Trichodesma Zeylanicum

Total Page:16

File Type:pdf, Size:1020Kb

Trichodesma Zeylanicum FACTSHEETS FOR FARMERS Created in Malawi, February 2016 www.plantwise.org Trichodesma zeylanicum Recognize the problem Flowering plant of African borage. Family: Boraginaceae (comfrey family) (Photo by Mark Marathon, Wikimedia) Common names: African borage, late weed, camel bush, cattle bush, jilarga, rough bluebell Chichewa: Chilungumwamba An annual herbaceous plant (30-40 cm tall) or occasionally a shrub (up to 100 cm high); with many side branches; bristly and hairy with a robust tap root; spiny hairs tend to break off in the skin when handled. Leaves: Green, covered in spiny hairs giving them a sandpapery feel; with conspicuous veins below; narrow and tapering to point to egg-shaped (3.5-10.5 cm long and 0.7-2 cm wide), narrowed towards the base, placed opposite each other near the base of the stem and alternate higher on the plant, inconspicuous leaf stalk. Flower of African borage. (Photo by Flowers: Pale blue (occasionally white), with a white centre, single (15-20 mm across) Pete Hill, www.flickr.com) with long pedicel, hanging from long flower stalks. Fruits: Capsules (dry fruit that opens at maturity), brown, four chambered, each containing one seed. Seeds: 3-angled, mottled with grey and brown on the outer surface (3-4 mm long). Background Origin: East and Southern Africa, Asia and Australasia. Introduction: Accidentally as a contaminant. Habitat: Tropical regions. Spread: Spread by animals and whirl wind. Invades: Crop fields, disturbed ground, roadsides and sandy river beds. Impacts: The weed competes with and replaces native vegetation. In crops, particularly beans, groundnuts, maize, cotton and sorghum, it competes in the latter stages of crop growth with serious infestations preventing harvest. Seeds germinate in very large numbers after the first weeding, forcing farmers to weed twice more in the growing season to ensure good yields. In parts of Zimbabwe, weeding African borage (together with a few other weedy species) increases yields by 52 and 117 kg/ha in cotton and maize, respectively. Land preparation is made considerably more difficult when this weed is present, due to the bulbous-based spiny hairs lodging in the skin during manual control activities. It is apparently also toxic to livestock. Scientific name(s) > Trichodesma zeylanicum The recommendations in this factsheet are relevant to: All Countries Authors: CABI. Edited by David Kamangira, Elisa Mazuma, Rose Mkandawire, Amos Ngwira Chitedze Agricultural Research Station Edited by Plantwise. tel: +265 1 767222 email: [email protected] Plantwise is a global initiative led by CABI Lose Less, Feed More ©CAB International. Published under a CC-BY-SA 4.0 licence. 110En.
Recommended publications
  • Additional Land South of the Gas Treatment Plant Vegetation and Flora Survey
    Additional Land South of the Gas Treatment Plant Vegetation and Flora Survey October 2013 Prepared for Chevron Australia Pty Ltd Astron Environmental Services 129 Royal Street East Perth WA 6004 Phone: (08) 9421 9600 Fax: (08) 9421 9699 Report Reference: 2609-13-BSR-1Rev0_131128 Email: [email protected] Doc. ID: G1-NT-REP00000220 Doc. ID: G1-NT-REP00000220 Additional Land South of the Gas Treatment Plant Vegetation and Flora Survey Prepared for Chevron Australia Pty Ltd Job Number: 2609-13 Reference: 2609-13-BSR-1Rev0_131128 Revision Status Rev Date Description Author(s) Reviewer A 01/11/2013 Draft Issued for Client Review N. Cadd J. Kruger B 08/11/2013 Revised Draft Issued for Client Review N. Cadd J. Kruger C 12/11/2013 Revised Draft Issued for Client Review N. Cadd V. Clarke 0 28/11/2013 Final Issued for Information N. Cadd V. Clarke Approval Rev Date Issued to Authorised by Name Signature A 01/11/2013 A. Smith/S. Finn S. Pearse B 08/11/2013 A. Smith/S. Finn S. Pearse C 12/11/2013 A. Smith/S. Finn S. Atkinson 0 28/11/2013 A. Smith/S. Finn S. Pearse © Copyright 2012 Astron Environmental Services Pty Ltd. All rights reserved. This document and information contained in it has been prepared by Astron Environmental Services under the terms and conditions of its Doc.contract ID: with G1-NT-REP00000220 its client. The report is for the clients use only and may not be used, exploited, copied, duplicated or reproduced in any form or medium whatsoever without the prior written permission of Astron Environmental Services or its client.
