915b: 1055.-Minn. 522.-\1. carolinensis. Ont., Minn., Tenn., N.C., S.C. pilipodex. S. Dak., Ark., Ala. Limnophyes carolinensis Srether, I97 5d: Hydrobaenus pilipodex Sether, 1976: 10-50.-s.c. 144.-5. Dak. coloradensis. Colo. scapulapilosus. Alas., Ont. Limnophyes coloradensis Sether, 197 5d: Hydrobaenus scapulapilosus Sether, L976: 1052.-Colo. 124.-Ont. eltoni. Grld.: PA. spinnatis. Ont., Qu6.; PA. Camptocladius eltoni Edwards, L922: Hydrobaenus spinnatis Sather, 1976: 203.-Bear Island. 152.-Qu€. fumosus. Ont., N.S., N.Y., N.C., S.C., Ga. travisi. Mass. Camptocladius fumosus Johannsen, 1905: Hydrobaenus travisi Sa.ther, 1989b: 261.-N.Y. 55.-Mass. Limnophyes cristatissimus Sether, 7975d: tumidistylus. N.W.T. 1032.-S.C. Hydrobaenus tumidistylus Sether, 1976: globifer. Y.T., N.W.T., Alta., Man., Ont., 160.-N.W.T. Grld.: PA. virgo. Ont. Camptocladius globifer LundstrOm, 1915: Hydrobaenus virgo Sather, L976: 16.-U.S.S.R. 128.-Ont. Limnophyes folliculatus Sether, L97 5d: 1050.-Man. groenlandiensis. Grld.

Genus KRENOSMITTIA Thienemann & Limnophy e s gr o e nl and ie nsis Andersen, Krtiger 1937. 70.-Grld. Krenosmittia Thienemann & KrUger, L939: hamiltoni. Alas. to N.W.T., B.C., Ont., Calif., 253. Type-species, Smittia gynocera Grld. Edwards, 1937, sensu Thienemann & Limnophyes hamiltoni Saether, 1969: Krriger (misident. = Spaniotoma 101.-B.C. CHIRONOMIDAE DE LA REGION NEARCTIQUE 29

hastulatus. B.C., Ont., S. Dak., Minn. hyporheicus. Pa., Va. Limnophyes hastulatus Saether, 1975d. Lopescladius hyporheicus Coffman & 1054.-S. Dak. Roback, 7984: l3l.-Pa.26 immucronatus. Widespread Can. s. to N. Mex., Nebr., N.C., S.C.; ?PA. Subgenus LOPESCLADIUS Oliveira Limnophyes immucronatus Sather, 1969: 103.-8.C. inermis. Kans. Limnophyes hudsoni Sether, 1975d: Lopescladius inermis Srether, 1983c: 1032.-Nebr. 293.-Kans. margaretae. Man. verruculosus. Ill., Mex.; NT. Limnophyes margaretae Srether, 797 5d: Lopescladius vetuculosus Sether, 1983c: 1053.-Man. 289.-Mex. minimus. Idaho, Colo., Ohio, N.Y.; PA. Chironomus minimus Meigen, 1818: 47.-Europe. Genus MESOCRICOTOPUS Brundin ninae. N.W"T., Man., Ont. Mesocricotopas Brundin, 1,956: l\4. Limnophyes ninae Snther, 1975d: Type-sp ecies, Tric h o cl adius 1048.-Man. thienemanni Goetghebuer (orig. des.). nudiradius. Man., S. Dak. Limnophyes nudiradius Srether, 197 5d:. thienemanni. N.W.T., Alta., Man.; PA. 1048.-5. Dak. Trichocladius thienemanni Goetghebuer, pilicistulus. Y.T., B.C., Oreg., S. Dak. 1940: 69.-Sweden. Limnophyes pilicistulus Sether, 197 5d: 1049.-5. Dak. prolongatus. Y.T., N.W.T., Man., Grld.; PA. Genus MESOSMITTIA Brundin C amptocladius pentaplastus v at. Mesosmittia Brundin, 19-56: 163. prolongatus Kieffer in Thienemann, Type-species: Spaniotoma flexuella 1921: 810.-Germany. Edwards, (orig. des.). (Spaniotoma pumilio. Grld.; PA. flexuella is type-species of Chironomus pumilio Holmgren, 1869: Pseudorthocladius Goetghebuer (q.u. by 41.-Spitsbergen. Edwards, 1932b: 141) ICZN ruling recisus. S. Dak., Nebr., N. Mex. required). Limnophyes recisus Srether, I975d: 10-52.-Nebr. acutistylus. N. Mex. Mesosmittia acutistylus Sether, 19859: 43.-N. Mex. Genus LIPUROMETRIOCNEMUS Sether lobiga. N. Mex. Lipurometriocnemus Sether, 1981b: 13. Mesosmittia lobiga Sather, 19859: 4-5.-N. Type-species, glabalus Seether (orig. Mex. des.). Spaniotoma .flexuella Edwards, 1929: 349 (sensu sublette & Sublette, 1979;77, vixlobatus. Y.T., S.C. misident.). Lipur ome triocnemus vixl ob atus Srther, mina. Ga. 1982:477.-5.C. Mesosmittia mina Snther, 19859: 46.-Ga. patrihortae. Ky., S.C., Ga. Mesosmittia patrihortae Sather, 1985g: Genus LOPESCLADIUS Oliveira 47.-S.C. Lopescladius Oliveira, 1967: 4I7. prolixa. Kans., Ill., Ky., Tenn., N. Mex., Ga. Type-species, minutis s imus Oliveira Mesosmittia prolixa Srether, 198-5g: (orig. des.). 48.-Kans. Cordites Brundin, 1966: 428 (nom. nud.). tora. S. Dak. Mesosmittia tora Sether, 19859: 50.-S. Subgenus CORDIELLA Coffman & Roback Dak. Cordiella Coffman & Roback, 1984: 130. Spaniotoma flexuella Edwards, 1929: 349 Type-species, hyporheicus Coffman & (sensu Sether, 1973a: 59, misident.). Roback (orig. des.). 30 NEARCTIC CHIRONOMIDAE

Genus METRIOCNEMUS Wulp ursinus. N.W.T., n. Qud., Grld.; PA. Metriocnemus Wulp, 1874: 136. Chironomus ursinus Holmgren, 1869: Type-species, Chironomus albolineatus 39.-Spitsbergen. Meigen (Coquillett, 1910: 569). yaquina. Oreg. Metriocnemus yaquina Cranston & Judd, abdominoflavatus. Ky., N.C., s. to Miss. & 1987: -534.-Ores. Fla.; NT. M e tr io cnemus ab domino -fl av atus Picado, I9l3: 284.-Costa Rica. Genus NANOCLADIUS Kieffer aequalis. Ill., N.Y., Ohio. Nanocladius Kieffer, 1913d: 31. Metriocnemus aequalis Johannsen, 1934: Type-species, vitellinus Kieffer (orig. 348.-N.Y. des.). albolineatus. Colo., N.Y., Ohio, Grld.; PA. Microcric otopns Thienemann & Chironomus albolineatus Meigen, 1818: Harnisch, 1932: 737. 39.-?Germany. Type-species, Cr i c ot opus Chironomus atratulus Zetterstedt. 1850: alb ic ornis Goetghebuer 3590.-Norway. (Goetghebuer in Goetghebuer & brusti. Mari. Lenz,1944: 114) = Chironomus Metriocnemus brusti Saether. 1989a: bicolor Zetterstedt. 407.-Man. edwardsi. B.C., Calif. Subgenus NANOCLADIUS Kieffer Metriocnemus edwardsi Jones, 1916: 385.-Calif. alternantherae. N. Mex. to N.C., s. to Fla. fuscipes. S. Dak., N.Y., Grld.; PA. Nanocladius alternantherae Dendy & Chironomus fuscipes Meigen, 1818: Sublette, 19-59: -510.-Ala. 49.-?Germany. anderseni, N.W.T., Sask., Man., S. Dak., knabi. Man. to Nfld., s. to Miss. & Fla.; Nebr., Ill. Bermuda. Nanocladius anderseni Srether, 7977b: Metriocnemus knabi Coquillett, 1904: 16.-Nebr. 11.-Mass. crassicornus. Sask., Nebr., Iowa. Iongipennis. Alas.; PA. Nanocladius crassicornus Sether. 1977b: Smittia longipennis Holmgren, 1883: 5 1 .-Iowa. 181.-Novaya Zemlya. distinctus. N.W.T., B.C. to. Ont., s. to Nebr., obscuripes. N.W.T., Minn., Ont., N.8., S.C., Ill., N.c., s.c., Fla. Grld.: PA. Orthocladius distinctus Malloch. 1915b: Chironomus obscuripes Holmgren, 518.-Ill. 1869: 38.-Spitsbergen. incomptus. Ont., N.C., S.C. Metriocnemus hygropetricus Kieffer, 79 72c: Nanocladius incomptus Sather. 1977b: 86.-Germany. 43.-S.C. Metriocnemus longitarsus Goetghebuer, mallochi. Ill.. Ohio. 1921: 77.-Belgium. Nanocladius mallochi Sublette, 1970: 70 perfuscus. N.W.T. (n. nam. for Orthocladius distinctus Metriocnemus perfuscus Malloch, 1934: var. bicolor Malloch, 1915b: 519, 18.-N.W.T. preocc. Zetterstedt, 1838).-Ill. picipes. Grld.; PA. minimus. N.C.. S.C. Chironomus picipes Meigen, 1818: Nanocladius minimus Sether, I977b: 52.-Not stated, ?Germany. 21.-S.C. polaris. N.W.T., Grld. rectinervis. Man., Ont., N.Y., Colo., Nebr., Metriocnemus polaris Kieffer, 1.926; N.C., S.C.; PA. 87.-N.W.T. Cricotopus rectinervis Kieffer, 1911c: tristellus. Grld.; PA. 199.-Germany. Metriocnemus tristellus Edwards, 1929: spiniplenus. Sask., Ont., N.B., Ohio, N.C., 312.-England. S.C. Nanocladius spiniplenus Sather, L977b: 37.-Ont. CHIRONOMIDAE DE LA REGION NEARCTIQUE 31

Subgenus PLECOPTERACOLUTHUS Steffan dubitatus. N.Y., Pa., Ohio. Plecopteracoluthus Steffan, 1965: 1330. Orthocladius dubitatus Johannsen, 1942: Type-species, downesi Steffan (orig. 72.-N.Y. des.). gelidus. N.W.T., Grld.; PA. Orthocladius gelidus Kieffer, 1922b: branchicolus. Sask., Ont., N.Y. 13.-Novaya Zemlya. Nanocladius branchicolus Srether, 1977b:. mixtus. N.W.T.; PA. 12.-Ont. Chironomus mixtus Holmgren, 1869: downesi. Ont., Qu6., Mich., N.C. 45.-Bear lsland. Plecopteracoluthus downesi Steffan, 1965: nanseni. N.W.T. 1331.-Qu6. Orthocladius nanseni Kieffer, 1926: 84.-N.W.T.

Genus OLMRIDIA Srether Subgenus EUORTHOCLADIUS Thienemann Oliveridia Sather, 1980a: 399 (n. nam. for Euorthocladius Thienemann, 1.93-5: 201 (as Oliveria Sather). genus). Type-species, Orthocladius Oliveria Srether, 1976: 48. (preocc. thienemanni Kieffer (orig. des.). Sutherland, L965). Type-species, Rhe orthocl adius'[hienemann, I 93-5: 205 Triss ocl adius tric ornis Oliver (as genus). Type-species, Orthocladius (orig. des.). saxicola Kieffer (orig. des.). Lapporthocladius Thienemann in hugginsi. Kans. Thienemann & Kriiger, 1,937: 266 (as Oliveridia hugginsi Ferrington & Sather, genus). Type-species, Orthocladius 1987: 452.-Kans. abiskoensis Edwards (orig. des.). tricornis. N.W.T., Grld.; PA. Trissocladius tricornis Oliver. 1976: abiskoensis. Can., U.S.A.; PA. 1055.-N.W.T. Orhocladius abiskoensis Edwards, 1937: 144.-Sweden. difficilis. N.W.T.. Grld. Genus OREADOMYIA Kevan & Chironomus difficilis Lundbeck, 1898: Cutten-Ali-Khan 282.-Grld. Oreadomyia Kevan & Cutten-Ali-Khan, rivicola. N.W.T., Sask., Grld.; PA. 1975: 856. Type-species, albertae Orthocladius rivicola Kieffer, 191 1c: Kevan & Cutten-Ali-Khan (orig. des.). 181.-Germany. rivulorum. Sask.: PA. albertae. Alta. Orthocladius rivulorum Kieffer, 1909: Oreadomyia albertae Kevan & 48.-Germany. Cutten-Ali-Khan. 197-5: 856.-Alta. saxosus. Can., U.S.A.; PA. Spaniotoma saxosus Tokunaga, 1939: 326.-Japan. Genus ORTHOCLADIUS Wulp thienemanni. Can., U.S.A., Grld.; PA. Orthocladius Wulp, 1874: 132. Orthocladius thienemanni Kieffer in Type-species, Tipula stercoraria De Kieffer & Thienemann, 1906: Geer, 1776:. 388 sensu Wulp (misident. 1.13.-Germany- = Chironomus oblidens Walker) (coquilleft, 1910: 581), (ICZN ruling Subgenus ORTIIOCLADIUS WulP required on conservation of misident. type-species (ICZN 70a)). annectens. N.W.T., B.C. to Ont., Minn., N.Y., N.J., Pa., s. to Miss. & Fla. Subgenus EUDACTYLOCLADIUS Orthocladius annectens Sether, 1969: Thienemann 61.-Man. Eudacrylocladias Thienemann, 193-5: 206 appersoni. Alas., Y.T., Mont., Colo, Calif., (as genus). Type-species, N.Y. Dactylocladius fuscimanus Kieffer Orthocladius appersoni Soponis, 1977: (orig. des.). 27.-Calif. 32 NEARCTIC CHIRONOMIDAE

barbicornis. Wash., S. Dak., Minn.; PA. knuthi. N.W.T., Grld.; PA. Tipula barbicornis Linnaeus, 1-767: Orthocladius knuthi Soponis, 1977: 974.-Europe. 58.-Grld. carlatus. Sask., Minn., Ohio, Pa., N.Y., Md., lapponicus. N.W.T., Ohio, Grld.; PA. Oreg., Calif., N. Mex., Ark. Orthocladius lapponicus Goetghebuer, Hydrobaenus carlatus Roback, 1957a: 1940: 60.-Sweden. 77.-Pa. mallochi. N.W.T., Wash. to Ont., s. to Calif., charensis. N.W.T., Grld. N. Mex., Ill., Md. Orthocladius charensis Soponis, 1977: Orthocladius mallochi Kieffer, 1919b: 191 33.-N.W.T. (n. nam. for Orthocladius lacteipennis clarkei. Ont., Minn., Iowa, Ill., Pa., Tex. Malloch, 1915b: 524, preocc. Orthocladius clarkei Soponis, 1977: Lundstrdm, 1910).-Mich. 37.-Ont. manitobensis. N.W.T., Man., Minn. cooki. Minn., Pa. Orthocladius manitobensis Sather, 1969: Orthocladius cooki Soponis, 1977: 69.-Man. 41.-Minn. nigritus. Sask. to Qu6., s. to Minn. & Va., curtiseta. B.C. Utah. Orthocladius curtiseta Srether, 1973a: 58 Orthocladius nigritus Malloch, 1915b: (n. nam. for Orthocladius breviseta 525.-Md. Sether, 1969: 65, preocc. Kieffer, oblidens. N.W.T., Qu6.; PA. 1923c).-B-C. Chironomus oblidens Walker, 1956a: decoratus. N.W.T., Grld.; PA. 180.-England. Chironomus decoratus Holmgren, 1869: obumbratus. Alas., widespread s. of N.W.T. 43.-Spitsbergen. Orthocladius obumbratus Johannsen, 1905: dentifer. Alas., N.W.T., Oreg. to Ont., Nev., 281.-N.Y. Wyo.; PA. Hydrobaenus paradorenus Roback, L957a: Orthocladius dentifer Brundin, L947: 79.-Pa. 21.-Sweden. oliveri. Oreg. to Qu6., s. to Tex., Ala., Pa. dorenus. Y.T., N.W.T., Oreg. to N.B., N.Y., Orthocladius oliveri Soponis, 1977: Colo., N. Mex., N.C., S.C. 87.-Qu€. Hydrobaenus dorenus Roback, 1957a: robacki. Sask., Pa. 78.-Pa. Orthocladius robackl Soponis, 1977: Orthocladius currani Sublette, 1967a:- 92.-Pa. 315.-N.Y. smolandicus. N.W.T., Alta., Man., Minn.; PA. excavatus. Alta.; PA. Orthocladius smolandicus Brundin, 1947: Orthocladius excavatus Brundin, 1947: 22.-Sweden. 20.-Sweden. subletti. Idaho, Mont., Calif., Utah & Colo. ferringtoni. Kans. Orthocladius subletti Soponis, 1977: Orthocladius ferringtoni Soponis, 1983: 9-5.-Idaho. 571.-Kans. trigonolabis. N.W.T., Colo., IJtah, Wyo., N. frigidus. Grld.; PA. Mex.; PA. Chironomus frigidus Zetterstedt, L838: Orthocladius trigonolabis Edwards, 1924b: 811.-Grld., ?Sweden. 170.-Spitzbergen. groenlandensis. Grld. wiensi. Man., S. Dak., Minn. Orthocladius groenlandensjs Goetghebuer, Orthocladius wiensi Sether, 1969: 1933: 26.-Grld. 73.-Man. hazenensis. N.W.T. Orthocladius hazenensis Soponis, 1977: Subgenus POGONOCLADIUS Brundin 56.-N.W.T. Pogonocladizs Brundin, 1956: 99. heflenthafi. Alas., N.W.T., Calif., N. Mex. Type-species, Chironomus consobrinus Orthocladius hellenthali Soponis, 1977: Holmgren (orig. des.). 57.-Calif. knabeni. Grld.; PA. consobrinus. N.W.T., Ont., Grld.; PA. Orthocladius knabeni Goetghebuer, L933: Chironomus consobrinus Holmgren, 1869: 26.-Grld. 44.-Spitsbergen. CHIRONOMIDAE DE LA REGION NEARCTIQUE -)-)

Subgenus SYMPOSIOCLADIUS Cranston alpicola. N.W.T., Labr.; PA. Symposiocladias Cranston, 1982: 419 (as Chironomus alpicola Zetterstedt, 1850: genus). Type-species, Orthocladius 3500.-Sweden. lignicola Kieffer (orig. des.). conversus. N. Mex.; PA. Chironomus conversus Walker, I956a: lignicola. Y.T., B.C. to Qu6., Oreg., Minn., 175.-England. Tex.; PA. quadrinodosus. Alas., N.W.T., Grld.; PA. Orthocladius lignicola Kieffer in Potthast, P ar acl adius qu adrino d o sas Hirvenoja, 1915: 273.-Germany. 1973: 98.-Sweden. Orthocladius /ryoni Soponis, 1977: 100.-Minn. Genus PARACRICOTOPUS Thienemann & Orthocladius unassigned to subgenus Harnisch Paracricotopus Thienemann & Harnisch, spitzbergensis. Grld.; PA. 1932: 136. Type-species, Cricotopus Dacrylocladius spitzbergensis Kieffer in niger Kieffer (Brundin, 1956: 119). Kieffer & Thienemann, 1919: 1 16.-Spitsbergen. glaber. S.C. Paracricotopus glaber Srether, 1980b: 138.-S.C. Genus PARACHAETOCLADIUS Wtilker millrockensis. N.C., S.C., Ga. Parachaetocladius 'tNilker, 1959: 44 (as Paracricotopus millrockensis Caldwell, subg. of Chaetocladius Kieffer, 1911c). 198-5: 161.-Ga. Type-species, abnobaeus Wtilker mozleyi. Ga. (mon.). Paracricotopus mozleyi Steiner, 1983: Habrobaenus Sether, 1,977 a:, 2354. 329.-Ga. Type-species, hudsoni Srether (orig. des.). Genus PARAKIEFFERIELLA Thienemann abnobaeus. Man., Ont., N.S., Pa., Tenn., Parakiefferiel/a Thienemann, 1936a: 19-5. N.C., S.C., Ga.; PA. Type-species, Spaniotoma coronata Chaetocladius abnobaeus WUlker, 1959: Edwards (orig. des.). 44.-Germany. Maricladius Sublette, t970:. 84. Habrobaenus hudsoni Srether, 1977 a: Type-species, C amPtocladius 2355.-S.C. subatetimus Malloch (orig. Parachaetocladius abnobeas (error), des.). Srether & Sublette, 1983: 39. hirtipectus. B.C. bathophila. Can., U.S.A.; PA. Parachaetocladius hirtipeclus Sether, Dactylocladius bathophilus Kieffer, I972c: 1969: 100.-8.C. 88.-Germany. imberbus. B.C., Calif. coronata. Can., U.S.A.; PA. Parachaetocladius imberbus Sather & Spaniotoma coronata Edwards, 1929: Sublette, 1983: 44.-Calif. 354.-Great Britain. gracillima. Alas.; PA. Camptocladius gracillimus Kieffer, L924b: Genus PARACLADIUS Hirvenoja 74.-Germany. Paracladius Hirvenoja, 1973:. 90- (n. nam. nigra. Alas., N.W.T.; PA. for P aratricho cl ad ius Thienemann). Parakiefferiella nigra Brundin, 1949: P ar atrichocladias Thienemann, 827.-Sweden. 1942: 3L4. (preocc. Santos subaterrima, N.W.T., Sask. to Qu6., Ill. Abreu, 1918). Type-species, Camptocladius sub aterrimrzs Malloch, Chironomus inserpens Walker 1915b:512.-Ill. (orig. des.). = Chironomus Parakiefferiella torulata Sether, 1969: conversus Walker. 138.-Man. -)+ NEARCTIC CHIRONOMIDAE

Genus PARALIMNOPHYES Brundin exagitans. S. Dak., Colo., Kans., Wis., Ill., Paralimnophyes Brundin, 1956: 129. Ohio, N.Y. Type-species, Camptocladius Metriocnemus exagitans Johannsen, 1905: hydrophilus Goetghebuer (orig. des.). 303.-N.Y. Metriocnemus brachyneura Malloch, 1915b: arcticus. Alas., N.W.T.; PA. 498.-Ill. Paralimnophyes arcticus Brundin, L956: impensus. Ont., Grld.; PA. 130.-Sweden. Chironomus impensus Walker, 19-56a: 184.-England. nasthecus. Man., S. Dak. Genus PARAMETRIOCNEMUS Goetghebuer Paraphaenocladius nasthecrs Sether, 7969: Parametriocnemus Goetghebuer, I932a: 22 1.23.-Man. (as subg. of Metriocnerzus Wulp, l8'l 4). Type-species, Metriocnemus stylatus Kieffer (mon.). Genus PARASMITTIA Strenzke Parasmittia Strenzke. 1950: 270. eoclivus. Qu6. Type-species, carinata Strenzke (orig. Parametriocnemus eoclivus Srether, 1969: des.). 108.-Qu6. graminicola. Y.T., N.W.T., Alta., Grld. carinata. N.S.; PA. Chironomus graminicola Lundbeck, 1898: Parasmittia carinata Strenzke, 19-50: 278.-Grld. 270.-Austria. Germany. hamatus. N.Y. to Maine, s. to Conn. Metriocnemus hamatus Johannsen, 1934: 349.-N.Y. Genus PARATRICHOCLADIUS Santos Abreu lundbeckii. Alta. to Qu6., s. to Fla., Calif. to Paratrichocladias Santos Abreu, 1978: 204 Tex., Grld. (as subg. of Orthocladius Wulp, 1874). Metriocnemus lundbeckii Johannsen, 1905: Type-species, Orthocladius tenuipes 302 (n. nam. for Chironomus nanus Becker, (Hirvenoja, 1973: 88) = Lundbeck, 1898: 28-5, preocc. Meigen, Chironomus rufiventris Meigen. 1818).-Grld. Synuicotopus Brundin, 19-56: 106. Metriocnemus innocuus Curran, 1930: Type-species, Chironomus rufiventris 33.-N.Y. Meigen (orig. des.). stylatus. Ont.; PA. Metriocnemus stylatus Kieffer, 1924b: nitidus. Man., Utah, Ill., Iowa, N.Y. 97.-Germany. Orthocladius nitidus Malloch, 1915b: vespertinus. Alta. 515.-Ill. Parametriocnemus vespertinus Srether, Trichocladius helis Roback, L957b: 1969: 119.-Alta. 1-5.-Utah. Syncricotopus fontinalis Sether, 1969: 77.-Man. Genus PARAPHAENOCLADIUS Thienemann rufiventris. Ont.; PA. Paraphaenocladius Thienemann in Spdrck Chironomus rufiventris Meigen, 1830: & Thienemann, 1924: 223. 249.-Middle Europe. Type-species, Metriocnemus skirwithensis. N.B., N.S.; PA. ampullaceus Kieffer (orig. des.) = Spaniotoma skirwithensis Edwards, 1929: Chironomus impensus Walker. 329.-England.

brevinervis. N.W.T. Chironomus brevinervis Holmgren, Genus PARATRISSOCLADIUS Zavrel 1869: 39.-N.W.T. Paratissocladius Zavrel, 1937: 10 (as Metriocnemus despectus Kieffer, 1926: subg. of Trissocladius Kieffer in 87.-N.W.T. Kieffer & Thienemann 1908). debilipennis. N.Y., Grld. Type-species, Trissocladius fluviatilis Chironomus debilipennis Lundbeck, 1898: Goetghebuer (orig. des.) = 286.-Grld. Chironomus excerptus Walker. CHIRONOMIDAE DE LA REGION NEARCTIQUE -1-')

Iawrencensis. Ont. obvius. N.W.T., n. Qu6.; PA. Paratrissocladius lawrencen.rir Cranston & Chironomus obvius Walker, 1956a: Oliver, 1988a: 445. -Ont. 174.-England. Subgenus MONOPSECTROCLADIUS Wtilker Genus PARORTHOCLADIUS Thienemann Monopsectrocladius Wtilker, 1956: 10. Parorthocladius Thienemann, 1935; 213. Type-species, Spaniotoma calcarata Type-species, Dactylocladius nudipennis Edwards (orig. des.). Kieffer (orig. des.). calcaratus. N.W.T., Alta.; PA. IJnnamed species.-B.C., Oreg., Mont., Spaniotoma calcarata Edwards, 1929: Alta., Ont. 333.-England.

Subgenus PSECTROCLADIUS Kieffer Genus PLATYSMITTIA Sether Platysmittia Srether, 1982: 503. barbatimanus. N.W.T., Grld.; PA. Type-species, fimbriata Sather (orig. Psectrocladius barbatimanus Kieffer, 7926: des.). 82.- N.w.T. barbimanus. N.W.T., Sask., S. Dak., Grld.; bilyji. Man. PA. Plarysmittia bilyji Srther, 1985a: Spaniotoma barbimanus Edwards, 1929: 528.-Man. 333.-England. fimbriata. Tenn. Psectrocladius armatus Andersen, 1937: Platysmittia fimbriata Srether, 1982: 56.-Grld. 50-5.-Tenn. fennicus. N.W.T., Labr., Grld.; PA. Psectrocladius fennicus StorE 1939: 24.-Finland. Genus PLIIUDSONIA Seether limbatellus. N.W.T., S. Dak., Grld.; PA. Plhudsonia Saether, 1982: 466. Chironomus limbatellus Holmgren, 1869: Type-species, partita Sether (orig. 44.-Spitzbergen. des.). Psectrocladius edwardsi Brundin, 1949: 816.-Sweden. partita. N.S., Tenn. psilopterus. Ohio. Plhudsonia partita Sather, 1982: Orthocladius psilopterus Kieffer in Kieffer 467.-Tenn. & Thienemann,1906:144.-Germany. semicirculatus. Ont. Psectrocladius semicirculatus Sr,ther, t969: Genus PSECTROCLADIUS Kieffer 79.-Ont. Psectrocladizs Kieffer, 1906a: 26 (as simulans. Alta., Sask., Ont., N.Y., Ohio; PA. subgenus of Orthocladirzs Wulp, 1874). Spaniotoma simulans Johannsen, 1937a: Type-species, Orthocladius psilopterus 67.-N.Y. Kieffer (Kieffer, 1906b: 356.). sordidellus. Not NE. Chironomus sordidellus Zetterstedt, 1838: Subgenus ALLOPSECTROCLADIUS Wiilker 814.-Sweden. Allopsectrocladius Wnlker, 1956:12. vernalis. Ont., Minn. to N.Y., s. to Miss. & Type-species, Chironomus dilatatus Fla., Wyo., Utah, Colo. Wulp (orig. des.). Orthocladius vernalis Malloch, 1915b: -520.-Ill. flavus. Sask., S. Dak., Ill., N.Y., Pa. zetterstedti. B.C., Ont.; PA. Orthocladius flavus lohannsen, 1905: Psectrocladius zetterstedtl Brundin, 1949: 270.-N.Y. 816.-Sweden. nigrus. Alta., Pa. Psectrocladius nigrus Roback, 1957 a: 88.-Pa. 36 NEARCTIC CFIIRONOMIDAE

Psectocladius unassigned to subgenus destitutus. Ont., N.B., N.S., S.C. Pseudorthocladius destitutus Sather & brevicosta. Wash. Sublette, 1983: 54.-S.C. Psectrocladius brevicosta Kieffer, 1917: dumicaudus. B.C., Calif. 362.-Wash. Ps eudorthocladius dumic audus Se,ther. elatus. N.Y. s. to Fla., Neb. s. to Miss. 1969: 726.-8.C. Psectrocladius elatus Roback, 1957a: lunatus. Mich. 89.-N.J. Pseudortocladius lunatus Sather & pilosus. S. Dak., N.J. Sublette. 1983: 63.-Mich. Psectrocladius pilosus Roback, L957a:- macrostomus. Fla. 88.-N.J. Pseudorthocladius macrostomus Soponis, polaris. N.W.T. 1980: 486.-Fla. Psectrocladius pol aris Kief.fer, 1,926: macrovirgatus. S.C., Ga. 83.-N.W.T. Pseudorthocladius macrovirgatus Saether & spinifer. Calif. Sublette. 1983: 87.-S.C. Orthocladius spinifer Johannsen, 1928: morsei. Qu6., N.B., S.C. 34.-Calif. Pseudorthocladius morsei Sether & Sublette, 1983: 85.-S.C. paravirgatus. S.C. Genus PSEUDORTHOCLADIUS Goetghebuer Pseudorthocladius paravirgatus Sether & Pseudorthocladius Goetghebuer, 1932a: 93 Sublette, 1983: 84.-S.C. (as subg. of Orthocladlus Wulp, 1874). pilosipennis. Ont.; PA. Type-species, Hydrobaenus curtistylus Pseudorthocladius pilosipennis Brundin, Goetghebuer (Goetghebuer in 7956: 139.-Sweden. Goetghebuer & Lenz, 1943: 73). rectilobus. Ga. (Technically nom. nud. as no Pseudorthocladius rectilobus Srether & type-species designated in original work Sublette, 1983: 58.-Ga. (ICZN 13b), see Crdnston, 197-5). tricanthus. N.I|., S.C. Pseudorthocladius tricanthus Sather & Subgenus LORDELLA Sather & Sublette Sublette, 1983: 67.-5.C. Lordella Saether & Sublette, 1983: 95. uniserratus. B.C., Ont., Calif., N. Mex., Type-species, comans Saether & Tenn., S.C. Sublette (orig. des.) = wingoi Sather Pseudorthocladius uniserratus Sa-ther & & Sublette. Sublette, 1983: lZ.-Tenn. virgatus, Ont., N.S., S.C., Tenn. wingoi. N.S., N.C., S.C. Pseudorthocladius virgatus Sether & Pseudorthocladius wingoi Sather & Sublette, 1983: 81.-S.C. Sublette, 1983: 97.-N.C. Pseudorthocladius comans Srether & Sublette, 1983: 9-5.-S.C. Genus PSEUDOSMITTIA Goetghebuer Pseudosmittia Goetghebuer, I932a: 726 (as Subgenus PSEUDORTHOCLADIUS subg. of Smittia Holmgren, 1869). Goetghebuer Type-species, Spaniotoma angusta Edwards (Edwards 1932c: 741). amplicaudus. S.C. (Technically invalid, no type-species Pseudorthocladius amplicaudus Sether & originally designated, ICZN 13(b)). Sublette, 1983: 56.-S.C. Prosmittia Brundin, 1956: 165. clavatosus. S.C. Type-species, Pseudosmittia jemtlandica Pseudorthocladius clavato.ra.r Sether & Brundin (orig. des.). Sublette, 1983: 58.-S.C. curticornus. Tenn. digitata. Fla.; NT. Pseudorthocladius curticornus Snther & Pseudosmittia digitata Sather, 1981b: Sublette, 1983: 74.-Tenn. 25.-St. Lucia. curtistylus. Ont., N.S.; PA. forcipatus. Ont", N.B., N.S., Vt., N. Mex.; P s eud ortho cladius curtisfy/us Goetghebuer, PA. 1921: 101.-Belsium. C amptochironomus forcipatus Goetghebuer, I92l: 87.-Belgium. CHIRONOMIDAE DE LA REGION NEARCTIQUE 3l

