Acta Geologica Polonica , Vol. 58 (2008), No. 4, pp. 407-423

Barremian and Early Aptian ammonites from the Godula Facies of the Silesian Unit in the Outer Western Carpathians, Czech Republic

ZDENĚK VAŠÍČEK

Institute of Geonics of Academy of Sciences of the Czech Republic, Studentská 1768, 708 00 Ostrava – Poruba, Czech Republic. E-mail: [email protected]

ABSTRACT:

VAŠÍČEK , Z. 2008. Barremian and Early Aptian ammonites from the Godula Facies of the Silesian Unit in the Outer Western Carpathians, Czech Republic. Acta Geologica Polonica , 58 (4), 407-423 . Warszawa.

The Early Cretaceous ammonites from the Hradiště Formation (Wernsdorfer Schichten in original concept) in the Czech part of the Silesian Unit of the Outer (Flysch) Western Carpathians are described. This rich material, collected over more than 40 years, includes many usually fragmentarily preserved ammonites. The majority of specimens are preserved as “half-ammonoids”; a small portion of them have pyritized inner whorls with preserved suture lines. 15 species-level taxa are described, partly in open nomenclature. One species , ? Valdedorsella ho - heneggeri , is described as new ; albeit its generic classification remains uncertain. The ammonites described span the late Early Barremian (Niclesia pulchella ammonite Zone) to basal Aptian ( Deshayesites oglanlensis ammonite Zone).

Key words: Outer Western Carpathians, Silesian Unit, Lower Cretaceous, Ammonites, Biostratigraphy.

INTRODUCTION (ROTH & M ATĚJKA 1953). In the Czech Republic , the centres of the industry and mining were situated in the The present paper reports on ammonites from the Oder River Basin (e.g. near Frýdlant n. O.); in Poland Lower Cretaceous of the Silesian Unit in the territory of they were in the Vistula River Basin (near Ustron). the Czech Republic . Most of the material was collected UHLIG (1883) was the first worker to monograph in the years 1962-74, primarily from nineteenth century the cephalopods yielded by the associated rocks . How - spoil heaps left during the extraction of low grade, nodu - ever, he did not provide details of the locations of the lar sedimentary iron ores (clay ironstones) . Only excep - mining sites but merely stated the name of the closest tionally are any of these spoil heaps still accessible . The municipality. VAŠÍČEK (1971) plotted on base maps the ammonites were yielded by dark -grey pelitic sediments spoil heaps with Lower Cretaceous ammonites in the of the Hradiště Formation ( VAŠÍČEK 1971, 1972; VAŠÍČEK Czech Republic and pinpointed the sites of mining at & al . 2004). The geographical location and geological that time. setting of ammonite source localities is shown in Text- The material studied herein supplements signifi - fig. 1; more details are to be found in VAŠÍČEK (1971, cantly the knowledge of the overall composition and 1972, 1981a, 1990) and VAŠÍČEK & al . (2004). distribution of Barremian-Early Aptian ammonite fau - Clay ironstone mining spoil heaps were left after nas in the Silesian Unit of the Outer (Flysch) Western the historical iron industry in the Podbeskydí area Carpathians and in the Tethyan Realm. 408 ZDENĚK VAŠÍČEK

Fig. 1. Geographical and geological position of places of occurrence of Lower Cretaceous ammonites referred to in the text

PRESERVATION OF THE MATERIAL suture may , however, also be observed by removing the original phragmocone shell. The most characteristic feature of the majority of The basic elements of the suture lines are desig - the ammonites from the dark grey, usually calcareous nated herein by the usual symbols : E – external lobe, claystones of the upper Hradiště Formation (as defined L – lateral lobe, U – umbilical lobe. Other dimensions by ELIÁŠ & al . 2003) is their half preservation with re - (in mm) are as follows: gard to the plane of symmetry. This is the “half-am - D – shell diameter, H – whorl height, U – umbili - monite preservation” or the “half-ammonoids” of cus width . Also given are simple ratios: H/D – whorl MAEDA & S EILACHER (1996, fig. 13 ). Half-ammonoids height to shell diameter , U/D – and umbilicus width to from the Beskydy Mts. have the original calcareous shell diameter. shells preserved only on the side immersed in the dysoxic sediment. Because of the strong compaction of the clay sedi - TAXONOMIC PART ment , up to as much as two-thirds of the original thick - ness, the majority of the buried shells were subse - The taxonomic classification applied herein follows quently pressed into the surface of the bedding plane. that of the Treatise ( WRIGHT & al . 1996) and of the Fos - In the course of this process, the shells were usually silium Catalogus ( KLEIN 2005, VERMEULEN & K LEIN crushed (see e.g. figs 5 and 6 in VAŠÍČEK 1972) , which 2006). The exceptions are the pulchelliids, which , in ac - is why the original whorl width has not been preserved. cordance with VERMEULEN (1996) and VERMEULEN & A small percentage of the spirally-coiled shells KLEIN (2006), are referred to the superfamily Endemo - have the inner whorls partially pyritized. In contrast to ceratoidea ; and the Macroscaphitidae, which , in accor - other specimens , such shells have their suture lines dance with BUSNARDO (in FISCHER &GAUTHIER 2006), partly preserved, mostly the juvenile sutures. The adult is placed within the superfamily Lytoceratoidea. EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 409

Order Ammonitida AGASSIZ , 1847 cepted in the new Treatise – WRIGHT & al . 1996) and Suborder Phylloceratina ARKELL , 1950 of KAKABADZE (2004) that the genera Costidiscus UH- Superfamily Phylloceratoidea ZITTEL , 1884 LIG , 1882, and Macroscaphites MEEK , 1876, form a di - Family Phylloceratidae ZITTEL , 1884 morphic pair, and that consequently the former is a Subfamily Phyllopachyceratinae COLLIGNON , 1937 synonym of the latter, is not followed herein (see also BUSNARDO in FISCHER & GAUTHIER 2006 , p. 153 ). Genus Phyllopachyceras SPATH , 1925 The hook-like morphotypes of non-uniform size il - lustrated and referred to Macroscaphites can be di - TYPE SPECIES : Ammonites infundibulum D´O R- vided into two basic morphological groups. The first BIGNY , 1841. includes shells with weak umbilical tubercles only or shells that do not bear any tubercles on the ribs. The second group contains shells with one or two rows of Phyllopachyceras vergonsense DELANOY & JOLY , tubercles, including the ventrolateral ones, which never 1995 appear in the first group. The dominant species of the (Pl. 1, Fig. 1 ) first group is Macroscaphites yvani (PUZOS ); it reaches a size of about 100 mm, the diameter of the coiled part 1995. Phyllopachyceras vergonsense nov. sp.; G. D ELANOY being about 45 mm, and it possesses constrictions (see & B. J OLY , pp. 132-133, pl. 2, figs 2, 3 a, b. e.g. UHLIG , 1883) [ Uhlig’s specimens of M. yvani, re - garded by AVRAM (1984) as belonging to the species MATERIAL: Single specimen M5/193 = B13808, M. perforatus , have , in my opinion , nothing in com - from spoil heap M5 near Malenovice. The final half of mon with the smaller shells of M. perforatus AVRAM , the shell comprises the body chamber, with a partly 1984]. corroded ventral area. Besides typical representatives of M. yvani , there occur similar, but markedly smaller, constricted forms, DESCRIPTION: The shell is involute, with a very nar - which are also usually referred to this species (DIM - row umbilicus. The body chamber possesses strong ITROVA , 1967, pl. 11, fig. 5) or to a separate subspecies , main ribs beginning at the umbilicus. The ribs may bi - M. y. striatisulcatus D´O RBIGNY (e.g. AVRAM , 1976, pl. furcate on the outer half of the flanks. One to two thin - 1, fig. 12). These smaller shells may represent micro - ner ribs are inserted between the main ribs, approxi - conchs of M. yvani, as already stated by KILIAN (1913) . mately at the bifurcation points of the main ribs . Secondary ribs may also bifurcate . Macroscaphites cf. abchasiensis M. K AKABADZE , REMARKS . The number of inserted ribs and the bi - 2004 furcation of the ribs are characteristic features of the (Pl. 1, Fig. 2 ) species, distinguishable even in poorly preserved speci- mens . 2004. Macroscaphites abchasiensis M. K AKABADZE , sp. nov .; M. V. K AKABADZE , p. 362, pl. 1, fig. 3 (cum syn.). DISTRIBUTION: Besides the specimen studied, P. 2005. Macroscaphites abchasiensis KAKABADZE ; M. V. K AK - vergonsense is known from a single specimen from the ABADZE in E. V. K OTETISHVILI & al ., p. 292, pl. 47, fig. 2. Gerhardtia sartousiana ammonite Zone of the Late Barremian, in southeast France ( DELANOY & J OLY MATERIAL: Two incomplete shells, in which only a (1995). section of the coiled part with the transition into the shaft is preserved (K3/003 = B13809 and M1/267 = B13810). The size of the specimen illustrated is esti - Suborder Lytoceratina HYATT , 1889 mated to reach about 55 mm. The shells are from the Superfamily Lytoceratoidea NEUMAYR , 1875 spoil heaps of Malenovice (M1) and Kozlovice (K3) Family Macroscaphitidae HYATT , 1900 respectively .