    [Show full text]
  • Ethnobotanical Study on Wild Edible Plants Used by Three Trans-Boundary Ethnic Groups in Jiangcheng County, Pu’Er, Southwest China
    Ethnobotanical study on wild edible plants used by three trans-boundary ethnic groups in Jiangcheng County, Pu’er, Southwest China Yilin Cao Agriculture Service Center, Zhengdong Township, Pu'er City, Yunnan China ren li ( [email protected] ) Xishuangbanna Tropical Botanical Garden https://orcid.org/0000-0003-0810-0359 Shishun Zhou Shoutheast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Liang Song Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Intergrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Ruichang Quan Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences & Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Huabin Hu CAS Key Laboratory of Tropical Plant Resources and Sustainable Use, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences Research Keywords: wild edible plants, trans-boundary ethnic groups, traditional knowledge, conservation and sustainable use, Jiangcheng County Posted Date: September 29th, 2020 DOI: https://doi.org/10.21203/rs.3.rs-40805/v2 License: This work is licensed under a Creative Commons Attribution 4.0 International License. Read Full License Version of Record: A version of this preprint was published on October 27th, 2020. See the published version at https://doi.org/10.1186/s13002-020-00420-1. Page 1/35 Abstract Background: Dai, Hani, and Yao people, in the trans-boundary region between China, Laos, and Vietnam, have gathered plentiful traditional knowledge about wild edible plants during their long history of understanding and using natural resources. The ecologically rich environment and the multi-ethnic integration provide a valuable foundation and driving force for high biodiversity and cultural diversity in this region.
    [Show full text]
  • Biosystematic Studies in Heliotropium Indicum, Trichodesma Indicum and T
    Int.J.Curr.Microbiol.App.Sci (2016) 5(4): 720-729 International Journal of Current Microbiology and Applied Sciences ISSN: 2319-7706 Volume 5 Number 4 (2016) pp. 720-729 Journal homepage: http://www.ijcmas.com Original Research Article http://dx.doi.org/10.20546/ijcmas.2016.504.083 Biosystematic Studies in Heliotropium indicum, Trichodesma indicum and T. zeylanicum of Boraginaceae Ashwini Kumar1 and Baidyanath Kumar2* 1PG Dept. of Botany, Gaya College, Gaya, India 2PG Department of Biotechnology, College of Commerce, Patna, Magadh University, India *Corresponding author ABSTRACT In the present observation biosystematics of three species of Boraginaceae viz., of Heliotropium indicum, Trichodesma indicum and Trichodesma zeylanicum was K eywo rd s studied. All the three species showed a marked difference in leaf and inflorescence Biosystematics, morphology, stomatal behaviour, epidermal trichome, pollen grain characteristics Heliotropium and stem anatomy, hypodromous and brochidodromous venation of their leaves. indicum. Leaves were landeolate to ovate. The inflorescence colour was pale blue to white. Trichodesma Length of inflorescence was found to be 4.0 to 6.5. The stomatal Index in these indicum, three species varied greatly. T. indicum and T. zeylanicum showed anomocytic Trichodesma stomata with 16mm in length and 12mm width and Stomatal Index was 34.88 and zeylanicum , 44.68 respectively. Heliotropium indicum exhibited anisocytic stomata which were Numerical analysis. 12mm long and 10mm wide. The Stomatal Index was 32.65. All the three species showed the presence of unicellular glandular hairs which differed in forms. All the three species showed tricolpate pollen grains, but differed in their form and size. Article Info All the three species showed more or less similar stem anatomical features with differences only in their hypodermal layers.