Smittia triappendiculata Goetghebuer, Rheocricotopus kenorensis Seether, 1969: l93l:. 217.-Belgium. 88.-Ont. nanseni. Alas. to N.W.T., Man., Ont., N.B., Calif., Grld. Subgenus RHEOCRICOTOPUS Thienemann Camptocladius nanseni Kieffer, L926: & Harnisch 85.-N.W.T. oxoniana. Grld.; PA. amplicristatus. S.C. Camptocladius oxonianus Edwards, 1,922: Rhe o cric o topus amplicris talas Saether, 204.-Bear Island. 1985i: 95.-S.C setavena. B.C. effusoides. S. Dak.; PA. Pseudosmittia setavena Sather. 1969: Rheocricotopus effusoides Srether, 1985i: 144.-B.C. 98.-S.D. effusus. Ill., N.C., S.C.; PA. Chironomus effusus Walker, 1956a: Genus PSILOMETRIOCNEMUS Sather 180.-England. Psilometriocnemus Srether, 1969: 105. Orthocladius striatus Malloch, 1915b: Type-species, tiannulatus Sether (orig. 517.-Ill. des.). eminellobus. Alta., Ont., Tenn., N.C., S.C. Rheocricotopus eminellobas Sether, 1969: triannulatus. Ont. to N.S., Minn., N.H., 85.-Alta. Kans., Tenn., S.C. pauciseta. 8.C., Alta. Psilometriocnemus tiannulatus Sether, Rheocricotopus pauciseta Srther, 1969: 1969: 107.-N.H. 89.-8.C. Psilometriocnemus cristatus Srether. 1982: tuberculatus. S.C., Ga. 480.-S.C. Rheocric otopus tub ercul a/as Caldwell, 1984: 84.-Ga. unidentatus. Ohio. Genus RHEOCRICOTPUS Thienemann & Rheocricotopus unidentatus Saether & Harnisch Schnell, 1988: 68.-Ohio. Rheocricotopas Thienemann & Harnisch. L932: 135. Type-species, Chir onomus Genus RHEOSMITTIA Brundin effusus Walker (Brundin, 1.956: Rheosmittia Brundin in Cranston & 1 18). S&ther, 1986: 33 (as subg. of P arakiefferiel/a Brundin 1956: 1-50, Subgenus PSILOCRICOTOPUS Sether invalid). Type-species, Parakiefferiella Psilocricotopas Sether, I985i: 72. spinicornis Brundin (Brundin in Type-species, Cricotopus atipes Kieffer Cranston & Sether, 1986: 33) (orig. des.). Unnamed species.-WidesPread NE. chapmani. Grld.; PA. Spanitoma chapmani Edwards, 1935: 471.-Grld. Genus SAETIIERIELLA Halvorsen conflusirus, S.C. Saetheriella Halvorsen, 1982: 132. Rheocricotopus conflusirus Saether, 1985i: Type-species, amplicristata (orig. des.). 72.-S.C. glabricollis. Man., Pa., Tenn., N.C., S.C.; PA. amplicristata. Tenn. Chironomus glabricollis Meigen, 1830: Saetheriella amplicristata Halvorsen, 1982: 248.-Europe. 133.-Tenn. Trichocladius gouini Goetghebuer in Gouin, 1936: 167.-France. Trichocladius extatus Roback, t957a: Genus SMITTIA Holmgren 84.-Pa. Smittia Holmgren, 1869: 47. Type-species, robacki. Ont., Mont. to N.Y., s. to Miss. & Chironomus brevipennis Boheman Fla. (mon.). Trichocladius robacki Beck & Beck. 1964: 204.-Fla. 38 NEARCTIC CHIRONOMIDAE

arctica. Alas. Genus SUBLETTIELLA Srether Smittia arctica Malloch, 1923: 775.-Alas. Sublettiella Sather, 1983b: 222. aterrima. Ont., N.Y., N.J., N. Mex., Ill., Ky., Type-species, calvata Srether (orig. N.C., s. to Miss. & Fla., Grld.; PA.; des.) AU. Chironomus aterrima Meigen, 1818: calvata. S.C. 47.-Europe. Sublettiella calva!a Sether. 1983b: crassicollis. Ont. 223.-5.C. Chironomus crassicollis Walker, 1848: 18.-Ont. edwardsi. N. Mex.: PA. Genus SYMBIOCLADIUS Kieffer Smittia edwardsi Goetghebuer, 1932a: Symbiocladius Kieffer, 1925a: 565. 123.-England. Type-species, Phae nocladius extrema. N.W.T., Grld.; PA. rhithogenae Zavrel (orig. des.). Chironomus extremus Holmgren, 1869: 40.-S pitsbergen. Subgenus SYMBIOCLADIUS Kieffer lasiophthalma. Ill.; PA. C ampto c I adius I a s iophth al mus Malloch, chattahoocheensis. Ga. 1915b:509.-Ill. Symbiocladius chattahoocheensis Caldwell, lasiops. Ill., Fla.; ?PA. 1984: 86.-Ga. Camptocladius lasiops Malloch, 191-5b: equitans. Utah, Calif., Colo., Vt. 508.-Ill. Trissocladius equitans Claassen, 1922: longitibia. Grld. 397.-Colo. Smittia longitibia Goetghebuer, 7933: 29.-Grld. nanseni. N.W.T. Genus SYNORTHOCLADIUS Thienneman Camptocladius nanseni Kieffer, 1926: Synorthocladius Thienemann, 1935: 211. 86.-N.W.T. Type-species, Dactylocladius semivirens parva. Grld. Kieffer (orig. des.). Chironomus parvus Lundbeck, 1898: 275.-Grld. semivirens. Widespread NE.; PA. polaris. N.W.T. Dactylocladius semivirens Kieffer, 1909: Camptocladius polaris Kieffer, 1926: 48.-Germany. 85.-N.W.T. polymorpha. N.W.T., Grld. Smittia polymorpha Andersen, 1937: Genus TETHYMYIA Wirth 76.-Grld. Tethymyia Wirth, 1949: 160- Type-species, roena. I]tah. aptena Wirth (orig. des.). Smittia roena Roback, 19-57b: 16.-Utah. subnudipennis. Grld. aptena. Calif. Smittia subnudipennis Goetghebuer, 1933: Tethymyia aptena Wirth, 1949: l6l.-Calif. 28.-Grld. velutina. N.W.T., Grld. Chironomus velutinus Lundbeck, 1898: Genus TIIALASSOSMITTIA Strenzke & 274.-Grld. Remmert Thalassosmittia Sfi enzke & Remmert. 1957 : 270. Type-species, Genus STILOCLADIUS Rossaro Camptocladius thalassophilus Bequaert Stilocladius Rossaro, I979l- 351. & Goetghebuer (orig. des.). Type-species, montanus Rossaro (orig. Saunderia Sublette. 1967a: 31.8. des.). Type-species, Camptocladius clavicornis Saunders (orig. des.). clinopecten. N.S., N.C., S.C. Stilocladius clinopecten Sather, 1982: clavicornis. B.C. s. to Calif. 494.-S.C. Camptocladius clavicornls Saunders, 1928: -528.-B.C. CHIRONOMIDAE DE LA REGION NEARCNQUE 39

marina. B.C. s. to Calif. Genus TRICHOCIIILUS Sather Camptocladius marinus Saunders, 1928: Trichochilus Sether, 1985f: 31. -526.-B.C. Type-species, Trichocladius lacteipennis pacifica. B.C. s. to Calif. Johannsen (orig. des.). Camptocladius pacificus Saunders, 1928: 523.-8.C. lacteipennis. Pa. Trichocladius lacteipennis Johannsen, 1908: 282.-Pa. Genus THIENEMANNIA Kieffer Thienemannia Kieffer. 7909: 49- Type-species, gracilis Kieffer (mon.). Genus TVETENIA Kieffer Tvetenia Kieffer, 1922b: 12. Type-species, pilinucha. Tenn. duodenaria Kieffer (orig. des.). Thienemannia pilinucha Sather, 198-5d: 127.-Tenn. calvescens. N. Mex.: PA. Spaniotoma calvescens Edwards, 1929: 353.-England. Genus THIENEMANNIELLA Kieffer duodenaria. Y.T., N.W.T.; PA. Thienemanniella Kieffer, 1911c: 201 (as Tvetenia duodenaria Kieffer, 1922b:. subg. of Corynoneura Winnertz, 1846). 12.-Novaya Zemlya. Type-species, Corynoneura clavicornis paucunca. Y.T., N.W.T., Alta., Man., Ont. Kieffer (Goetghebuer in Goetghebuer Eukiefferiella. paucunca Saether, 1.969: & Lenz, 1939b:7). 46.-Alta. vitracies. Sask., Ont. cubitus. B.C. Eukiefferiella vitracies Sether, 1969: Corynoneura cubitus Garrett, t925: 49.-Ont. 10.-B.C. elana. I-Itah. Corynoneura elana Roback, 19-57b: Genus UNNIELLA Sether 10.-Utah. Unniella Sether, 1982: 502- Type-species, majuscula. Ont.; PA. multivirga Sether (orig. des.). Corynoneura majuscula Edwards, 1924c: 183.-England. multivirga. S.C., Ga., Fla. mallochi. N. Mex., Tex. Unniella multivirga Sather, 1982: Thienemanniella mallochi Sublette, 1970: 503.-Fla. 88.-Tex. similis. N.Y., Ohio., Ill., N. Mex., Tex. Corynoneura similis Malloch, 1915b: Genus VMCRICOTOPUS Schnell & Sether 413.-Ill. Vivacricotopas Schnell & Sether, xena. Ont., Ill. to N.Y., s. to Fla. 1988: 49. Type-species, ablusus Corynoneura rena Roback, 1957a: 61.-Pa. Schnell & Saether (orig. des.).

piloculus. N.W.T. Genus TOKUNAGAIA Sether Hydrobaenus piloculus Cranston & Oliver, Tokunagaia Sather, L973a: 56. 1988a: 435.-N.W.T. Type-species, Spaniotoma kibunensis Tokunaga (orig. des.). Genus XYLOTOPUS Oliver excellens. B.C.; PA. Xylotopus Oliver, 1982: 167. Type-species, Eukiefferiella excellens Brundin, 1956: Orthocladius par Coquillett (orig. des.). 89.-Sweden. obriaini. N.W.T. par. Ont. to N.S., s. to Iltah, Tex. & Fla. Tokunagaia obriaini Hayes & Murray, Orthocladius par Coquillett, 1901b: 1988: 361.-N.W.T. 608.-N.J. rowensis. Alta., Ont. Eurycnemus scitulus Coquillett, 1901b: Adactylocladius rowensis Srther, 1969: 608.-N.J. 58.-Alta. 40 NEARCTIC CHIRONOMIDAE

Metriocnemus par Johannsen, 1905: Orthocladius bifasciatus Malloch, 1918: 301.-N.Y. 42.-nt. Orthocladius clepsydrus Coquillett, 1.902b: 92.-N. Mex. Genus ZALUTSCHIA Lipina Trichocladius ater Goetghebuer, 1955: Zalutschia Lipina, 1939: 106. 167.-Grld. Type-species, zalutschicola Lipina (orig. Trichocladius septis Roback, 1957b: des.). 14.-Utah.

briani. N.C., S.C., Fla. Nomina dubia in Orthocladiinae Zalutschia briani Soponis, 1979: 1.25.-Fla. Camptocladius fulvipluma Kieffer, 7926: furcarca. N.W.T. 85.-N.W.T. Zalutschia furcarca Sether, 1976: Camptocladius glacialis Kieffer, L926: 217.-N.W.T. 8-5.-N.W.T. fingulata. 8.C., Ont., Qu6., Ind. Chironomus minutus Zetterstedt, 1850: Zalutschia lingulata Sether, 1976: 3522.-Sweden. 222.-Ont- Chironomus pubitarsis Zetterstedt, 1850: Zalutschia lingulata lingulata Saether, 3514.-Sweden. 1976: 223.-Ont. Metriocnemus similis Kieffer, 1922b:. Zalutschia lingulata pauca Sa,ther, 1976: ll.-Novaya Zemlya. 228.-8.C. Orthocladius sordens Johannsen. L905: obsepta. Ont., Qu6. 272.-N.Y. Orthocladius obseptus Webb, 1969: 91.-Ont. pusa. Man. Zalutschia pusa Satber, 1976: 234.-Man. Subfamily CHIRONOMINAE sivertseni. N.B., N.S.; PA. Eukiefferiella sivertseni Aagard, 1979: Tribe CHIRONOMINI 95.-Norway. tatrica. Alas.; PA. Genus ACALCARELLA Shilova Spaniotoma tatrica Pagast in Zavrel & Acalcarella Shilova, 1955: 319. Pagast, 193-5: 156.-Czechoslovakia. Type-species, Cryptochironomus nucus tornetraeskensis. Not NE. Pankratova (orig. des.). Orthocladius tornetriiskensis Edwards in Thienemann, I94I:2l1.-Sweden. IJnnamed species. Ont. trigonacies. Alas., N.W.T. -N.W.T., Zalutschia trigonacies Sather, 1976: 242.-N.W.T. Genus APEDILUM Townes vockerothi. Qu€. Apedilum Townes, 1,945: 32. Zalutschia vockerothi Sather, 1976: Type-species, subcinctum Townes 219.-Qu6. (orig. des.). xethis. Utah. Hydrobaenus xethis Roback, 19-57b: elachistus. Mass. to Va., Calif. to N.C. & l3.-Utah. Fla. zalutschicola. Alas. to N.W.T., s. to Man., Apedilum elachistus Townes, 1945: N.H. & S.C.; PA. 33.-Tex. Zalutschia zalutschicola Lipina, 1939: subcinctum. Ont., Calif., Nev. to N. Mex., 96.-U.S.S.R. Okla., Fla. Apedilum subcinctum Townes, 1945: Incerta sedis in Orthocladiinae 33.-Nev. Camptocladius flavibasis Malloch, 1915b: Paralauterborniella subcincta alamedensis 511.-Ilf. Sublette, 7960: 203.-Calit. Hydrobaenus anteilis Roback, 1957b: Paralauterborniella subcincta subcincta 14.-Utah. (Townes), 1945: 33.-Calif. Orthocladius albidohalteralis Malloch, 191.5b:528.-Ill- CHIRONOMIDAE DE LA REGION NEARCTIQUE 4T

Genus ASHEUM Sublette & Sublette Genus BECKIDIA Sether Asheum Sublette & Sublette. 1983: Beckidia S€ether, 1979: 315 (n. 34 (n. nam. for Pedionomus nam. for Beckiella Sether). Sublette). Beckiella Srether, 1977b: ll9 Pedionomus Sublette, 1964b: 137 (preocc. Grandjean, 1964). (preocc. Gould, 1840). Type-species, Harnischia tethys Type-species, beckae Sublette Townes (orig. des.). (orig. des.). tethys. Sask., Ont., Qu6., S. Dak., N. Mex.; beckae. N. Mex. to N.C., s. to Fla. PA. Pedionomus beckae Sublette. t964b: Harnischia tethys Townes, 1945: 137.-La- 152.-S. Dak.

Genus AXARUS Roback Genus CHERNOVSKIIA Sether Axarus Roback, 1980b: 32 (n. nam. Chernovskiia Sether. t977b: lO7. for Anceus Roback). Type-species, Harnischia orbicus Anceus Roback, 1963a:237 (preocc. Townes (orig. des.). Risso, 1816). Type-species, Chironomus festivus Say (orig. amphitrite. Alta., Sask., S. Dak., Kans., Neb., des.). Iowa, N.C., S.C. Harnischia amphitrite Townes, 1945: dorneri. Tex., Fla. 1-51.-S. Dak. Chironomus dorneri Malloch, 1915b: orbicus. S. Dak., Neb., Iowa, Kans., N.C., 471.-Tex. S.C., Calif.; PA. festivus. Qu6. s. to Kans. & Tex. Harnischia orbicus Townes. 194-5: Chironomus festivus Say, 1823: 151.-Kans. 13.-Iil. Chironomus lineatus Say, 1823: 14.-Pa. Genus CHIRONOMUS Meigen Chironomus lineola Say, emend. Chironomus Meigen, 1803: 260. Wiedemann, 1828. Type-species, Tipula plumosa Chironomus lasiomerus Walker. Linnaeus (Latreille, 1810: 442). 1848: 19.-Ont. Tendipes Meigen, 1800: 17. rogersi. Ky., N.C., s. to Miss. & Fla. Tendipes suppressed by ICZN, Xenochironomus rogersi Beck & 1963b: 339 (Opinion 678). Beck, 19-58: 27.-Fla. Holtedahlia Kieffer, 1922b: 5. scopula. Alta. to N.Y., s. to Kans. & Ohio. Type-species, borealis Kieffer Xenochironomus scopula Townes, (orig. des.). 1945: 93.-Qu6. taenionotus. Iowa to Mich., Pa., s. to Tex. & Subgenus CAMPTOCHIRONOMUS Kieffer Fla. Camptochironomus Kieffer, 1918a: Chironomus taenionotus Sav. 1829: 45. Type-species, Chironomus 149.-Ind. aprilinus Meigen, 1830 sensu Kieffer (misident. = Chironomus tentans Fabricius) (Goetghebuer Genus BEARDIUS Reiss & Sublette in Goetghebuer & Lenz, 1937'. Beardius Reiss & Sublette, 1985: 2r). 179. Type-species, parcus (orig. des.). grandivalva. Alas., N.W.T.; PA. Tendipes grandivalva Shilova, 19-57: truncatus. Tex. 226.-U.S.S.R. Beardius truncatus Reiss & Sublette. 1985: 188.-Tex. 42 NEARCTIC CHIRONOMIDAE

pallidivittatus. Sask., Ill.; PA. crassicaudatus. Man., Ont., S. Dak., Minn. to Chironomus tentans var. N.J., s. to N. Mex. & Fla. pallidivittatus Malloch, 1915b: Chironomus crassic audatas Malloch, 445.-Ilt. 191-5b: 4-53.-Ill. tentans. Alas., N.W.T., B.C. to Que., s. to S. cucini. Ont., Ind., Calif. Dak., Ohio, N.Y., N. Mex.; PA. Chironomus cucini\N ebb, 7969: Chironomus tentans Fabricius. 1805: 105.-Ont. 38.-Europe. decorus. N.W.T. to Qu6., Wash., Nebr., S. vockerothi. Alta. Dak., Iowa, Ill., N.Y., Ohio, Kans., N. Chironomus vockerothi Rasmussen, Mex., Fla. 1984: 1643.-Alta. Chironomus decorus Johannsen, 1905: 239.-Wash., Nebr., Kans., Iowa, Ill., Subgenus CIIAETOLABIS Townes Ohio, N.Y. Chaetolabis Townes, 1945: 114 (as Chironomus similis Johannsen, 1905: subg. of Tendipes Meigen, 1800). 236--Ill., S. Dak., N.Y. Type-species, Tendipes atroviridis Chironomus cayugae Johannsen in Tilbury, Townes (orig. des.). 1913: 308.-N.Y. Chironomus distinguendus Kieffer, 1917: atroviridis. B.C. to Ont., N.Y., Mass., s. to 346.-Wash. Iowa & Va. decumbens. N.W.T., Man. Tendipes atoviridis Townes, 1945: Chironomus decumbens Malloch, 1934: 114.-N.Y. 16.-N.W.T. ochreatus. Qu6., Mich., Maine, N.J., s. to frommeri. Oreg., Calif., lftah, N. Mex. Ark. & Fla. Chironomus frommeri Atchley & Martin, Tendipes ochreatus Townes, 1945: 197l: 3I9.-Calif . 115.-N.J. hyperboreus. Alas., N.W.T., Alta., Sask., Colo., Grld.; PA. Subgenus CHIRONOMUS Meigen Chironomus hyperboreus Staeger, 184-5: 349.-Grld. anonymus. Calif., Tex.; NT. major. Okla., Ga. Chironomus anonymus Williston, 1896: Chironomus major Wilker & Butler, L983: 274.-St. Vincent. 128.-Ga. anthracinus. 8.C., Alta., Sask., Wis. to N.Y., maturus. Alta., Man. to N.Y., S. Dak., Ind., Mass., Calif.; PA. Calif., N. Mex., La. Chironomus anthracinus Zetterstedt, 1860: Chironomus maturus Johannsen, 1908: 6499.-Sweden. 279.-N.Y. Chironomus rempelii Thienemann, 194L: pilicornis. Alas., N.W.T., Alta. to Man.; PA. 234.-Sask. Tipula pilicornis Fabricius, 1794: Chironomus hyperboreus authors, not 243.-Europe. Staeger. Chironomus polaris Kirby, 1824: athalassicus. B.C. 218.-Amer. within the Arctic Circle. Chironomus athalassicus Cannings, plumosus. Widespread NE.; PA.; NT. 1975:447.-8.C. Tipula plumosa Linnaeus, 1758: atrella. B.C. to P.E.I., s. to Calif. & N. Mex. 587.-Europe. Tendipes atrella Townes, 1945: Tipula annularis De Geer, 7776: 124.-Nev. 379.-Europe. atritibia. N.W.T., B.C., Ont., Qu€., N.Y. Chironomus cristatus Fabricius. 180-5: Chironomus atritibia Malloch, 1934: 39.-N.Y. 16.-N.W.T. Chirono mus ferrugine ovittatus Zetterstedt biseta. Hudson Bay Territory. 1850: 3492.-Sweden. Tendipes biseta Townes, 1945: Chironomus imperator Walley, I926a: 127.-Hudson Bay Territory. 64.-Ont. brevipalpis. N.W.T. prior. Alas. Hol te d ahl ia brevip alpis Kieffer, Chironomus prior Bttler, 1982: 1926: 78.-N.W.T. 222.-Alas. CHIRONOMIDAE DE LA REGION NEARCTIQUE 43

pungens. D.C., Fla. longipes. Ont.; PA. Tendipes pungens Townes, 1945: 119.-Fla. Chironomus longipes Staeger, L839: riparius. Widespread NE., Grld.; PA. -570.-Denmark. Chironomus riparius Meigen, 1804: Chironomus dorsalis authors, not Meigen. 13.-Europe. mendax. Calif.; PA. Chironomus albistria Walker, 1848: Chironomus mendax Stor5, 1936: 17.-Ont. 124.-Finland. Chironomus seras Malloch, 1915b: 481.-Ill. Nomina dubia in Chironomus Chironomus militaris Johannsen, I937b: Chironomus attenuatus Walker, 1848: 46.-N.Y. 20.-Ont. sanctipauli. Alas. Chironomus bifimb riatus Kieffer, Chironomus sanctipauli Sublette, 1966: 1922f: I45.-"Iex. 604 (n. nam. for Chironomus conformis Chironomus lasiopus Walker, 1848: Malloch, 1923: 772, preocc. Holmgren, 19.-Ont. 1869).-Alas. Chironomus redeuns Walker. 1956b: staegeri. Widespread Can. s. to Calif., Ala., 42L-U.S.A. N.C., S.C., Grld.; PA. Chironomus staegeri Lundbeck, Incerta sedis in Chironomus 1898: 271.-Grld. Chironomus bimacula Walker, 1848: Chironomus fasciventris Malloch, 191-5b: 15.-Ont. 438.-il1. Chironomus borealis Curtis. 1835: stigmaterus. Wash. to N.Y., s. to Calif. & lxxvii.-Arctic Amer. Fla. Chironomus excavatus Kieffer, 1917: Chironomus stigmaterus Say, L823: 348.-N.Y. 15.-?Pa. Chironomus excisus Kieffer, 1917: Chironomus glaucurus Wiedemann, 1828: 346.-Pa. 15.-Pa. Chironomus halophilus Packard in Verrill, tardus. Alas. 1873: 539.-Mass. Chironomus tardus Butler, 1,982: Chironomus harti Malloch. 1915b: 222.-Alas. 457.-l1l. tuberculatus. Alta., S. Dak., Hudson Bay Chironomus melanderi Kieffer, 19 17: Territory. 347.-Wash. Tendipes tuberculatus Townes, 1945: Chironomus nigritibia Walker, 1848: 128.-Hudson Bay Territory. 16.-Ont. tuxis. Man., Ont., Minn. to Maine, s. to Ill. Chironomus trichomerus Walker, 1848; & N.J., Fla. 21.-Ont. Chironomus tuxis Curran, 1930: 31.-N.Y. utahensis. Sask., Man. to Oreg. & Minn., s. to Calif.. N. Mex. Genus CLADOPELMA Kieffer Chironomus utahens is Malloch. 191-5b: Cladopelma Kieffer, l92lc: 274. 438.-Utah. Type-species, Chironomus virescens vancouveri. B.C. Meigen, (Harnisch, 1923: 304). Chironomus vancouveri Michailova & Cryptocladopelma Lenz, 1947a:. 37 (nom. Fischer, 1986: 99.-B.C. nud.). whitseli. Calif.. N. Mex. Chironomus whitseli Sublette & Sublette, amachaerus. Ont., Qu6., Minn., Ill., N.H., s. I97 4: 7 .-Calif. to Miss. & Fla., Calif. Harnischia amachaerus Townes, 1945: Subgenus LOBOCHIRONOMUS Ryser, 168.-N.Y. Wtilker & Scholl boydi. Ky., N.C., s. to Miss. & Fla. Mex.; NT. Lobochirononus Ryser, Wtilker & Scholl, Chironomus boydi Beck, t962: 91.-Fla. 1985: 386. Type-species, montuosus collator. S. Dak., Minn., Ont., Qu6., Ohio, (orig. des.). N.Y., Okla., Fla. Harnischia collator Townes, 1945: 169.-N.Y. AA NEARCTIC CHIRONOMIDAE

edwardsi. 8.C., Ont., N.Y., Calif., Nev., Ariz., parafulvus. Fla. Ky., N.C. s. to Miss. and Fla.; PA. Chironomus parafulvus Beck & Beck, Tendipes edwardsi Kruseman, 1933: 1964: 2OI.-Fla. 194.-Netherlands. ponderosus. S. Dak., Ohio, N. Mex., Okla., galeator. Iowa, Ohio, N. Mex. to N.C., s. to Ark., La., N.C., S.C. Fla. Chironomus ponderosus Sublette, 1964b: Harnischia galeator Townes, 1945: I29.-La. 170.-Fla. psittacinus. Alas. to N.Y., s. to Oreg. & Ky.; spectabilis. Ont. PA. Harnischia spectabilis Townes, 1945: Chironomus psittacinus Meigen, 1830: 168.-Ont. 2{l.-Europe. viridula. Alas., N.W.T., Sask., Minn. to N.Y., Chironomus stylifera Johannsen, L908: Calif. to N.C., s. to Fla.; PA. 281.-N.Y. Tipula viridula Linnaeus, 1767: Chironomus obtusilobtts Malloch, 1923: 975.-Sweden. 171.-Alas. Chironomus farinalis Walley, 1926b: 205.-Alta. Genus CRYPTOCHIRONOMUS Kieffer ramus. Sask. Cryptochironomus Kietfer, 1.918a: 46 (as Cryptochironomus ramus Mason, 1986: subg. of Chironomus Meigen, 1803). 409.-Sask. Type-species, Chironomus chlorolobus scimitarus. Alta., Ont., Mich., N.Y., Md., Fla. Kieffer (orig. des.) = Chironomus Cryptochironomus scimitarzs Townes, supplicans Meigen. 194-5: 98.-N.Y. sorex. Mich., N.Y., Okla., Tex., N.C., S.C., argus. Pa. Fla. Cryptochironomus argus Roback, 1957al. Cryptochironomus sorex Townes, 1945: 106.-Pa. 97.-N.Y. blarina. AIta. to Ont., Minn. to N.Y., s. to N. Mex. & Fla. Cryptochironomus blarina Townes, 194-5: Genus CRYPTOTENDIPES Lenz 99.-N.Y. Cryptotendipes Lenz, 7947a: 34. conus. Sask., Ont. Type-species, Chironomus usmaensis Cryptochironomus conus Mason, 1986: Pagast (Beck & Beck, 1969: 294). 400.-Sask. curryi. Sask. ariel. B.C., S. Dak., Kans., Calif. Cryptochironomus curryi Mason, 1986: Tendipes ariel Sublette, 1960: 222.-Calit. 404.-Sask. casuarius. Alta., Man., Ont., N.Y., Kans., digitatus. Y.T., N.W.T., Sask. to Ont., Nfld., Ky., N.C., s. to Fla. Mass. s. to N. Mex. to Ga. Harnischia casuaria Townes, 194-5: Chironomus digitatus Malloch, 191-5b: 162.-N.Y. 483.-Ill. darbyi. Man., S. Dak., Calif., Nev., Ariz., N. eminentia. Sask. Mex.; PA. Cryptochironomus eminentia Mason, 1986: Tendipes darbyi Sublette, 1960: 406.-Sask. 221--Calit. fulvus. Mont., Sask., Man. to N.B., s. to emorsus. Man., Qu6., N.Y., N.J., D.C., S. Calif. & Fla. Dak., Kans., Iowa, Ohio, Ky., N.C., s. Chironomus fulvus Johannsen, 1905: to Fla. 224.-N.Y. Harnischia emorsa Townes, 194-5: Chironomus mallochi Kieffer, 1919b: i91 161.-N.Y. (n. nam. for Chironomus abbreviatus pilicuspis. Man. Mafloch, 1915b: 451, preocc. Kieffer, Cryptotendipes pilicuspis Sather, I977b: 1913a).-Ill. 98.-Man. Chironomus parvilamellalas Malloch, pseudotener. S. Dak., Mich., N.Y., Ohio, Pa., 1915b: 479.-Ill. D.C.; PA. Cryptochironomus ps eudotener Goetghebuer, 1922: 38.-Belgium. CHIRONOMIDAE DE LA REGION NEARCTIQUE 45