Genus Macroscaphites MEEK , 1876 DESCRIPTION: The shells are evolute, small. The sculpture is composed of simple ribs , somewhat TYPE SPECIES : Scaphites Yvani PUZOS , 1836. stronger on the shaft than on the coiled part and arched convexly towards the aperture. Constrictions do not oc - The opinion of AVRAM (1976, 1978, 1984) ( ac - cur . 410 ZDENĚK VAŠÍČEK

REMARKS: The incomplete specimens studied herein partly preserved sutures (T5/60, T5/333 = B13811, are very close in size and ribbing to the Georgian T5/409 = B13812). All specimens are from the spoil species. The only marked difference is a striking ar - heap T5 and the outcrop T9, near the municipality cuation of the shaft in M. abchasiensis. of Tichá. UHLIG (1883, p. 207) mentioned an interesting specimen from the locality of Veřovice, which reached DESCRIPTION: The shells are semi-involute, with a size of less than 51 mm. He stated that this specimen slightly arched whorl flanks and a relatively wide um - was either a dwarf M. yvani (typical representatives of bilicus. The sculpture is variable . The outer shell sur - which also occur here), an independent species, or an face bears slightly S-shaped main ribs with shallow individual affected by a disease. constrictions on their posterior sides. The main inter- rib spaces bear growth lines (in juveniles) or thin ribs DISTRIBUTION: The Georgian specimens of M. (in adults) , which are sometimes well developed. abchasiensis are from the Lower Aptian ( KAKABADZE Marked constrictions on external moulds correspond 2004) . The spoil heaps, from which the studied mate - to the main ribs and their accompanying slight con - rial comes , yielded Late Barremian and Early Aptian strictions. The constrictions are quite abundant, 8 to 11 ammonites, including M. yvani . per whorl, and start already in juveniles, at a shell di - ameter about 10 mm.

Suborder Ammonitina HYATT , 1889 SUTURE LINE: The most complete outer suture lines Superfamily Desmoceratoidea ZITTEL , 1895 (B13812) are similar to those illustrated by UHLIG Family Desmoceratidae ZITTEL , 1895 (1883) (Text-fig. 2). The suture lines are complex. Subfamily Puzosiinae SPATH , 1922 The lateral lobe is deep, somewhat asymmetrically trifid. The subsequent lobe towards the line of coiling Genus Melchiorites SPATH , 1923 is also trifid . The next umbilical lobe does not have this character any more. Saddles are asymmetrically two- TYPE SPECIES : Ammonites Melchioris TIETZE , 1872. branched.

MEASUREMENTS: The shells are variable not only Melchiorites blayaci (KILIAN , 1900) in sculpture but also in size . The measured values en - (Pl. 1, Figs 3, 4 ; Text-fig. 2 ) compass, on the one hand, biological variability in the species and, on the other hand, the results of shell 1900. Desmoceras Blayaci ; W. K ILIAN in J. B LAYAC , p. 22. deformation. Biological variability is here heavily 1972. Melchiorites blayaci (KILIAN in BLAYAC ); Z. V AŠÍČEK , overprinted by post-mortem deformation. Besides pp. 76 -77 , pl. 13, figs 4, 5, pl. 16, fig. 4 (cum syn.). forms with a whorl height greater than their umbilicus width (see e.g. VAŠÍČEK 1972, p. 77), there also occur MATERIAL: 15 shells, laterally compressed (e.g. shells in which the whorl height and the umbilicus T9/56, T5/260, 314). In three specimens it was pos - width are approximately equal, as e.g. B13812, where sible to expose the pyritic internal mould, with at D = 36 mm, H = 13.0 (0.36) and U = 13.3 (0.37). The diameter of the largest preserved shells is about 45 mm.

REMARKS: The present description amplifies the characteristics of the species given by VAŠÍČEK (1972), which were based mostly on the holotype. The mate - rial studied revealed a considerable variability of M. blayaci in sculpture and measured parameters . Ac - cordingly , in contrast to the synonymy in VAŠÍČEK (1972), it is possible to refer to M. blayaci some of the shells included by UHLIG (1883) in Haploceras char - rierianum. A species close to M. blayaci is M. melchioris (TI- Fig. 2. Melchiorites blayaci (KILIAN ); B13812. The greater part of ETZE , 1872), which is, however, usually preserved as the outer suture line at H = 9.5 mm internal moulds . The latter differs especially in the EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 411 shape of the constrictions, which are not S-shaped, and comes from spoil heap KZ2 in the vicinity of Ko - in their later appearance, at a diameter of about 25 mm. zlovice. The two species differ also in their relative ages: M. melchioris is a Late Barremian species whereas M. DESCRIPTION: The specimen is evolute to semi- blayaci is Early Barremian. evolute, rather small, with continuously arched flanks on the ultimate whorl. On the oldest preserved part OCCURRENCE: So far the species is known exclu - only fine and dense growth lines are present on the sur - sively from the Lower Barremian ( K. compressissima face of the original shell. Irregularly arranged S-shaped Zone , based on the co-occurrence of Kotetishvilia cf. bulged ribs with convex curvature toward the aperture compressissima in the studied section) of the Silesian appear close to them. On the posterior side the ribs are Unit. cut more clearly; a depression occurs behind them , to which an indistinctly limited constriction on the inter - nal mould could correspond in some cases. The ante - Genus Valdedorsella BREISTROFFER , 1947 rior side of the rib passes continuously into an inter - costal space. TYPE SPECIES : Desmoceras akuschaense ANTHULA , 1899. MEASUREMENTS: On the holotype, at the maxi - mum shell diameter, H. I MMEL measured: D = 71 mm, H = 26.0 (0.37), U = 39.0 (0.39). On the imperfectly ?Valdedorsella hoheneggeri sp. nov . preserved specimen B13813, at D = 29.0 mm, H = 12.0 (Pl. 1, Fig. 5 ) (0.41) and U = 10.3 (0.355).

1883 Lytoceras n. sp. aff. Agassizianum PICT . (?); V. U HLIG , REMARKS: Besides UHLIG ’s (1883) specimen , re - pp. 200-201, pl. 14, fig. 8 ferred to Lytoceras n. sp. aff. Agassizianum , the spec - imen described herein represents the second find of the TYPE: The holotype is the original of UHLIG (1883 , pl. species . As the suture line, which is important for ac - 14, fig. 8 ), referred by him to Lytoceras n. sp. aff. curate genus - and family-level classification, is not Agassizianum . It is deposited in the collections of the known in either of the specimens, assignation of this Bayerische Staatssammlung für Paläontologie und his - species to the genus Valdedorsella is left in open torische Geologie in Munich , under collection number nomenclature . AS III 93 (in the set “Belegmaterial Uhlig, 1883, UHLIG (1883, p. 200) referred his specimen to the Wernsdorfer Schichten”). genus Lytoceras and, in open nomenclature, to the species L. agassizianum PICTET , 1847 or L. ven - STRATUM TYPICUM and LOCUS TYPICUS : ?Bar - trocinctum QUENSTEDT , 1849. Based on the illustra - remian, Hradiště Formation, Straconka locality, tions in PICTET & R OUX (1847, fig. 4 ), the former cer - Poland. tainly appears to be morphologically close to the Silesian material. However, both L. agassizianum and DERIVATIO NOMINIS: In honour of the eminent L. ventrocinctum are, according to their authors, of Al - Austrian geologist Ludwig HOHENEGGER , who first bian (Gault) age , and belong to the genus Kossmatella recognized the Silesian Unit succession of the Outer JACOB , 1907. Western Carpathians and defined its basic lithos - Neither the holotype nor the second specimen stud - tratigraphy . From his ammonite collection, studied ied herein is well preserved. The umbilicus in the holo - subsequently by V. UHLIG , comes a single incomplete type is not clean, so the measurements were taken on shell of the species described herein . the smaller specimens. The umbilicus width (U/D) of the smaller specimen indicates a semi-evolute than DIAGNOSIS: Shell evolute, with irregularly arranged rather than an evolute shell, a feature which is also char - S-shaped bulged ribs. acteristic of the Albian Kossmatella, including PICTET ‘s K. agassiziana. The latter are , however , clearly distinct MATERIAL: A single semi-juvenile specimen , of stratigraphically from the Silesian specimens. which less than half is preserved as original shell. The The S-shaped ribs resemble somewhat the ribbing area near the aperture is largely incomplete. The other pattern in Silesites vulpes (COQUAND in MATHERON , half of its ultimate whorl is indicated only by its outer 1880 ). However, in contrast to silesitids, accentuated contour (spec. KZ2/65-52 = B13813). The specimen constrictions are missing in the newly described species. 412 ZDENĚK VAŠÍČEK