    [Show full text]
  • Palatability of Plants to Camels (DBIRD NT)
    Technote No. 116 June 2003 Agdex No: 468/62 ISSN No: 0158-2755 The Palatability of Central Australian Plant Species to Camels Dr B. Dorges, Dr J. Heucke, Central Australian Camel Industry Association and R. Dance, Pastoral Division, Alice Springs BACKGROUND About 600,000 camels (Camelus dromedarius) are believed to inhabit the arid centre of Australia, mainly in South Australia, Western Australia and the Northern Territory. Most of these camels are feral. A small camel industry has developed, which harvests selected animals for domestic and export markets, primarily for meat. Camels can eat more than 80% of the common plant species found in Central Australia. Some plant species are actively sought by camels and may need to be protected. METHOD Observations of grazing preferences by camels were made periodically for up to 12 years on five cattle stations in Central Australia. Where camels were accustomed to the presence of humans, it was possible to observe their grazing preferences from a few metres. Radio transmitters were fitted on some camels for easy detection and observation at any time. These evaluations were used to establish a diet preference or palatability index for observed food plants. Table 1. Palatability index for camels Index Interpretation 1 only eaten when nothing else is available 2 rarely eaten 3 common food plant 4 main food plant at times 5 preferred food plant 6 highly preferred food plant 7 could be killed by camel browsing More information can be obtained from the web site of the Central Australian Camel Industry Association http://www.camelsaust.com.au 2 RESULTS Table 2.
    [Show full text]
  • Pyrrolizidine Alkaloids in Plants Used in the Traditional Medicine of Madagascar and the Mascarene Islands
    REVIEW Pharmazeutisches Institut der Rheinischen Friedrich-Wilhelms-Universität, Bonn, Germany Pyrrolizidine alkaloids in plants used in the traditional medicine of Madagascar and the Mascarene islands E. Roeder, H. Wiedenfeld Received April 26, 2011, accepted May 20, 2011 Prof. Dr.Dr.h.c. Erhard Roeder,Dr.Helmut Wiedenfeld, Pharmazeutisches Institut der Universität, An der Immenburg 4, D-53121 Bonn, Germany [email protected], [email protected] Dedicated to Prof. Dr. med. Dr. rer. nat. Drs. h.c. Ernst Mutschler on the occasion of his 80th birthday Pharmazie 66: 637–647 (2011) doi: 10.1691/ph.2011.1572 Pyrrolizidine alkaloids (PAs) can be hazardous to the health of humans and animals. Although their toxicity has been known for a long time, PA containing plants are still in use in many traditional medicines. Traditional healing systems have become of increasing interest as many people believe that they can be used without any risk and side effects. This also applies to the traditional medicine of Madagascar and the Mascarene island (Mauritius, Réunion, Rodriguez). Recent literature reports have recommended this traditional medicine because of its good efficacy and pharmacological properties. However, several plants are listed there which have already been described to contain toxic PAs or are suspected of containing them. 1. Introduction play a growing role in the treatment of diseases in the form of self-medication. Pyrrolizidine alkaloids (PAs)are naturally occurring compounds In addition, increased tourism has made it possible that people which are contained in about 3% of all flowering plants (Bull come into contact with different traditional healing systems all et al.