Genus CYPHOMELLA Sether Subgenus IRMAKIA Reiss Cyphomella Sather, 1977b: l0l. Irmakia Reiss, 1988: 1. Type-species, Type-species, gibbera Sether (orig. schachti Reiss (orig. des.). des.). cuneatus. Man., Qu6., s. to Iowa & N.J., argentea. Ont., N.J. N.C.. S.C. Harnischia argentea Townes, 1945: Harnischia cuneata Townes, 1945: 164.-Ont. 163.-N.J. cornea. S. Dak., Kans.; PA. fastigatus. Qu6., N.B., N.Y., Mich., Ill. Cyphomella cornea Sather, I97'7b: Harnischia fastigata Townes, 194-5: 105.-5. Dak. 162.-N.Y. gibbera. Sask., S. Dak. Cyphomella gibbera Sather, 1977b:- 103.-5. Dak. Genus DICROTENDIPES Kieffer grisea. Mont., Mich., Ill., Okla., Tex. Dicrotendipes Kieffer, I9t3d: 23. Chironomus griseus Malloch, 1915b: Type-species, pictipennis Kieffer 468.-Mich. (preocc. Philippi, 1865) = Polypedilum qu atuor d e c impunc tatum Goetghebu er, 7936 = Dicrotendipes pilosimanus Genus DEMEIJEREA Kruseman Kieffer, l9l4= Chironomus Demeijerea Kruseman, 1933: I54. septemmaculatus Becker, 1908 (mon.). Type-species, Tipula rufipes Linnaeus Limnochironomus Kieffer, 1920a; t66. (orig. des.). Type-species, Tendipes falciformis Kieffer (orig. des.) = Chironomus abruptus. N.Y., Mo. nervosus Staeger. Glyptotendipes abruptus Townes, 1945: 138.-N.Y. adnilus. Ariz. atrimana. Alta., Ont., N.Y., Conn., Md., Mo. Dicrotendipes adnilus Epler, 1987: to N.C., s. to Fla. 28.-Ariz- Chironomus atrimanus Coquillett, 1902b: aethiops. Utah, Ariz., N. Mex., Fla., Mex. 94.-Mo. Tendipes aethiops Townes, 1945: 107.-N. brachialis. Man., Ont., N.Y. to N.J., Oreg., Mex. IJtah, Ark., Ky., N.C., s. to Fla. botaurus. Man., Mont., Mich., Kans., Okla. Chironomus brachialis Coquillett, 1901b: Tendipes botaurus Townes, 1945: 607.-N.J. 109.-Mont. obreptus. N.Y., N.C., S.C. californicus. Oreg. to Tex., Mex.; NT. Glyptotendipes obreptus Townes, 1945: Chironomus californicus Johannsen, 190-5: 139.-S.C. 277.-Calif. crypticus. Calif., N. Mex., Kans.; NT. Dicrotendipes crypticus Epler, 1987: Genus DEMICRYPTOCHIRONOMUS Lenz 39.-N. Mex. Demicryptochironomus Lenz, 194Ia:. 34. fumidus. Ont., Nfld., Wash., Oreg. to N.H., s. Type-species, Chironomus vulneratus to N. Mex. & Ala. Zetterstedt, 1838: 814 (mon.). (Junior Chironomus fumidus Johannsen, 1905: synonym of Schadinia Lipina, 1939: 221.-N.Y. 107, for stability it is recommended Chironomus incognitus Malloch, 1915b: tbat Demicryptochironomus be used 480.-Ill. until a ruling is given by the ICZN). leucoscelis. Ont. to N.8., Minn., Mich. to N.J., Idaho, Ky., N.C., s. to Miss. & Subgenus DEMICRYPTOCIIIRONOMUS Fla. Tendipes leucoscelis Townes, 1945: vulneratus. Not NE. 104.-Idaho. Chironomus vulne ratus Zetter stedt, lobiger. Alas., N.W.T., Man., Idaho, Wyo., 1838: 814.-Sweden. Ind., Mich., N.Y.; PA. Limnochironomus lobiger Kieffer, l92la: 71.-Germany. 46 NEARCTIC CHIRONOMIDAE

lobus. Ky., N.C., s. to Miss. & Fla. pagana. B.C. to Qu€., N.Y., Mich., S. Dak. Chironomus lobus Beck, 7962: 89.-Fla. Miss.; PA. lucifer. Sask., Ont., widespread U.S.A. Chironomus paganus Meigen, 1838: Chironomus lucifer Johannsen, 1907b: 7.-Belgium. 110.-Kans. Einfeldia synchrona Oliver, 1971: Chironomus indistinctus Malloch, 1915b: 1591.-Ont. 477.-I!].. milleri. Man., Ont., Oreg., Minn., Mich., Ind., N.Y. Genus ENDOCHIRONOMUS Kieffer Tendipes milleri Townes, 1945: 110.-N.Y. Endochironomus Kieffer, 1918b: 69. modestus. Widespread NE., Grld.; PA. Type-species, Chironomus alismatis Chironomus modestus Say, 1823: 13.-Pa. Kieffer (orig. des.) = Tipula tendens neomodestus. N.W.T. to Qud., s. to Tex. & Fabricius. Fla. Chironomus neomodestus Malloch, 1915b: dispar. Not NE. 475.-\t. Chironomus dispar Meigen, 1830: Chironomus tenellus authors, not 247.-Not stated. Zetterstedt. oldenbergi. Grld.; PA. nervosus. Alas., N.W.T., B.C. to N.B., s. to Endochironomus oldenbergi Goetghebuer, Calif. & Fla., Grld.; PA. I932b: 130.-Europe. Chironomus nervosus Staeger, 1839: nigricans. N.W.T., B.C. to N.B., s. to Calif. 567.-?Denmark. & Fla. Chironomus brevitibialis Zetterstedt, 18-50: Chironomus nigricans Johannsen, 190-5: 3-535.-Sweden. 219.-N.Y., N.J. simpsoni. N.Y., Ga., Fla. Chironomus johnsoni Kieffer, l9}6a: 19 Dicrotendipes simpsoni Epler, 1987: (n. nam. for Chironomus albipennis 84.-N.Y. Meigen of Johnson, 1900, nom. nud.). thanatogratus. Fla. Chironomus albipennis authors, not Dicrotendipes thanatogratrzs Epler, 1987: Meigen. 88.-Fla. subtendens. B.C. to Qu6. & Nfld., s. to N. tritomus. Alas., Sask., Mont. to N.H., Okla., Mex. & Fla. Ky., N.C., s. to La. & Fla.; pA. Tanytarsus subtendens Townes, 1.94-5: Chironomus tritomus Kieffer in 65.-N.Y. Thienemann & Kieffer, 1916: Chironomus viridis authors, not Macquart. -524.-Sweden. Chironomus incurvus Sublette, 1964b: 126.-La. Genus ENDOTRIBELOS Grodhaus Endotribelos Grodhaus, 1987a: 237. Type- species, Tendipes hesperius Sublette Genus EINFELDIA Kieffer (orig. des.). Einfeldia Kieffer, I924a: 393. Type-species, pectoralis Kieffer (mon.). hesperium. Calif., Fla. Tendipes hesperius Sublette, 1960: austini. N. Mex. to N.C., s. to Fla. 217.-Calif. Einfeldia austini Beck & Beck, 1970: 40.-Fta. brunneipennis. Minn., Iowa to Mass., N.J., Genus GILLOTIA Kieffer Ky., N.C., s. to Miss. & Fla. Gillotia Kieffer, 1927c: 272. Type-species, Chir o nomus brunne ipe nnis Johannsen, Gillotia fuscipes Kieffer, 1922d (sub. 1905: 20-5.-N.Y. mon.; preocc. Kieffer, 1909) = chelonia. N.Y., N.J., D.C., N. Mex. to La. Chironomus trifidus Freeman, 1957 Tendipes chelonia Townes, 194-5: (replacement nam.). 114.-N.Y. natchitocheae. La. alboviridis. Iowa, Ill., Kans., N. Mex., Okla. Chironomus natchitocheae Sublette, 1964b: Chironomus alboviridis Malloch. 191-5b: 124.-La. 482.-Tl/. CHIRONOMIDAE DE I-A REGION NEARCTIQUE A1

Genus GLYPTOTENDIPES Kieffer Siolimyia Fittkau, t968b:- 260. Glyptotendipes Kieffer, I9l3e:. 255. Type-species, amazonica Fittkau (orig. Type-species, sigillatus Kieffer (orig. des.). des.) = ?Glyptotendipes foliicola Kieffer. amazonicus. Fla., Mex.; NT. Siolimyia amazonica Fittkau, 1968b: amplus. Sask., N.J. 26Q.-Brazil. Glyptotendipes amplus Townes, 194-5: carus. Ky., N.C. s. to Tex. & Fla.; NT. 145.-N.J. Tendipes carus Townes, 1945: barbipes. Alas., B.C. to N.B., s. to N. Mex. 118.-Venezuela. & S.C.; PA.; OR. devineyae. N.C. s. to Miss. & Fla. Chironomus b arbipes Staeger, 1839: Chironomus devineyae Beck, 1961: 561..-not stated. 726.-Fla. dreisbachi. N.W.T., Sask., Mich., Ont. holoprasinus. Md., Calif. to N.C., s. to Fla.; Glyptotendipes dreisbachl Townes, 1945: NT. 145.-Mich. Chironomus holoprasinus Goeldi, 190-5: lobiferus. B.C. to Qud.. s. to Calif. & Fla. 135.-Brazil. Chironomus lobiferus Say, 1823: 1,2.-?Pa. Chironomus fulvipilus Rempel, 1939: Chironomus ithacanensis Johannsen, 1908: 210--Brazil. 279 (n. nam. for Chironomus pictus. Se. U.S.A.; NT. caliginosus Johannsen, 1905: 205, Goeldichironomus pictus Reiss, 1974: preocc. Meunier, 1904).-N.Y. 86.-Brazil. Glyptotendipes americanus Kieffer, 1917: 355.-Wash. meridionalis. Man., S. Dak. to N.C., s. to N. Genus GRACEUS Goetghebuer Mex. & Fla. Graceus Goetghebuer, 1928: 15. Glyptotendipes meridionalis Dendy & Type-species, ambiguus Goetghebuer Sublette, 1959: 517.-Okla. (mon.). pallens. Not NE. Chironomus pallens Meigen, 1.804: Unnamed species.-Ont. 12.-Not stated. Chironomus polytomus Kieffer, 1804: -56.-Germany. Genus HARNISCHIA Kieffer paripes. Alta. to Nfld., s. to Miss. & Harnischia Kieffer, 1921.c: 273. Fla.; PA. Type-species, fuscimanus Kieffer, (sub. Chironomus paripes Edwards, 1929: mon.; Kieffer, I92Ia: 69). 392.-England. seminole. Fla. curtilamellata. Widespread NE.; PA.; AF.; Glyptotendipes seminole Townes, 1945: OR.; AU. 146.-Fla. Chironomus curtil amell atus Malloch, senilis. Ont., N.Y., Mich., Iowa. 1915b: 474.-Mich. Chironomus senilis I ohannsen, 1937b: incidata. S. Dak., N. Mex., Okla., Tex. 37.-N.Y. Harnischia incidata Townes. 194-5: testaceus. Okla. to N.Y., s. to La. & Fla. 166.-Okla. Glyptotendipes testaceus Townes, 1945: 140.-N.Y. unacus. Sask., Qu€. Genus HYPORIIYGMA Reiss Glyptotendipes unacus Townes, 1945: Hyporhygma Reiss, 1982: 289. 146.-Qu6. Type-species, Chir onomus quadripunctarus Malloch (mon.).

Genus GOELDICHIRONOMUS Fittkau quadripunctatum. Ont. to Nfld., N.H., Mass., Goeldichironomus Fittkau , 1,965:- 210. Wis. to Ohio, Fla. Type-species, Chironomus holoprasinus Chironomus quadripunctaras Malloch, Goeldi (orig. des.). 191-5b: 437.-Wis. 48 NEARCTIC CHIRONOMIDAE

Genus KIEFFERULUS Goetghebuer Genus MICROCHIRONOMUS Kieffer Kiefferulus Goetghebuer, L922: 40. Microchironomus Kieffer, 1918b: 113. (as Type-species, Tanytarsus subg. of Chironomus Meigen, 1803). tendipediformis Goetghebuer (orig. Type-species,lendli Kieffer (orig. des.). des.). Leptochironomus Pagast, 1931: 210 (as subg. of Chironomus Meigen, 1803). Subgenus KIEFFERULUS Goetghebuer Type-species, balticus Pagast (Townes, 1945: 152) = Chironomus tener dux. Sask., Man., Ont., Qud., Oreg. to R.I., s. Kieffer. to N. Mex. & Fla. Chironomus dux Johannsen, 1.90-5: nigrovittatus. S. Dak. to Ill., N.Y. s. to La. 231.-N.Y. & Fla., Calif., N. Mex., Kans. Chironomus nigrovittatus Malloch, 1915b: Subgenus WIRTHIELLA Sublette 456.-Ill. Wirthiella Sublette, 1960: 216 (as subg. of Tendipes Meigen, 1800). Type-species, modocensis Sublette (orig. des.). Genus MICROTENDIPES Kieffer Microtendipes Kieffer, I9l5a:. 70. modocensis. Calif. Type-species, Tendipes abbreviatus Tendipes modocensis Sublette, 1960: Kieffer (orig. des.) = Chironomus 276.-Calif. chloris Meieen.

anticus. coastal t". ," Ga., Tex. Genus KLOOSIA Kruseman Chironomus anticus Walker. 1848: Kloosia Kruseman, 1933: 152. Type- 21.-Ga. species, Chironomus pusillus Linnaeus caducus. N.B., Minn. to N.Y., s. to Calif. & (orig. des.). Ga. Oschia Seether, 1983a: 395. Type-species, Microtendipes caducus Townes, 1945: 24. dorsenna Sether (orig. des.). (n. nam. for Chironomus pallidus Johannsen, I9O5: 230, preocc. dorsenna. S.C. Fabricius, 180-5).-N.Y. Oschia dorsenna Sether, 1983a: caelum. Ont., Qu6., N.B., N.Y., Ohio, Mass. 396.-S.C. Microtendipes caelum Townes, 1945: 23.-Mass. pedellus. N.W.T., B.C. to N.B. & Maine, s. Genus LAUTERBORNIELLA Thienemann & to Calif. & Fla.; PA. Bause Tipula pedella De Geer, 1776: Lauterborniella Thienemann & Bause in 378.-Sweden. Bause, l9l3: 120. Type-species, Microtendipes pedellus var. aberuans Tanytarsus agrayloides Kieffer (Kieffer, (Jghannsen), 1905: 221.-N-Y-, Pa., I92tb: 28). Wash., N.J. Microtendipes pedellus var. pedellus (De agrayloides. Widespread Can., Calif. to N.C. Geer), 1776: 378.-B.C. to Qu6., s. to & Fla.; PA. R.I.; PA. Tanytarsus agrayloides Kieffer, 1911a: Microtendipes pedellus var. stygius Townes, 51.-Germany. 1945: 26.-N.Y. Chironomus fascipes Coquillett, 1908: 14-5.-N.J. Genus LIPINIELLA Shilova Chironomus deflexa Walley, 1926b: Lipiniella Shilova, t96I: L9. Type-species, 206.-Ont. araenicola .Shilova (mon.). Ilnnamed species.-N.W.T. Genus NILOTHAUMA Kieffer Nilothauma Kieffer, I921c: 27Q. Type-species, pictipenne Kieffer (sub. mon.; Kieffer, 1.92Lb: 37). Kribioxenus authors. not Kieffer. CHIRONOMIDAE DE LA REGION NEARCTIQUE 49

babiyi. Man. to Qu6., N.Y., Mich., Ohio, directus. Kans., Ky., N.C., s. to La. & Fla. Okla., Tex., N.C., S.C., Fla. Tendipes directus Dendy & Sublette, 1959: Chironomus babiyi Rempel, I937b: -514.-Ala. 274.-N.Y. elodeae. Qu6., N.Y. bicorne. N.C., S.C., Fla. Harnischia elodeae Townes. 1945: Kribioxenus bic ornis Townes, 194-5: 156.-N.Y. 35.-S.C. forceps. Man., Mich., Mass. mirabile. Qud., N.Y., Fla. Harnischia forceps Townes, 194-5: Krib ioxenus mirabilis Townes, 194-5: 157.-Mass. 35.-N.Y. frequens. Widespread NE.; PA. Chironomus frequens Johannsen, 190-5: 230.-N.Y. Genus OMISUS Townes hirtalatus. Ill. to Pa.. s. to Miss. & Fla. Omisus Townes, 1945: 27. Type-species, Chironomus hirtalatus Beck & Beck, 1964: pica Townes (orig. des.). 203.-Fla. monochromus. Sask., Qu6., Mich., Ill. to pica. Idaho, Man. to Qu6., N.H. to Conn., Mass., s. to Calif. & Fla.; PA. Ky., N.C., s. to Miss. & Fla. Chironomus monochromus Wulp, 1874: Omisus pica Townes, 1945:. 27.-N.H. 129 (n. nam. for Chironomus unicolor Wulp, 1858: 162, preocc. Walker, 1848).-Netherlands. Genus PAGASTIELLA Brundin pectinatellae. Ill. to Pa., s. to Miss. & Fla. Pagastiella Brundin, 1949:. 840. Cryptochironomus pectinatellae Dendy & Type-species, Chironomus orophilus Sublette, 19-59: -516.-Ala. Edwards (orig. des.). potamogeti. Man. to Qu6., s. to N. Mex. & Fla. orophila. Maine to Conn., Ky., N.C., s. to Harnischia potamogeti Townes, 1945: Miss. & Fla. 159.-N.Y. Chironomus orophilus Edwards, schneideri. Ky., N.C., s. to Miss. & Fla. 1929: 405.-Great Britain. Parachironomus schneideri Beck & Beck, ostansa. N.W.T., B.C. to Ont., N.H., N.C., 1969: 287.-Fla. S.C. sublettei. Fla. Paralauterborniella ostansa Webb, 1969: Chironomus sublettei Beck, 1961: 102.-Ont. 127.-Fla. Paralauterborniella ostensa (error), tenuicaudatus. B.C., Sask. to Qu6., Boerger et al., 1982: 917.-Alta- widespread U.S.A.; PA. Chirono mus tenuic aud atus Malloch. 1915b:475.-I].l. Genus PARACHIRONOMUS Lenz varus. Man., N.Y.; PA. Parachironomus LeDz, 1927: 160. Chironomus varus Goetghebuer, 1921: Type-species, Chironomus cryptotomus 162.-Belgium. Kieffer (Townes, L945: 152)-

abortivus. Ont., N-Y., Pa., Ill., Oreg. to S. Genus PARACLADOPELMA Harnisch Dak., s. to N. Mex. & Okla. Paracl adopelma Harnisch, 1923: 304. Chironomus abortivus Malloch. 191-5b: Type-species, Tendipes camptolabis 465.-Ill. Kieffer (orig. des.). alatus. Ky., N.C., s. to Miss. & Fla. Chironomus alatus Beck. 1962: 91.-Fla. alphaeus. Calif.; PA. carinatus. S. Dak. to N.Y., s. to N. Mex. & Tendipes alphaeus Sublette, 1960: Fla. 222.-Calit. Harnischia carinata Townes. 1945: camptolabis. Ont.; PA. 158.-Ark. Tendipes camptolabis Kieffer, l9l3a: chaetaolus. Calif. to Fla., Kans., Ind. 40-Germany. Tendipes chaetaola Sublette, 1960: tloris. S. Dak., Iowa, Okla. 220.-Calit. Harnischia doris Townes. 1945: 151.-Okla. -50 NEARCTIC CHIRONOMIDAE

galaptera. B.C., Man., Qu6., Ont., S. Dak. to duplicatus. Ont., S. Dak., Mich. to Okla., N.Y. & III. R.I., N.C., S.C. Harnischia galaptera Townes, 1945: 148 Chironomus duplicatus Johannsen, I937b: (n. nam. for Chironomus claripennis 28. (n. nam. for Chironomus basalis Malloch, 191-5b: 439, preocc. Malloch, 1915b: 441, preocc. Staeger, Lundbeck, 1898).-Mich. 1845).-N.Y. loganae. Kans., Ohio, Ky., N.C., s. to Miss. & Fla. fuscitibia. Calif. Paracladopelma loganae Beck & Beck, Paratendipes fuscitibia Sublette, 1960: 1969: 303.-Fla. 200.-Calif. nais. Ont., N.Y., Mich., Pa. nitidulus. Ohio, Mich., N.J., Pa., Del., Md. Harnischia nais Townes, 1945: 149.-N.Y. Chironomus nitidulus Coquillett, 190lb: nereis. Sask., Minn., Mich., Kans., N.C., S.C., 608.-N.J. Ga.; PA. Chironomus nitidellus (error), Malloch, Harnischia nereis Townes, 1945: 1915b: 468. 150.-S.C. nudisquama. Ont.; PA. nixe. Colo. Chironomus nudisquama Edwards, 1929: Harnischia nrxe 'fownes, \945: 149.-Colo. 396-England, Ireland. undine. Alta. to N.Y., Oreg., S. Dak., Kans., subaequalis. Man. to Qu6., s. to Kans. & Ky., N.C., s. to Miss. & Fla.; PA. Fla., Calif.; PA. Harnischia undine Townes, 194-5: Chironomus subaequalis Malloch, 191-5b: 149.-Oreg. 440.- Ill. winnelli. Y.T., N.W.T., Ont., Wis., Mich. thermophilus. Calif., Nev., Ariz., N. Mex. Paracladopelma winnelli Jackson, 1977: Paratendipes thermophilus Townes, 1.945: 1334.-Mich. 31..-Ariz.

Genus PARALAUTERBORNIELLA Lenz Genus PIIAENOPSECTRA Kieffer Paralauterborniella Lenz, l94lb: 48. Phaenopsecta Kieffer, 7921c: 274- Type-species, Chironomus brachylabis Type-species, Chironomus leucolabis Edwards (orig. des.) = Chironomus Kieffer (Goetghebuer in Goetghebuer nigrohalteralis Malloch. & Lenz, 1938: 80) = Chironomus flavipes Meigen. nigrohalterale. Y.T. to Ont., Colo., N.Y., s. to Tex. & Fla.; PA. dyari. Mont., Calif., Ohio, Fla. Chironomus nigrohalteralls Malloch, Tanytarsus dyari Townes, L945: 7-5.-Calif. 1915b: 440.-Ill. flavipes. Wash. to Qud., s. to Calif. & Fla.; PA. Chironomus flavipes Meigen, 1818: Genus PARATENDIPES Kieffer -5O.-Europe. Paratendipes Kieffer, I9lla: 41,. Tanytarsus flavicauda Malloch, 1915b: Type-species, Chironomus albimanus 493.-Ill. Meigen (orig. des.). incompta. Grld.; PA. Chironomus incomptus Zetterstedt, 1838: albimanus. Alta., Ont., Qu6., N.I}., Oreg. to 816.-Sweden. N.Y., s. to Calif. & S.C.; PA. mortensoni. Calif. Chironomus albimanus Meigen, 1818: 40 Phaenopsectra mortensonl Grodhaus, (n. nam. for Chironomus annularis 1987b: 137.-Calif. Meigen, 7804:77, preocc. De Geer, obediens. Wash., Sask. to N.8., s. to Colo. & 1776).-Europe. S.C.; PA. basidens. Iowa, N.J., N. Mex., Ala. Tanytarsus obediens Johannsen, 190-5: Paratendipes basidens Townes, 194-5: 286.-N.Y. 30.-N.J. pilicellata. Calif. Phaenopsectra pilicellata Grodhaus, 1976: 405.-Calif. CHIRONOMIDAE DE LA REGION NEARCTIQUE 51

profusa. Wash. to Mont., s. to Calif. & N. aviceps. Alta., Ont., Qu6. to N.H., Mont. s. Mex. to Calif. & Tex., Fla. Tanytarsus profusus Townes, 1945: Polypedilum aviceps Townes, 1945: 73.-Nev. 61.-N.Y. punctipes. Sask. to Ont., Qu6., N.Y., Idaho, braseniae. N.W.T. to Ont., N.Y., Ohio, N.J., N.C., S.C.; PA. Wash., Oreg., S. Dak., Ky., Va. to Fla. Chironomus punctipes Wiedemann, 1817: Chironomus bras eniae Leathers. L922: 65.-Germany. 8.-N.Y. vittata. Ont., Qu6., N.J., N.C., S.C. californicum. Calif. Tanytarsus vittatus Townes, 1945: Polypedilum californicum Sublette, 1960: 77.-N.J. 204.-Calif. calopterus. Ont., Qu€., Mich., N.Y., Md., Va. Chironomus calopterus Mitchell, 1908: Genus POLYPEDILUM Kieffer 8.-Md. Polypedilum Kieffer, l9l2b: 4I. Polypedilum walleyi Townes, 1945: Type-species, pelostolum Kieffer = 52.-Qu6. Chironomus nubifer Skuse (Ashe, 1981: cinctum. Oreg., Calif., Nev., Va. 58). Polypedilum cinctum Townes, 1.94-5: 59.-Nev. Subgenus PENTAPEDILUM Kieffer convictum. Sask., Ont., Qu6., s. to Kans. & Pentapedilum Kieffer, l9I3b: 25. Fla.: PA. Type-species, Pentapedilum stratiotale Chironomus convictus Walker, 1956a: Kieffer (Edwards, 1929: 376) = 161.-England. Chironomus tritus W alker. Chironomus flavus Johannsen, 1905: 225.-N.Y. sordens. Qu6., Mich. to N.Y., R.I., Va.; PA. Chironomus flaviventris Johannsen, 1907b: Tanytarsus sordens Wulp, 1874: 111.-Kans. 1.41.-Netherlands. Polypedilum obtusum Townes, 1945: Tanytarsus fulvescens Johannsen, 1905: 60.-N.Y. 293.-N.Y. fallax. Alta., Sask., Qu6., s. to Utah & Fla. Calopsecffa americ ana Kieffer, 1917: Chironomus fallax Johannsen, 190-5: 357.-N.Y. 210.-N.Y. tritum. B.C. to N.8., s. to Calif., Tex. & Fla.; Polypedilum fallax var. fallax Johannsen, PA. t905: 210.-Ont., Qu€., Ill., Ind., Chironomus titus Walker, 1956a: Mich., Vt., N.Y., Tenn. 342.-England. fuscipenne. 8.C., Wash., Oreg., Mont. to Polypedilum albulum Townes, 1945: Colo.; PA. 63.-N.Y. Chironomus fuscipennis Meigen, 1818: 35.-Europe. Subgenus POLYPEDILUM Kieffer illinoense. N.W.T. to Qu6., s. to Calif. & Fla.; PA. albicorne. B.C., Ont., Qu€., Nfld., Wis. to Chironomus illinoensis Malloch. 1915b: Ohio & Mass., Idaho, Calif. to La., 471.-ilL Fla. laetum. Alta. to Qu6., s. to Calif., N. Mex., Chironomus albicornis Meigen, 1838: N.C., S.C.; PA. 6.-Europe. Chironomus laetus Meigen, 1818: Polypedilum nigritum Townes, 1,945: 38.-Europe. 56.-N.Y. nubeculosum. N.W.T. to Qu6., Minn. to angustum. 8.C., Ont., Qu6., Nfld., Minn. to N.Y., Idaho, Ind., Colo.; PA. Va., Calif., Ark., Fla. Chironomus nubeculosus Meigen, 1804: Polypedilum angustum Townes, L94-5: 18.-Europe. 56.-N.J. ontario. Ont., Qu6. s. to N. Mex., La. & artifer. Wash., Calif. to Qu6., N.Y., Ohio, N. D.C. Mex.. Va. Chironomus ontario Walley, 1926b: Chironomus artifer Cvrran, 1930: 206--Ont- 32.-N.Y. Chironomus hirtipes Mitchell, 1908: 9 (preocc. Macquart, 1834).-Md. 52 NEARCTIC CHIRONOMIDAE

ophioides. N.W.T., Qu6., Idaho, Minn. to Polypedilum simulans Townes, 1945: Mass., s. to Va., Calif. 43.-N.Y. Polypedilum ophioides Townes, 1945: isocerus. Calif., Nev. -57.-N.Y. Polypedilum isocerus Townes, 1.945: parascalaenum. Fla. 42.-Calif. Polypedilum parascalaenum Beck, 1,962: labeculosum. Calif., Nev., Ariz., N. Mex. 91.-Fla. Chironomus labeculosus Mitchell. 1908: pedatum. 8.C., Ont., Qu6., N.S., Mich., N.Y., t4.-Ariz. Ya., La-, Wash., Mont., Calif. pardus. Tex., S.C. Polypedilum pedatum pedatum Townes, Polypedilum pardus Townes, 1945: 1945: -55.-N.Y- 41.-Tex. Polypedilum pedatum excelsius Townes, paryum. Md. to Calif., Tex., Fla. 1945: 55.-Wash. Polypedilum parvum Townes, 194-5: subcultellatum. Calif. 40.-Miss. Polypedilum subcultellatun Sublette, 1960: pterosopilus. Ariz., N. Mex., Tex., Fla. 204.-Calif. Polypedilum pterospilus Townes, 1945: sulaceps. Wash. to S. Dak., Calif. to N. Mex. 40.-Tex. Polypedilum sulaceps Townes, 1945: scalaenum. Widespread NE.; PA. 58.-Idaho. Tipula scalaena Schrank, 1803: trigonus. Ont. to N.B., Idaho, S. Dak. to 73.-Germany. N.Y., s. to N. Mex. & Fla. Chironomus needhamii Johannsen, 1908: Polypedilum trigonus Townes, 194-5: 278.-N.Y., Ind., Kans., Wash.; PA. 49.-N.Y. vibex. Ont., Qu6., N.J., Wash., Nev., N.C., S.C., Fla. Genus ROBACKIA Saether Polypedilum vibex'fownes, 1945: 52.-Ont. Robackia Sether, 1977b: 123. Type-species, Tendipes demeij erei Subgenus TRIPODURA Townes Kruseman (orig. des.). Tripodura Townes, 1945: 36. Type-species, simulans Townes (orig. claviger. Wash., Sask., S. Dak. to Ind., s. to des.). N. Mex. & Fla. Harnischia claviger Townes, 1945: acifer. Man., Mich., Ohio; PA. 158.-S.C. Polypedilum acifer Townes, 1945: demeijerei. Y.T., N.W.T., Alta. to Wyo. & 46.-Mich. Ohio, s. to Miss. & Fla.; PA. albinodus. Ont., Qu6., Idabo, Ohio, Va., Tendipes demeijerei Kruseman, 1933: Calif., N.C., S.C. 19-5.-Europe. Polypedilum albinodus Townes, 1945: 41.-Idaho. apicatum. Calif., Nev., Ariz., N. Mex., Ill. Genus SAETHERIA Jackson Polypedilum apicatum Townes, 194-5: Saetheria Jackson, 1977: 1325. 39.-N. Mex. Type-species, Harnischia fylus Townes Chironomus octopunctatus Loew, 1861b: (orig. des.). 33 (of Malloch 191-5b: 427).-il. floridense. Miss., Fla. hirta. S.C. Polypedilum floridense Townes, 1945: Saetheria hirta Satber, 1983a: 400.-S.C. 43.-Miss. tylus. Y.T., N.W.T., Ind., Kans., N.C., S.C., gomphus. Alta., Ont., Mass., N.C., S.C., Fla. Ga. Polypedilum gomphus Townes, 1945: Harnischia tylus Townes, 1945: 150.-Ind. 42.-Mass. halterale. Widespread NE. Chironomus halteralis Coquillett, 1901a: Genus SERGENTIA Kieffer 17.-D.C. Sergentia Kieffer, 1922a: 288. Polypedilum digitifer Townes, 194-5: 4-5.-S. Type-species, profundorum Kieffer Dak. (orig. des.) = Chironomus coracinus Chironomus griseopunctatas Malloch, Zetterstedt. 191-5b: 428.-lll- CHIRONOMIDAE DE I-A REGION NEARCTIQUE -)--t