DISTRIBUTION: UHLIG ’s (1883) specimen comes strongly deformed body chamber , has H = 10.7 (0.36) from the locality of Straconka , Poland. The specimen and U = 11.1 (0.38) at D = 29.5 mm. Its maximum di - studied herein comes from the latest Barremian of ameter affected by deformation reaches 34 mm. The spoil heap KZ2 in the vicinity of Kozlovice. body chamber begins at a shell diameter of 24 mm.

SUTURE LINE: The suture line in B13816, does not ?Family Silesitidae HYATT , 1900 have preserved an external lobe and is imperfectly pre - served on the outer flank . However, it is evident that the Genus Haplobrancoceras AVRAM , 1990 suture line is quite simply and faintly articulated (Text- fig. 3). The lateral lobe is simple, trifid, and the saddles TYPE SPECIES : Haplobrancoceras subquadratum are only slightly articulated. A similarly simple suture AVRAM , 1990. line is also preserved in the largest shell studied .

Haplobrancoceras subquadratum AVRAM , 1990 REMARKS: H. subquadratum is characterised by (Pl. 1, Figs 6-8 ; Text-fig. 3 ) rather evolute coiling and relatively simple suture lines. The material from the Beskydy Mts. extends 1990. Haplobrancoceras subquadratum n. gen., n. sp .; E. Avram’s knowledge of the species, through preserva - AVRAM , pp. 29-31, pl. 1, figs 1 -6, text-fig. 1 a, b . tion of the surface sculpture of the original shell and of 1995. Haplobrancoceras subquadratum AVRAM ; E. A VRAM , the external morphology of the shell, in spite of being pl. 19, figs 8 a, b . affected by strong deformation of the body chamber. Several ribs preserved on the termination of the largest MATERIAL: Three specimens represented by original specimen resemble secondary ribs of Silesites vulpes shells, the smallest of which (M5/046 = B13816) has (COQUAND ). From H. subquadratum the latter species the whorls filled with pyrite. After removing the orig - differs , however, in better developed constrictions on inal shell, several outer, incomplete suture lines were the juvenile whorls as well as in the ribbing of the shell exposed. The remaining two shells (T1/81 = B13817 between them. The Beskydy Mts. specimens differ and the largest one T1/137 = B13818) are laterally slightly from AVRAM ’s (1990) types in a narrower um - compressed . bilicus ; however, this may have resulted from sec - ondary deformation of the Silesian shells . DESCRIPTION: The shells are semi-involute, with Because of the relatively evolute coiling and , first slightly arched whorl flanks and a narrow, flat outer of all, because of its simple suture line, AVRAM (1990) side that is distinctly separated from the flanks. The regarded Haplobrancoceras to be the oldest represen - umbilicus is relatively wide and, similarly to the outer tative of the Late Cretaceous subfamily Brancocerati - side, separated from the flanks by a low and steep um - nae SPATH , 1934 (superfamily Acanthoceratoidea bilical wall. The shell surface bears fine, S-shaped GROSSOUVRE , 1894). He (AVRAM 1990) derived his growth lines. At the termination of the largest shell Haplobrancoceras from haploceratids (and this is in - (T1/137), remains of three blunt ribs are clearly seen cluded as the first half in its name), based on the form on the flanks. Weak, periodical constrictions probably of the suture line in the umbilical area in Neolissoceras occur on the moulds. (formerly Haploceras ) grasianum (see WIEDMANN , 1966, text-fig. 42). This interpretation is no longer sup - MEASUREMENTS: The least deformed juvenile ported by the present stratigraphical knowledge of both specimen B13816 has H = 5.0 (0.36) and U = 5.4 lineages. Namely, the last haploceratids ( N. sub - (0.37) at D = 13.7 mm. The largest specimen , with a grasianum ) disappeared in the latest Hauterivian (Pseudothurmannia ohmi Zone), whereas H. subquad - ratum did not appear before the late Early Barremian. The first true brancoceratids appeared with the begin - ning of the Albian. Any transitional representatives are not known. Consequently (M. Company , personal com - munication ), Haplobrancoceras is not considered a representative of the superfamily Acanthoceratoidea. Based on the presence of constrictions in the juveniles Fig. 3. Haplobrancoceras subquadratum AVRAM ; B13816 . and especially on the silesitid rib type, preserved in the A substantial part of the outer suture line at H = 4.2 mm largest shell of the material studied, I regard Haplo - EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 413 brancoceras to be a representative of the family Sile - compressissima figured by VERMEULEN (2003, pl. 22, sitidae (superfamily Desmoceratoidea), with a simpli - figs 1 -3). fied suture line. DISTRIBUTION: The species K. compressissima , the DISTRIBUTION: In Romania, according to AVRAM index of the eponymous zone, is known widely from (1990), H. subquadratum occurs in the K. compres - the Lower Barremian of the Mediterranean area, sissima to Barrancyloceras barremense ammonite mainly in France. Zones of the late Early to early Late Barremian. The Beskydy specimens are from spoil heap T1 (late Early Barremian) and M5 (early Late Barremian). Kotetishvilia ex gr. sauvageaui (HERMITE , 1879) (Pl. 1, Figs 11, 12 )

Superfamily Endemoceratoidea SCHINDEWOLF , 1966 2003. Kotetishvilia sauvageaui (HERMITE ); J. V ERMEULEN , Family Pulchellidae DOUVILLÉ , 1890 pp. 133-136 . Subfamily Psilotissotiinae VERMEULEN , 1995 2003. Kotetishvilia sp. gr. sauvageaui (HERMITE ); J. V ER - MEULEN , pl. 19, figs 11, 12, pl. 28, figs 5 -7. Genus Kotetishvilia VERMEULEN , 1997 2006. Kotetishvilia sauvageaui (HERMITE ); J. V ERMEULEN & J. K LEIN , pp. 206-207 (cum syn.) . TYPE SPECIES : Pulchellia changarnieri SAYN , 1890. MATERIAL: A single specimen , preserved as an al - most complete outer mould and part of its original Kotetishvilia cf. compressissima (D´O RBIGNY , 1841) shell, embracing a 45º sector of a single whorl (T3/106 (Pl. 1, Figs 9, 10 ) = B13815) , from spoil heap T3, where it occurs to - gether with Late Barremian Silesites seranonis, Euly - 1841. Ammonites compressissimus , D´O RBIGNY ; A. D´O RBI - toceras phestum , and other taxa. GNY , pp. 210-211, pl. 61, figs 4, 5 . 2003. Kotetishvilia compressissima (D´O RBIGNY ); J. V ER - DESCRIPTION: The shell is involute, small. On the MEULEN , pp. 123-126, pl. 2, figs 7, 8, pl. 20, figs 1 -12, circumference , part of the ventral area is preserved in pl. 21, figs 1 -11, pl. 22, figs 1 -7. the form of a narrow, smooth zone delimited by a con - 2006. Kotetishvilia compressissima (ORBIGNY ); J. V ERMEU - spicuous marginal edge against the sides . The shell sur - LEN & J. K LEIN , pp. 202-204 (cum syn.) . face bears growth lines, initially fine and dense, S- shaped , which strengthen somewhat in the final half MATERIAL : A single external mould of a juvenile near the circumference. As demonstrated by the orig - specimen (T9/77 = B13814), with the ventral region inal shell , it is evident that the strengthened ribs di - missing, from the T9 outcrop . verge slightly in the vicinity of the shell termination. The maximum shell diameter reaches about 21 mm . DESCRIPTION: The shell is involute, small. The um - bilicus is very narrow, with flat flanks. Wide and flat REMARKS: The specimen is a juvenile with well pre - ribs separated by narrow and deep intercostal furrows served sculpture. Initially uniform fine growth lines occur on more than a half of the ultimate whorl. The strengthen later into stronger ribs near the circumfer - furrows disappear at about mid-height of the ultimate ence. The fine sculpture and the marginal edge that sep - whorl . 11 -12 ribs occur on a half whorl at the maxi - arates the whorl flanks from the outer side, and the fact mum shell diameter. that it is a juvenile, make its taxonomic interpretation uncertain. It is very similar to , and possibly conspecific MEASUREMENTS: At the maximum diameter D = with, either Subpulchellia oehlerti (NICKLĖS , 1894 ) or 18 mm, the whorl height is 10.3 mm (H/D = 0.57). Kotetishvilia savageaui. The assignment to K. ex gr. sauvageaui , suggested herein, is based on stratigraph - REMARKS: The morphology of the ventral region, ical grounds: the Beskydy Mts. specimen is dated as which is very important for taxonomic determination, Late Barremian, and thus a determination as the Early is not known. The presence of wide, flat ribs on the cir - Barremian S. oehlerti does not fit. The specimens il - cumference of the shell, and the overall morphology of lustrated by VERMEULEN (2003, pl. 19, figs 11, 12 and the specimen , suggest that it belongs to the genus pl. 28, figs 5 -7) as Kotetishvilia sp. gr. sauvageaui are Kotetishvilia. It most resembles the juvenile of K. very close to the the specimen described here. 414 ZDENĚK VAŠÍČEK