    [Show full text]
  • Two Newly Naturalized Plants of the Boraginaceae in Taiwan: Trichodesma Indicum (L.) Lehm
    Taiwan J For Sci 29(2): 149-56, 2014 149 Research note Two Newly Naturalized Plants of the Boraginaceae in Taiwan: Trichodesma indicum (L.) Lehm. and Trichodesma zeylanicum (Burm.f.) R. Br. Chiu-Mei Wang,1) Kun-Cheng Chang2,3) 【Summary】 Trichodesma indicum (L.) Lehm., an herb native to Asia and Africa, and Trichodesma zeylani- cum (Burm.f.) R. Br., one native to Africa, tropical Asia, and Australasia, were recently found in west-central Taiwan. Both species are considered to be newly naturalized. This report gives taxo- nomic descriptions, illustrations, and distribution maps. Photographs are also provided to help with identification. Key words: Trichodesma indicum, Trichodesma zeylanicum, Boraginaceae, naturalized plant, Tai- wan. Wang CM, Chang KC. 2014. Two newly naturalized plants of the Boraginaceae in Taiwan: Trichodesma indicum (L.) Lehm. and Trichodesma zeylanicum (Burm.f.) R. Br.. Taiwan J For Sci 29(2):149-56. 1) Department of Biology, National Museum of Natural Science, 1 Guancian Rd., Taichung 40453, Taiwan. 國立自然科學博物館生物學組,40453台中市館前路1號。 2) Department of Forestry and Natural Resources, National Chiayi Univ., 300 Xuefu Rd., Chiayi City 60004, Taiwan. 國立嘉義大學森林暨自然資源學系,60004嘉義市鹿寮里學府路300號。 3) Corresponding author, e-mail:[email protected] 通訊作者。 Received September 2013, Accepted February 2014. 2013年9月送審 2014年2月通過。 150 Wang and Chang─Newly naturalized plants of the Boraginaceae in Taiwan 研究簡報 兩種新歸化的紫草科植物: 印度碧果草及斯里蘭卡碧果草 王秋美1) 張坤城2,3) 摘要 原產於熱帶亞洲及非洲的印度碧果草與斯里蘭卡碧果草(亦見於澳洲)最近發現於臺灣中部。這兩 種紫草科植物,均為新近歸化於臺灣的外來種。本文提供此兩物種的分類特徵描述,手繪圖,以及在 臺灣的分布點以供區分鑑定。 關鍵詞:印度碧果草、斯里蘭卡碧果草、紫草科、歸化植物、臺灣。 王秋美、張坤城。2014。兩種新歸化的紫草科植物:印度碧果草及斯里蘭卡碧果草。台灣林業科學 29(2):149-56。 INTRODUCTION The Boraginaceae comprises about 156 lia (Weigend et al. 2013), and is characterized genera and 2500 species which are distrib- by the connective of the anthers prominently uted in temperate and tropical regions, and subulate and spirally twisted.
    [Show full text]
  • Approved Conservation Advice for the Monsoon Vine Thickets on the Coastal Sand Dunes of Dampier Peninsula
    Environment Protection and Biodiversity Conservation Act 1999 (EPBC Act) Approved Conservation Advice for the Monsoon vine thickets on the coastal sand dunes of Dampier Peninsula 1. The Threatened Species Scientific Committee (the Committee) was established under the EPBC Act and has obligations to present advice to the Minister for Sustainability, Environment, Water, Population and Communities (the Minister) in relation to the listing and conservation of threatened ecological communities, including under sections 189, 194N and 266B of the EPBC Act. 2. The Committee provided its advice on the Monsoon vine thickets on the coastal sand dunes of Dampier Peninsula ecological community to the Minister as a draft of this approved conservation advice. In 2013, the Minister accepted the Committee’s advice, adopting it as the approved conservation advice. 3. The Minister amended the list of threatened ecological communities under section 184 of the EPBC Act to include the Monsoon vine thickets on the coastal sand dunes of Dampier Peninsula ecological community in the endangered category. It is noted that the ecological community is also listed as the Monsoon vine thickets on the coastal sand dunes of Dampier Peninsula on the Western Australian list of threatened ecological communities endorsed by the Western Australia Minister for the Environment. 4. The nomination and a draft description for this ecological community were made available for expert and public comment for a minimum of 30 business days. The Committee and Minister had regard to all public and expert comment that was relevant to the consideration of the ecological community. 5. This approved conservation advice has been developed based on the best available information at the time it was approved; this includes scientific literature, advice from consultations, existing plans, records or management prescriptions for this ecological community.