Subgenus SERGENTIA Kieffer colei. B.C., Alta., s. to Calif. & Colo. Chironomus colei Malloch in Cole & albescens. B.C., Alta., Wash. s. to Calif. & Lovett, l9l9: 255.-Oreg. Colo. fuscipatellus. B.C., Oreg., Calif. Tanytarsus albescens Townes, 1945: Stenochironomus fuscipatellus Borkent, 73.-Wash. 1984: 64.-8.C. coracina. N.W.T. to Sask., Ont., Mont., Minn. hilaris. Y.T., B.C to Nfld., s. to N. Mex. & to ohio, Grld.; PA.; oR. Fla., Mont., Idaho. Chironomus coracinus Zetterstedt. 1850: Chironomus hilaris Walker, 1848: 3508.-Sweden. 17.-Unknown. Chironomus taeniapennis Coquillett, Genus STELECHOMYIA Reiss 1901b: 607.-Mass. Stelechomyia Reiss, 1982: 294. Chironomus nephopterus Mitchell, 1908: Type-species, Chironomus perpulcher 7.-Md. Mitchell (orig. des.). Chironomus exquisitus Mitchell, 1908: 11.-Md. perpulchra. Ont. to Maine, s. to Tex. & Fla. Chironomus zonopterus Mitchell, 1908: Chironomus perpulcher Mitchell, 1908: 12.-N.J. 13.-Md. macateei. Man. to Qu6., s. to Fla. & Tex. Chironomus macateei Malloch, 1915a: 45.-Md. Genus STENOCHIRONOMUS Kieffer maculatus. N. Mex., Tex. Stenochironomus Kieffer in Kieffer & Stenochironomus maculatas Borkent, 1984: Thienemann, l9l9: 44. Type-species, 52.-N. Mex. fasciatus Geoffroy (Kieffer, 1922d: 56) niger. Ariz. = gibbus Fabricius. Stenochironomus niger Borkent, 1984: 57 --Ariz. Subgenus PETALOPHOLEUS Borkent poecilopterus. Ont., Qu6., s. to Colo. & Fla. Petalopholeus Borkent, 1984: 92. Chironomus poecilopterus Mitchell, 1908: Type-species, Stenochironomus cinctus 10.-Md. Townes (orig. des.). pulchripennis. Ont., Qud., Nfld., s. to Pa. & Mich., Colo. aestivalis. N.J. to Fla. Chironomus pulchripennis Coquillett, Stenochironomus aestivalis Townes. 194-5: 1902b:94.-N.H. 86 (in part).-N.J. unictus. Qu6. to Miss. & Fla. browni. Fla. Stenochironomus unictus Townes. 1945: Stenochironomus browni Townes. 1945: 89.-N.J. 86.-Fla. woodi. Ont. to Nfld., s. to Va. cinctus. Ont., Qud., s. to Fla. Stenochironomus woodi Borkent. 1984: Stenochironomus cinctus Townes. 194-5: 91.-Qu6. 87.-Fla. totifuscus. Calif., Ariz. Stenochironomus totifuscus Sublette, 1.960: Genus STICTOCHIRONOMUS Kieffer 210.-Calif. Stictochironomus Kieffer in Kieffer & Thienemann, 1919: 44. Type-sPecies, Subgenus STENOCHIRONOMUS Kieffer Chironomus pictulus Meigen (Townes, 1945: 77). albipalpus. Pa., Md., Va. Stenochironomus albipalpns Borkent, 1984: albicrus. Kans. 66.-Md. Tanytarsus albicrus Townes, 1945: annettae. Tex. 79.-Kans. Stenochironotnus annettae Borkent. 1984: annulicrus. Ont., N.Y., S. Dak., Tenn. 54.-Tex. Tanytarsus annulicrus Townes, 1945: bisetosus. Ariz. 83.-N.Y. Stenochironomus bisetosus Borkent. 1984: 61.-Ariz. 54 NEARCTIC CHIRONOMIDAE

devinctus. Ont., Minn. to Maine, s. to Tex. & luski. Man., Calif. Fla. Synendotendipes luski Grodhaus, 1987 a: Chironomus devinctus Say, 1829: 202.-Calif. 150.-Ind. Chironomus compes Coquillett, 1908: 14.5.-Md. Genus TRIBELOS Townes Chironomus ornatipes Kieffer, 1917: Tribelos Townes, '1,945: 66 (as subg. of 343.-N.Y. Tanytarsus Wulp, 1874 sensu Townes). flavicingula. Ont.; PA. Type-species, Chironomus dimorphus Chironomus flavicingula Walker, 1848: Malloch (orig. des.) = Chironomus 20.-Ont. jucundus Walker. lutosus, Mass. Tanytarsus lutosus Townes, 1945: atrum. Man., Ont., Qu6., Minn., Mich., Va., 83.-Mass. N.C., S.C., Fla. marmoreus, Sask., Qu6., N.Y., S.Dak., Mo., Tanytarsus ater Townes, 1945: 68.-Va. N. Mex. fuscicorne. Minn. to Maine, s. to Tex. & Fla. Tanytarsus marmoreus Townes, 1945: Chironomus fuscicornis Malloch, 191-5b: 81.-Sask. 466.-I!l. naevus. Calif., N. Mex. jucundum. B.C., Oreg. to N.B., s. to Okta. & Chironomus naevus Mitchell, 1908: Fla. 14.-N. Mex. Tanytarsus jucundus Walker, 1848: palliatus. Wis. to N.Y. & D.C., s. to Tex. & 76.-Ga. Ala. Chironomus dimorphus Malloch, 191-5b: Chironomus palliatus Coquillett, 7902b: 464.-ln. 95.-D.C. Chironomus fusciventris Malloch, 1915b: pictulus. Man.; PA. 465.-Wis. Chironomus pictulus Meigen 1830: 244 (n. Tanytarsus protextus Townes, 194-5: nam. for Chironomus sticticus Meigen, 69.-N.Y. 1818: 37, preocc. Fabricius, subatrum. Calif. 1781).-Not stated. Tribelos subatrum Grodhaus. L987a: quagga. Sask., Calif. 227 --Calif . Tanytarsus quagga Townes, 1945: subletteorum, Calif. 81.-Calif. Tribelos subletteorum Grodhaus, 1987 a: rosenschoeldi. Alas., N.W.T., Sask.; PA. 232.-Calif. Chironomus rosenschoeldi Zetterstedt, 1838: 811.-Europe. sticticus. Man.; PA. Genus XENOCIIIRONOMUS Kieffer Tipula stictica Fabricius, 1781: Xe nochiro nomus Kieffer, l92lc: 27 3. 407.-Germany. Type-species, Chironomus xenolabis Tipula histrio Fabricius, 1794: Kieffer (Goetghebuer in Goetghebuer 244--Germany. & Lenz, 7937l. 32). unguiculatus. N.W.T., Grld. Chironomus unguiculatus Malloch, 1934: xenolabis. B.C. to Qu6., s. to N. Mex. & 16.-N.W.T. Fla., Oreg. to Mo.; PA. varius. Minn. to N.Y., Iowa to Ohio, Colo., Chironomus xenolabis Kieffer in Kans., N. Mex. Thienemann & Kieffer, 1916: Tanytarsus varius Townes, 1945: 82.-N.Y. 526.-Sweden. virgatus. Mont., Ill. Tanytarsus virgatus Townes, 194-5: 84.-Mont. Genus XESTOCHIRONOMUS Sublette & Wirth Xestochirononas Sublette & Wirth. 1972: Genus SYNENDOTENDIPES Grodhaus 7 (n. nam. for Insulanus Sublette). Synendotendipes Grodhaus, 1987a: 198. Insulanus Sublette, 1967b: 546 (preocc. Type-species, luski Grodhaus (orig. Fleutiaux, 1933). Type-species, des.). Chironomus furcata Johannsen (orig. des.). CHIRONOMIDAE DE LA REGION NEARCTIQUE 55

brunneus. N. Mex. miildlekauffi. Man., Ont., N.Yi Ill., Kans., Xestochironomus brunneus Borkent, 1984: Fla. 28.-N. Mex. Pseudochironomus middlekauffi'Iownes, subletti. N. Mex., Tex., S.C., Fla. 1945: 18.-N.Y. Xestochironomus subletti Borkent, 1984: netta. Ont., Mich., N.Y., Mass., N.J. 25.-Tex. Pseudochironomus netta Townes, 1945: 19.-N.Y. pseudoviridis. Man., S. Dak., Iowa to N.Y., Genus ZAVRELIELLA Kieffer Colo. to Tex. Zavreliella Kieffer, l92}b: 334. Chironomus pseudoviridis Malloch, 191-5b: Type-species, Chironomus clavaticrus 450.-Ill. Kieffer (orig. des.) = Chironomus rex. Man., Iowa, Fla. marmoratus Wulp. Pseudochironomus rex Hauber, 7947: 458.-Iowa. varipennis. Ont., Qud., Mich., Mass., Pa., s. richardsoni. Ont. s. to Calif. & Fla. to N. Mex. & Fla. Pseudochironomus richardsoni Malloch, Chironomus varipennis Coquillett, 1902b: 1915b: 500.-Ill. 94.-N. Mex.

Tribe TANYTARSINI Tribe PSEUD O CHIRONOMINI Genus CLADOTANYTARSUS Kieffer Genus PSEUDOCHIRONOMUS Malloch Cladotanytarsas Kieffer, I92lc: 277. Pseudochironomus Malloch, 191-5b: -500. Type-species, pallidus Kieffer (sub. Type-species, richardsoni Malloch (orig. mon.; Kieffer, 1922e: 100). des.). Lenziella Kieffer, 1922c: 36I. Type-species, bicornuta Kieffer (orig. anas. Qu6., N.Y. des.). Pseudochironomus anas Townes. 7945:. 17.-N.Y. aeiparthenus. Ont. articaudus. Man., Ont., Qu6. Cladotanytarsus aeiparthenus Bilyj, Pseudochironomus articaudus Seether, 1989: 948.-Ont. 1977b: 81.-Man. atridorsum. Man.; PA. aureus. Kans. Cladotanytarsus atridorsum Kieffer, 1924b: Psectrocladius aureus Johannsen, 1908: 38.-Germany. 283.-Kans. crusculus. S. Dak., N. Mex. badius. Man. Lenziella cruscula Sether, 1971: 1825.-5. Pseudochironomus badius Sether 1977b: Dak. 69.-Man. daviesi. Ont. banksi. Ont., N.J., Mass., Fla. Cladotanytarsus daviesi Bilyj' 1989: Pseudochironomus banksi Townes, 1945: 9-51.-Ont. 17.-Mass. elaensis. Ont. chen. Ont., Md., N. Mex. to La. Cladotanytarsus elaensis Bilyj, 1989: Pseudochironomus chen Townes. 1945: 955.-Ont. 18.-Md. fusiformis. Ont. crassus. N.W.T., Man. Cladotanytarsus fusiformis Bilyj, Pseudochironomus crassus Townes, 1945: 1989: 9-54.-Ont. 15.-Hudson Bay Territory, N.W.T. mancus. Iowa: PA. fulviventris. Man. to N.8., S. Dak. to N.Y., Chironomus mancus Walker, I956a: s. to Tex. & Fla. 161.-England. Chironomus fulviventris Johannsen, 1.905: muricatus. Ont. 229.-N.Y. Cladotanytarsus muricatus Bilyj' julia. Qu€., Mich., N.Y., Ohio, N. Mex. to 1989: 956.-Ont. La. pinnaticornis. Ont. Orthocladius julia Cwran, 1930: 34.-N.Y. Cladotanytarsus pinnaticornis Bilyj, Pseudochironomus alx Townes. 1945: 1989: 953.-Ont. 19.-Mich. 56 NEARCTIC CHIRONOMIDAE

tribelus. Ont. apposita. Ont.; PA. Cladotanytarsus tribelus Bilyj, 1989: Chironomus apposita Walker, 1956a. 957.-Ont. 191.-England. viridiventris. Alas., N.W.T., Alta., Ont., Qu6., attenuata. Ont.; PA. Mich., N.Y., Ohio, Calif. to N.C., s. to Micropsectra attenuata Reiss, 1969c: Fla. 432.-Germany. Tanytarsus viridiventis Malloch, 1915b: borealis. N.W.T., Novaya Zemlya. 491.-Mich. Oeklandia borealis Kieffer. 1922b: 6.-Novaya Zemlya. brundini. Y.T., N.W.T., Grld.; PA. Genus CONSTEMPELLINA Brundin Micropsectra brundini Sdwedal, 1979: Constempellina Brundin, 1947 :, 82. 134.-Grld. Type-species, Tanytarsus brevicosta connexa. Sask., Ont., Qu6., Maine. Edwards (orig. des.). Chironomus connexus Kieffer, 7906a: l7 (n. nam. for Chironomus confinis brevicosta. Alas. to N.W.T., Man. to Qu6.; Walker, 1848: 15, preoccc. Meigen, PA. 1830).-Ont. Tanytarsus brevicosta Edwards, 1937: Chironomus brunneus Walker. 1848: 146.-Sweden. 21.-Ont. deflecta. N.Y., Ohio. Tanytarsus deflectus Johannsen, 1905: Genus CORYNOCERA Zetterstedt 288.-N.Y. Corynocera Zettersted,t, 1838: 856. dives. Mont., Wis. to Maine, Colo., Ohio, Ky., Type-species, ambigua Zetterstedt N.Y. (mon.). Tanytarsus dives Johannsen, 1905: 288.-N.Y. ambigua. N.W.T.; PA. groenlandica.' Y.T., Grld.; PA. Corynocera ambigua Zetterstedt, 1838: Micropsectra groenlandica Andersen, 1937: 856.-Sweden. 33.-Grld. insignilobus. Grld., N.W.T.; PA. Micr ops e c tr a ins ignil obus Kieffer, Genus KRENOPSECTRA Reiss t924c: 86.-Norway. Krenopsectra Reiss, 1969b: 435. junci. Mich., Grld.; PA. Type-species, fallat Reiss (orig. des.). Chironomus junci Meigen, 1818: 50.-Germany. IJnnamed species.-W. Can. Chironomus brunnipes Zetterstedt, 1850: 3518.-Sweden. lindrothi. Grld.: PA. Genus MICROPSECTRA Kieffer Micropsectra lindrothi Goetghebuer in Micropsectra Kieffer, 1909: -50 (as subg. of Goetghebuer & Lindroth, 1931: Tanytarsus Wulp, 1874). Type-species, 278.-Iceland. Tanytarsus inermipes Kieffer (Kieffer, logani. Utah., N. Mex. l92lb: 35, as inermis) - Chironomus Tanytarsus logani Johannsen, 1928: notescens Walker, . 33.-Utah. Lauterbornia Kieffer, t9tla: 42. nigripila. Sask., Man., Qu6., N.Y. s. to Calif. Type-species, Chironomus c oracinus & La. Zetterctedt, 1850 sensu Kieffer, Tanytarsus nigripilus Johannsen, 1905: (misident., orig. des.) 287.-N.Y. Lundstroemia Kieffer, t92lc: 276. polita. Alta., Sask., Qu6., Pa., Md., S. Dak., Type-species, Tanytarsus roseiventris Ill., Oreg. s. to Ariz. Kieffer, 1909 (sub. rnon.; Kieffer, Tanytarsus politus Malloch, 1915b: r921b: 36). 493.-Iil. Oeklandia Kieffer, L922b: 6. recurvata. N.C., S.C., Grld.; PA. Type-species, borealis Kieffer (mon.). Micropsectra recurvata Goetghebuer, 1928: 117.-Belgium. CHIRONOMIDAE DE LA REGION NEARCTIQUE )/

sedna. N.W.T. intricatus. Man.; PA. Lauterbornia sedna Oliver, 1976: Tanytarsus intricatus Goetghebuer, t92l: 1061.-N.W.T. 121.-Belgium. tori. Grld. laccophilus. Man.; PA. Micropsectra rori Sdwedal, 1981: Tanytarsus laccophilus Edwards, 1929: 28.-Grld. 409.-England. xantha. Alta., Mass., Ky., N.C., s. to Miss. & natvigi. N.W.T., Grld.; PA. Fla. Micropsectra natvigi Goetghebuer, 1933: Calopsectra xantha Roback, L955: Zl.-Grld. 4.-Mass. penicillatus. Alas., N.W.T.; PA. Tanytarsus pencillatus Goetghebuet, 7928: 143.-England. Genus NIMBOCERA Reiss setosimanus. N.W.T., Grld.; PA. Nimbocera Reiss, 1972: 54. Type-species, Ditanytarsus setosimanus Goetghebuer, patagonica Reiss (orig. des.). 1933: 2t.-Grld. similatus. Alas.. Wis. pinderi. S.C., Ga., Fla. Tanytarsus similatus Malloch, 191-5b: Nimbocera pinderi Steiner & Hulbert, 494.-Wis. 1982: 228--Fla, tenuis. Grld.: PA. Chironomus tenuis Meigen, 1830: 255.-?Germanv. Genus PARAPSECTRA Reiss Parapsectra Reiss, 1969a: 192. Type-species, Chironomus nanus Genus RHEOTANYTARSUS Thienemann & Meigen (orig. des.). Bause Rheotanytarsas Thienemann & Bause in Unnamed species.-Maine, N.C. Bause, I9l3: l2O (as subg. of Syntanytarsus Thienemann & Bause in Bause, 1913). Type-species, Tanytarsus Genus PARATANYTARSUS Thienemann & pentapoda Kieffer (Goetghebuer in Bause Goetghebuer & Lenz, 1954: 132). Paratanytarsas Thienemann & Bause in Bause, l9l3: 120. Type-species, akrina. Kans. Tanytarsus lauterborni Kieffer (Reiss & Calopsectra akrina Roback, 1960: Sdwedal, l98l: 74). l.-Kans. Stylotanytarsus Kieffer, t92Ic: 276. distinctissimus. Alta., N.Y., Ohio; PA. Type-species, Tanytarsus Stempellinella distinctissima Brundin, L947: bauseellus (Langton et al., 1988: 88.-Sweden. 318). exiguus. Qu6., Nfld., Mont. to N.Y., s. to Miss. & Fla. dissimifis. Ont., S. Dak. to Iowa, N.Y., Tanytarsus exiguus Johannsen, 1905: Ohio, Okla., Tex., N.C., S.C.; 294.-N.Y. PA, perhaps widespread. magnini. Qud. Tanytarsus dissimilis Johannsen, 190-5: Rheotanytarsus magnini Cloutier & 292.-N.Y. Harper, 1986: 1.-Qu6. Paratanytarsus confusus Palmdn, pellucidus. Ont. 1960: 283.-Finland. Chironomus pellucidus Walker, 1848: dubius. Alta., Ill., Ky., N.C., s. to Miss. & 21.-Ont. Fla. Tanytarsus dubius Malloch, 1915b: 496.-Ill. Genus SKUTZIA Reiss grimmii. Wash., Ont., N.B., Maine, Skutzia Reiss, 1985: 173. Type-species N.Y. inopinata Reiss (orig. des.). Chironomus grimmii Schneider, 1885: 301.-U.S.S.R. inopinata. B.C. Tanytarsus boiemicus Bause, 1913: Skutzia inopinata Reiss, 1985: I74.-8.C. 97.-Czechoslovakia. 58 NEARCTIC CHIRONOMIDAE

Genus STEMPELLINA Thienemann & Bause allicis. La. Stempellina Thienemann & Bause in Tanytarsus a//icis Sublette, L964b: Bause, l9l3: 720. Type-species, 146.-La. Tanytarsus bausei Kieffer (mon.). anderseni. Alta., Man., Grld. Tanytarsus anderseni Reiss & Fittkau, bausei. Not NE. I97I: 129.-Grld. Tanytarsus bausei (originally as bauri, brundini. N.Y.; PA. typographical error) Kieffer, 1911a: Tanytarsus brundini Lindeberg, 1963: 46.- Germany. 127.-Finland. johannsenii. N.Y" buckleyi. Alta., N.Y., Ill., D.C., N. Mex. to Tanytarsus johannsenii Thienemann & N.C., s. to Fla. Bause in Bause, 1913: 67.-N.Y. Tanytarsus buckleyi Sublette, 1964b: leptocelloides. Alta., Ont. 143.-La. Stempellina leptocelloides Webb, 1969: confusus. N.Y., D.C., Ill., Okla., Ky., N.C., s. 94.-Ont. to Tex. & Fla. ranota. Ont. Tanytarsus confusus Malloch, 1915b: Stempellina ranota Webb, 1969: 96.-Ont. 490.-Ill. rodesta. Ont. curticornis. Alta.; PA. Stempellina rodesta Webb, 1969: 9-5.-Ont. Tanytarsus curticornis Kieffer, l9lla: 53.-Germany. debilis. Man.; PA. Genus STEMPELLINELLA Brundin Chironomus debilis Meigen, 1830: Stempellinella Brundin, 1,947: 87. 25-5.-Germany. Type-species, Tanytarsus saltuum dendyi. N. Mex., Ky., N.C., s. to Tex. & Fla. Goetghebuer (orig. des.). Tanytarsus dendyi Sublette, 1,964b: 147.-La. brevis, Man., Ont.; PA. dispar. Alta., Man., Ont.; PA. Tanytarsus brevis Edwards, 1929: Tanytarsus dispar Lindeberg, 1967: 420.-Scotland, England, Ireland. 55.-Finland. minor. Alas., Man.; PA. fatigans. Colo., N.Y. Tanytarsus minor Edwards, 1929: 420.-N. Tanytarsus fatigans Johannsen, 190-5: Wales, England. 292.-N.Y. fimbriatus. N. Mex.; PA. Tanytarsus fimbriatus Reiss & Fittkau, Genus SUBLETTEA Roback 1971: 118.-Spain. Sublettea Roback, 7975:- 7l (as subg. of gracilentus. Alas., N.W.T., B.C., Oreg.; PA. Tanytarsus Wulp, 1874). Type-species, Chironomus gracilentus Holmgren, 1883: Tanytarsus coffmani Roback (orig. 18l.-Novaya Zemlya. des.). Chironomus deviatus Malloch, 1923: 772.-Alas. coffmani. N.B., Vt., N.Y., Ohio, Pa., s. to guerlus. Man., N.Y., Pa., N.J., Fla. Ga. Calopsectra guerla Roback, I957a: Tanytarsus coffmani Roback, 1975: 133.-Pa. 72.-N.8. inaequalis. Alas., N.W.T., Man.; PA. Tanytarsus inaequalis Goetghebuer, l92l: 118.-Belgium. Genus TANYTARSUS Wulp islandicus. N.W.T. Tanytarsus Wulp, 1874: 134. Tanytarsus islandica Malloch, 1934: Type-species, Chironomus signatus 17.-N.W.T. Wulp, 1858 (ICZN 196I: 361, Opinion lapponicus. Man.; PA. 616). Tanytarsus lapponicus Lindeberg, 1970: Calopsectra Kieffer, 1909: -50. 310.-Finland. Type-species, Calopsectra limneticus. Alta., La. gregaria Kieffer (Kieffer, l92lb: Tanytarsus limneticus Sublette, 1964b: 36). 146.-La. CHIRONOMIDAE DE LA REGION NEARCTIQUE -59

mendax. B.C., Man., N. Mex. to La., N.C., Genus ZAVRELIA Kieffer S.C.; PA. Zavrelia Kieffer in Bause l9l3: 73. Tanytarsus mendax Kieffer, t925b: Type-species, pentatoma Kieffer (mon.). 2Z3--Germany. Tanytarsus holochlorus Edwards, 1929: pentatoma. Not NE. 414.-England. Zavrelia pentatoma Kieffer in Bause, 1913: Tanytarsus xanthus Sublette, 1,964b: 75.-Czechoslovakia. 142.-La. IJnnamed species.-B.C., Ont., N.8., muticus. N.Y., Wis., Ill., Ohio, Colo. N.S. Tanytarsus muticus Johannsen, 1905: 294.-N.Y. nemorosus. Man.; PA. Tanytarsus nemorosus Edwards, 1929: 416.-England. neoflavellus, Man., S. Dak., Okla., N.Y., Ill., Ohio. s. to Tex. & Fla. Tanytarsus neoflavellus Malloch, 191-5b: 489.-Ill. niger. N.W.T., Grld.; PA. Tanytarsus niger Andersen, 1937 : 40.-Grld. norvegicus. N.W.T.; PA. Fournieria norvegica Kieffer, 1924a: 394.-Norway. pallidicornis. N. Mex.; PA. Chironomus pallidicornis Walker, 1956a: 169.-England. quadratus. N. Mex. to N.C., s. to Fla. Tanytarsus quadratus Sublette, 1,964b: 144.-La. quadridentatus. Man.; PA. Tanytarsus quadridentatus Brundin, 1947: 75.-Sweden. recens. N. Mex. to N.C.. s. to Fla. Tanytarsus recens Sublette, 1964b. 743.-La. signatus. Ont., N.Y.; PA. Chironomus signatus Wulp, 1858: 169.-Netherlands. tibialis. Ont. Tanytarsus tibialis Webb, 1969: 100.-Ont. trilobus. Ont. Tanytarsus tilobus Webb, 1969: 99.-Ont. usmaensis. Man.; PA. Tanytarsus usma€nsis Pagast, 193 1: 232.-U.S.S.R. varelus. Pa. Calopsectra varela Roback, 1957a: 128.-Pa.

Incerta sedis in Tanytarsus Chironomus flab ell atus Meigen, 1818: 51.- Not NE. Chironomus flavellus Zetterstedt, 1838: 816.- Probably not NE. Chironomus pusio Meigen, 1830: 2-56.-Not NE. 60 NEARCTIC CHIRONOMIDAE

REFERENCES\BIBLIO GRAPHIE from Southern Indian Lake. Canada. Can. J. Fish. Aquat. sci. 41: 659-671. Aagard, K. 1979. Eukiefferiella sivertseni n. sp. Bilyj, B. & I.J. Davies. 1989. Descriptions and from Norway (Diptera: Chironomidae). ecological notes on seven new species of Entomologica scand. Suppl. l0: 95-97. Cladotanytarsus (Chironomidae: Diptera) Andersen, F.S. 1937. Uber die Metamorphose collected from an experimentally acidified der Ceratopogoniden und Chironomiden lake. Can. J. Zool. 67; 948-962. Nordost-Gr0nlands. Meddr Grdnland Boerger, H.J., H.F. Clifford & R.W. Davies. 116(1):1-95. 1982. Density and microdistribution of Ashe, P. 1981. A taxonomic note on the genus chironomid larvae in an Alberta brown-water Polypedilum Kieffer (Diptera: Chironomidae) stream. Can. J. Zool. 60:. 9t3-920. with an earlier date for the genus and Boesel, M.W. 1983. A review of the genus designation of a type-species. Aquat. Insects Cricotopus in Ohio, with a key to adults of 3-. 57 -59. species of the northeastern United States Ashe, P. 1983. A catalogue of chironomid (Diptera, Chirqnomidae). Ohio J. Sci. 83: genera and subgenera of the world including 7 4-90. synonyms (Diptera: Chironomidae). Boesel, M.W. & R.W. Winner. 1980. Entomologica scand. Suppl. I7: l-68. Corynoneurinae of northeastern United Ashe, P. & D.A. Murray. 1980. Nosfococladius, States, with a key to adults and observations a new subgenus of Cricotopus (Diptera: on their occurrence in Ohio (Diptera: Chironomidae). Pp. 105-111. /n: Murray, Chironomidae). J. Kans. ent. Soc. 53: D.A. (Ed.): Chironomidae - Ecology, 501--508. Systematics, Cytology & Physiology. Boheman, C.H. 1865. Spetsbergen insekt-fauna. Pergamon Press, Oxford. 3-54 pp. K. svenska VetenskAkad. Ofvers. Fdrhandl. Atchley, W.R. & J. Martin. 1971. A 22: 563-57'7. morphometric analysis of differential sexual Borkent, A. 1984. The systematics and dimorphism in larvae of Chironomus phylogeny of the Stenochironomus complex (Diptera). Can. Ent. lO3:319-327. (Xe s t ochiron o mus, H arrisias, and Bause, E. 1913. Die Metamorphose der Stenochironomus) (Diptera: Chironomidae). Gattung Tanytarsus und einiger verwandter Men. ent. Soc. Can. 128: I-269. Tendipedidenarten. Ein beitrag zur Brimley, C.S. 1938. The insects of North Systematik der Tendipediden. Arch. Carolina. N.C. Dep. of Agriculture, Raleigh, Hydrobiol. Suppl. 2: I-241. 560 pp. Beck, E.C. 1961. Two new Chironomidae Brundin, L. 1947. Zur Kenntnis der (Diptera) and additional state records from schwedischen Chironomiden. Ark. Zool. 39:. Florida. Fla Ent. 44: t25-128. 1-95. Beck. E.C. 1962. Five new Chironomidae Brundin. L. 1.949. Chironomiden und andere (Diptera) from Florida. Fla Ent. 45: 89-92. Bodentiere der sridschwedischen Beck, E.C. & W.M. Beck. t969. Chironomidae Urgebirgsseen. Ein Beitrag zur Kenntnis der (Diptera) of Florida. 111. The Harnischia bodenfaunistischen Charakterziige complex (Chironominae). Bull. FIa St. Mus. schwedischer oligotropher Seen. Rep. Inst. biol. Sci. 13: 277-313. Freshwat. Res. Drottningholm 30: 1-915. Beck, W.M. & E.C. Beck. 1958. A new species Brundin, L. 19-56. Zur Systematik der of. Xenochironomus from Florida (Diptera: Orthocladiinae (Dipt., Chironomidae). Rep. Chironomidae). Fla Ent. 4l: 27-28. Inst. Freshwat. Res. Drottningholm 37:5-185. Beck. W.M. & E.C. Beck. 1964. New Brundin, L. 1966. Transantarctic relationships Chironomidae from Florida (Diptera). Fla and their significance, as evidenced by Ent. 47:2Ot-207. chironomid midges with a monograph of the Beck. W.M. & E.C. Beck. 1966. Chironomidae subfamilies Podonominae and Aphroteniinae (Diptera) of Florida. I. Pentaneurini and the Austral Heptagyiae. K. svenska (Tanypodinae). Bull. Fla St. Mus. biol. Sci. VetenskAkad. Handl. ll:. 7-472. 10: 305-379. Butler, M.G. 1982. Morphological and Beck, W.M. & E.C. Beck. L970. The immature phenological delimitation of. Chironomus stages of some Chironomini (Chironomidae). prior sp. n. and C. tardus sp. n. (Diptera: Q. Jl Fla Acad. Sci. 33: 29-42. Chironomidae), sibling species from Arctic Bilyj, B. 1984. Descriptions of two new species Alaska. Aquat. Insects 4: 219-235. of Tanypodinae (Diptera: Chironomidae) Caldwell, B.A. 1984. Two new species and records of other chironomids from Georgia CHIRONOMIDAE DE LA REGION NEARCNQUE 61