DISTRIBUTION: According to VERMEULEN (2003), equally strong . Similar fragments figured by DELANOY K. sauvageaui occurs in an interval near the (1992) as Audouliceras sp. (forme 4) appear to be Lower /Upper Barremian boundary, and is known from morphologically close. However, in these specimens the Coronites darsi , Holcodiscus uhligi and Toxancy - the ribbing is poorly preserved and I have conse - loceras vandenheckii ammonite Zones. The specimen quently excluded them from the synonymy. studied comes from the Upper Barremian. DISTRIBUTION: According to KOTETIŠVILI (1970 ) and DELANOY (1992), Audouliceras (forme 4 ) occurs Suborder Ancyloceratina WIEDMANN , 1966 in the Late Barremian, probably in the Imerites giraudi Superfamily Ancyloceratoidea GILL , 1871 Zone. In the study area ?Audouliceras sp. occurs Family Ancyloceratidae GILL , 187 within assemblages dated as Late Barremian to earli - Subfamily Ancyloceratinae GILL , 1871 est Aptian.

Genus Audouliceras THOMEL , 1964 Subfamily Helicancylinae HYATT , 1894 TYPE SPECIES : Ancyloceras audouli ASTIER , 1851. Genus Lytocrioceras SPATH , 1924

?Audouliceras sp. TYPE SPECIES : Ancyloceras jauberti ASTIER , 1851. (Pl. 2, Fig. 4 )

?1955. Jauberticeras cf. dubius (forme no1); S. S. S ARKAR , Lytocrioceras cf. jauberti (ASTIER , 1851) p. 99, pl. 4, fig. 10, pl. 8, fig. 2 . (Pl. 2, Fig. 5 ) ?1970. Crioceratites aff. elegans D´O RBIGNY ; E. V. KOTETIŠVILI , pp. 69-70, pl. 5, fig. 3 . 1851. Ancyloceras Jauberti, ASTIER ; J.-E. A STIER , pp. 455- 456, pl. 23, fig. 17 . MATERIAL: A single, badly preserved and deformed 1992. Lytocrioceras jauberti (ASTIER ); G. D ELANOY & A. whorl fragment (only the flanks are preserved) loosely POUPON , pp. 368-372, pl. 1, figs 1 – 3, pl. 2, figs 1 -4, coiled in a spiral (M1/012 = B13819), from spoil heap M1 . pl. 3, fig. 1 . 1999. Lytocrioceras jauberti (ASTIER ); L. E BBO & al., pp. 2- DESCRIPTION: The specimen is represented by ap - 4, pl. 2, fig. 2, pl. 4, fig. 2, pl. 5, figs 1, 3, pl. 6, fig. 3 proximately a quarter of a whorl, with a rapidly in - (cum syn.) . creasing , albeit strongly deformed whorl height. Strong, trituberculate main ribs occur on the whorl MATERIAL: Five fragments of strongly deformed flanks. Their shape is heavily obscured by deformation, adult hook-like parts represented by either original so that all the tubercles seem to fuse with each other. shells or their external moulds (M5/162, 165, 250, Almost straight secondary ribs occur between the main M1/317, M1/320 = B13820) ; in none of them has the ribs. It cannot be excluded that some, otherwise sim - juvenile spiral part been preserved. M1/320 = B13820 ple ribs, bifurcate in the vicinity of the line of coiling is the largest and most complete specimen. All are from (in one case they bifurcate on the flank). On the cir - spoil heaps M1 and M5 . cumference, 5 to 6 secondary ribs occur in the intervals between the main ribs. The deformed maximum whorl DESCRIPTION: The largest fragment represents an height is about 27 mm. about 10 cm long, negligibly arcuate, i.e. almost straight shaft passing into a hooked part (flexus). This REMARKS: The fragment is probably a part of a passes into a short, probably incomplete shaft of the loosely coiled spiral followed by the hook of a large terminal part of the shell forming with the foregoing shell. According to the ribbing and literature data shaft an angle of about 40 degrees. The longer (first) (above all in DELANOY 1992), it can be deduced that shaft is covered with dense and thin, slightly S-shaped, the fragment could belong to the genus Audouliceras . proverse ribs. Sporadically, some of the ribs bifurcate Among the most characteristic features of the speci - somewhere at about shaft mid-height. Near and on the men are the rapid growth in whorl height and the fact flexus the ribs are slightly convex towards the aperture. that the main ribs, between which a relatively high Ribs on the terminal shaft are straight. On the transi - number of shorter secondary ribs can be found, are tion from the first shaft to the hook, a single rib occurs EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 415 that is markedly stronger than the others. The de - DISTRIBUTION: According to DELANOY (1998), H. formed shell height in the flexus axis is 16.5 mm. coulleti is known only from the Upper Barremian of southeast France, namely from the uppermost part of REMARKS: The sculpture, size and overall shape of the Hemihoplites feraudianus Zone and the lower the hook-like fragments are closest to those of Ly - part of the Imerites giraudi Zone. The specimen is tocrioceras jauberti. However, in contrast to the Early from the Upper Barremian of the area of Kozlovice- Barremian age of the type material of this species, the Žáry. fragments described herein are of Late Barremian age. Although no apparent morphological differences can be identified, I designate the Silesian specimens on ?Heteroceras ex gr. emerici (D´O RBIGNY , 1842) stratigraphical grounds only as L. cf. jauberti . (Pl. 2, Fig. 1 )

DISTRIBUTION: According to EBBO & al . (1999), the 1842 Turrilites Emericianus D´O RBIGNY , pp. 580-582, pl. type material of L. jauberti comes from the Lower Bar - 141, figs 3 -6 remian ( Kotetishvilia nicklesi Zone) of southeast 1998 Heteroceras emerici D´O RBIGNY ; G. D ELANOY , pp. 51- France. The specimens studied herein come from as - 78 (cum syn.) semblages dated as Toxancyloceras vandenheckii to Imerites giraudi ammonite Zones of the Late Bar - MATERIAL: A single strongly deformed shaft frag - remian (spoil heap M5) and as Late Barremian to ear - ment (M5/375 = B13822) from spoil heap M5 near liest Aptian (spoil heap M1) . Malenovice.