    [Show full text]
  • Volume 2 Book with NUMBER 1-402
    FLORA OF KARNATAKA A Checklist Volume - 2 : Gymnosperms & Angiosperms CITATION Karnataka Biodiversity Board, 2019. FLORA OF KARNATAKA, A Checklist, Volume – 2: Gymnosperms and Angiosperms. 1 - 1002 (Published by Karnataka Biodiversity Board) Published: December, 2019. ISBN - 978-81-939228-1-1 © Karnataka Biodiversity Board, 2019 ALL RIGHTS RESERVED • No part of this book, or plates therein, may be reproduced, stored in a retrieval system or transmitted, in any form or by any means, electronic, mechanical, photocopying recording or otherwise without the prior permission of the publisher. • This book is sold subject to the condition that it shall not, by way of trade, be lent, re-sold, hired out or otherwise disposed of without the publisher’s consent, in any form of binding or cover other than that in which it is published. • The correct price of this publication is the price printed on this page. Any revised price indicated by a rubber stamp or by a sticker or by any other means is incorrect and should be unacceptable. DISCLAIMER • THE CONTENTS INCLUDING TEXT, PLATES AND OTHER INFORMATION GIVEN IN THE BOOK ARE SOLELY THE AUTHOR’S RESPONSIBILITY AND BOARD DOES NOT HOLD ANY LIABILITY. PRICE: ` 2000/- (Two thousand rupees only). Printed by : Peacock Advertising India Pvt Ltd. # 158 & 159, 3rd Main, 7th Cross, Chamarajpet, Bengaluru – 560 018 | Ph: 080 - 2662 0566 Web: www.peacockgroup.in FOREWORD About 60% of the Western Ghats is present in Karnataka State, with this large part of the peninsular green cover coupled with inland forest plateau enriched by the seven river systems, the State of Karnataka showcases a wider floral wealth harboring highest number of apex predators all of which is conserved by a framework of various statutes.
    [Show full text]
  • Bardi Plants an Annotated List of Plants and Their Use
    H.,c H'cst. /lust JIus lH8f), 12 (:J): :317-:359 BanE Plants: An Annotated List of Plants and Their Use by the Bardi Aborigines of Dampierland, in North-western Australia \!o\a Smith and .\rpad C. Kalotast Abstract This paper presents a descriptive list of the plants identified and used by the BarcE .\borigines of the Dampierland Peninsula, north~\q:stern Australia. It is not exhaust~ ive. The information is presented in two wavs. First is an alphabetical list of Bardi names including genera and species, use, collection number and references. Second is a list arranged alphabetically according to botanical genera and species, and including family and Bardi name. Previous ethnographic research in the region, vegetation communities and aspects of seasonality (I) and taxonomy arc des~ cribed in the Introduction. Introduction At the time of European colonisation of the south~west Kimberley in the mid­ nineteenth century, the Bardi Aborigines occupied the northern tip of the Dam­ pierland Peninsula. To their east lived the island-dwelling Djawi and to the south, the ~yulnyul. Traditionally, Bardi land ownership was based on identification with a particular named huru, translated as home, earth, ground or country. Forty-six bum have been identified (Robinson 1979: 189), and individually they were owned by members of a family tracing their ownership patrilineally, and known by the bum name. Collectively, the buru fall into four regions with names which are roughly equivalent to directions: South: Olonggong; North-west: Culargon; ~orth: Adiol and East: Baniol (Figure 1). These four directional terms bear a superficial resemblance to mainland subsection kinship patterns, in that people sometimes refer to themselves according to the direction in which their land lies, and indeed 'there are.
    [Show full text]
  • Boraginaceae), and the Phylogeny of Boraginoideae
    !" #$ % " "& '()*"'+ (,-./01 ** -)2'/)*) %*()'-) %%*(* 3443 Dissertation for the Degree of Doctor of Philosophy in Systematic Botany presented at Uppsala University in 2002 Abstract Långström, E. 2002. Systematics of Echiochilon and Ogastemma (Boraginaceae), and the phylogeny of Boraginoideae. Acta Univ. Ups. Comprehensive Summaries og Uppsala Disserta- tions from the Faculty of Science and Technology 693. 34 pp. Uppsala. ISBN 91-554-5257-4. Echiochilon, Ogastemma and Sericostoma are revised resulting in the recognition of 15 spe- cies of Echiochilon and one Ogastemma species. Several species are placed in synonymy and three new species are described, E. baricum, E. callianthum and E. cyananthum. The single species of Sericostoma is shown to be nested within Echiochilon. The plastid atpB gene was sequenced for Echiochilon and Ogastemma from the Old World and Antiphytum from the New World, plus for a selection of 33 other Boraginaceae taxa. They were analysed together with selected outgroup taxa to give a framework of the tribes of Boraginoi- deae. The analysis gave support for establishing the new tribe Echiochileae for Antiphytum, Echio- chilon and Ogastemma, and for merging the traditionally accepted tribe Eritrichieae with Cyno- glosseae. The ITS region was sequenced for all but one species of Echiochilon and for representa- tives of Antiphytum and Ogastemma. Phylogenetic analysis of Echiochilon revealed that the strongly zygomorphic-flowered species form a paraphyletic group. The morphological data gave results fairly congruent with the ITS phylogeny. Biogeographic interpretations of the ITS and atpB phylogenies indicated a trans-Atlantic dispersal of Antiphytum as the most plausible explanation to the Old/New World disjunction. Analyses using DIVA (Dispersal Vicariance Analysis) of the distributions of the Echiochilon spe- cies indicated an ancestor to Echiochilon with a wide distribution over northern Africa and Arabia to India.