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Dendy, J.S., & J.E. Sublette. 19-59. The Fabricius, J.C. 178I. Species insectorum Chironomidae (= Tendipedidae: Diptera) of exhibentes eorum differentias specificas, Alabama with descriptions of six new species. synonyma, auctorum, loca natalia, Ann. ent. Soc. Am. 52: 506-519. metamorphosin. 517 pp. Ifamburgi et Eaton, A.E. 1875. Breves dipterarum uniusque Kilonii [=Hamburg and Kiel]. lepidopterarum Insulae Kerguelensi Fabricius, J.C. I794. Entomologia systematica indigenarum diagnoses. Entomologist's mon. emendata et aucta. Secundum classes, Mag. 12: 58-61. ordines, genera, species, adjectis synonymis, Edwards, F.W. 1922. Results of the Oxford locis, descriptionibus, observationibus. 4, University Expedition to Spitzbergen, 1921. 472 pp. Copenhagen. Ann. Mag. nat. Hist. (9)lO: t93-2t5. Fabricius, J.C. 180-5. Systema antliatorum Edwards, F.W. I924a. A note on the genus secundum ordines, genera, species. 373 pp-, Protanypus Kieffer (Diptera: Chironomidae). + 30 pp. Brunsvigae [=Brunswick]. Annfs Biol. lacustre 13: 7I9-I22. Ferrington, L.C. & O.A. Sether. 1987. Male, Edwards, F.W. I924b. Results of the Merron female, pupa and biology of. Oliveridia College Expedition to Spitzbergen, 7923, hugginsi n. sp. (Chironomidae: Diptera) from No. 4. Diptera . Ann. Mag. nat. Kansas. J. Kans. ent. Soc. 6O: 457-461 Hist. (9)14: 162-174. Fitch, A. 7847. Winter insects of eastern New Edwards, F.W. 1924c. Some British species of York. Am. J. Agric. Sci. 5: 274-284. C orynoneura (Diptera-Chironomidae). Fittkau, E.J. 1957. Thienemannimyia und Entomologist's mon. Mag. 60: 182-189. Conchapelopia, zstei neue Gattungen Edwards, F.W. 7929. British non-biting midges innerhalb der Ablabesmyia-Costalis-Gruppe (Diptera, Chironomidae). Trans. R. ent. (Diptera, Chironomidae). Soc. Lond. 7'7: 279-430. Chironomidenstudien VII. Arch. Hydrobiol. Edwards, F.W. 1931. Diptera of Patagonia and 53: 313-322. South Chile. 11, 5, Chironomidae: 233-331.. Fittkau, E.J. 1962. Die Tanypodinae (Diptera: Trustees of the British Museum, London. Chironomidae). Die Tribus , Edwards, F.W. 1932a. Notes on Highland Macropelopiini und Pentaneurini. Abh. Diptera, with descriptions of six new species. Larvalsyst. Insekten 6: 7-453. Scott. Nat. 794: 43-52. Fittkau, E.J. 7965. Revision der von E. Goeldi Edwards, F.W. I932b. Recent literature, Faune aus dem Amazonasgebiet beschriebenen de France: 23, Diptdres: Chironomidae, IV. Chironomiden (Diptera). Par M. Goetghebuer. Paris (Lechavalier), Chironomidenstudien X. Beitr. neotrop. 1932. Entomologist 65: 140-141. Fauna 4: 209-226. Edwards, F.W. 1933. Oxford University Fittkau, E.J. 1968a. Eine neue Expedition to Hudson's Strait, 1931: Diptera Tanypodinae-Gattung, Dj almab atis ta Nematocera. With notes on some other (Chironomidae, Diptera), aus dem species of the genus Diamesa. Ann. Mag. brasilianischen Amazonasgebiet. Amazoniana nat. Hist. 10(12): 6Il-620. l:327-349. Edwards, F.W. 1935. Diptera Nematocera from Fittkau, E.J. 1968b. Siolimyia amazonica n. East Greenland. Ann. Mag. nat. Hist. gen. n. spec., eine flugfdhige Chironomide 10(15):467-473. (Diptera) mit linem Hypopygium inversum. Edwards, F.W. 1937. Chironomidae (Diptera) Amazoniana L: 259-265. collected by Prof. Aug. Thienemann in Fittkau, E.J. & D.A. Murray. 198-5. Swedish Lappland. Ann. Mag. nat. Hist. Ser. Radotanypus a new genus of Tanypodinae 10(20): 140-148. from the Nearctic (Diptera, Chironomidae). Edwards, F.W. 1938. I. A new genus of the Spixiana Suppl. (Muench.) 17: 209-273. subfamily Podonominae. Pp. 152-154. In: Fittkau, E.J. & D.A. Murray. 1988. Edwards, F.W. & A. Thienemann. Neuer Bethbilbeckia floridensis: a new genus and Beitrag zur Kenntnis der Podonominae species of Macropelopiini from the South (Dipt. Chironomidae). Chironomidae aus Eastern Nearctic (Diptera: Chironomidae). Lappland IV. Zool. Anz. 122: 752-758. Spixiana Suppl. (Muench.) 74: 253-259. Enderlein, G. 1972. Die Insekten des Antarkto- Fleutiaux, E. 1933. Genre nouveau et especes Archiplata-Gebietes. K. svenska nouvelles d'Eucnemididae de Madagascar VetenskAkad. Handl. 48: 106-110. (Col.). Bull. Soc. ent. Fr. (1932): 286-288. Epler, J.H. 1987. Revision of the Nearctic Freeman, P. 1957. A study of the Dicrotendipes Kieffer, 1913 (Diptera: Chironomidae (Diptera) of south of Chironomidae). Evol. Monogr. 9: i.-1.02. CFIIRONOMIDAE DE LA REGION NEARCTIQUE

the Sahara. Part III. Bull. Br. Mus. nat. Goetghebuer, M. 195-5. Deux Chironomides Hist. (Ent.) 5: 321-426. nouveaux du Groenland (Dipt. Nemat.). Fries, B.F. 1830. Beskrifning 6fver ett nytt Expeditions polaires francaises (Groenland): sllgte Hydrobaenus hOrande till Tipulariae. (Collection A. Bauer du Musee Zoologique K. svenska VetenskAkad. Handl. 1829: de Strasbourg). Bull. Ass. philomath. Als. 176-187. Lorr. 9: 767. Garrett, C.B.D. 1925- Seventy new Diptera. Goetghebuer, M. & F. Lenz. 7936. Key to Pseudoleria, Helomyzidae, Tipulidae, Tendipedidae-Pelopiinae (Chironomidae- Chironomidae, Dixinae, Orphnephilidae, Tanypodinae). Fliegen palaearkt. Reg. 13b: Mycetophilidae. Cranbrook, B.C., 16 pp. 1-81. Gistl, J. 1835. Neue Arten von Dipteren aus Goetghebuer, M.& F.Lenz. 1937. der Umgegend von Mrinchen, benannt und Tendipedidae-Tendipedidae beschrieben von Meigen, aufgefunden von (Chironomidae-Chironominae). [part]. Dr. J. Waltl. Faunus 2: 66-72. Fliegen palaearkt. Reg. 13c: 1-48. Goeldi, E.A. 190-5. Os mosquitos no Para. Goetghebuer, M. & F. Lenz- 1938. Mems Mus. paraense Hist. nat. Ethnogr. 4: Tendipedidae-Tendipedidae 734-r39. (Chironomidae-Chironominae). [part]. Goetghebuer, M. 1921. Chironomides de Fliegen palaearkt. Reg. 13c: 73-128. Belgique et sp6cialement de la zone des Goetghebuer, M. & F. Lenz. 1939a. Flandres. M6m. Mus. r. Hist. nat. Belg. 8: Tendipedidae-Diamcsinae r-211. (Chironomidae-Diamesinae). Fliegen Goetghebuer, M. 1922. Nouveaux matdriaux palaearkt. Reg. 13d: 1-30. pour l'6tude de la fauna des Chironomides Goetghebuer, M. & F. Lenz. 1939b. de Belgique. Annls Biol. lacustre ll:. 38-62. Tendipedidae-Corynone urinae Goetghebuer, M. 1927 - Les Cricotopus de (Chironomidae-Corynoneurinae). Fliegen Belgique (Dipt. Chironomides). Bull. Annls palaearkt. Reg. 13f: 1-39. Soc. r. ent. Belg. 67: 5l-54. Goetghebuer, M. & F. Lenz. 1940. Goetghebuer, M. 1928. Diptdres Tendipedidae-Orthocladiinae (Ndmatocdres): Chironomidae. IIL (Chironomidae-Orthocladiinae) [part]. Chironomariae. Faune Fr. 18: 1-174. Fliegen palaearkt. Reg. 13g: 1-24. Goetghebuer, M. 1931. Ceratopogonidae et Goetghebuer, M. & F- Lenz. 1943. Chironomidae nouveaux d'Europe. Bull. Tendipedidae-Orthocladiinae Annls Soc. r. ent. Belg. Tl; 217-218. (Chironomidae-Orthocladiinae) [part]. Goetghebuer, M. 1.932a. Diptdres Fliegen palaearkt. Reg. 13g: 65-t1,2- Chironomidae 1V (Orthocladiinae, Goetghebuer, M. & F. Lenz. 1,944. Corynoneurinae, Clunioninae, Diamesinae). Tendipedidae-Orthocladiinae Faune Fr.23: 1.-204. (Chironomidae-Orthocladiinae) fpart]. Goetghebuer, M. 7932b. Ceratopogonidae et Fliegen palaearkt. Reg. 13g: 113-144. Chironomidae nouveaux ou peu connus Goetghebuer, M. & F. Lenz. 1954. d'Europe. Bull. Annls Soc. r. ent. Belg. 72: Tendipedidae-Tendipedidae t25-r30. (Chironomidae-Chironominae). [part]. Goetghebuer, M. 1933. Chironomides du Fliegen palaearkt. Reg. 13c: 128-168. Groenland oriental, du Svalbard et de la Goetghebuer, M. & C.H. Lindroth. 1931. Fam. terre de Frangois Joseph. Skr. Svalbard Ceratopogonidae und Chironomidae. Pp. Ishavet 53: 19-31. 274-285. In: Lindroth, C.H. (Ed.). Die Goetghebuer, M. 193-5. Ceratopogonidae et Insektenfauna Islands und ihre Problem. Chironomidae rdcoltds pour la premidre fois Zool. Bidr. Upps. 13: 105--599. en Belgique. Bull. Annls. Soc. r. ent. Belg. Gouin, F. t936. M6tamorphoses de quelques 75: 413-418. Chironomides d'Alsace et de Lorraine avec Goetghebuer, M. 1936. Chironomides du Congo la description de trois espdces nouvelles par Belge. Revue Zool. Bot. atu.28 453-492- M. Goetghebuer. Revue fr. Ent. 3: l5l-173- Goetghebuer, M. L940. Chironomides de Gould, J. 1840. Birds of Australia, 1-80. Laponie Su6doise. Bull. Annls Soc. r. ent. (Aves). Belg. 80: 55-72. Gowin, F. 1943. Orthocladiinen aus Lunzer Goetghebuer, M. 1944. Ceratopogonidae et Fliessengewdssern. II. Arch. Hydrobiol. 40: Chironomidae nouveaux ou peu connus 114-122. d'Europe (douziOme note). Biol. Jaarb. 11: 35-44. NEARCTIC CHIRONOMIDAE

Grandjean, F. 1964. Oribates mexicains (1'e sequentibus in insulis Waigatsch et Novaja sdrie) Dampfiella Selln. et Beckiella n.g. Semlia, anno 187-5 collecta. Ent. Tidskr. 4; Acarologia 6; 694-77I. 762-190. Grodhaus, G. 1976. Two species of Jackson, G.A. 1,977. Nearctic and Palaearctic Phaenopsectra with drought resistant larvae Paracladopelma Harnisch and Saetheria n. (Diptera: Chironomidae). J. Kans. ent. Soc. gen. (Diptera: Chironomidae). J. Fish. Res. 49: 405-418. Bd Can. 34:.1327-1359. Grodhaus, G. 1987a. Endochironomus Kieffer, Jacobson, G. 1898. Compte rendu de Tribelos Townes, Synentotendipe.r, n. gen., I'exp6dition envoyde par l'Acaddmie and Endotibelos, n. gen. (Diptera: Impdriales des Sciences a Novaia Zemlia en Chironomidae) of the Nearctic region. J. erc 1896- IV. Imp. Akad. Nauk St. Kans. ent. Soc. 60: 167-247. P6tersbourg, Z?p., Fiz.-Mat. Otd. ser. 8, 8: Grodhaus, G. 1987b. Phaenopsectra mortensoni l7t-244. n. sp. and its relationship to other Johannsen, O.A. 1903. Part 6. Aquatic Chironomidae (Diptera) of temporary pools. Nematocerous Diptera. Pp. 328-44I. In: Entomologica scand. Suppl. 29 137-145 Needham, J.G., A.D. MacGillivray, O.A. Haliday, A.H. 1855. Descriptions of insects Johannsen & K.C. Davis. Aquatic Insects of figured, and reference to plates illustrating New York State. Bull. 68 [=Ent. 18]: 197- the notes on Kerry insects. Nat. Hist. Rev. 517 (= N.Y. St. Univ. Bull. 295). 2: 59-64. Johannsen, O.A. 1905. Aquatic nematocerous Halvorsen, G.A. 1982. Saetheriella amplicristata Diptera II. Chironomidae. Pp. 76-37. In: gen. n., sp. n., a new Orthocladiinae Needham, I.G., K.I. Morton, & O.A. (Diptera: Chironomidae) from Tennessee. Johannsen. May and midges of New Aquat. Insecrs 4: 131-136. York. Third report on aquatic insects. Bull. Ilansen, D.C. & E.F. Cook. I976. The 86 1=911. 231: 7-352 (=N.Y. St. Educ. Dep. systematics and morphology of the Nearctic Bull. 343). species of Diamesa Meigen, 1835 (Diptera: Johannsen, O.A. 7907a. Notes on the Chironomidae). Mem. Am. ent. Soc. 30. 203 Chironomidae. Pp. 400-401. .In: Skinner, H. pp. (Ed.). Notes and news. Ent. News 18: Harnisch, O. 1923. Metamorphose und System 400-402. der Gattung Cryptochironomus. K s.l. Zool. Johannsen, O.A. 7907b. Appendix. Some new Jb. (Abt. Syst. Geogr. Biol. Tiere) 47: species of Kansas Chironomidae. Pp. 27r-308. 109-172. .[re: Tucker, E.S. Some results of Hashimoto, H. 1973. A new species of the desultory collecting of insects in Kansas and marine chironomid Paraclunio (Diptera) Colorado. Kans. Univ. Sci. Bull. 4: 5l-112. from southern California. Annotnes zool. Johannsen, O.A. 1908. New North American jap. 46: 266-273" Chironomidae. Pp. 264-285. In: Felt, E.P. Hashimoto, H. 1974. A new species of 23rd report of the State Entomologist on Clunionine from North America. Ent. injurious and other insects of the State New Tidskr. Suppl. 95: 108-114. York, 7907. N.Y. St. Mus. Bull. 124: 5- Hauber, IJ.A. 1947. The Tendipedinae of Iowa -541. (= N.Y. St. Univ., Educ. Dep. Bull. (Diptera). Am. Midl. Nat. 38: 456-465. 433). Ilayes, B.P. & D.A. Murray. 1988. Tokunagaia Johannsen, O.A. 1921,. The genus Diamesa obriaini-a new species of Orthocladiinae Meigen (Diptera, Chironomidae). Ent. News (Diptera: Chironomidae) from Northwest 32: 229-232. Territories, arctic Canada. Entomologica Johannsen, O.A. 1926. The genus scand. 18: 361-366. Trichotanypus Kieffer (Chironomidae, Hirvenoja, M. t973. Revision der Gattung Dipt.). Can. Ent. 58: 99-100. Cricotopus van der Wulp and ihrer Johannsen, O.A. 1928. Two new species of Verwandten (Diptera, Chironomidae). Annls western chironomids (Diptera). J. Ent. Zool. zool. Fenn. 10: 1-363. 20: 33-35. Holmgren, A.E. 1869. Bidrag till Kdnnedomen Johannsen, O.A. 1934. New species of North om Beeren Eilands och Spetsbergens America Ceratopogonidae and Chironomidae. Insekt-Fauna. K. svenska VetenskAkad. Jl N.Y. ent. Soc. 42: 343-352. Handl. (n.s.) 8(-5): 1-56. Johannsen, O.A. 1937a. Aquatic Diptera. Part Holmgren, A.E. 1883. Diptera. Insecta a viris III. Chironomidae: Subfamilies Tanypodinae, doctissimis Nordenskidld illun ducem Diamesinae and Orthocladiinae. CHIRONOMIDAE DE LA REGION NEARCTIQUE

Mem. Cornell Univ. agric. Exp. Stn 205: nouveau. Bull. Soc. Hist. nat. Metz 27: 1-84. 103- 105. Johannsen, O.A. t937b. Aquatic Diptera. Part Kieffer, J.J. 191,1c. Nouveaux Tendipddides du IV. Chironomidae: Subfamily Chironominae. groupe Orthocladius (Dipt.). (1'note); (2* Mem. Cornell Univ. agric. Exp. Stn 210: 3- note). Bull. Soc. ent. Fr. 1911: 181-187; 56. r99-202. Johannsen. O.A. 1942. Immature and adult Kieffer, J.J. 19t2a. Nouveaux Chironomides stages of new species of Chironomidae (Tendipedidae) de Ceylan. Spolia zeylan. 8: (Diptera). Ent. News 53: 7O-75- l-24. Johannsen. O.A. 7943. Adult and immature Kieffer, J.J. I9l2b. Tendipedidae (Dipt.). In: stages of Cricotopus elegans n. sp. M. Sauter's Formosa Ausbeute. Supplta. (Chironomidae, Diptera). Ent. News 54: ent. l.: 27-43. 77-79. Kieffer, J.J. 191.2c. Quelques nouveaux Johannsen, O.A. 7946. Revision of the North Tendip6dides (Dipt.) obtenus d'dclosion. I. American species of the genus Pentaneura note. Bull. Soc. ent. Fr. 17: 86-88. (Tendipedidae: Chironomidae, Diptera). Jl Kieffer. J.J. l9l3a. Nouvelle contribution a la N.Y. ent. Soc. 54: 267-289. connaissance des Tendip6dides D'Allemagne. Johannsen, O.A. 1947. A new species of Bull. Soc. Hist. nat. Metz. 3: 37-44. Hydrob aenus (Chaetocladius) fr om Kieffer, J.J. 1913b. Nouveaux Chironomides Connecticut with notes on related forms (Tendip6dides) d'Allemagne. Bull. Soc. Hist. (Diptera, Chironomidae). Ent. News 58: nat. Metz 28:. 7-35. I7l-174. Kieffer, J.J. 1913c. Nouvelle 6tude sur les Johnson, C.W. 1900. Order Diptera. Pp. 617- Chironomides de l'Indian Museum de '700. In: Smith, J.B. Insects of New Jersey. Calcutta. Rec. Indian Mus. 9: ll9-I97. A list of the species occurring in New Jersey Kieffer. J.J. 1913d. Chironomidae et with notes on those of economic importance. Cecidomyidae. Result. scient. Voy. Ch. Bull. N.J. agric. Ann. Rept. (1889) 27 Alluaud et R. Jeanell en Afrique Orientale (Supp.): 1-75-5. (1e11-1012) (Dipt.) I: 1-43. Johnson, C.W. 1925. Fauna of New England. Kieffer, J.J. 1913e. Dasyhelea halophila n. sp., 15. List of the Diptera or two-winged flies. eine neue halophile Zuckmticke. Pp. 255- Occ. Pap. Boston Soc. nat. Hist. 7: l-326. 256. In: Kieffer, J.J., & C. van Douwe Jones, F.M. 1916. Two associates of the (Eds). Zur Flora und Fauna der California pitcher plant, Darlingtonia Strandtrimpel von Rovigno (in Istrien). Biol. californica (Dipt.). Ent. News 27: 385-392. Zbl. 33: 254-258. Karunakaran, L. 1969. A new genus of the Kieffer. J.J. 1914. South African subfamily Tanypodinae (Diptera, Chironomidae (Diptera). Ann. S. Afr. Mus. Nematocera: Chironomidae) from Singapore. l0: 259-270. Proc. R. ent. Soc. Lond. 38: 75-79. Kieffer, J.J. 1915a. Neue Chironomiden aus Kellogg, V.L. 1900. An extraordinary new Mitteleuropa. Boteria Ser. zool. 13: 6-5-87. maritime fly. Biol. Bull. 1: 81-87. Kieffer. J.J. 1915b. Fauna faeroensis: 2. Kevan, D.K.McE. & F.E.A. Cutten-Ali-Khan. Tendipedidae (Chironomidae). Zool. Jb. 39:. 1975. Canadian Nymphomyiidae (Diptera). 103-720. Can. J. Zool. 53: 8-53-866. Kieffer, J.J. 1917. Chironomides d'Amdrique Kieffer, J.J. 1906a. Diptera Fam. conservds au Mus6e National Hongrois de Chironomidae. Genera Insect. 42: I-78. Budapest. Budapest Magyar Nemzeti Muz., Kieffer, J.J. 1906b. Description d'un genre Ann. Hist. Nat. 15: 292-364. nouveau et de quelques espBces nouvelles Kieffer, J.J. 1918a. Beschreibung neuer, auf de Dipt0res de I'Amdrique du Sud. Annls Lazarettschiffen des 0stlichen Soc. scient. Brux. 30: 349-358. Kreigsschauplatzes und bei Ignalino in Kieffer, J.J. 1909. Diagnoses de nouveaux Litauen von Dr. W. Horn gessammelter Chironomides d'Allemagne. Bull. Soc. Hist. Chironomiden mit Uebersichtstabellen einer nat. Metz 26:. 37-56- Gruppen von paldarktischen Arten (Dipt.). Kieffer, J.J. 1911a. Nouvelles descriptions de Ent. Mitt. 7: 35-53;94-IIO; 163-l7O; 177- Chironomides obtenus d'dclosion. Bull. Soc. 188. Hist. nat. Metz 27: l-60. Kieffer, J.J. 1918b. Chironomides d'Afrique et Kieffer, J.J. 1911b. Description d'un d'Asie conservds au Mus6e National Chironomide d'Am€rique format un genre Hongrois de Budapest. Annls hist.-nat. Mus. natn. hung. 16: 31-139. 66 NEARCTIC CHIRONOMIDAE

Kieffer, J.J. 1,919a. Chironomides d'Europe l'Europe. Annls Soc. scient. Brux. 43: 390- conserv6s au Mus6e National Hongrois de 397. Budapest. Budapest Magyar Nemzeti Muz., Kieffer, J.J. 1924b. Chironomides nouveaux ou Ann. Hist. Nat. 17: 1-160. rares de l'Europe centrale. Bull. Soc. Hist. Kieffer, J.J. 1919b. Observations sur les nat. Moselle 30 [-ser. 3,6]: 11-110. Chironomides (Dipt.) ddcrits par J.R. Kieffer, J.J. 1924c. Quelques nouveaux Malloch. Bull. Soc. ent. Fr. 7979: l9l-794. Chironomides de Scandinavie. Annls Soc. Kieffer, J.J. 1,920a. Tableau synoptique des scient. Brux. 44: 80-86. Chironomides pal6arctiques appartenant aux Kieffer, J.J. 1,925a- Deux genres nouveaux et genres Polypedilum et Limnochironomus. plusieurs espCces nouvelles du groupe des Annls Soc. scient. Brux. 39: 759-767. Orthocladiariae (Dipteres, Chironomidae). Kieffer, J.J. I920b. IJn nouveau genre de Annls Soc. scient. Brux. 44: 55-5-566. Chironomide (Dipt.). Bull. Soc. ent. Fr. Kieffer, J.J. 7925b. Nouveaux reprdsentants du I9l9: 333-334. groupe Tanytarsus (Diptdres Chironomides) Kieffer, J.J. 1921,a. Chironomides nouveaux ou de I'Europe centrale. Annls Soc. scient. peu connus de la rdgion paldarctique. Bull. Brux. 44: 278-227. Soc. Hist. nat. Moselle 29 [-ser.3,-5]: -51-109. Kieffer, J.J. 1.925c. Description de deux Kieffer, J.J. 1921b. Chironomides de l'Afriqu6 Chironomides remarquables d'Estonie 6quatoriale (1'" partie). Annls Soc. ent. Fr. (Dipt.). Bull. Soc. ent. Fr. 30: 15-16. 90:1-56 Kieffer, J.J. 1926. Chironomiden der 2. Kieffer, J.J. 1921,c. Synopse de la tribu des Fram-Expedition (1898-1902). Norsk ent. Chironomariae (Diptdres). Annls Soc. scient. Tidsskr. (1925) 2: 78-89. Brux. 40: 269-277. Kieffer, J.J. & W. Lundbeck. 1911. Diptera. Kieffer, J.J. I922a. Diagnoses de nouveaux Pp. 272-275. In: Koenig, A. Avifauna genres et espCces de Chironomides (Dipt.). spitzbergensis. Bonn. 294 pp. Bull. Soc. ent. Fr. 7921,: 287-289. Kieffer. J.J. & A. Thienemann. 1906. Uber die Kieffer, J.J. 1922b. Cbironomides de la Chironomidengattung Orthocladius. Z. wiss. Nouvelle-Zemble. Rep. scient. Results InsektBiol. 2: L43-156. Norw. Exped. to Novaya Zemlya 1927. 2: 1,- Kieffer, J.J. & A. Thienemann. 1908. I. Neue 24. und bekannte Chironomiden. Pp. 1-10, 33- Kieffer, J.J. 1922c. Nouveau Chironomides a 39, 78-84. In: Kieffer, J.J. & A. larves aquatiques. Annls Soc. scient. Brux. Thienemann. Neue und bekannte 4r-. 355-367. Chironomiden und ihre Metamorphose. Z. Kieffer, J.J. I922d. Chironomides de l'Afrique wiss. InsektBiol. 4: 1-10, 33-39,78-84, 724- Equatoriale (2' partie). Annls Soc. ent. Fr. I28, 184-190, 21,4-219, 256-259, 277 -286. 91: l-72. Kieffer, J.J. & A. Thienemann. I9I9. Kieffer, J.J. 1922e. Chironomides nouveaux ou Chironomiden gesammelt von Dr. A. Koch peu connus de la rdgion paldartique. Annls (Munster i.W.) auf den Lofoten, der Soc. scient. Brux. 42: 71,-128. Bareninsel und Spitzbergen (Dipt.). Kieffer, J.J. 1922t. Notice sur quelque ((Vorwort von A. Kock: 38-39)). Chironomides d'Amdrique et de Nouvelle- Chironomiden der nordlichen Polarregion Z6.lande. Annls Soc. linn. Lyon (1921) 68: (Kieffer, J.J.: 40-48; 110-120). 145-r48. Chironomidenlarven und -puppen von Kieffer, J.J. 1923a- Diagnose de quelque Bareninsel un Spitzbergen. (Thienemann, A.: nouveaux Tanypodines (Dipt.). Bull. Soc. 120-124). Ent. Mitt. 8: 38-48; 710-724. ent. Fr. 296-297. Kruseman, G. 1933. Tendipedidae Kieffer, J.J. 1923b. Chironomides de l'Afrique Neerlandicae. Pars I. Genus Tendipes cum E,quatorial (3'partie). Annls Soc. ent. Fr. generibus finitimis. Tijdschr. Ent- 76: ll9- 92: 149-204. 216. Kieffer, J.J. 7923c. Nouvelle contribution A Langton, P.H., P.S. Cranston & P. Armitage. l'6tude des Chironomides de la 1988. The parthenogenetic of water Nouvelle-Zemble. Rept. scient. Results supply systems, Paratanytarsus grimmii Norw. Exped. Nova Zemlya 9: 3-11. (Schneider) (Diptera: Chironomidae). Bull. Kieffer, J.J. I923d. Chironomides nouveaux ou ent. Res. 78:. 3I7-328. peu connus de la r€gion paldarctique. Annls Latreille, P.A. 1810. Considdrations gdndrales Soc. scient. Brux. 42: 138-180. sur l'ordre naturel des animaux composant Kieffer, J.J. 1924a. Quelques Chironomides les classes des Crustaces, des Arachnides et nouveaux et remarquables du Nord de des Insectes; avec un tableau m6thodique de CHIRONOMIDAE DE LA REGION NEARCTIQUE 67

leurs genres, dispos6s en familles. Paris, 444 Loew, II. 1866. Diptera Americae pp- septentrionalis indigena. Centuria septima. Leathers, A.L. 1922. Chironomus braseniae Berl. ent. Z. 10:. l-54. new species (Dip., Chironomidae). Ent. Lundbeck, W. 1898. Diptera groenlandica. News 33: 8. Naturhist. for. Kj@benhavn, Vidensk. meddel. Lenz, F. L921. Chironomidenpuppen und lO: 236-314. -larven. Bestimmungstabellen. Dt. ent. Z. LundstrOm, C. 1910. Beitrdge zur Kenntnis der l92l: 148-162. Dipteren VI. Chironomidae. Acta Lenz, F. l94la. Die Metamorphose der Soc. Fauna Flora fenn. 33:. 7-47. Chironomidengattung Cryptochironomus. Lundstrdm, C. 1915. Resultats scientifiques de Zool. Anz.133:.29-41. I'Expddition Polaire Russe en 1900-1903, Lenz, F. 1941b. Die Jugenstadien der Sectio sous la direction du Baron E. Toll. Sect. E: Chironomariae (Tendipedini) connectentes Zoologie 2, 8- Diptera Nematocera aus den (Subf. Chironominae = Tendipedidae). arctischen Gegenden Sibiriens. [Akad. Nauk Zusammenfassung und Revision. Arch. S.S.S.R.l Imp. Akad. Nauk Petrograd, Zap., Hydrobiol. 38: 1-69. Fiz.-Mat. Otd. (Acad. Imp. des Sci. de Leonard, M.D. 1928. List of the insects of New Petsrograd, Me'm. Cl. Phys.-Math.) ser. 8, York with a list of the spiders and cerain 29: l-33. other allied groups. Mem. Cornell agric. Macquart, J. 1826. Insectes Diptdrds du Nord Exp. Stn. (1926) 101:1-1121. de la France. I. Tipulaires. Soc. d'Amateurs LeSage, L. & A.D. Harrison. 1980. Taxonomy Sci., Agr. Arts, Lille, Recueil Trav. of Cricotopus species (Diptera: 182311824: 59-224. Chironomidae) from Salem Creek, Ontario. Macquart, J. 1834. Historie Naturelle des Proc. ent. Soc. Ont. lI7:. 57-714. Insectes. tome 1. 2. Diptdres. 578 pp. Lindeberg, B. 1963. Taxonomy, biology and Macquart, J. 185-5. Diptdres exotiques biometry of Tanytarsus curticornis Kieff. and nouveaux ou peu connus. Libr. Encycl. T- brundini n. sp. (Dipt. Chironomidae). Roret. Suppl. 5: 35-1-56. Suom. hydnt. Aikak. 29: 118-1.30. Makarchenko, E.A. 1982. Khironomidy roda Lindeberg, B. 1967- Sibling species delimitation Protanypus Kieffer (Diptera, Chironomidae) in the Tanytarsus lestagei aggregate (Diptera, dal'nego Vostoka SSSR. Pp. 124-744. In: Chironomidae). Annls zool. Fenn. 4: 45-86. Biologiya presnovodnykh Zibotnykh dal'nego Lindeberg, B. 1970. Tanytarsini ((Diptera, vostoka. Akad. NAUK. SSSR. biol. pochv. Chironomidae) from northern Fennoscandia. Inst., Vladivostok. Annls zool. Fenn. 7: 303-312- Makarchenko, E.A. 1983. Arctodiamesa gen. Linnaeus, C. 1758. Systema naturae per regna nov. - Novyi rod khironomid podsemeistva tria naturae, secundum classes, ordines, Diamesinae (Diptera, Chironomidae). In: genera, species, cum caracteribus, Vsesoyuzneji simposium, Biologicheskie differentiis, synonymis, locis. Ed. 1.0. 1., 824 problemy severa, Magadan 2: 264-265 . pp' Holmiae (=Stockholm). Makarchenko, E.A. 1,986. Diamesa subletti sp. Linnaeus, C. I767- Systema naturae, per regna n., a new species of chironomid (Diptera, tria naturae, secundum classes, ordines, Chironomidae) from Canada. Aquat. Insects genera, species, cum caracteribus, 8:155-1-57. differentiis, synonymis, locis. Ed. 13, Malloch, J.R. 1915a. Four new North America revised. 1(2): -533-1327- Holmiae Diptera. Proc. biol. Soc. Wash. 28; 45-48. ( =Stockholm). Malloch, J.R. 1915b. The Chironomidae, or Lipina, N.N. 1939. Novye formy molodykh midges, of Illinois, with particular reference stadii Chironomidae podopitnykh ozer to the species occurring in the Illinois River. sapropelovoi stantsii v Zaluchie. Trudy Lab. Bull. Ill. St. Lab. nat. Hist. L0: 275-543. sapropel. Otlozh. 1: 89-107. Malloch, J.R. 1918. A new species of Loew, H. 1861a. Diptera. Americae Orthocladius (Chironomidae, Diptera). Bull. septentrionalis indigena. Centuria prima. Brooklyn ent. Soc. 13:. 42. Berl. ent. Z- 5: 307-359. Malloch, J.R. 1919. Part C. Diptera (excluding Loew, tI. 1861b. Diptera aliquot in insula the Tipulidae and Culicidae). The Diptera Cuba collecta. Wien. ent. Mschr. 5: 33-43. collected by the Canadian Arctic Expedition, Loew, II. 1864. Diptera Americae 1913-1918. Rep. Can. Arctic Expedition septentrionalis indigena. Centuria quinta. 1913-1918. Vol. III: Insects. pp. 34-90 Berl. ent. Z. 8: 49-104. Malloch, J.R. 1923. Insects, arachnids, and chitopods of the Pribilof Islands, Alaska. NEARCTIC CHIRONOMIDAE