DESCRIPTION: The shaft is slightly arcuate, with a Family Heteroceratidae SPATH , 1922 small increase in height. The fragment bears medium- strong, rather dense ribs of uniform type. They are Genus Heteroceras D´O RBIGNY , 1850 slightly prorsiradiate, straight to finely S-shaped. The fragment is about 100 mm long and 52.5 mm high. TYPE SPECIES : Turrilites Emericianus D´O RBIGNY , 1842. REMARKS: In its overall morphology and, among other features, slight arcuation of the shaft , the Silesian specimen resembles H. emerici, especially specimens Heteroceras aff. coulleti DELANOY , 1994 designated by DELANOY (1998) to as H. emerici mor - (Pl. 2, Fig. 2 ) phe tardieui . The fragmentary preservation of the spec - imen does not allow an unequivocal determination 1994. Heteroceras coulleti sp. nov.; G. D ELANOY , p. 547, (see DELANOY 1998 for detailed discussion on the con - text-fig. 2 . cept and variability of H. emerici ). 1998. Heteroceras coulleti DELANOY ; G. D ELANOY , pp. 44- Similar fragments of arcuate shafts were referred 51, pl. 10, figs. 1-4, pl. 11, figs. 1-5, pl. 12, figs. 3, 4, by NIKOLOV (1964) to Anahamulina silesiaca (UH- 6, pl. 13, figs. 1, 2, pl. 14, figs. 1-3, pl. 40, fig. 2, pl. LIG ) (his pl. 4, fig. 4) and A. picteti (EICHWALD ) (his pl. 48, fig. ?1, 3 (cum syn.) . 4. fig. 5; pl. 5, fig. 3; see also DIMITROVA , 1967, pl. 40, figs. 3, 4). All these specimens may be conspecific with MATERIAL: A single fragment of a straight to slightly the Silesian form. arcuate shaft (KZ2/65-15 = B13821) from spoil heap KZ2 . DISTRIBUTION: Unequivocal reports of H. emerici are known only from France and Georgia. According DESCRIPTION: The shaft has strong, relatively to DELANOY (1998), it occurs in the Late Barremian sparsely arranged, slightly arcuate simple ribs. The Imerites giraudi Zone. shaft height is 19 mm.

REMARKS: In spite of its fragmentary preservation , ?Heteroceras sp. it is possible to assume that the shaft fragment with its (Pl. 2, Fig. 3 ) strong ribs, which follows the trochospiral juvenile part, belongs to the group of H. coulleti (compare 1972. Ptychoceras morloti OOSTER ; Z. V AŠÍČEK , pp. 65-66, with the synonymy). pl. 10, figs. 4, 5 . 416 ZDENĚK VAŠÍČEK

?1933. Heteroceras vermiforme n. sp.; I. R OUCHADZÉ , p. remains problematic. Nevertheless, I believe that my 233, pl. 13, fig. 5 . material does not belong to the genus Ptychoceras ?1938. Ptychoceras sp. ind.; I. M. R UCHADZE , p. 163, pl. 1, but represents instead a small form of the genus Het - fig. 5 . eroceras. It probably represents a new species but, in ?1998. Heteroceras baylei REYNES ; G. D ELANOY , pp. 95-99, the absence of the juvenile part, it must remain in pl. 16, fig. 3, pl. 50, fig. 3 . open nomenclature as ?Heteroceras sp.

MATERIAL: Three incomplete shelly specimens, in DISTRIBUTION: The type of Ptycoceras morloti comes which the most juvenile parts are missing (Ce3/12 = from the Veveyse section, spanning the uppermost B13823; KN10/2, 34). They come from small out - Valanginian to the lowermost Barremian (BUSNARDO & crops in the Bílý potok stream near Čeladná (Ce3) and al . 2003) . No ammonites of Late Barremian age are from Kunčice p. O. (KN10). The finds of VAŠÍČEK known from this locality. French specimens of Hetero - (1972) are from the old spoil heaps K5 and V5 near ceras baylei come from the Late Barremian Imerites gi - Kozlovice and Veřovice respectively. raudi ammonite Zone ( DELANOY 1998). ROUCHADZÉ (1933) gave an Early Aptian age for H. vermiforme but, DESCRIPTION: The specimens are relatively small, from the data of KAKABADZE (1989, p. 556), its age can characterized by hook-like shells with straight shafts be inferred to be approximately latest Barremian . The in contact. The first shaft (proversum) bears simple Silesian material is dated as Late Barremian. proversal oblique ribs; the terminal shaft (retrover - sum) bears simple, sparser retroversal ribs. The flexus that connects the shafts is covered with ribs that bi - Family Hemihoplitidae SPATH , 1924 furcate at the base or , at the termination of the flexus , somewhat higher. The shell illustrated has a length of Genus Pachyhemihoplites DELANOY , 1992 36 mm; the flexus height is about 8 mm. TYPE SPECIES : Pachyhemihoplites thomeli DE- REMARKS: The specimen described as Ptychoceras LANOY , 1992. morloti by VAŠÍČEK (1972) is very similar to the ones described herein. Also very similar is OOSTER ‘s spec - imen of P. morloti (OOSTER , 1860) , which, however, is Pachyhemihoplites sp. far larger. P. morloti , which has never been found (Pl. 3, Fig. 2 ) complete , has three shafts. The specimens from the Beskydy Mts. would thus represent the first and the ?1992 Pachyhemihoplites contei sp. nov.; G. D ELANOY , pp. second shaft of this species. 105-107, pl. 9, figs. 2 – 5, 7 In size, shape and sculpture, the small specimens ?2002 Spinocrioceras sp.; Z. V AŠÍČEK & P. S KUPIEN , p. 69, from the Beskydy Mts. resemble some shells desig - pl. 2, fig. 2 nated by DELANOY (1998) as H. baylei ; only shells with shafts in contact with each other are included into the MATERIAL: A single incomplete and deformed orig - synonymy. The most complete specimen of DELANOY inal shell with part of the penultimate and ultimate (illustrated in his pl. 50, fig. 3 ) has an initial turriliti - whorls preserved (M5/105 = B13824). Both this spec - cone juvenile stage . However, no such juvenile stage imen and the specimen referred to Spinocrioceras sp . has been found preserved in the material from the in VAŠÍČEK & S KUPIEN (2002), come from spoil heap Beskydy Mts. M5 near Malenovice. In the size and sculpture of the first shaft (prover - sum), H. vermiforme ROUCHADZÉ also appears to be DESCRIPTION: The shell is evolute, small (maximum close. It has the helically-coiled beginning of the shell, diameter about 50 mm). The whorl section is unknown. but the second shaft (retroversum) has not been pre - On the penultimate whorl, trituberculate main ribs al - served. Thus, it is not known if both shafts are in con - ternate with one to two secondary ribs. The main ribs of tact or form an open flexus. Ptychoceras sp. ind. (in this whorl are remarkable in that a narrow furrow occurs RUCHADZE 1938) is similar in both size and sculpture in their axes between the umbilical and lateral tubercles, of both shafts; however, the juvenile part of the shell giving the the impression of the eye of a needle (fibu - is not preserved. lation). In a single case, a shorter inserted rib starts at the Because the juvenile part of the specimens from the level of the umbilical tubercle. The other inserted ribs Beskydy Mts. is not known, their generic classification begin already on the line of coiling. EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 417

On the ultimate whorl, the main trituberculate ribs MATERIAL: A single incomplete shelly specimen alternate with one or , exceptionally , two simple in - with a corroded and , on the internal side , unpreserved serted (secondary ) ribs. Both the umbilical tubercles, part of the proversum (spec. K7/3 = B123825) , from which lie rather high above the line of coiling, and the a Barremian ammonite assemblage collected from lateral tubercles, which lie close to the ventrolateral tu - spoil heap K7, between the municipalities of Ko - bercles, are conical. The ventrolateral tubercles are zlovice and Tichá. clavate. The secondary ribs usually lack umbilical tu - bercles. Ventrolateral tubercles are usually indicated on DESCRIPTION: The specimen possesses two shafts in them; in one case a lateral tubercle is developed. contact. Both the proversum and the retroversum bear wide and blunt simple ribs. Ribs on the proversum are REMARKS: The imperfect preservation and the un - oblique, those on the retroversum subhorizontal. On the known whorl section make both species and generic retroversum the ribs are rather sparsely arranged. The determination difficult. With regard to the character - flexus is covered with different, i.e. thinner ribs, some istic arrangement of the tubercles on the main ribs of which bifurcate at the base of the flexus. Others are (the lower row may be designated as periumbilical tu - shorter, split or inserted on the outer half of the flexus. bercles, the lateral tubercles are well separated from the The shaft height in the flexus reaches 12 mm. The dis - lower ones, and the ventrolateral tubercles are clavate), tance between the preserved beginning of the prover - Pachyhemihoplites is the most appropriate genus . The sum and the circumference of the flexus is 60 mm. Silesian specimen appears to be close, but not identi - cal , to P. contei DELANOY . A similar tubercle spacing REMARKS: The specimen from the Beskydy Mts. is can also be observed in the phragmocone that was very close to the holotype of IMMEL (1987) in dimen - figured by VAŠÍČEK & S KUPIEN (2002, pl. 2, fig. 2 ) and sions, close distance between both shafts, and in sculp - designated as Spinocrioceras sp. However, in this ture. IMMEL ’s species glemmbachensis differs, how - specimen periumbilical tubercles are also indicated ever, in having oblique ribs on the holotype retroversum ; on the usually bituberculate secondary ribs . the ribs in the Beskydy Mts. specimens are subhori - zontal. A close species is Duyeina boutini (MATHERON , DISTRIBUTION: Representatives of Pachyhemiho - 1880 ), from which the described specimen differs in plites were known so far only from southeast France having the shafts in contact and in having bifurcated or (DELANOY 1992), from the upper Barrancyloceras inserted ribs in the area of the flexus. D. boutini also re - barremense Zone ( =Toxancyloceras vandenheckii and sembles Duyeina kleini VERMEULEN , 2005 . Gerhardtia sartousiana Zones) of the Late Barremian (actually Pachyhemihoplites is mostly characteristic of DISTRIBUTION: IMMEL (1987) reported the species the latter Zone - VERMEULEN 2003). The Silesian spec - from the Upper Barremian of the Northern Calcareous imen comes from an assemblage dated roughly as Alps ; VERMEULEN (2005 a, p. 162) from the uppermost Toxancyloceras vandenheckii to Imerites giraudi am - part of Subsaynella sayni Subzone and the Gerhardtia monite zones of the Late Barremian. sartousiana Zone states of the Late Barremian of France; and VERMEULEN (2005 b) from only the Late Barremian Toxancyloceras vandenheckii Zone. Family Hamulinidae GILL , 1871