    [Show full text]
  • Laurent Garcin, Mdfrs
    LAURENT GARCIN, M.D. F.R.S.: A FORGOTTEN SOURCE FOR N. L. BURMAN’S FLORA INDICA (1768) ALEXANDRA COOK1 Abstract. Laurent Garcin (ca. 1681–1751), a Dutch East India Company ship’s surgeon, Fellow of the Royal Society and correspond- ing member of the Académie royale des sciences (Paris), has largely vanished from the annals of botanical and medical science. Yet data presented in this article demonstrate that ca. 1740 he gave some or all of his plant collections from his Asian travels in the 1720s to J. Burman, a correspondent in Amsterdam. Those collections in turn greatly enriched Flora Indica by N. Burman (hereafter Burman fil.) to the tune of 98 specimens. Burman’s work is an important historical source for the botany not only of modern-day India, as the title suggests, but also of Sri Lanka, Indonesia and Iran—the “Indies” as they were understood in the eighteenth century. So far only a handful of Garcin’s specimens have come to light (G-Burman). These few extant specimens testify to Garcin’s collecting zeal and keen eye for materia medica. Keywords: Asia, Johannes Burman, Cinnamomum, Garcinia, materia medica, Salvadora Laurent Garcin (ca. 1681–1751),2 a Franco-Swiss botanist, of the Swiss Confederation) (Chambrier 1900: 251; Bridel Dutch East India Company (hereafter VOC) ship’s surgeon, 1831: 99). Upon joining the VOC Garcin himself reported Fellow of the Royal Society and corresponding member that he came from Nyon, a town in the canton of Vaud not of the Académie royale des sciences (Paris), has largely far from Geneva.
    [Show full text]
  • Boraginaceae
    J. Adelaide Bot. Gard. (15)2: 93-99(1993) NEW TAXA AND COMBINATIONS IN THE BORAGINACEAE B.R.Randell State Herbarium, Botanic Gardens, North Terrace, Adelaide, South Australia 5000 Albstract Two new taxa, Ehretia grahamii Randell and Trichodesma zeylanicum var. grandiflorum Randell, as well as a new combination Ehretia saligna var. membranifolia (R.Br.) Randell, are here published. In addition, the nomenclature of Australian material of Anzsinckia is discussed. 1. Ehretia saligna R.Br. var. membranifolia (R.B .) Randell, comb. nov. Basionym: Ehretia membranifolia R.Br., Prodr. 497 (1810). Type: Broad Sound, RBrown 102, 25.ix.1802, n.v. Shrub or tree to 8 m tall. Leaves: blades elliptic, broadly lanceolate to ovate, not falcate, 4-15 cm long, 6-60 mm wide, rarely drooping, ± chartaceous, green; petioles 5-20 mm long. Inflorescences terminal on short branchlets, spreading. Flowers: specimens either with hermaphrodite flowers, with exserted stigma but anthers almost enclosed in corolla tube; or with male flowers, having exserted anthers but ovary (aborted and drying black), style and stigma enclosed in corolla tube. Fruit red, succulent, drying black, splitting into 4 1-seeded nutlets. Peach bush. Distribution and ecology Occurs in northem Northern Territory, north-western Queensland and along the eastern highlands from northem Queensland to central New South Wales; usually restricted to vine scrubs, fringing forests or rainforest margins, in contrast to drier habitats occupied by the type variety. Notes Robert Brown described two species, E. saligna and E. membranifolia, based largely on leaf morphology. Bentham (1868) accepted these, but put emphasis on floral differences, describing one species with exsertal anthers, the other with enclosed anthers.
    [Show full text]