Diptera (except Tipulidae, Rhyphidae and Chironomidae) from Japan. Konty0, Tokyo Calliphoridae). N. Am. Fauna 46: 170-227. 55: 459-476. Malloch, J.R. 19-14. The exploration of Nowicki, M.S. 1873. Beitrlige zur Kenntniss der Southampton Island, Hudson Bay, by George Dipterenfauna Galiziens. 35 pp. Krakau, Miksch Sutton, sponsored by Mr. John Poland. Bonner Semple 1929-1930. Part II. Oliveira, S.J. de. 1967. Novo genero de Zoologie. Section 4. Spiders and insects (in Chironomidae da Amazdnia (Insecta, part) of Southampton Island. III. Diptera). Atas Simp. Biota amazon. 5 Chironomidae, Sciaridae, Phoridae, (Zool.): 4r7 -479. Syrphidae, Piophilidae, Helomyzidae, Oliver, D.R. 1959. Some Diamesini Calliphoridae, Oestridae, and Tachinidae. 1. (Chironornidae) from the Nearctic and Mem. Carneg. Mus. L2, pt. II, sect. 4: 1,3-32. Palaearctic. Ent. Tidskr. 90:. 48-64. Mason, P.G. 1986. Four new species of the Oliver, D.R. 1971,. Description of Einfeldia Cryptochironomus fulvus (Johannsen) species synchrona n. sp. (Diptera: Chironomidae). complex (Diptera: Chironomidae). Can. Ent. 103: 1591-1595. Entomologica scand. L6: 399-473. Oliver, D.R. 1976. Chironomidae (Diptera) of Meigen, J.W. 1800. Nouvelle classification des Char Lake, Cornwallis Island, N.W.T., with mouches A deux ailes (Diptera L.) d'apres un descriptions of two new species. Can. Ent. plan tout nouveau. Paris, 40 pp. 108:10-53-1064. Meigen, J.W. 1803. Versuch einer neuen Oliver, D.R. 1.977. Bicinclus-group of the genus Gattungseinteitung der europAischen Cricotopus van der Wulp (Diptera: zweiflrigetigen Insekten. Magazin Insektenk. Chironomidae) in the Nearctic with a (Illiger) 2: 259-287. description of a new species. J. Fish. Res. Meigen, J.W. 1804. Klassifikazion und Bd Can. 34:.98-104. Beschreibung der europdischen zweiflrigeligen Oliver, D.R. 1981. Description of Euryhapsis Insecten (Diptera Linn.). Erster Band. Abt. new genus including three new species I., pp. I-I52. Abt. II, pp. 153-314. (Diptera: Chironomidae). Can. Ent. 113: Braunschweig. 7lt-122. Meigen, J.W. 1818. Systematische Beschreibung Oliver, D.R. 7982. Xylotopus, a new genus of der bekannten europ5ischen, zweiflUgeligen Orthocladiinae (Diptera: Chironomidae). Insekten. 1, xxxvi + 332 pp. Forstmann, Can. Ent. L1.4: 167-1,68. Aachen. Oliver, D.R. 1.984. Description of a new species Meigen, J.W. 1830. Systematische Beschreibung of Cricotopus van der Wulp (Diptera: der bekannten europdischen zweiflrigeligen Chironomidae) associated with Myriophyllum Insekten. 6: 1-401 pp. Schulz, Hamm. spicatum. Can. Ent. 116:. 1287-1292. Meigen, J.W. 1838. Systematische Beschreibung Oliver, D.R. & M.E. Dillon. 1988. Review of der bekannten europtiischen zweiflrigeligen Cricotopus (Diptera: Chironomidae) of the Inseckten. 7: 7-434. Schulz, Hamm. Nearctic zone with descriptions of two new Meunier, F. 1904. Monographie des species. Can. Ent. 120. 464-496. Cecidomyidae, des Sciaridae, des Oliver. D.R. & M.E. Roussel. 1983. Mycetophilidae et des Chironomidae de Redescription of Brillia Kieffer (Diptera: I'ambre de la Baltique. Annls. Soc. scient. Chironomidae) with descriptions of Nearctic Brux. 28: 1,2-274. species. Can. Ent. Il5:. 257-279. Michailova. P. & J. Fischer 1986. Chironomus Osten Sacken, C.R. 7877. Western Diptera: vancouveri sp. n. from Canada (Diptera, Descriptions of new genera and species of Chironomidae). Reichenbachia 23: 99-106. Diptera from the region west of the Mitchell, E.G. 1908. Descriptions of nine new Mississippi and especially from California. species of gnats. Jl N.Y. ent. Soc. t6: 7-I4. Bull. U.S. geol. geogr. Surv. Terr. 3: Murray, D.A. & E.J. Fitrkau. 1985. Hayesomyia 189-354. a new genus of Tanypodinae from the Packard, A.S. 1869. On insects inhabiting salt Holarctic (Diptera: Chironomidae). Spixiana water. Proc. Essex Inst. 6: 41-51 Suppl. (Muench.) lI: 195-207. Pagast, F. 1931. Chironomiden aus der Muttkowski. R.A. 191-5. New insect life Bodenfauna des Usma-Sees in Kurland. histories. L Bull. Wis. nat. Hist. Soc. 13: Fofia zool. hydrobiol. 3: L99-248. L09-r22. Pagast, F. 1947. Systematik und Verbreitung Nishida, H. 1987. Pseudocricotopus, a new der um die Gattung Diamesa gruppierten subgenus of the genus Cricotopus (Diptera, CHIRONOMIDAE DE LA REGION NEARCTIQI]E

Chironomiden. Arch. Hydrobiol. 41: 435- Chironomidengattung Goeldichironomus 596. Fittkau (Diptera, Insecta). Stud. neotropical Palm6n, E. 1960. Paratanytarsus-Arten (Dipt., Fauna 9: 95-122. Chironomidae) aus dem -mesohalinen und Reiss, F. 1982. Hyporhygma n. gen. und oligohalinen Brackwasser der Finnischen Stelechomyia n. gen. aus Nordamerika Meerbusens. Suom. hyOnt. Aikak. 26: 280- (Diptera, Chironomidae). Spixiana 29r. (Muench.) 5: 289-302. Panzer, G.W.F. 1813. Faunae insectorum Reiss, F. 1985. Die panamarikanisch verbreitete germanicae initiae oder Deutschlands Tanytarsini-Gattung Skutzia gen. nov. Insekten. H. 109, 24 pp.Nurnberg. (Diptera, Chironomidae). Spixiana Suppl. Petch, C.E. & J.B. Maltais. 1932. A (Muench.) II: 173-178. preliminary list of the insects of the Province Reiss, F. 1988. Irmakia, ein neues Subgenus of Quebec. Part II. Diptera, by A.F. Winn von Demicryptochironomus Lenz, 1941, mit & G. Beauliesu. Revised and supplemented. der Beschreibung von vier neuen Arten Que. Soc. Protect. Plants, Ann. Rept. 23124: (Diptera, Chironomidae). Spixiana 1-100 (Muench.) 1l: t-12. Philippi, R.A. 1865. Aufztihlung der Reiss, F. & E.J. Fittkau. I971. Taxonomie und chilenischen Dipteren. Verh. zool.-bot. Ges. 6kologie europiiisch verbreiteter Wien 15: 595-782. Tanytarsus -Arten (Chironomidae, Diptera). Picado, C. 1913. Les bromeliac6es epiphytes, Arch. Hydrobiol. Suppl. 40: 75-200. consid6r6es comme une milieu biologique. Reiss, F. & L. Siiwedal. 1981. Keys to males Bulf. scient. Fr. Belg. 47: 2I5-360. and pupae of the Palaearctic (excl. Japan) Pinder, L.V.C. 1983. The larvae of Paratanytarsas Thienemann & Bause, 1913, Chironomidae (Diptera) of the Holarctic n. comb., with descriptions of three new region-Introduction. Pp. 7 -10. In: species (Diptera: Chironomidae). Wiederholm, T. (Ed.). Chironomidae of the Entomologica scand. Suppl. 15: 73-104. Holarctic Region. Keys and diagnoses. Part 1 Reiss. F. & J.E. Sublette. 1985. Beardius new Larvae. Entomologica scand. Suppl. 19: genus with notes on additional Pan-American -t-437. taxa. (Diptera, Chironomidae). Spixiana Potthast, A. 191-5. Uber die Metamorphose der Suppl. (Muench.) lt 179-193. Orthocladius-Gruppe. Ein Beitrag zur Remy, P. 1.928. Arthropodes terrestres rdcoltds Kenntnis der Chironomiden. Arch. au Groenland au cours de la croisidre du Hydrobiol. Suppl. 2: 243-376- "Pouquoi-Pas?" en 1926 (1' liste). Bull. Rasmussen, J.B. 1984. Chironomus bimens. Soc. linn. Lyon 7: 51--53. (Camptochironomus) vockerothi n. sp. Rempel, J.G. 1937a. Notes on the genus (Diptera: Chironomidae) from Alberta, Chasmatonotas with descriptions of three Canada. Can. Ent. 716: t643-1646. new species (Diptera: Chironomidae). Can. Reiss, F. t969a. Die neue, europdisch Ent. 69:. 250-255. verbreitete Chironomidengattung Parapsectra Rempel, J.G. t937b. A new species of the mit einem brachypteren Artvertreter aus subgenus Krib ioxenus (Diptera, Mooren (Diptera). Arcb. Hydrobiol. 66: Chironomidae). Can. Ent. 69:. 274-275. r92-2tr. Rempel, J.G. t939. Neue Chironomiden aus Reiss, F. 1969b. Krenopsecta fallax gen- n. sp. Nordostbrasilien. Zool. Anz- 127: 209-216. n. (Diptera, Chironomidae) aus den Alpen Risso, A. 1816. Historie Naturelle des und Pyrenden. Annls zool. Fenn. 6: 435-442. Crustaces des environs de Nice. A La Reiss, F. L969c. Revision der Gattung Libraie Greque-Latin-Allemande, Paris: 1- Micropsectra Kieff., 1.909 (Diptera, 175 Chironomidae). 1. Die attenuata-Gruppe Roback, S.S. 19-55. The Tendipedid Fauna of a der Gattung Micropsectra. Beschreibung 5 Massachusetts cold spring (Diptera: neuer Arten aus Mitteleuropa und Tendipedidae). Notu. Nat. 270: l-8. Nordafrika. Dt. ent. Z. 16:. 431-449. Roback, S.S. 1957a. The immature tendipedids Reiss, F. 1972. Die Tanytarsini (Chironomidae, of the Philadelphia area. Monogr. Acad. Diptera) Stidchiles und Westpatagoniens. nat. Sci. Philad. 9: I-152 Mit Hinweisen auf die Tanytarsini-Fauna der Roback, S.S. I957b. Some Tendipedidae from Neotropis. Stud. neotropical Fauna 7: 49-94. Utah. Proc. Acad. nat. Sci. Philad. 109: 1- Reiss, F. F. I974. Die in stehenden GewAssern 24. der Neotropis verbreitete NEARCTIC CHIRONOMIDAE

Roback, S.S. 7959a. The subgenus Ablabesmyia (Diptera: Chironomidae: Tanypodinae). of Pentaneura (Dipteia: Tendipedidae: Proc. Acad. nat. Sci. Philad. 135: 254-260. Pelopiinae). Trans. Am. ent. Soc. 85: Roback, S.S. 1984. Tanypodinae (Diptera: 113-13-5. Chironomidae) from Afognak and Kodiak Roback, S.S. 1959b. Some Tendipedidae from Islands, Alaska. Proc. Acad. nat. Sci. Philad. Montana. Notul. Nat. 315: 1-4. 136: 12-23. Roback, S.S. 1960. A new Calopsectra from Roback, S.S. 1985. The immature chironomids Kansas (Diptera: Tendipedidae: of the eastern United States VI. Calopsectrini). Notul. Nat. 328: 1-4. Pentaneurini. Genus Ablabesmyia. Proc. Roback, S.S. 1,963a. The genus Acad. nat. Sci. Philad. 137:. 153-212. Xe no c hir onomus (Diptera: Tendipedidae) Roback, S.S. 1986a. The immature chironomids Kieffer, taxonomy and immature stages. of the eastern United States VIII. Trans. Am. ent. Soc. 88: 235-245. Pentaneurini - Genus Nilotanypus, with the Roback, S.S. 1963b. New neotropical description of a new species from Kansas. Coelotanypus (Diptera, Tendipedidae, Proc. Acad. nat. Sci. Philad. I38: 443-465. Pelopiinae). Ent. News 74: 169-7'76. Roback, S.S. 1986b. Reomyia a new genus of Roback, S.S. 1966. A new Procladius species Tanypodinae-Pentaneurini. Spixiana with description of the immature stages (Muench.) 9: 283-284. (Dipt.: Chironomidae). Ent. News 77: Roback, S.S. & R.P. Rutter. 7988. Denopelopia 777 -184. atria, a new genus and species of Roback, S.S. I97 1. The adults of the subfamily Pentaneurini (Diptera: Chironomidae: Tanypodinae (= Pclopiinae) in North Tanypodinae) from Florida. Spixiana Suppl. America (Diptera: Chironomidae). Monogr. (Muench.) 14: I17 -12'7. Acad. nat. Sci. Philad. 17: 1-410 pp. Rossaro, B. 1979. Stilocladius montanus n. gen. Roback, S.S. 1975. A new subgenus and n. sp.: Descrizione di un nuovo genere e di species of the genus Tanytarsus una nuova specie di Orthocladiinae delle (Chironomidae: Chironominae: Tanytarsini). Alpi Italiane (Diptera, Chironomidae). Boll. Proc. Acad. nat. Sci. Philad. 727:7I-80. Mus. civ. Stor. nat. Verona 6: 347-352. Roback, S.S. 1,978a. The immature chironomids Ryser, H.M., W. WUlker & A. Scholl. 198-5. of the eastern United States III. Revision der Gattung Chironomus Meigen Tanypodinae-Anatopyniini, Macropelopiini (Diptera) l0- Lobochironomus n. subg. (C. and Natarsiini. Proc. Acad. nat. Sci. Philad. montuosus n. sp., C. storai Goetgh., C. I29: I5I-202. mendax Stori). Revue suisse Zool. 92: 385- Roback, S.S. 1978b. New name for Brundinia 404. Roback nec Brundinia Tottenham. Ent Sether, O.A. 1968. Chironomids of the Finse News 89: 141. Area, Norway, with special reference to their Roback, S.S. 19'79. Hudsonimyia karelena, a distribution in a glacier brook. Arch. new genus and species of Tanypodinae, Hydrobiol. 64: 426-483. Pentaneurini. Proc. Acad. nat. Sci. Philad. Saether, O.A. 1969. Some Nearctic 131:1-8. Podonominae, Diamesinae, and Roback, S.S. 1980a. The immature chironomids Orthocladiinae (Diptera: Chironomidae). of the eastern United States IV. Bull. Fish. Res. Bd Can. 170: 1-154. Tanypodinae-Procladiini. Proc. Acad. nat. Sether. O.A. I971. Four new and unusual Sci. Philad. 132: I-63. Chironomidae (Diptera). Can. Ent. 103: Roback, S.S. 1980b. New name for Anceus r799-1827. Roback nec lnceus Risso. Ent. News 91: Saether, O.A. 1.973a. Four species of -1L- Bryophaenacladius Thien, with notes on Roback, S.S. 1981. The immature chironomids other Orthocladiinae (Diptera: of the eastern United States V. Chironomidae). Can. Ent. 10-5: -51-60. Pentaneurini-Thienemannimyia group. Proc. S Acad. nat. Sci. Philad. 133: 73-128. three new species of Monodiamesa Kieffer, Roback, S.S. 1,982. The Tanypodinae (Diptera: with keys to Nearctic and Palaearctic species Chironomidae) of Australia II. Proc. Acad. of the genus (Diptera: Chironomidae). J. nat. Sci. Philad. 1,34: 80-112. Fish. Res. Bd Can. 30:. 665-679. Roback, S.S. 1983. Krenopelopia hudsoni: a Saether, O.A. 1.975a. Nearctic and Palaearctic new species from the eastern United States Heterotrissocladius (Diptera: Chironomidae). Bull. Fish. Res. Bd Can. 193: 67 pp. CHIRONOMIDAE DE LA REGION NEARCTIQUE 7l

Sather, O.A. 1975b. Two new species of (Diptera: Chironomidae) from South Heterotanytarsas Spflrck, with keys to Carolina. Aquat. Insects 3: 193-198. Nearctic and Palaearctic males and pupae of S the genus (Diptera: Chironomidae). J. Fish. Chironomidae) from SE U.S.A., with Res. Bd Can. 32: 259-270. descriptions of Plhudsonia, Unniella and Sether, O.A. 1975c. Two new species of Platysmittia n. genera and Atelopodella n. Protanypus Kieffer, with keys to Nearctic and subgen. Entomologica scand. 13: 46-5-510. Palaearctic species of the genus (Diptera: Seether, O.A. 1983a. Oschia dorsenna n. gen., Chironomidae). J. Fish. Res. Bd Can- 32: n. sp. and Saetheria hirta n.sp., two new" 367-388. members of the Harnischia comPlex Sether, O.A. 1975d. Twelve new species of (Diptera: Chironomidae). Entomologica Limnophyes Eaton, with keys to Nearctic scand. 14: 395-404. males of the genus (Diptera: Chironomidae). Srther, O.A. 1983b. A review of Holarctic Can. Ent. 107: 1.029-1056. Gymnometriocnemus Goetghebuer, 1932, with Sather, O.A. I976. Revision of Hydrobaenus, the description of Raphidocladius subgen. n. Trissocladius, Zalutschia, Paratrissocladius, and Sublettiella gen. n. (Diptera: and some related genera (Diptera: Chironomidae). Aquat. Insects 5: 209-226. Chironomidae). Bull. Fish. Res. Bd Can. Sather, O.A. 1983c. Three new species of 195:287 pp. Lopescladius Oliveira, 1967 (syn. "Cordites" Sether. O.A. 1977a. Habrobaenus hudsoni n. Brundin, 19-56, n. syn.), with a phylogeny of gen., n. sp. and the immatures of Baeoctenus the Parakiefferiella grotp. Mem. Am. ent. bicolor Saether (Diptera: Chironomidae). J. Soc. 34: 279-298. Fish. Res. Bd Can. 34: 2354-2361. Srther, O.A. 198-5a. Male and female imagines Saether, O.A. 1977b. Taxonomic studies on of Platysmittia bilyji sp. n. (Diptera: Chironomidae: Nanocladius, Chironomidae) from Manitoba, Canada. Pseudochironomus, and the Harnischia Entomologica scand. 15: 527-531. complex. Bull. Fish. Res. Bd Can. 196:. 143 Saether, O.A. 1985b. Heleniella parva sp. n. pp. (Diptera: Chironomidae) from South Sather, O.A. 19'79. New name for Beckiella Carolina and Tennessee, U.S.A. Saether, 1977 (Diptera: Chironomidae) nec Entomologica scand. 1-5: 532-535. Beckiella Grandjean, 1964 (Acari: Oribatei). Sether, O.A. 198-5c. Apometriocnemus Entomologica scand. 10: 31-5. fontinalis n. gen., n. sp. (Diptera: Srether, O.A. 1980a. New name for Oliveria Chironomidae) from Tennessee, U.S.A. Saether, 1976 (Diptera: Chironomidae) nec Entomologica scand. 15: -536-539. Oliveria Sutherland, 196-5 (+Cnidaria: Sather, O.A. 1985d. Redefinition and review of Anthozoa) with a first record for the Thienemannia Kieffer, 1909 (Diptera: European continent. Entomologica scand. Chironomidae), with the description of Z. 11: 399-400. pilinucha sp. n. Aquat. Insects 7: III-I3I. Sether. O.A. 1980b. The females and Srether, O.A. 198-5e. A review of Odontomesa immatures of Paracricofopus Thienemann Pagast, 1947 (Diptera, Chironomidae, and Harnisch, 1932, with the description of Prodiamesinae). Spixiana Suppl. (Muench.) a new species (Diptera: Chironomidae). tt; 15-29. Aquat. Insects 2: 1,29-145. Srether, O.A. 198-5f. Trichochilus, a new genus Sather, O.A. 1981a. Doncricotopus bicaudatus of Orthocladiinae (Diptera, Chironomidae). n. gen., n. sp. (Diptera: Chironomidae, Spixiana Suppl. (Muench.) 11: 31-36. Orthocladiinae) from the Northwest Sather, O.A. 198-59. The imagines of Territories, Canada. Entomologica scand. Mesosmittia Brundin, 1956, with description 12: 223-229. of seven new species (Diptera, Saether, O.A. 1981b. Orthocladiinae Chironomidae). Spixiana Suppl. (Muench.) (Chironomidae: Diptera) from the British ll:37-54. West Indies with descriptions of Saether, O.A. 198-5h. Diplosmittia carinata spec. Antillocladius n. gen., Lipurometriocnemus n. nov. from Michigan (Diptera, Chironomidae). gen., and Diplosmittia n. gen. Entomologica Spixiana Suppl. (Muench.) 11: -5-5-57. scand. Suppl. 16: l-46. Srether, O.A. 1985i. A review of the genus Sether, O.A. 1981c. Compteromesa oconeensis Rheocricotopas Thienemann & Harnisch, gen. n., sp. n., a new Prodiamesinae 1932, with the description of three new 72 NEARCTIC CHIRONOMIDAE

species (Diptera, Chironomidae). Spixiana mitgebrachten Dipteren. Verh. zool.-bot. Suppl. (Muench.) 1t: -59-108. Ges. Wien 76: 927-934. Sether, O.A. 1989a. Metriocnemu.s van der Schlee, D. 1968. Vergleichende Wulp: a new species and a revison of species Merkmalsanalyse zur Morphologie und described by Meigen, Zetterstedt, Staeger, Phylogenie der Corynoneura-Gruppe Holmgren, Lundstrdm and Strenzke (Diptera, Chironomidae). Zugleich eine (Diptera: Chironomidae). Entomologica allgemeine Morphologie der scand. 19: 393-430. Chironomiden-Imago ( ). Stuttg. Beitr. Sether, O.A. 1989b. Two new species of Naturk. 180:1-150. Hydrobaenus Fries from Massachusetts, Schneider, A. 1885. Chironomus grimmii und U.S.A., and Japan (Diptera: Chironomidae). seine Parthenogenese. Zool. Beitr. 1: 301- Entomologica scand. 20: -5-5-63. 302. Sether, O.A. & O.A. Schnell. 1988. Two new Schnell, O.A. & O.A. Srther. 1988. species of the Rheocricotopus (R.) effusus Vivacricotoplr.r, a new genus of group (Diptera, Chironomidae). Spixiana Orthocladiinae from Norway. Spixiana Suppl. (Muench.) 14:. 65-74. Suppl. (Muench.) 14:49-55 - Sather, O.A. & J.E. Sublette. 1983. A review Schrank. F. von P. 1803. Fauna Boica. of the genera Doithrix n. gen., Durchgedachte Geschichte der in Baiern Georthocladlas Strenzke, Parachaetocladius einheimischen und zahmen Thiere. 3: l-272. Wtilker and Pseudorthocladius Goetghebuer Landshut. (Diptera: Chironomidae, Orthocladiinae). Serra-Tosio, B. 1968. Taxonomie Entomologica scand. Suppl. 20: 1-100. phylogenetique des Diamesini: les genres Sether, O.A. & E. Willassen. 1988. A review Potthastia Kieffer, Sympotthastia Pagast, of Lappodiamesa Serra Tosio, with the Parapotthastia n. g. et Lappodiamesa n. g. description of L. boltoni spec. nov. from (Diptera, Chironomidae). Trav. Lab. Ohio, USA. Spixiana Suppl. (Muench.) 14: Hydrobiol. Piscic. Univ. Grenoble 59-60: 75-84. t77 -t64. Santos Abreu, E. 1918. Ensayo de una Serra-Tosio, B. I971. Deux Diamesini Monografia de los Tendipedidos de las Islas nouveaux d'Espagne (Diptera, Canarias. Mems R. Acad. Cienc. Artes Chironomidae). Trav. Lab. Hydrobiol. Barcelona 74: 159-326. Piscic. Univ. Grenoble 62: 147-L67. Saunders, L.G. 1928. Some marine insects of Shilova, A.I. 1955. Nekotorye massovya vidy the Pacific coast of Canada. Ann. ent. Soc. tendipedid (Diptera, Tendipedidae) basseina Am.2t: 52t-545. Amudarja. Ent. Obozr. 34:. 313-322. Sfiwedaf , L. 1979. Micropsectra brundlni n. sp. Shilova, A.I. 1957. Palaearktischeskie vidy (Diptera: Chironomidae) and geographical podroda Camptochironomus Kieff. roda variation in the Micropsectra insignilobus Tendipes Mg. (Diptera, Tendipedidae). Ent. aggregate. Entomologica scand. 10: 133-138. Obozr. 36:. 224-230. Sdwedal, L. 1981. Description of Micropsectra Shilova, A.I. 7961. Noryi rod i vid tendipedid tori n. sp. from Greenland, with notes on the (Diptera, Tendipedidae) Lipiniella Shilova, recurvata-group (Diptera: Chironomidae). gen. n. Byull. Inst. Biol. Vodokhran. 11: Entomologica scand. 12: 27-30. 19-23. Say, T. 1823. Descriptions of dipterous insects Skuse, F.A.A. 1889. Diptera of Australia. Part of the United States. J. Acad. nat. Sci. V1. Chironominae. Proc. Linn. Soc. Philad. 3: 9-54. N.S.W.-The 4: 215-3ll- Say, T. t829-I930. Descriptions of North Soponis, A.R. L977. A revision of the Nearctic American dipterous insects. J. Acad. nat. species of Orthocladius (Orthocladius) van Sci. Philad. 6: 149-t78, (1929); 183-188, der Wulp (Diptera: Chironomidae). Mem. (1830). ent. Soc. Can. lO2: 187 pp. Schiner, J.R. 1856. Ammerkungen zu dem Soponis, A.R. t979. Zalutschia briani n. sp. Bande V, Pag. 13 diesser Verhandlungen from Florida (Diptera: Chironomidae). abgedruckten Aufsdtze Frauenfeld's: Beitrag Entomologica scand. Suppl. l0: 125-13I. zur Insektengeschichte. Verh. zool.-bot. Soponis, A.R. 1980. Pseudorthocladius Ges. Wien 6: 215-224. macrostomus, a new species of chironomid Schiner, J.R. 1865. Bericht tiber die von (Diptera) with a long proboscis. Fla Ent. Weltumseglungsreise der k. Fregatte Novara 63: 485-490. CHIRONOMIDAE DE LA REGION NEARCTIQUE

Soponis, A.R. 1983. Orthocladius and Diamesinae. Proc. U.S. natn. Mus. 115: (Orthocladius) ferringtoni n. sp., from Kansas 85- 136. (Diptera: Chironomidae). J. Kans. ent. Soc. Sublette, J.E. L964b- Chironomidae (Diptera) 56:571-571. of Louisiana I. Systematics and immature Splirck, R. L922-7923. Beitrlige zur Kenntnis stages of some lentic chironomids of der Chironomidenmetamorphose I-IV. Ent. west-central Louisiana. Tulane Stud. Zool. Meddr. L4: 32-48 11922); 49-109, 11923). 11:109-150. Sptirck, R. & A. Thienemann. 1924. Sublette, J.E. 1966. Type specimens of "Metriocnemus" ampullaceus vat. austriacus. Chironomidae (Diptera) in the U.S. National Verh. int. Verein. theor. angew. Limnol. 2: Museum. J. Kans. ent. Soc. 39: 580-607. 222-223. Sublette, J.E. 1,967a. Type specimens of Staeger, C. 1839. Systematisk fortegnelse over Chironomidae (Diptera) in the Canadian de i Danmark hidtil fundne Diptera. National Collections, Ottawa. J. Kans. ent. Naturhist. Tidsskr. (1838-1839) 2: 549-600. Soc. 40: 290-331. Staeger, C. 1845. Gr{nlands antliater. Sublette, J.E. 1,967b. Type specimens of Naturhist. Tidsskr. (1844-1845) ser. 2,7: 346- Chironomidae (Diptera) in the Cornell 369. University Collection. J. Kans. ent. Soc. 40: Steffan, A.W. 1965. Plecopteracoluthus downesi 477 -564. gen. et sp. nov. (Diptera: Chironomidae), a Sublette, J.E. l9'70. Type specimens of species whose larvae live phoretically on Chironomidae (Dipt.) in the Illinois Natural larvae of Plecoptera. Can. Ent. 97: History Survey Collection, IJrbana. J. Kans. t323-t344. ent. Soc. 43:. 44-95. Steiner, B. t983. Paracricotopus mozleyi n. sp. sublette, J.E. & M.s. Sublette. 1965. Family from Georgia (Diptera: Chironomidae). Chironomidae. Pp- 142-181.. In; Stone, A., Mem. Am. ent. Soc. 34: 329-335. C.W. Sabrosky, W.W. Wirth, R.H. Foote, & Steiner, J.W. & J.L. Hulbert. 1982. Nimbocera J.R. Coulson (Eds). A catalogue of the pinderi, a new species (Diptera: Diptera of America North of Mexico. U.S. Chironomidae) from the Southeastern United Dep. Agric. Handb. No. 276: I-1696. States. Fla Ent. 65: 228-233. Sublette, J.E. & M.S. Sublette. I971. The Stone, A. & Wirth, W.W. 1947. On the marine Orthocladiinae (Chironomidae, Dipt.) of midges of the genus Clunio Haliday California. I. The Cricotopus infuscatus (Diptera, Tendipedidae). Proc. ent. Soc. group. Ent. News 82:. 85-102. Wash. 49: 2OL-224. Sublette. J.E. & M.S. Sublette. I974. A review Stori, R. 1936. FOr faunan nya nematocerer. of the genus Chironomus (Diptera, Notul. ent. 16: 124-125. Chironomidae) V. The maturus complex. - Stor6, R. 1939. Mitteilungen riber die Stud. nat. Sci. (Portales, NM) 1, 8: 41 pp. Nematoceren Finnlands II. Notul. ent. 19: Sublette, J.E. & M.S. Sublette. L979. Part II. 16-30. A synopsis of the Chironomidae of New Strenzke, K. 194I- Terrestrische Chironomiden Mexico. Pp. 54-128. In: Sublette, J.E. (Ed.). X. Georthocladius luteicornls Goetgh. (mit Evaulation of long term effects of thermal einem Beitrag von Dr. M. Goetghebuer). effluents on stream biota. Final Rep. N. Zool. Anz. 132: 177 -185. Mex. Energy Inst. 32: 1-156. Strenzke, K. 1950. Systematik, Morphologie Sublette, J.E. & M.S. Sublette. 1983. Asheum, und Okologie der terrestrischen a replacement name in Cbironomidae Chironomiden. Arch. Hydrobiol. Suppl. 18: (Diptera). Ent. News 94. 34. 207-4r4. Sublette, J.E. & W.W. Wirth- 1972. New Strenzke, K. & H. Remmert. 1,957. genera and species of West Indian Terrestrische Chironomiden. XVII. Chironomidae (Diptera). Fla Ent. -55: 1-17. Thal ass osmittia thalas s ophila (Bequ. u. Srilc, K. & J- ZavreL 7924. O epoikickych a Goetgh.). Kieler Meeresforsch. I3:, 263-273. parasitickych larvdch Chironomidu. lUber Sublette, J.E. 1960. Chironomid midges of epoikische und parasitische California. I. Chironominae, exclusive of Chironomidenlarven). Acta Soc. Sci. nat. Tanytarsini (=Calopsectrini). Proc. U.S. moravo-siles. 1: 3-53-391. natn. Mus. 112: L97-226. Thienemann, A. t921. Die Metamorphose der Sublette, J.E. 1964a. Chironomid midges of Chironomidengattungen Camptocladius, California. II. Tanypodinae, Podonominae, Dyscamptocladius und Phaenocladias, mit Bemerkungen tiber die Artdifferenzierung NEARCTIC CHIRONOMIDAE