Genus Duyeina VERMEULEN , 2005 Superfamily Deshayesitoidea STOYANOW , 1949 Family Deshayesitidae STOYANOW , 1949 TYPE SPECIES : Anahamulina glemmbachensis IM- MEL , 1987. Genus Paradeshayesites KEMPER , 1967

TYPE SPECIES : Hoplites laevisculus von KOENEN , Dueyina aff. glemmbachensis (IMMEL , 1987) 1902. (Pl. 3, Fig. 1 ) Paradeshayesites semenovi (BOGDANOVA , 1999) 1987. Anahamulina glemmbachensis n. sp .; H. I MMEL , pp. (Pl. 3, Fig. 3 ) 124-125, pl. 14, fig. 1. 2005a. Dueyina glemmbachensis (IMMEL ); J. V ERMEULEN , p. 1999. Deshayesites semenovi BOGDANOVA , sp. nov.; T. N. 164. BOGDANOVA , p. 47-49, pl. 1, figs 1-3. 418 ZDENĚK VAŠÍČEK

MATERIAL: A single strongly deformed shelly specimen in the new Treatise ( WRIGHT & al . 1996), is charac - (Os/1 = B13826) from an outcrop of the upper Hradiště terised by arched whorls. The laterally strongly com - Formation on the left bank of the Ostravice River besides pressed Silesian specimen bears several ribs with so- the sports field in the municipality of Ostravice. called fibulation (“needle eye”) on the penultimate whorl, a feature so far unknown in representatives of DESCRIPTION : The shell is semi-involute, large, this genus. with high whorls and a relatively narrow umbilicus. Of significance among the ammonites from the The umbilical wall is low and steep, and passes con - Beskydy Mts. is the genus Kotetishvilia, and espe - tinuously into the flanks of the ultimate whorl. The cially K. compressissima. A juvenile specimen , deter - strong, slightly S-shaped ribs pass across the outer mined in this work as K. ex gr. sauvageaui , resembles side without interruption and weakening. The ribs be - in its sculpture Subpulchellia oehlerti (NICKLES ). An - gin in concavely arched bulges on the umbilical wall other specimen from the Beskydy Mts., determined on the line of coiling. The majority of ribs are simple, earlier as Psilotissotia aff. chalmasi by VAŠÍČEK (1972, bifurcating rarely at about whorl mid-height. Between p. 83, pl. 14, fig. 6), also probably belongs to S. oehlerti . the simple ribs traversing the whole whorl, single ribs Pulchelliids occur in the Silesian Lower Creta - are inserted, beginning at about whorl mid-height. ceous only sporadically, but several species have been recorded. According to the survey of VERMEULEN MEASUREMENTS: The maximum shell diameter in (2003) and others, in addition to the above-presented the axis of elongation reaches 125 mm. At a shell di - species, the Silesian Unit also yielded Kotetishvilia ameter of 123.5 mm (measured between the major brevicostata (KOTETISHVILI , 1980) [= Psilotissotia sp. axes of deformation), H = 54 mm (0.44) and U = 29 ind. in VAŠÍČEK 1972, pl. 14, fig. 5] , Nicklesia pulchella (0.235). At this diameter, there are 29 ribs at the um - (D´O RBIGNY , 1841) [from VAŠÍČEK 1979, pl. 1, fig. 5; bilicus and 65 ribs at the the circumference per whorl. and probably Pulchellia Karsteni of UHLIG , 1883, pl. 20, fig. 1], and possibly also Gerhardtia provincialis REMARKS: The narrow umbilicus and character of (D´O RBIGNY ) [most probably Pulchellia lindigi the ribbing make the specimen very similar to Pa - KARSTEN in UHLIG 1883, pl. 20, fig. 6 ]. radeshayesites weissi (NEUMAYR & UHLIG , 1881 ). Besides the forms described herein and those re - However, on the basis of the rare rib bifurcation, the ported by VAŠÍČEK (1972), there are several additional higher position of the inserted ribs and the rib density , problematic fragments in my collection which extend it is referred to Paradeshayesites semenovi. the taxonomic composition of the Early Cretaceous ammonite assemblage of the Beskydy Mts. Among DISTRIBUTION: Besides the studied specimen, from these is Rugacrioceras martinsi (REYNES ), which is the Deshayesites oglanlensis ammonite Zone [dating probably the correct name for the fragmentary speci - based on the co-occurring Paradeshayesites weissi - men illustrated by VAŠÍČEK (1972, pl. 3, fig. 3) as formis (BOGDANOVA , 1983 )] the species P. semenovi is Acrioceras (A. ) aff. tabarelli . Another species is known from the Lower Aptian of Mangyshlak. Manoloviceras saharievae (MANOLOV , 1962), based on the specimen referred formerly to Veleziceras uh - ligi (VAŠÍČEK , 1972) and re-interpreted subsequently by DISCUSSION VAŠÍČEK &WIEDMANN (1994) [I do not agree with WRIGHT & al . 1996, p. 226, fig. 177 a, b, who put The incompleteness and bad preservation of the Manoloviceras into the synonymy of Orbignyceras material studied make final determination of several ROYO Y GOMEZ , 1945 b] . taxa impossible. There were considerable problems Many of UHLIG ’s species from the Barremian of the with the ancyloceratids , in which the juvenile parts Silesian Unit , described in his monograph of 1883 and were usually not preserved. Similarly, the turriliticone not re-studied for many years , recently became types of juvenile whorls are missing in the case of the hetero - a number of newly-defined genera, especially those es - ceratids (the only exception is represented by a frag - tablished by Jean VERMEULEN (2000, 2005, 2006). ment referred to as ? Heteroceras sp. juv. by VAŠÍČEK These are: Barrancyloceras VERMEULEN , 2000 (type- & S KUPIEN 2002, pl. 1, fig. 3), which makes their species Crioceras hammatoptychum UHLIG , 1883), Pty - generic classification uncertain . chohamulina VERMEULEN , 2005 (type-species Very interesting but also highly problematic is the Hamulina ptychoceroides UHLIG , 1883), Vasicekina form described herein as Pachyhemihoplites sp. This VERMEULEN , 2005 (type-species Hamulina paxillosa genus which , for unknown reasons , was not included UHLIG , 1883), Leroyoceras VERMEULEN , 2006 (type- EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 419 species Crioceras Hoheneggeri UHLIG , 1883), Terrest - scribed so far belong to the Mediterranean bioprovince. rina VERMEULEN , 2006 (type-species Hamulina Quen - Consequently, the Barremian ammonite zonation as stedti UHLIG , 1883) and Leptohamulina VERMEULEN , defined in this area (after REBOULET , H OEDEMAEKER & 2006 (type-species Hamulina n. f. ind. UHLIG , 1883 = al . 2006) is here applied (see Text-fig. 4). Nevertheless, Hamulina distans HOHENEGGER ). with the exception of Nicklesia pulchella , known from Ostravice, or Kotetishvilia cf. compressissima , from Tichá, the majority of the zonal /subzonal am - STRATIGRAPHIC CONTRIBUTION monite indexes of the Mediterranean scheme do not occur in the Silesian Unit. However, indirect bios - The ammonites from the upper Lower Cretaceous tratigrapical correlations are possible based on com - of the Godula (basinal) facies of the Silesian Unit de - plete ammonite assemblages .