bei den Chironomiden riberhaupt. Arch. Tokunaga, M. 7937. Chironomidae from Japan I{ydrobiol. Suppl. 2: 809-850. (Diptera), IX Tanypodinae and Diamesinae. Thienemann, A. 1,926. I{ydrobiologische Philipp. J. Sci. 57: 2l-65. Untersuchungen an den kalten Quellen und Tokunaga, M. 1939. Chironomidae from Japan Bdcken der Halbinsel Jasmund auf Rtigen. (Diptera). XI. New or little-known midges, Arch. Hydrobiol. 17: 221-336. with special reference to the metamorphosis Thienemann, A. 1934. Chironomiden- of torrential species. Philipp. J. Sci. 69: Metamorphosen. VIII. " Phaenocladius" 297 -345. Encycl. ent. Diptera 7: 29-46. Tottenham, C.E. 1949. New generic names in Thienemann,A. 1935. Chironomiden- Staphylinidae (Coeloptera). Proc. R. ent. Metamorphosen. X. "Orthocladius Soc. Lond. 18: 78. -Dactylocladizs" (Dipt.). Stettin. ent. Ztg Townes, H.K. 1945. The Nearctic species of 96: 2OI-224. Tendipedini (Diptera, Tendipedidae Tbienemann, A. 1936a. Alpine Chironomiden. (=Chironomidae)). Am. Midl. Nat. 34: Ergebnisse von lJntersuchungen in der t-206. Gegend von Garmisch-Partenkirchen, Verrall, G.H. 79t2. Another hundred new Oberbayern. Arch. Hydrobiol. 30: L67-252. British species of Diptera. Entomologist's Thienemann, A. 1936b. Chironomiden- mon. Mag. 48: 20-27, 56-59, 1.44-147; Metamorphosen. XI. Gattung Eukiefferiella. 190-197. Stettin. ent. Ztg 9'7: 43-65. Verrill, A.E. 1873. Report upon the Thienemann, A. 7939. Dritter Beitrag zur invertebrate of Vineyard Sound and Kenntnis der Podonominae (Dipt. the adjacent waters, with an account of the Chironomidae). (Chironomiden aus physical characters of the region. Pp.295- Lappland VI). Zool. Anz. 728: 761-176. 178. In: Baird, S.F., Report on the Thienemann, A. 7947. Lappliindische condition of the sea fisheries of the south Chironomiden und ihre WohngewAsser. coast of New England in 1871 and 7872. (Ergebnisse von lJntersuchungen im U.S. Comn. Fish Fisheries. 1: 1-852. Abiskogebiet in Schwedisch-Lappland). Walker, F. 1848. List of the specimens of Arch. Hydrobiol. Suppl. 77:- 7-253. dipterous insects in the collection of the Thienemann, A. 7942. Trichocladius-Arten aus British Museum. Part 1: l-229. London. dem Lunzer Seen. (Chironomiden aus dem Walker, F. 1856a. Insecta Britannica: Diptera. Lunzer Seengebiet. V.). Arch. Hydrobiol. 3: l-352. London. 39:294-315. Walker, F. 1856b. Diptera. 1. Pp. 415-474. Thienemann, A. & O. I{arnisch. 1932. In. Saunders, W.W. (Ed.). 1850-1869. Chironomiden-Metamorphosen. IV. Die Insecta Saundersiana: Or characters of Gattung Cricotopus v.d.W. Zool. Anz. 99: undescribed insects in the collection of 13-5- 143. William Wilson Saunders, Esq., F.R.S., Thienemann, A. & J.J. Kieffer. 1916. F.L.S., &c. 4 vols. London. Schwedische Chironomiden. Arch. Walley, G.S. 1925. New Canadian llydrobiol. Planktonk. Suppl. 2: 483-554. Chironomidae of the genus Tanypus (Dipt.). Thienemann, A. & F. Kriiger. 1,937. Can. Ent. 57: 271-278. "Orthocladius" abiskoensis Edwards und Walley, G.S. 1926a. New Canadian rubicundus Mg., zwei "Puppen-Species" der Chironomidae. Can. Ent. 58: 64-65. Chironomiden. (Chironomiden aus Lappland Walley, G.S. 1926b. Four new Canadian II.) Zool. Anz. 1I7: 257-267. Chironomidae. Can. Ent. 58: 205-207. Thienemann, A. & F. Krriger. 1939- Walley, G.S. 7928a. A new species of Terrestrische Chironomiden II. Zool. Anz. Cricotopus with a key to the genus (Dipt., 127: 246-258. Chironomidae). Can. Ent. 6O: 2l-22. Tilbury, M.R. 1913. Notes on the feeding and Walley, G.S. 1928b. The genus Tanypus in rearing of the midge, Chironomus cayugae Canada, with a key to the North American Johannsen. Jl N.Y. ent. Soc. 21: 305-308. species (Diptera, Chironomidae). Ann. ent. Tokunaga, M. 1933. Chironomidae from Japan Soc. Am. 2L: 581-593. (Diptera). I. Clunioninae. Philipp. J. Sci. Webb, D.W. 1969. New species of chironomids 5l: 87 -99. from Costello Lake, Ontario (Diptera: Tokunaga, M. 1936. Chironomidae from Japan Chironomidae). J. Kans. ent. Soc. 42: (Diptera), VI Diamesinae. Philipp. J. Sci. 91-108. 59: 526-552. CHIRONOMIDAE DE LA REGION NEARCNQUE

Wiedemann, C.R.W. 1817. Neue Zweiflrigler und Verbreitung. Arch. Hydrobiol. Suppl. (Diptera Linn.) aus der Gegend um Kiel. 24: l-66. Wiedemanns zool. Mag. 1: 61-86. WLilker, W. 1959. Drei neue Wiedemann, C.R.W. 1828. Aussereuropiiische Chironomiden-Arten (Dipt.) und ihre zweifltigelige Insekten. 1 lIamm, 32 + 608. Bedeutung fr-ir das Konvergenzproblem bei Wiederholm, T. I975. Description of Imagines und Puppen. Arch. Hydrobiol. Protanypus saetheri n. sp. from Alaska Suppl.25: 44-64. (Diptera: Chironomidae). Entomologica Wtilker W.F. & M.G. Butler. 1983. scand. 6: 224-228. Karyosystematics and morphology of Wiederholm, T.(Ed.) 1983. Chironomidae of Northern Chironomus (Diptera: the Holarctic Region. Keys and diagnoses. Chironomidae): Freshwater species with Part 1 Larvae. Entomologica scand. larvae of tbe salinarius-type. Entomologica Suppl. -19:- l-437. scand. 14: 127-1.36. Wiederholm, T.(Ed.) 1986. Chironomidae of Wulp, F.M. van der. 1858. Beschrijving van the Holarctic Region. Keys and diagnoses. eenige nieuwe of twijfelachtige soorten van Part 2 Pupae. E,ntomologica scand. Diptera uit de familie der Nemocera. Suppl. -28: l-482. Tijdschr. Ent. 2: 159-185. Wiederholm, T.(Ed.) 1989. Chironomidae of Wulp, F.M. van der. 1861. Over Nieuwe the Holarctic Region. Keys and diagnoses. soorten van Diptera. Tijdschr. Ent. 4: 1-16. Part 3 - Adult males. Entomologica scand. Wutp, F.M. van der. 1867. Eenige Noord- Suppl. 34: l-532. Americaansche Diptera. Tijdschr. Ent. 12; Willassen, E. 1985. A review of Diamesa davisi t25-764. Edwards and the davisi group (Diptera, Wulp, F.M. van der. 1874. Dipterologische Chironomidae). Spixiana Suppl. (Muench.) aanteekneningen. Tijdschr. Ent. I7: 1.04- 17:109-137. 148. Williston, S.W. 1896. On the Diptera of St. Zavrel, I. L937. Eine neue Trissocladius art. Vincent (West Indies). Trans. Roy. Spisy vyddv. prir. Fak. Masaryk. Univ. 239:. Entomol. Soc. London. III: 253-446. 1.-1.2. Winnertz, J. 1846. Beschreibung einiger neuer Zavrel, J. 1941. Chironomidarum larvae et Gattungen aus der Ordnung der Zweiflrigler. nymphae llI. ("Pseudokiefferiella"). Ent. Stettin. ent. Ztg 7: ll-20. Listy 4: 9-14. Winnertz, J. 1,852. Dipterologisches. Zavrel, J. & F. Pagast. 193-5. Zwei neue Stettin. ent. ZtE i3: 49-58. Orthocladiinen-Arten aus Hoher Tatra. Cas. Wirth, W.W. 1949. A revision of the clunionine csl. Spol. ent.32: 156-160. midges with descriptions of a new genus and Zetterstedt, J.W. 1838. Diptera. Insecta four new species (Diptera: Tendipedidae). Lapponica 3: 477-868. Leipzig. Univ. Calif. Publs Ent. 8: 151-182. Zetterstedt, J.W. 18-50. Diptera Scandinaviae. Wirth, W.W. t957. The species of Cricotopus Diposita et descripta. 9: 3367-3710. midges living in the blue-green alga Nostoc Zetterstedt, J.W. 1860. Diptera Scandinaviae. in California (Diptera: Tendipedidae). Diposita et descripta. 14: 6191-6609. Pan-Pacif. Ent. 33: 12I-126. Wirth, W.W. & J.E. Sublette. t970. A review of the Podominae of North America with descriptions of three new species of Trichotanypus (Diptera: Chironomidae). J. Kans. ent. Soc. 43: 33-5-354. Wrubleski, D.A. & S.S. Roback. 1987. Two species of Procladius (Diptera: Chironomidae) from a northern prairie marsh: descriptions, phenologies and mating behaviour. J. N. Am. Benthol. Soc. 6: 198- 2r2. Wrilker, W. 1956. Zur Kenntnis der Gattung Psectrocladias Kieff. (Dipt., Chironom.). Individuelle Variabilitdt. Grenzen und Mdglichkeiten der Artentrennung, Okologie 76 NEARCTIC CHIRONOMIDAE

INDEX alboviridis (Malloch), Gillotia 46 albulum Townes, Polypedilum 51 A aleta Roback, Conchapelopia 11 abanus Curran, Cricotopus LJ algens (Coquillett), Apsectrotanypus .9 abbreviatus (Malloch), Cryptochironomus 44 allicis Sublette, Tanytarsus 58 abdominoflavalus Picado, Metriocnemus 30 Allopseclrocladius Wiilker 35 aberrans (Johannsen), Microtendipes 48 Alolanypus Roback .8 aberrala Lundbeck. Diamesa 11 afphaeus(Sublette),Paracladopelma .... - 49 abetusRoback,Procladius ...... 15 alpicola (Zetterstedt), Paracladius JJ abiskoensis Edwards, Orthocladius ...... 31 alpina Tokunaga, Diamesa AbiskomyiaEdwards ....20 alternantherae Dendy & Sublette, Nanocladius 30 AblabesmyiaJohannsen ...... 10 amachaerus (Townes), Cladopelma 43 abnobaeus (Wiilker), Parachaetocladius . . . . 33 amazonicus (Fittkau), Goeldichironomus 47 abnobeus Sether & Sublette, ambigua Zetterstedt, Corynocera 56 Parachaetoclad i us -t -t americana Fittkau, Ablabesmyia 10 abortivus (Malloch), Parachironomus 49 americana (Fittkau), Meropelopia 13 abrupta (Garrett), Procladius l6 americ ana (Kieffer), Po.lypedilurn 51 abruplus (Townes). Demeijerea A< americanus Beck & Beck, Nilotanypus IJ absurdus (Johannsen), Cricotopus L3 americanus Kieffer, Glyptotendipes ia Acalcarella Shilova 40 amphitrite (Townes), Chernovskiia 4I Acamptocladius Brundin 20 amplexivirilia Hansen, Diamesa l7 acifer Townes, Polypedilum 52 amplicaudus Sather & Sublette, acigus Snther, Gymnometriocnemus 27 Pseudorthocladius 36 aclines Roback, Derotanypus 9 amplicristata Halvorsen, Saetheriella ...... 37 aclines (Sublette), Derotanypus 9 amplicristatus Srcther, Rheocricotopus 37 acra (Roback), Rheopelopia 13 amplus Townes, Glyptotendipes ...... 47 Acricolopus Kieffer 20 anas Townes, Pseudochironomus 55 aculislylus S&ther, Mesosmittia 29 Anceus Roback 4I A dac tylo c ladizs Sather 26 ancysla Roback, Diamesa l7 Adiamesa Kieffer 17 anderseni Sather, Nanocladius 30 adnilus Epler, Dicrotendipes A< anderseni Reiss & Fittkau, Tanytarsus 58 adsimilis (Goetghebuer), Chaerocladius ')') angustum Townes, Polypedilum 51 adumbratus Johannsen, Procladius 15 annectens Sather, Orthocladius 31 aeiparthenus Bilyj, Cladotanytarsus JJ annellae Borkent, Stenochironomus 53 aequalis Johannsen, Metriocnemus 30 annularis (De Geer), Chironomus 42 aequifasciata (Dendy & Sublette), annularis (Meigen), Paratendipes 50 Ablabesmyia 10 annulata (Say), Ablabesmyia 10 aestivalis Townes, Stenochironomus 53 annulator Goetghebuer, Cricotopus 23 aestivus (Brundin), Bryophaenocladius 2l annulicrus (Townes), Stictochironomus . 53 aethiops (Townes), Dicrotendipes 45 annuliventris (Malloch), Euryhapsis 26 agrayloides (Kieffer), Lauterborniella . . . . . 48 anomala Beck & Beck, Paramerina ...... 13 alx Townes, Pseudochironomus 55 anonymus Williston, Chironomus 42 akrina (Roback), Rheotanytarsus 57 anteilis Roback, (Hydrobaenus) ...... 40 alamedensis (Sublette), Apedilum 40 anthracinus Zetterstedt, Chironomus ...... 42 alaskensis Brundin, Trichotanypus 7 anlicus (Walker), Microtendipes ...... 48 alaskensis Coquillett, Telmatogeton 7 Anlillocladius Sather 20 alaskensis (Malloch), Derotanypus 9 Apedilum Townes 40 alaskensis Roback, Ablabesmyia 10 Apelopia Roback 76 alatus (Beck), Parachironomus 49 apicalis (Walley), Djalmabatista 14 a/ba (Roback), Meropelopia 13 apicatum Townes, Polypedilum 52 albertae Kevan & Cutten-Ali-Khan, Apomelriocnemus Sether 20 Oreadomyia 31 AponleusRoback ...... 8 albescens (Townes), Sergentia 53 appendiculata (Lundstr6m), Arctodiamesa 16 albicorne (Meigen), Polypedilum 51 appersoni Soponis, Orthocladius 31 albicrus (Townes), Stictochironomus . . 53 apposita (Walker), Micropsectra 56 albidohalleralis Malloch, (Orthocladius) . . . 40 ApseclrolanypusFittkau .....8 albimanus (Meigen), Paratendipes 50 aptena Wirth, Tethymyia 38 albinodus Townes, Polypedilum 52 aratus Roback, Cricotopus 23 albipalpus Borkent, Stenochironomus 53 arctica(Boheman), Diamesa ...... l7 albipennis (Meigen), Endochironomus 46 arctica Kieffer, Corynoneura ...... 22 albiplumus Sether, Cardiocladius 2I arclica (Malloch), Pseudodiamesa . . 18 albistria Walker, Chironomus +J arclica Malloch, Smittia 38 albolineatus (Meigen), Metriocnemus 30 arclicus Brundin, Paralimnophyes ...... 34 CHIRONOMIDAE DE LA REGION NEARCTIQUE 77

Arclodiamesa Makarchenko 76 basidens Townes, Paratendipes 50 Arclopelopia Fittkau 11 bathyphila (Kieffer), Monodiamesa l9 arcuata (Garrett), Procladius I6 bathophila (Kieffer), Parakiefferiella JJ arcualus Sather, Antillocladius 20 baueri (Goetghebuer), Acricotopus 20 argentea (Townes), Cyphomella A< bauri (Kieffer), Stempellina 58 argus Roback, Cryptochironomus 44 b ausei (Kieffer), Stempell ina )5 ariel (Sublette), Cryptotendipes . 44 Beardius Reiss & Sublette 41, arielina (Coquillett), Lasiodiamesa 7 beckae (Sublette), Asheum : 4l aris Roback, Alotanypus 8 becki Roback, Labrundinia 12 armatus Andersen, Psectrocladius 35 Beckidia Saether 4l articaudus Sether, Pseudochironomus l) Beckiella Srther 41 artifer (Curran), Polypedilum 51 belkini Dendy & Sublette, Cricotopus ...... 25 arlisia (Roback), Pagastia 18 bellus (Loew), Procladius 15 AsayiaRoback... 10 beringensis Oliver & Dillon, Cricotopus 23 asetus Roback, Procladius 15 beringensis Cranston & Oliver, Asheum Sublette & Sublette 47 Apometriocnemus . . : . . . 20 aspera (Roback), Ablabesmyia . . . 10 berneri Beck & Beck, Larsia IL asquamatus Andersen, Limnophyes 28 bertrami Edwards, Diamesa 77 astis (Roback), Bryophaenocladius 2I Bethbilbeckia Fittkau and Murray 9 Atelopodella Sather 26 biappendiculata Goetghebuer, Diamesa tt ater Goetghebuer, (Trichocladius) 40 bicaudatus Seether, Doncricotopus 25 aterrima (Meigen), Smittia 38 bicinctus (Meigen), Cricotopus 23 athalassicus Cannings, Chironomus qL bicolor (Malloch), Nanocladius 30 atomarius (Zetterstedt), Limnophyes 28 bicolor Rempel, Chasmatonotus ...... 22 atra Winnert4 Corynoneura 1a bicolor Sather, Baeoctenus 20 atratulus (Zetterstedt), Metriocnemus 30 bicorne (Tnwnes), Nilothauma 49 atrella (Townes), Chironomus 42 bifasciata (Coquillett), Zavrelimyia 74 atria Roback & Rutter, Denopelopia 11 lrifascialus Malloch, (Orthocladius) ... . 40 atridorsurn Kieffer, Cladotanytarsus - f) bifida (Garrett), Procladius L6 atrimana (Coquillett), Demeijerea A< bifimbriatus Kieffer, Chironomus 43 atripes Rempel, Chasmatonotus 22 bifurcatus Cranston & Oliver, Cricotopus 24 atritibia Malloch, Chironomus bilobatus Brundin, Diplocladius 25 atroviridis (Townes), Chironomus A1 bilyji Sather, Platysmittia 35 atrum (Townes), Tribelos 54 bimacula Walker, Chironomus 43 atlenuata Reiss, Micropsectra )o bimaculatus Osten Sacken, Chasmatonofis . . . 22 attenuatus Walker, Chironomus {J biseta (Townes), Chironomus 42 atus Roback, Coelotanypus 8 bisetosus Borkent, Stenochironomus 53 aurea Johannsen, Ablabesmyia . . . t4 blarina Townes, Cryptochironomus ...... 44 aureus (Johannsen), Pseudochironomus 55 bohemani Goetghebuer, Diamesa 7'7 aureus Roback, Clinotanypus 8 boiemicus (Bause), Paratanylarsus 57 autiensis (Roback), Ablabesmyia 10 boliekae Beck & Beck, Monopelopia ...... 13 austini Beck & Beck, Einfeldia 4() boltoni Srether & Willassen, Lappodiamesa 18 aviceps Townes, Polypedilum 51 borealis Curtis, Chironomus ...... 43 Axarus Roback A1 borealis Garrett, Diamesa L7 borealis Goetghebuer, Limnophyes 28 B borealis (Kieffer), Micropsectra 56 babiyt (Rempel), Nilothauma 49 boreoafpina (Goetghebuer), Krenosmittia . . . . . 28 bacilliger (Kieffer), Cricotopus 24 BOREOCHLINI .... 7 badius Sether, Pseudochironomus 55 BoreochlusEdwards .-.-....7 Baeoclenus Sather 20 Boreoheptagyia Brundin 16 baltimoreus (Macquart), Natarsia 10 BOREOIIEPTAGYIINI 16 banana Garrett, Diamesa LI botaurus (Townes), Dicrotendipes 45 banksl Townes, Pseudochironomus i) boydi (Beck), Cladopelma 43 baptlstensis Boesel, Cricotopus LJ brachialis (Coquillett), Demeijerea 45 barbatimanus Kieffer, Psectrocladius 35 brachyneura (Malloch), Paraphaenocladius 34 barbatulus Sublette, Procladius 15 branchicolus Sether, Nanocladius 31 barberl (Coquillett), Thienemannimyia I4 branickii (Nowicki), Pseudodiamesa 18 barberi Sether & Sublette, Doithrix 25 braseniae (Leathers), Polypedilum 51 barbicornls (Linnaeus), Orthocladius JL brevicosta (Edwards), Constempellina 56 barbimanus (Edwards), Psectrocladius 35 brevicosla Kieffer, Psectrocladius 36 barbipes (Staeger), Glyptotendipes 47 brevinervis (Holmgren) Paraphaenocladius 34 b as alis (Malloch), Para tendipes 50 brevinervis (Malloch), Eukiefferiella 26 b as alis (Staeger), Cricotopus brevipalpis (Kieffer), Chironomus 42 basalis (Walley), Ablabesmyia 10 brevis (Edwards), Stempellinella . 58 78 NEARCTIC CHIRONOMIDAE

breviseta S&ther, Orthocladi us 3Z ceri,r Roback, Cricotopus 23 brevitibialis (Zetterstedt), Dicrotendipes 40 chaetaolus (Sublette), Parachironomus 49 briani Soponis, Zalutschia 40 Chaetocladius Kieffer ZL Brillia Kieffer 20 Chaetolabis Townes ^a browni Kellogg, Eretmoptera 26 changi Sather, Heterotrissocladius 27 browni Townes, Stenochironomus 53 chapmani (E,dwards), Rheocricotopus 37 brumalis (Edwards), Gymnometriocnemus 27 charensis Soponis, Orthocladius AL a) bruna Roback, Conchapelopia . . . 11 Chasmatonotus Loew brundini Lindeberg, Tanytarsus )t' chaltahoocheensis Caldwell, Symbiocladius 38 brundini Siiwedal, Micropsectra 56 cheimatophila Hansen, Diamesa 17 Brundinia Roback 9 chelonia (Townes), Einfeldia 46 Brundiniella Roback 9 chen Townes, Pseudochironomus 55 brunnea (Roback), Macropelopia 9 Chernovskiia Sether 4T brunneipennis (Johannsen), Einfeldia . . . ., 46 chiobates Hansen, Diamesa 17 brunneus Borkent, Xestochironomus fJ CHIRONOMINAE . 40 brunneus Roback, Procladius 15 CHIRONOMINI 40 brunneus (Walker), Micropsectra 56 Chironomus Meigen 4l brunnicans Walley, Cricotopus chorea Lundbeck, Diamesa I7 brunnipes (Zetterstedt), Micropsectra )o chrysos (Sublette). Hayesomyia 11 brusti Sather, Lasiodiamesa ,7 cilium Oliver, Euryhapsis z6 brusti Srether, Metriocnemus 30 cinctipes (Johannsen), Ablabesmyia 10 Bryophaenocladius Thienemann )1 cinctum Townes, Polypedilum 51 buckleyl Sublette, Tanylarsus 58 cinctus Townes, Stenochironomus 53 bureni Wirth, Thalassomya '7 Cladopelma Kieffer 43 byssina (Schrank), Camptocladius a1 Cladotany(arsus Kieffer 55 claripennis (Lundbeck), Eukiefferiella 26 c claripennis (Malloch), Paracladopelma 50 caducus Townes, Microtendipes 48 claripennis (Malloch), Procladius 15 caelum Townes, Microtendipes 48 clarkei Soponis, Orthocladius 5Z caena Roback, Diamesa 77 clavata Edwards, Diamesa t7 calcaralus (Edwards), Psectrocladius 35 clavatosus Szether & Sublette. californicum Sublette, Polypedilum 51 Pseudorthocladius 36 californicus (Johannsen), Dicrotendipes 45 clavalusBeck,Tanypus ...... 16 californiensis Hashimoto, Clunio 22 clavicornis (Saunders), Thalassosmittia 38 caliginosus (Johannsen), Clinotanypus 8 claviger(Townes), Robackia ...... 52 caliginosus (Johannsen), Glyptotendipes 47 clavigera Sather, Compterosmittia 22 Calopsectra Kieffer 58 clavus Roback, Procladius 15 calopterus (Mitchell), Polypedilum 51 clepsydrus Coquillett, (Orthocladius) ...... 40 Calotanypus Roback I4 Clinocladius Sublette 2I calvata Sather, Sublettiella 38 clinopecten Sather, Stilocladius 38 calvescens (Edwards), Tvetenia 39 ClinotanypusKieffer ...... 8 calvescens Sether, Hydrobaenus 27 Clunio Haliday 22 Camptochironomus Kieffer 4T COELOTANYPODINI ...... 8 Camptocladius Wulp 2T CoelolanypusKieffer ...... 8 camptolabis (Kieffer), Paracladopelma 49 coffmani (Roback), Sublettea 58 camptophleps (Edwards), Krenosmittia 28 colei (Malloch), Stenochironomus 53 cana Roback, Arctopelopia 11 colenae Hansen, Diamesa 17 canadensis Bilyj, Larsia l2 collator (Townes), Cladopelma 43 Cantopelopia Roback 11 coloradensis Srcther, Limnophyes 28 Cardiocladius Kieffer a1 comans Sather & Sublette, Pseudorthocladius 36 carinala Sather, Diplosmirtia 25 comosaSublette,Pentaneura ...... 13 carinata Strenzke, Parasmittia Jq compes (Coquillett), Stictochironomus 54 carinatus Sublette, Tanypus t6 Compteromesa Sether 19 carinatus (Townes), Parachironomus 49 Compterosmiltia S€ther 22 carlalus (Roback), Orthocladius 3L concovus Roback, Tanypus 16 carnea (Johannsen), Zavrelimyia . t4 Conchapelopia Fittkau 11 carneosa Filtkau, Zavrelimyia 14 concinnus (Coquillett), Coelotanypus ...... 8 carolinensis Sether, Limnophyes 28 confinis (Walker), Micropsectra 56 carus (Townes), Goeldichironomus A1 conflusirus Sather, Rheocricotopus 37 casuarius (Townes), Cryptotendipes 44 conformis (Holmgren), Hydrobaenus 27 caudalus Edwards, Protanypus 19 conformis Malloch, Chironomus 43 cayugae Johannsen, Chironomus /a confusa (Garrett), Diamesa l7 celeripes Winnertz, Corynoneura 22 confusus Malloch, Tanytarsus 58 cenlralis (Malloch), Derotanypus 9 confusus Palm€n, Paratanytarsus ...... 57 CHIRONOMIDAE DE IA REGION NEARCTIQUE 79

connexa (Kieffer,), Micropsectra 56 decolorata (Malloch), Larsia l2 consobrinus (Holmgren), Orthocladius )L decoralus (Holmgren), Orthocladius 32 Conslempellina Brundin 56 decorus Johannsen, Chironomus 42 conus Mason, Cryptochironomus decumbens Malloch, Chironomus 42 conversus (Walker), Paracladius 33 deflecta (Johannsen), Micropsectra 56 conviclum (Walker), Polypedilum )l deflexa (Walley), Microtendipes ...... 48 cooki Sather, Heterotrissocladius 27 deltaensis Roback, Procladius 15 cooki Soponis, Orthocladius JL Demeiierea Kruseman 45 coracina (Zetterstedt), Sergentia 53 demeijerei (Kruseman), Robackia 52 Cordiella Coffman & Roback 29 Demicryptochironomus Lenz , 45 Cordites Brundin 29 dena (Roback), Macropelopia ...... 9 cornea Sether, Cyphomella 45 dendyl Sublette, Tanytarsus 58 1a cornulicaudata (Walley), Helopelopia . . Denopelopia, Roback & Rutter 11 coronata (Edwards), Parakiefferiella JJ denticulatus Sublette, Procladius 15 coronatus Hirvenoja, Cricotopus z) dentifer Brundin, Orthocladius 32 Corynocera Zetterstedt 56 dentolatens (Srether), Acamptocladius 20 Corynoneura Winnertz 22 dentus Roback, Procladius 15 crassicaudalus Malloch, Chironomus A' depectinata Sether, Monodiamesa 19 crassicollis (Walker), Smittia 38 DerotanypusRoback ...... 9 crassicornus Sather, Nanocladius 30 desis Roback, Procladius 15 crassinervis (Zetterstedt), Procladius l) despectus (Kieffer), Paraphaenocladius 34 crassus Townes, Pseudochironomus )) deslitulus Srther & Sublette, cretis Roback, Procladius l) Pseudorthoclad ius 36 Cricotopus Wulp . L3 deviatus (Malloch), Tanytarsus 58 cristatissimus Srether, Limnophyes 28 devinclus (Say), Stictochironomus 54 cristatus Fabricius, Chironomus 42 devineyae (Beck), Goeldichironomus cristatus Sether, Psilometriocnemus devonica (Edwards), Eukiefferiella 26 crusculus (Sather), Cladotanytarsus 55 Diamesa Meigen T6 cryplicus Epler, Dicrotendipes 45 DIAMESINAE 1,6 Cryptochlronomus Kieffer DIAMESINI 76 Cryptocladopelma Lenz CJ diara Roback, Corynoneura 22 Cryptotendipes Lenz 44 diastena (Sublette), Sympotthastia 19 cubila Garrett, Prodiamesa 20 Dicrolendipes Kieffer 45 cubitus (Garrett), Thienemanniella ...... 39 difficilis (Lundbeck), Orthocladius 3l cucini Webb, Chironomus 42 digila{a Sether, Pseudosmittia 36 culiciformis (Linnaeus), Procladius 15 digitalus (Malloch), Cryptochironomus 44 cultriger Kieffer, Diplocladius 25 digiiatus Sether, Bryophaenocladius 27 cumulatus Hirvenoja, Cricotopus zt digitifer Townes, Polypedilum 52 cuneatus (Townes), Demicryptochironomus 45 diflonae Cranston & Oliver, Doithrix ...... 25 currani Beck & Beck, Guttipelopia . AI dimorphus (Malloch), Tribelos 54 currani Sublette, Orthocladius Diplocladius Kieffer 25 currani (Walley), Conchapelopia 11 Diplomesa Pagast . 18 curryi Mason. Cryptochironomus 44 Diplosmit(ia Sather 25 curticornis Kieffer, Tanytarsus 58 directus (Dendy & Sublette), Parachironomus 49 curticornus Sether, Georthocladius 26 discolor (Coquillett), Psectrotanypus . . . . . 9 curticornus Sether & Sublette, dispar Lindeberg, Tanytarsus 58 P seudorthocla d ius 36 dispar (Meigen), Endochironomus 46 curtilamellala (Malloch), Harnischia 47 dissimilis (Johannsen), Paratanytarsus 57 curtisela Sether, Orthocladius distinctissimus (Brundin), Rheotanytarsus . . . . 57 curtistila Sether, Heleniella 27 distinctus (Malloch), Nanocladius 30 curtistylus Goetghebuer, Pseudorthocladius 36 distinguendus Kieffer, Chironomus 42 curtus Hirvenoja, Cricotopus z) dfversus Boesel, Cricotopus 25 curlus Roback, Procladius 15 dives (Johannsen), Micropsectra 56 cyanea (Thienemann), Eukiefferiella . . 26 Djalmabatista Fittkau L4 cylindraceus (Kieffer), Cricotopus L3 Doithrix Sether & Sublette 25 Cyphomella Srether 45 domus (Sublette), Hydrobaenus 28 Doncricolopus Sather 25 D dorenus (Roback), Orthocladius 32 darbyt (Sublette), Cryptotendipes 44 doris (Townes), Paracladopelma 49 daviesi Bilyj, Cladotanytarsus 55 dornerl (Malloch), Axarus 4L davisl Edwards, Diamesa I7 dorsalisMeigen, Chironomus ...... 43 debilipennis (Lundbeck), Paraphaenocladius J+ dorsenna (Srether), Kloosia 48 debllis (Meigen), Tanytarsus )6 downesi (Steffan), Nanocladius 31 decedens (Walker), Macropelopia 9 drelsbaclri Townes, Glyptotendipes 47 80 NEARCTIC CFIIRONOMIDAE