Fig. 4. Uppermost Hauterivian, Barremian and lowermost Aptian ammonite zonation (according to REBOULET & al . 2006) 420 ZDENĚK VAŠÍČEK

The stratigraphically oldest finds described here Early Aptian. The oldest fossiliferous deposits corre - come from the the Kotetishvilia compressissima am - spond to the Early Barremian Nicklesia pulchella Zone monite Zone of the late Early Barremian . These are and the youngest to the Early Aptian Deshayesites Kotetishvilia cf. compressissima and Melchiorites oglanlensis Zone. Above a macrofaunally barren in - blayaci from the Tichá 9 outcrop. From the same or ap - terval , another ammonite horizon , dated as the Late proximately the same stratigraphical level, comes Hap - Aptian Acanthoplites nolani Zone ( VAŠÍČEK , 1981 b) , lobrancoceras subquadratum , known from spoil heap was found in the uppermost part of the Hradiště For - T1 near Tichá, and from spoil heap M5, near Malen - mation in the late 1980 s. ovice . Spoil heap M5 is characterised by a rich am - The 15 species level taxa described in this paper are monite assemblage (e.g. Eulytoceras phestum, Cos - largely species that were previously unrecorded from tidiscus recticostatus, Macroscaphites yvani, the Silesian Unit. Some Late Barremian ancylocer - Leptohamulina distans ) which , together with Phyl - atids, especially Lytocrioceras cf. jauberti , and larger lopachyceras vergonense, ?Heteroceras ex gr. emerici , representatives of the genus Heteroceras, preserved and Pachyhemihoplites sp. , indicates its early Late only in fragments, appear to be the most interesting. Barremian age ( Toxancyloceras vandenheckii , Ger - The generic affiliation of the newly described hardtia sartousiana up to a lower part of the Imerites species ?Valdedorsella hoheneggeri remains uncertain. giraudi ammonite Zones ). In the Western Carpathians , as well as in the whole According to its stratigraphical position in France, Mediterranean Tethys, the Silesian Unit represents a the species Heteroceras aff. couleti , known from spoil specific Early Cretaceous sedimentation area , in which heap KZ2, near Kozlovice , indicates the boundary in - a more than 1 km thick set of dark coloured, dysoxic, terval of the Hemihoplites feraudianus and the I. gi - largely pelitic deposits have accumulated. The Bar - raudi zones . remian and Early Aptian part of it has been famous for Particularly noteworthy is the occurrence of Pa - rich and specifically diverse ammonite assemblages for radeshayesites semenovi (not reported in my former re - a long time. ports ). It comes from an outcrop near a sports field in A dysoxic environment in the Hradiště Formation is the Ostravice River bank in the municipality of the indicated be its dark colour, often with an increased con - same name. It was found co-occurring with P. weissi - tent of pyrite in the sediment or with the pyritization of formis (BOGDANOVA ) ( see VAŠÍČEK & al . 2004), a inner whorls of ammonite shells, by usually higher species characteristic of the Early Aptian oglanlensis TOC content and above all by the fact that the ammonite ammonite Zone. assemblages are usually accompanied by no or only spo - UHLIG (1883), on the basis of the rich collection of radic benthos. The ammonites usually occur in thin, L. HOHENEGGER and those of his other collaborators, de - about 10 -15 cm thick fossiliferous horizons. This, to - scribed more than 70 ammonite species from equivalent gether with the frequent fragmentary preservation of the strata of the Silesian Unit (Wernsdorfer Schichten in the ammonite shells, indicates that shells from the deep original conception). These were largely new genera and shelf environment were redeposited by distal turbidite species, usually without accurate stratigraphical posi - currents into deeper-water parts of the basin where there tions. On the basis of the revision of UHLIG ’s material , was a lack of oxygen at the bottom. Here, the unburied supplemented by my own collections ( VAŠÍČEK , 1972, upper halves of shells were often dissolved, leading to 1973), as well as other collections from the Silesian Unit their designation as “half-ammonoids”. in the Czech Republic (given e.g. in VAŠÍČEK , 1979, 1981 a, 1990, VAŠÍČEK & K LAJMON , 1998, VAŠÍČEK & SKUPIEN , 2002, V AŠÍČEK & al. 2004), and this paper, it Acknowledgements can be stated that in the Hradiště Formation of the Sile - sian Unit in the Czech Republic more than 30 species of The author’s thanks go to G. DELANOY (Nice) and J.-L. ammonites occur, including three new genera and ten LATIL (Grenoble) for constructive discussion about the de - new species that were unknown to UHLIG . termination of some genera and species as well as for the pro - vision of some old ammonite works. The author is very grate - ful to H. IMMEL (Munich) for information on the deposition CONCLUSIONS of Uhlig’s original of Lytoceras n. sp. aff. Agassizianum and for the measurement of its parameters. Mrs K. MEZI - The ammonite-bearing deposits of the Hradiště HORÁKOVÁ (Ostrava) is thanked for taking high quality pho - Formation in the Silesian Unit , in the Godula facies , tographs. The contribution was prepared thanks to support belong to an interval spanning the Early Barremian to provided by the Institute of Geonics of Academy of Sci - EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 421 ences of the Czech Republic in the framework of research — 1994. Nouvelles conceptions génériques au sein de la fa - plan Z 308 60518. Moreover, the author is obliged to both the mille Heteroceratidae Spath, 1922 (Ammonidea, Ancy - reviewers for their remarks and recommendations. I am grate - loceratina). Comptes Rendus de l´Académie des Scien - ful to M. COMPANY (Granada) and R. MARCINOWSKI (Warsaw) ces Paris, Série II, 318 , 543-548. for stimulating ideas and proposals, which I have largely ac - — 1998. Biostratigraphie des faunes d´ammonites a la limité cepted, and to Chris J. WOOD and Ireneusz WALASZCZYK , for Barrémien-Aptien dans la région d´Angles-Barreme- linguistic corrections. Castellane. Étude particulière de la famille des Hetero - ceratina Spath, 1922 (Ancyloceratina, Ammonoidea). Annales du Muséum d´Histoire Naturelle de Nice , 12 REFERENCES (1997), 1-270. DELANOY , G. & J OLY , B . 1995. Sur quelques Phylloceratidae ASTIER , J.-E . 1851. Catalogue descriptif des Ancyloceras ap - (Phylloceratina, Ammonoidea) du Barrémien supérieur partenant a l´étage Néocomien d´Escragnolles et des Bas - et de l´Aptien inférieur du Sud-Est de la France. Méso - ses Alpes. Annales des Sciences Physiques et Naturelles, gée, 54 , 121-138. d´Agriculture et d´Industrie Lyon, (série 2), 3, 435-456. DELANOY , G. & P OUPON , A. 1992. Sur le genre Lytocrioce - AVRAM , E. 1976. Les fossiles du flysch éocrétacé et des cal - ras Spath, 1924 (Ammonoidea, Ancyloceratina). Geo - caires tithoniques des hautes vallées de la Doftana et du bios, 25 , 367-382. Tirlung (Carpates Orientales). Mémoires de l´Institut de DIMITROVA , N. 1967. Les fossiles de Bulgarie IV. Crétacé in - Géologie et de Géophysique Bucuresti , 24, 5-74. férieur – Cephalopoda (Nautiloidea et Ammonoidea). — 1978. Une nouvelle espece de céphalopode dans le Bar - Académie Bulgare des Sciences, 1-424. Sofia. [In Bul - rémien supérieur de la Roumanie: Costidiscus tardus n. garian with French summary ] sp. Dari de seama ale sedintelor Institutul de Geologie EBBO , L., P OUPON , A., D ELANOY , G. & G ONNET , R. 1999. si Geofizica Bucuresti, 3. Paleontologie , 64 (1976- Nouvelles données sur le genre Lytocrioceras Spath, 1977), 5-8. 1924 dans le Barrémien inférieur du Sud-Est de la — 1984. Correspondent species of the genera Macroscap - France. Annales du Muséum d´Histoire Naturelle de hites Meek and Costidiscus Uhlig. 75 years of the La - Nice, 14 , 1-25. boratory of Paleontology (University of Bucharest), Spe - ELIÁŠ , M ., SKUPIEN , P. & VAŠÍČEK , Z . 2003. A proposal for cial Volume, 67-80. the modification of the lithostratigraphical division of the — 1990. Haplobrancoceras n. g., ammonite barrémienne à lower part of Silesian Unit in the Czech area (Outer ligne cloisonnaire simplifiée. Dari de seama ale sedin - Western Carpathians). Sborník Vědeckých Prací Vysoké telor Institutul de Geologie si Geofizica Bucuresti, 3. Pa - Školy báňské – Technické Univerzity Ostrava, Řada Hor - leontologie, 74 (1987), 27-32. nicko-Geologická, Monografie 8, 49, 7-13. [ In Czech — 1995. Lower Cretaceous (Valanginian-Early Aptian) am - with English summary ] monite succession in the Svinita region (SW Rumania). FISCHER , J.-C. & G AUTHIER , H. 2006. Révision critique de la Géologie Alpine, Mémoire hors serie , 20 (1994), 113-167. Paléontologie Francaise d´Alcide d´Orbigny, vol. 4, Cép - BOGDANOVA , T.N . (1983). Zona Deshayesites tuarkyricus – halopodes Crétacés. 292 p . Backhuys Publishers ; Leiden nizhnjaja zona apta Turkmenii. Ezhegodnik Vsesoju - – Pays-Bas. znogo Paleontologicheskogo Obshchestva , 26 , 128-147. HERMITE , H . 1879. Études géologiques sur les Iles Baléares. — 1999. Nizhnij apt Gornogo Mangyshlaka (stratigrafija, I. Majorque et Minorque. 359 p. Pichon & Savy ; Paris. korreljacija, ammonity). Stratigrafija, Geologicheskaja IMMEL , H . 1987. Die Kreideammoniten der Nördlichen Korreljacija , 7 (4), 40-53. Kalkalpen. Zitteliana, 15 , 3-163. BUSNARDO , R . in J.C. F ISCHER & H. G AUTHIER 2006. Révi - JACOB , C H. 1907. Études paléontologiques et stratigraphiques sion critique de la Paléontologie Francaise d´Alcide sur la partie moyenne des terrains crétacés dans les d´Orbigny, vol. 4, Céphalopodes Crétacés, pp. 152-153 . Alpes francaises et les régions voisines. Annales de Backhuys Publishers ; Leiden – Pays-Bas. l´Université de Grenoble, 19 , 221-534. BUSNARDO , R., C HAROLLAIS , J.-J., W EIDMANN , M. & CLA - KAKABADZE , M.V . 1989. The Barremian biostratigraphical VEL , B. 2003. Le Crétacé inférieur de la Veveyse de subdivisions of Georgia (USSR) and comparison with Châtel (Ultrahelvétique des Préalpes externes; canton de some Western Mediterranean Regions. In : J. WIEDMANN Fribourg, Suisse). Revue de Paléobiologie, 22 (1), 1-174. (Ed .), Cretaceous of the Western Tethys. Proceedings 3 rd DELANOY , G . 1992. Les ammonites du Barrémien supérieur International Cretaceous Symposium, Tübingen 1987, de Saint-Laurent de l´Escarène (Alpes Maritimes), Sud- pp. 551-560. E. Schweizerbart´sche Verlagsbuchhand - Est de la France. Annales du Muséum d´Histoire Natu - lung ; Stuttgart. relle de Nice, 9, 1-148. — 2004. New and less known ammonites of the family 422 ZDENĚK VAŠÍČEK