dubitatus Johannsen, Orthocladius -11 fascipes (Coquillett), Microtendipes 48 dubius (Malloch), Paratanytarsus 57 fasciventris Malloch, Chironomus 43 dub ius (Meigen), Nilotanypus l-f fastigatus (Townes), Demicryptochironomus 45 dumicaudus Sather, Pseudorthocladius 36 fastuosa (Johannsen), Natarsia 10 duodenaria Kieffer, Tvetenia 39 fatigans Johannsen, Tanytarsus 58 duplicatus (Johannsen), Paratendipes 50 fennicus Stori, Psectrocladius 35 dusena Roback, Conchapelopia 11 ferringtoni Sather, Odontomesa l9 dux (Johannsen), Kiefferulus 48 ferringtoni Soponis, Orthocladius 32 dyari (Coquillett), Psectrotanypus 9 ferrugineovittatus Zetrcrstedt, Chironomus 42 dyari (Townes), Phaenopsectra JI' festivellus (Kieffer), Cricotopus festivus (Meigen), Cricotopus LJ E feslivus (Say), Axarus aa edurus Sublette & Sublette, Cricotopus z5 festivus (Holmgren), Chaetocladius edwardsi Brundin, Psectrocladius J) fimbriata Sather. Platysmittia 35 edwardsi Goetghebuer, Smittia 38 fimbriatus Reiss & Fittkau, Tanytarsus 58 edwardsi Jones, Metriocnemus 30 fimbriatus (Walker), Nilotanypus 13 edwardsi (Kruseman), Cladopelma 44 fimbriosus Sether & Sublette effusoides Sather, Rheocricotopus Georthocladius 26 effusus (Walker), Rheocricotopus al fittkaui (Beck & Beck), Meropelopia ...... 13 Einfeldia Kieffer 46 fitlkauiSchlee, Corynoneura ...... 22 elachislus Townes, Apedilum 40 FittkauimyiaKarunakaran .,..9 elaensis Bilyj, Cladotanytarsus )) flabellatus (Meigen), Tanytarsus 59 elana (Roback), Thienemanniella 39 flannagani Srther, Cricotopus 23 elatus Roback, Psectrocladius 36 flavellus (Zetterstedt), Tanytarsus 59 elegans Johannsen, Cricotopus z5 flavens (Malloch), Epoicocladius 25 elodeae (Townes), Parachironomus 49 flavibasis Malloch, (Camptocladius) 40 elongatus Roback, Procladius 15 flavibasis Malloch, Cricotopus 24 elloni (Edwards), Limnophyes 28 flavicauda (Malloch), Phaenopsectra 50 eminellobus Sether, Rheocricotopus 5t flavicinctus(Loew), Clinotanypus ..... 8 eminentia Mason, Cryptochironomus 44 flavicingula (Walker), Stictochironomus 54 emorsus (Townes), Cryptotendipes 44 flavidus Kieffer, Procladius t5 Endochironomus Kieffer 46 flavifrons (Johannsen), Brillia . 2l Endotribelos Grodhaus 40 flavifrons (Johannsen), Meropelopia 13 ensifer Saether & Sublette, Doithrix 2J flavipes Johannsen, Cricotopus 24 eoclivus Sether, Parametriocnemus J9 flavipes (Meigen), Phaenopsectr^ . 50 ep hemerae (Kieffer), Epoicocladius 26 flaviventis (Johannsen), Polypedilum 51 ephippium (Zetterstedt), Cricotopus LJ flavocinctus (Kieffer), Cricotopus 23 Epoicocladius Zavrel 25 flavoscutellatus(Malloch),Bryophaenocladius 2L equitans (Claassen), Symbiocladius -J|' flavus (Johannsen), Psectrocladius 35 Sretmoptera Kellogg 26 flavus (Johannsen), Polypedilum . 51 Eudactylocladius Thienemann 31 flavusKieffer,Coelotanypus ...." 8 Eukiefferiella Thienemann 26 flexuella (Edwards), Mesosmittia 29 eumorpha (Sublette), Brundiniella 9 florens (Johannsen), Apsectrotanypus ...... 9 Euorthocladius Thienemann 31 f-loridana Beck & Beck, Labrundinia ...... 12 Euryhapsis Oliver . 26 floridense Townes, Polypedilum 52 Eutanypus Coquillet 1,7 floridensis Fittkau &Murray, Bethbilbeckia .... 9 exagitans (Johannsen), Paraphaenocladius -1+ fluminalis (Sublette), Zavrelimyia 74 excavatus Brundin, Orthocladius -)L foliaceus Wirth & Sublette, Trichotanypus . . 7 excavatus Kieffer, Chironomus 4J folliculatus S&ther, Limnophyes 28 excellens (Brundin), Tokunagaia 39 fonticola Sether, Diamesa L7 excelsius Townes, Polypedilum )z fontinalis Saether, Apometriocnemus ...... 20 excisus Kieffer, Chironomus 43 fontinalis (Sather), Paratrichocladius ...... 34 exiguus (Johannsen), Rheotanytarsus 57 forceps (Townes), Parachironomus 49 exilis Johannsen, Cricotopus 24 forcipalus (Goetghebuer), Pseudosmittia 36 exquisitus (Mitchell), Stenochironomus . . . 53 fragilis (Walley), Paramerina ...... 13 exratus (Roback), Rheocricotopus -)t freemani Sublette, Procladius 15 exlrema (Holmgren), Smittia 38 frequens (Johannsen), Parachironomus 49 frigidus (Zetterstedt), Orthocladius 32 F frommeri Atchley & Martin, Chironomus 42 fallax (Johannsen), Polypedilum . .tl fugax (Johannsen), Cricotopus 23 farinalis (Walley), Cryptochironomus 44 fulva (Johannsen), Sympotthastia 19 fasciala Beck & Beck, Conchapelopia 11 fulva (Kieffer), Odontomesa 19 fasciger (Curran), Djaknabatista I4 fulvescens (Johannsen), Polypedilum 51 fascipennis Coquillett, Chasmatonotus 22 fulvipilus (Rempel), Goeldichironomus 47 CHIRONOMIDAE DE LA REGION NEARCTIQUE 81

fulvipluma Kieffer, Camptocladius 40 grisea (lvfalloch), Cyphomella 45 fulviventris (Johannsen), Pseudochironomus )l griseopunctatus (Maltoch), Polypedilum 52 fulvus (Johannsen), Cardiocladius )1 grodhausiSublette,Tanypus ...... 16 fulvus (Johannsen), Cryptochironomus 44 groenlandensis Goetghebuer, Orthocladius . ., . 32 fumidus (Johannsen), I)icrotendipes A< groenlandica Andersen, Micropsectra ...... 56 fumosinus (Curran), Bryophaenocladius )1 groenlandiensis Andersen, Limnophyes 28 fumosus (Johannsen), Limnophyes 28 guerlus (Roback), Tanytarsus 58 furcarca Sather, Zalutschia 40 Guttipelopia Fittkau 11 furcata Edwards, Diamesa l7 gultipennis (Wulp), Guttipelopia 11 furcatus (Kieffer), Bryophaenocladius 2l guttularis (Coquillett), Psectrotanypus .... 9 furtivus Sublette & Sublette, Cricotopus 25 Gymnome(riocnemus Goetghebuer 26 /asca (Johannsen), Cardiocladius 2l fuscatus Wirth, Cricotopus 25 H fusciceps (Edwards), Thienemannimyia I4 Habrobaenus Srether 33 fuscicorne (Malloch), Tribelos 54 Ilalocladius Hirvenoja 27 fuscipalellus Borkent, Stenochironomus 53 halophilus Packard, Chironomus 43 fusclpenne (Meigen), Polypedilum 51 halterale (Coquitlett), Polypedilum 52 fuscipes (Meigen), Metriocnemus 30 halvorseni (Cranston & Sether), fuscitibia Sublette, Paratendipes )1, Krenosmittia 28 fusciventris (Malloch), Tribelos 54 hamatus (Johannsen), Parametrio;nemus 34 fuscus (Kieffer), Cricotopus L' hamiltoni (Srether), Hydrobaenus 28 fusiformis Bilyj, Cladotanytarsus f) hamiltoni Srether, Limnophyes 28 fusistylus (Goetghebuer), Hydrobaenus 28 hamiltoniSather,Protanypus ...... 19 futilis Wulp, (Tanypus) 74 hamiltoni Srther & Sublette, Doithrix 25 hanseniRoback,Paramerina ...... 13 G hanseni Wirth & Sublette, Trichotanypus 7 gaedii (Meigen), Potthastia l6 IlarnischiaKieffer ...... 47 galaptera (Townes), Paracladopelma 50 harti Malloch, Chironomus 43 galeator (Townes), Cladopelma 44 hastulatus Sather, Limnophyes 29 garretti Sublette & Sublette, Diamesa T7 hauberi Beck & Beck, Ablabesmyia 10 taftetti (Walley), Psectrotanypus 9 haydaki Hansen, Diamesa L7 gelidus (Kieffer), Cricotopus ?3 Ilayesomyia Murray & Fittkau 11 gelidus Kieffer, Orthocladius JI hazenensis Soponis, Orthocladius 32 geminala Kieffer, Diamesa 11 Heleniella Gowin 27 geminatus (Say), Cricotopus 25 &e/is (Roback), Paratrichocladius 34 Georthocladius Strenzke 26 hellenthali Soponis, Orthocladius 32 gera (Roback), Ablabesmyia 10 IlelopelopiaRoback ...... L2 gesla Roback, Cantopelopia 11 hesperium (Sublette), Endotribelos 46 gesra (Roback), Meropelopia IJ Hesperodiamesa Sublette 18 gibbera Srcther, Cyphomella A< heteropus (Coquillett), Diamesa I7 Gillotia Kieffer 46 Ileterotanytarsus Spiirck 27 glatrer Srether, Paracricotopus JJ Heterotrissocladius Spiirck 27 glabricollis (Meigen), Rheocricotopus 3l hilaris (Walker), Stenochironomus 53 glacialis Edwards, Cricotopus 4+ hirta Srether, Heleniella 27 glacialis Kieffer, (Camptocladius) ...... 40 hirta S&ther, Saetheria 52 glaucurus Wiedemann, Chironomus qJ hirtalatus (Beck & Beck), Parachironomus 49 globlfer (Lundstrdm), Limnophyes 28 hfrtapex Sether, Heterotrissocladius ...... 27 globislylus Roback, Cricotopus 25 hirtipectus Sether, Parachaetocladius . . . . 33 Glyptotendipes Kieffer 47 hirtipennis(Loew), Macropelopia ..... 9 Goeldichironomus Fittkau 47 hirtipes (Mitchell), Polypedilum 51 gomphus Townes, Polypedilum 52 histrio (Fabricius), Stictochironomus 54 goniodes (Sublette), Conchapelopia 11 holmgreni (Jacobson), Chaetocladius ...... 22 gouin i (G oetghebuer), Rheocricotopus at holochlorus Edwards, Tanytarsus 59 Graceus Goetghebuer 47 holoprasinus (Goeldi), Goeldichironomus 47 gracilentus (Holmgren), Tanytarsus 58 Hololanypus Roback 15 gracilistylus Brundin, Boreochlus 7 Holtedahlia Kieffer ...... 4L gracillima (Kieffer), Parakiefferiella JJ hospita (Edwards), Eukiefferiella 26 graminicola (Lundbeck), Parametriocnemus 34 hudsoni Roback, Krenopelopia 12 grandilobus Brundin, Chaetocladius aa hudsoni Srether, Hydrobaenus 28 grandls Roback, Procladius l) hudsoniSather, Limnophyes ...... 29 grandivalva (Shilova), Chironomus A1 hudsoni (Sather), Parachaetocladius 33 gregsonl Edwards, Diamesa tt HudsonimyiaRoback ...... Lz gretis Roback, Procladius 15 hugginsi Ferrington & Sether, Oliveridia 31 grlmmil (Schneider), Paratanytarsus 57 humeralis (Zetterstedt), Cricotopus 23 82 NEARCTIC CHIRONOMIDAE

hyalinus Coquillett, Chasmatonotus 22 jucundum (Walker), Tribelos 54 Hydrobaenus Fries 27 julia (Curran), Pseudochironomus 55 hygropetricus Kieffer, Metriocnemus 30 junci (Meigen), Micropsectra 56 hyperborea Holmgren. Diamesa 11 junus Roback, Cricotopus 25 hyperboreus Staeger, Chironomus ^a hyporheicus Coffman & Roback, K Lopesclad i us 29 kansensis Roback, Nilotanypus 13 Hyporhygma Reiss 4'7 karelena Roback, Hudsonimyia 72 Karelia Roback 10 I kenorensis Sather, Rheocricotopus 37 iberica Serra-Tosio, Potthastia 18 kiefferi (Garrett), Parochlus ...... 8 idei (Walley), Ablabesmyia 10 Kiefferulus Goetghebuer 48 illinoense (Malloch), Polypedilum 51 Kloosia Kruseman 48 illinoensis (Malloch), Ablabesmyia 10 knabeni Goetghebuer, Orthocladius 32 illoba Oliver, Euryhapsis z6 knabi (Coquillett), Metriocnemus 30 imberbus Sether & Sublette, knuthi Soponis, Orthocladius 32 Parachaetocla dius JJ Krenopelopia Fittkau 12 immucronalus Srether, Limnophyes 29 Krenopsectra Reiss 56 impeclinus Saether, Bryophaenocladius 2l Krenosmittia Thienemann & Krtiger 28 impensus (Walker), Paraphaenocladius J4 Kribioxenus Kieffer ...... 48 imperator Walley, Chironomus A1 imperialis Sublette, Tanypus l6 L inaequalis Goetghebuer, Tanytarsus )n Iabeculosum (Mitchell), Polypedilum 52 incallida (Walker), Diamesa I7 labradorensis Sather, Hydrobaenus 27 incidata Townes, Harnischia A1 Labrundinia Fittkau 72 incognitus (Malloch), Dicrotendipes A< laccophilus (E,dwards), Paratanytarsus 57 incompta (Zetterstedt), Phaenopsectra 50 lacteipennis Johannsen, Trichochilus 39 incomplus Sather, Nanocladius 30 lacteipennis Malloch, Orthocladius 32 inconspicua (Malloch), Pentaneura 13 laculata Oliver & Roussel, Brillia 27 inculta Beck & Beck, Pentaneura IJ lacustris Edwards, Corynoneura 22 incurvus (Sublette), Dicrotend ipes 46 laetum (Meigen), Polypedilum 51 indistincta Beck & Beck, Larsia 1.2 laetus Hirvenoja, Cricotopus 24 indistinctus (Malloch), Dicrotendipes 46 Lappodiamesa Serra-Tosio 18 inermis Srether, Lopescladius 29 lapponicus Goetghebuer, Orthocladius 32 infuscatus (Malloch), Cricotopus LJ lapponicus Lindeberg, Tanytarsus 58 innocuus (Curran), Parametriocnemus 34 Lapporthocladias Thienemann 37 inopinala Reiss, Skutzia JI laricomalis Edwards, Cricotopus 24 insignilobus Kieffer, Micropsectra 56 LarsiaFittkau -...... 12 insignipes Kieffer, Diamesa 17 LasiodiamesaKieffer ...... 7 Insulanus Sublette

lindrothi Goetghebuer, Diamesa 1'7 mancus (Walker), Cladotanytarsus 55 lindrothi Goetghebuer, Micropsectra 56 manitobensis Srether, Orthocladius 32 lineatus (Say), Axarus 4l marcidus (Walker), Heterotrissocladius . . 27 lineola (Say), Axarus rnargarelae Sather, Limnophyes 29 lingulata Sether, Zalutschia 40 marginelfa (Malloch), Larsia ...... 12 Lipiniella Shilova 48 mariae Wirth & Sublette, Trichotanypus ...... 8 Lipurometriocnemus Srcther 29 Maricladius Sublette .....-.33 lobata Edwards, Corynoneura 22 marina (Saunders), Thalassosmittia 39 lobiferus (Say), Glyptotendipes 4'7 marionensis Sether, Gymnometriocnemus 26 lobiga Sather, Mesosmittia 29 marmoreus (Townes), Stictochironomus 54 Iobiger (Kieffer), Dicrotendipes A< marshalli Stone & Wirth, Clunio 22 Lobochironomus Ryser, Wtilker & Scholl +J martini Sether, Hydrobaenus 28 lobus (Beck), Dicrotendipes 46 maturus Johannsen, Chironomus 42 loganae Beck & Beck, Paracladopelma 50 melanderi Kieffer, Chironomus 43 logani (Johannsen), Micropsectra 56 melanops (Wiedemann), Conchapelopia 11 longifurc a Kieffer, Brillia )1 melanosoma (Goetghebuer), Arctopelopia . . . . 11 longimana Kieffer, Potthastia 18 mendax Kieffer, Tanytarsus 59 longipennis (Holmgren), Metriocnemus 30 mendax Stori, Chironomus 43 longipes Staeger, Chironomus 43 mendotae Muttkowski, Diamesa l7 longitarsus Goetghebuer, Metriocnemus 30 mera Roback, Conchapelopia 11 longitibia Goetghebuer, Smittia 38 meridionalis Dendy & Sublette, Lopescladius Oliveira 29 Glyptotendipes 47 Lordella Sather & Sublette 36 Meropelopia Roback 12 lucens (Zetterstedt), Acricotopus 20 Merotanypus Roback ...... 9 luciae LeSage, Cricotopus z5 Mesocricolopus Brundin 29 lucifer (Johannsen), Dicrotendipes 46 Mesopelopia Roback ...... 11 lunatus Sether & Sublette, Mesosmiltia Brundin 29 Pseudorthocladius 36 Metrlocnemus Wulp 30 lundbeckii (Johannsen), Parametriocnemus 34 Microchironomus Kieffer 48 Lundstroemia Kieffer 56 Microcricotopas Thienemann & Harnisch 30 lundstromi Goetghebuer, Procladius 16 Micropsectra Kieffer 56 lupus Willassen, Diamesa I7 Microtendipes Kieffer 48 lurida Beck & Beck, Larsia t2 middlekauffi Townes, Pseudochironomus 55 lurida (Garrett), Boreoheptagyia I6 militaris Johannsen, Chironomus 43 luskl Grodhaus, Synendotendipes 54 milleri (Townes), Dicrotendipes 46 lutosopra (Garrett), Odontomesa 20 millrockensis Caldwell, Paracricotopus 33 lutosus (Townes), Stictochironomus

N o naelis Roback, Coelotanypus 8 obediens (Johannsen), Phaenopsectra 50 naevus (Mitchell), Stictochironomus 54 oblidens (Walker), Orthocladius 3L nais (Townes), Paracladopelma 50 obreptus (Townes), Demeijerea 45 Nanocladius Kieffer 30 obriaini Hayes & Murray, Tokunagaia 39 nansenl Kieffer, Orthocladius 31 obscurifuscus Sublette & Sublette, nanseni (Kieffer), Pseudosmittia a! Cricotopus 25 nanseni (Kieffer), Smittia J6 obscuripes (Holmgren), Metriocnemus 30 n anus (Llndbeck), Pa rametriocnemus J4 obscurus (Johannsen), Cardiocladius 2l narda Roback, Krenopelopia 1,2 obsepta (Webb), Zalutschia 40 nasthecus Sather, Paraphaenocladius 34 obtusilobus (Malloch), Cryptochironomus 44 Natarsia Fittkau 10 obtusum Townes, Polypedilum )l NATARSIINI 10 obumbratus Johannsen, Orthocladius 3L nalchitocheae (Sublette), Einfeldia 46 obvius (Walker), Psectrocladius 35 natvigi (Goetghebuer), Hydrobaenus 27 occidentalis (Coquillett), Prodiamesa 20 nalvigi (Goetghebuer), Paratanytarsus 57 oceanicus (Packard), Cricotopus 25 nearctic a Brundin, Lasiodiamesa ,l ochrealus (Townes), Chironomus nearctica Sether, Odontomesa 19 oconeensis Sether, Compteromesa . t9 needhamii (Johannsen), Polypedilum 52 octomerus Srether, Hydrobaenus 28 nemorosus Edwards, Tanytarsus 59 octopunctatus (Loew), Polypedilum 52 neofl avellus Malloch, Tanytarsus 59 Odontomesa Pagast 19 neomodestus (Malloch), Dicrotendipes 46 Oeklandia Kieffer 56 neopilosella Beck & Beck, Labrundinia . . . . L2 ogemawi Roback, Trissopelopia 74 neopunctipennis Sublette, Tanypus I6 okoboji (Walley), Telopelopia I4 nephopterus (Mitchell), Srenochironomus 53 oldenbergi Goetghebuer, Endochironomus 46 nereis (Townes), Paracladopelma 50 olivacea (Meigen), Prodiamesa 20 nerius (Curran), Compterosmittia zz oliveri Se,ther, Chaetocladius ')) nervosus (Staeger), Dicrotendipes 46 oliveri Srether, Heterotrissocladius 27 netta Townes, Pseudochironomus 55 oliverl Soponis, Orthocladius nietus Roback, Procladius 15 Oliveria Sather JI niger Andersen, Tanytarsus 59 Oliveridia Sather JI niger Borkent, Stenochironomus )5 olivetus Boesel, Cricotopus 'r< nigra Brundin, Parakiefferiella . . J5 Omisus Townes 49 nigra Roback, Ablabesmyia 10 ontario (Walley), Polypedilum 51 nigricans (Johannsen), Endochironomus 46 onteona Roback, Diamesa I7 nigripila (Johannsen), Micropsectra 56 ophioides Townes, Polypedilum 52 nigrilibla Walker, Chironomus 43 orbicus (Townes), Chernovskiia 4l nigriturn Townes, Polypedilum fl Oreadomyia Kevan & Cutten-Ali-Khan 3l nigri(us Malloch, Orthocladius 5Z ornata Beck & Beck, Ablabesmyia 10 nigrohalterale (Malloch), ornatipes (Kieffer), Stictochironomus

par (Coquillett), Xylotopus 39 pellucidus (Walker), Rheotanytarsus )/ par (Johannsen), Xylotopus 40 Pelopia Meigen I6 Paraboreochlus Thienemann . 7 penicillatus (Goetghebuer), Paratanytarsus 57 Parachaelocladius Wiilker JJ Pentaneura Philippi IJ Parachironomus Lenz 49 PENTANEURINI . . 10 Paracladlus Hirvenoja JJ Pentapedilum Kieffer 51 Paracladopelma Harnisch 49 pentatoma Kieffer, Zavrelia 59 Paraclunio Kieffer 7 perda (Roback), Rheopelopia l4 Paracrlcotopus Thienemann & Harnisch JJ peregrinus (Edwards), Parochlus 8 paradorenus (Roback), Orthocladius aa perennls (Meigen), Chaetocladius 1., parafulvus (Beck & Beck), perennis Sather, Heterotanytarsus 27 Cryptochironomus 44 perfuscus Malloch, Metriocnemus 30 parafuscatus Sublette & Sublette, perpulchra (Mitchell), Stelechomyia 53 Cricotopus 25 persimilis (Johannsen), Boreochlus 7 paragrelis Roback, Procladius 15 pertinax (Garrett), Pseudodiamesa Id parajanta Roback, Ablabesmyia 10 Petalopholeus Borkent 53 Parakiefferiella Thienemann 5J Phaenopseclra Kieffer 50 Paralaulerborniella Lenz 50 philosphagnos Beck & Beck, Ablabesmyia 10 Paralimnophyes Brundin 34 Phycoidella Sather 20 paramaculipennis (Roback), Rheopelopia . . l4 pica Townes, Omisus 49 paramelanops Roback, Conchapelopia 11 piclpes (Meigen), Metriocnemus 30 Paramerina Fittkau IJ pictipennis (Zetterstedt), Psectrotanypus 9 Parametriocnemus Goetghebuer 34 plclulus (Meigen), Stictochironomus

polaris Kieffer, Psectrocladius 36 quadralus Sublette, Tanytarsus 59 polaris (Kieffer), Smittia 38 quadridentatus Brundin, Tanytarsus 59 polaris Kieffer, (Syndiamesa) \9 quadrinodosus Hirvenoja, Paracladius ...... 33 polila (Malloch), Micropseclra . 56 quadripunctatum (Malloch), Hyporhygma . . . . 47 politus (Coquillett), Cricotopus quagga (Townes), Stictochironomus 54 polymorpha Andersen, Smittia 38 Polypedilum Kieffer )l R polytomus (Kieffer), clyptotendipes . . . 47 RadotanypusFittkau &Murray ...... 9 ponderosus (Sublette), Cryptochironomus 44 ramosusSather,Protanypus ...... 19 posticalis (Lundbeck), Trichotanypus 8 ramus Mason, Cryptochironomus 44 potamogeti (Townes), Parachironomus 49 ranota Webb, stempellina Potthastia Kieffer 18 Raphidocladius Sather 27 prlor Butler, Chironomus rasha Roback, Ablabesmyia 10 PROCLADIINI .... 74 rawsoniBrundin,Lasiodiamesa ...... 7 Procladius Skuse 15 recens Sublette, Tanytarsus 59 Prodiamesa Kieffer 20 recisusSether,Limnophyes ...... 29 PRODIAMESINAE . T9 rectilobus Sather & Sublette. profusa (Townes), Phaenopsectra 51 Pseudorthocladius . 36 prolilobata Sether, Monodiamesa 19 rectinervis (Kieffer), Nanocladius 30 prolixa Srther, Mesosmittia 29 recurvata Goetghebuer, Micropsectra 56 prolongatus Roback, Procladius 15 redeuns Walker, Chironomus 43 prolongatus (Kieffer), Limnophyes 29 rempelii Thienemann, Chironomus Aa Prosmittia Brundin 36 /emus Sublette, Cricotopus 1A PROTANYPINI l9 Reomyia Roback LJ Prolanypus Kieffer I9 retifinis Sather, Brillia 2l Protenthes Johannsen 16 rex Hauber, Pseudochironomus 55 protextus (Townes), Tribelos

scitulus (Coquillett), Xylotopus 39 rtictus (Meigen), Stictochironomus 54 scopula (Townes), Axarus 4t Sticlochironomus Kief fer 53 scutellata Winnertz, Corynoneura 23 stigmaterus Say, Chironomus 4J sedna (Oliver), Micropsectra )/ Stilocladius Rossaro 38 semicirculatus Sather, Psectrocladius 35 striatus (Malloch), Rheocricotopus 5I seminole Townes, Glyptotendipes 47 stygius Townes, Microtendipes 48 semivirens (Kieffer), Synorthocladius 38 stylatus (Kieffer), Parametriocnemus J+ senata (Walley), Hayesomyia . . . . 11 stylifera (Johannsen), Cryptochironomus 44 s enex (J ohannsen), Acricotopus 20 S tylo tanytarcus Kieffer JI senilis (Johannsen), Glyptotendipes subaequalis (Malloch), Paratendipes 50 septris Roback, (Trichocladius) . . . 40 subaterrima (Malloch), Parakiefferiella JJ sequax (Garrett), Pagastia 18 subalrum Grodhaus, Tribelos 54 sequoiaensis (Sublette), Larsia l2 subcincla (Townes), Apedilum 40 sera Roback, Brillia . LL subcinctum Townes, Apedilum 40 Sergenlia Kieffer 52 subcultellatum Sublette, Polypedilum

tenuis (Meigen), Paratanytarsus 57 tuberculala Srcther, Monodiamesa t9 lesta Roback, Paramerina IJ luberculatus Caldwell, Rheocricotopus 5t testaceus Townes, Glyptotendipes 4'1 luberculatus (Townes), Chironomus 43 Tethymyia Wirth . 38 tumidistylus Sather, Hydrobaenus 28 tethys (Townes), Beckidia 4T turp is (Zetterstedt), Prodiamesa 20 Thalassomya, Schiner 7 luxis Curran, Chironomus 43 Thalassosmittia Strenzke & Remmert 38 Tvelenia Kieffer 39 thanatogralus Epler, Dicrotendipes 46 tylus (Townes), Saetheria 52 thermophilus Townes, Paratendipes 50 thienemannl (Goetghebuer), Mesocricotopus 29 U lhienemanni Kieffer, Orthocladius 37 unacus Townes, Glyptotendipes ...... 47 Thienemannia Kieffer 39 undine (Townes), Paracladopelma . 50 Thienemanniella Kieffer 39 unguiculatus (Malloch), Stictochironomus 54 Thienemannimyia Fittkau I4 unicalcar (Srcther), Eukiefferiella 26 thoracicus (Loew), Clinotanypus 8 unicolor (Walker), Hayesomyia 11 thryptica (Sublette), Zavrelimyia l4 unicolor (Wulp), Parachironomus ...... 49 libialis (Meigen), Cricotopus 24 unictus Townes, Stenochironomus 53 tibialis (Sraeger), Parochlus 8 unidenlatus Sether & Schnell, Rheocricotopus 37 tibialis Webb, Tanytarsus 59 unimaculatus Loew, Chasmatonotus 22 tillandsia Beck & Beck, Monopelopia . . . . 13 uniserratus Sather & Sublette, lincta Roback, Xenopelopia I4 Pseudorthocladius 36 Tokunagaia Sether 39 univiltatus Coquillett, Chasmatonotus 22 tora Srether, Mesosmittia 29 Unniella Sather 39 tori Sdwedal, Micropsectra 57 ursinus (Holmgren), Metriocnemus 30 tornetraeskensis (Edwards), Zalutschia 40 arsus (Kieffer), Diamesa 17 torulata Sather, Parakiefferiella JJ usmaensis Pagast, Tanytarsus 59 lotifuscus Sublette, Stenochironomus 53 utahensis Malloch, Chironomus 43 traenis (Roback), Bryophaenocladius 2l travlsl Sather, Hydrobaenus 28 v tremulus (Linnaeus), Cricotopus 24 vancouveri Michailova & Fischer, triannulatus (Macquart), Cricotopus .A Chironomus 43 triannulalus Srether, Psilometriocnemus 3t varelus (Roback), Tanytarsus ...... 59 triappendiculard (Goerghebuer), varlabilis (Staeger), Halocladius 27 Pseudosmittia 37 varipennis (Coquillett), Zavreliella 55 Tribelos Townes

vockerothlHansen,Diamesa ..... 18 vockerothl Rasmussen, Chironomus ...... 42 vockerothiSrether,Zalutschia .... 40 vulneratus (Zetterstedt), Demicryptochironomus ...... 45 w walleyiTownes,Polypedilum ..... 51 waltliiMeigen, (Diamesa) ...... 19 wartinbei(Roback),Reomyia .... 13 whltseli Sublette & Sublette, Chironomus . . 43 wienslSrether, Orthocladius .....32 wilhmiRoback,Procladius ...... 15 wingoi Sather & Sublette, Pseudorthocladius 36 winnelll Jackson, Paracladopelma . . . . 50 wirlhi Roback, Clinotanypus 8 wirlhl Sather & Sublette, Georthocladius . . 26 WlrthiellaSublette .....48 woodi Borkent, Stenochironomus ...... 53 x xantha(Roback),Micropsectra ...57 xanthusSublette,Tanytarsus ..,.. 59 xena (Roback), Thienemanniella ...... 39 XenochironomusKieffer ...... 54 xenolabis (Kieffer), Xenochironomus . . . . . 54 XenopelopiaFittkau ....14 Xestochironomus Sublette & Wirth ...... 54 xcthis (Roback), Zalutschia ...... 40 XylolopusOliver ...... 39

Y yaquina Cranston & Judd, Metriocnemus 30 z ZalutschiaLipina ...... 40 zalulschicolaLipina,Zalutschia ... 40 zavrellPagast,Sympotthastia ..... 19 Zavrelia Kieffer ...... 59 ZavrellellaKieffer ...... 55 ZavrelimyiaFittkau .....14 zetterstedtlBrundin, Psectrocladius . . . 35 zonopterus (Mitchell), Stenochironomus 53