Macroscaphitidae Hyatt, 1900 from the Barremian-Apt - QUENSTEDT , F.A . 1845-1849. Petrefactenkunde Deutsch - ian of Georgia and Crimea. Trudy Geologicheskogo In - lands. Der ersten Abtheilung. Erster Band. Cephalopo - stituta Akademii Nauk Gruzii, Novaja Serija , 119, 360- den. 581 p . L. F. Fues ; Tübingen. 367. [ In Russian ] PICTET , F.J. & R OUX , W . 1847. Description des mollusques KAKABADZE , M.V . in KOTETIŠVILI , E.V., K VANTALIANI , I.V., fossill qui se trouvent dans les Grės Verts des environs KAKABADZE , M.V. & CIKERIDZE , L. R. 2005. Atlas of de Genėve. Mémoires de la Société Physique et d´His - Early Cretaceous fauna of Georgia. Family Macroscap - toire Naturelle Genėve, 11 , (2), 247-412. hitidae Hyatt, 1900. Trudy Akademii nauk Gruzii, Geo - REBOULET , S., H OEDEMAEKER , P.J. (reporters), AGUIRRE -U R- logicheskij Institut im. A.I. Dzhanelidze, Novaja serija, RETA , M.B., A LSEN , P., A TROPS , F., BARABOSHKIN , E.Y., 120, 291-293. Tbilisi. [In Russian with English sum - COMPANY , M . & al . 2006. Report on the 2 nd interna - mary ] tional meeting of the IUGS Lower Cretaceous ammonite KILIAN , W . in BLAYAC , J . 1900. Sur le Crétacé inférieur du working group, the “Kilian Group” (Neuchâtel, Switzer - bassin de l´Oued Cherf (Algérie). Travaux du Labora - land, 8 September 2005). Cretaceous Research, 27, toire de Géologie de la Faculté des Sciences de l´Uni - 712-715. versité de Grenoble, 5, 19-27. ROTH , Z. & M ATEJKA , A . 1953. The pelosiderites of the KILIAN , W . 1910-1913. Erste Abteilung: Unterkreide Moravosilesian Beskydy, their historical importance, (Palaeocretacicum). Lieferung 1 – 3. In : F. F RECH geological occurrence, petrographical and chemical na - Lethaea Geognostica. II. Das Mesozoicum, Band 3 ture, and their origin . Geotechnica, 16, 1-111. [In Czech (Kreide). 398 p. Schweitzerbart ; Stuttgart. with English summary ] KLEIN , J . 2005. Lower Cretaceous Ammonites I. Perisphinc - ROUCHADZÉ , I. 1933. Les ammonites Aptiennes de la Géor - taceae 1. Himalayitidae, Olcostephanidae, Holcodisci - gie occidentale. Bulletin de l´Institut Géologique de dae, Neocomitidae, Oosterellidae. Fossilium Catalogus Géorgie, 1 (1932), 165-273. I: Animalia ( Ed . W. RIEGRAF ). Pars 139. 484 p. Backhuys RUCHADZE , I.M . 1938. Quelques céphalopodes nouveaux Publishers ; Leiden. ou peu connus de l´Aptien de la Géorgie . Bjulleten´ Ge - KOTETIŠVILI , E.V . 1970. Stratigrafija i fauna kolchiditogo i ologicheskogo Instituta Gruzii, 3 (2), 129-189. [ In Geor - smezhnych gorizontov zapadnoj Gruzii. Mecniereba, 1- gian, with Russian and French summaries ] 116. Tbilisi. [In Russian ] SARKAR , S.S. 1955. Révision des ammonites déroulées du — 1980. Semejstvo Pulchelliidae H. Douville (iz nizhne - Crétacé inférieur du Sud-Est de la France. Mémoires de melovych otlozhenij Juga SSSR). Trudy Akademii Nauk la Société Géologique de la France (NS), 72 , 1-176. Gruzinskoj SSR, Geologicheskij Institut imeni A. I. Dzha - TIETZE , E . 1872. Geologische und paläontologische Mit - nelidze, Novaja serija , 67 , 1-110. [In Russian ] theilungen aus dem südlichen Theil des Banater Gebirg - MAEDA , H. & S EILACHER , A . 1996. Ammonoid taphonomy. stockes. Jahrbuch der Kaiserlichen-Königlichen Geol - In : N.H. L ANDMAN , K. T ANABE , R .A. D AVIS : Ammonoid ogischen Reichsanstalt, 22 , 35-142. paleobiology, 13, pp. 543-578 . Plenum Press ; New York. UHLIG , V. 1882. Die Wernsdorfer Schichten und ihre Äqui- MATHERON , P . 1880. Recherches paléontologiques dans le valente. Sitzungsberichte der Kaiserlichen Akademie der Midi de la France. 1-12. Marseille. Wissenschaften in Wien, Mathematisch-Naturwis - NEUMAYR , M. & UHLIG , V . 1881. Über Ammonitiden aus den senschaftliche Classe, 86 , 1-32. Hilsbildungen Norddeutschlands. Palaeontographica, — 1883. Die Cephalopodenfauna der Wernsdorfer Schichten. 27 , 129-203. Denkschriften der Kaiserlichen Akademie der Wis - NICKLĖS , R . 1894. Contributions á la paléontologie de Sud- senschaften Wien, Mathematisch-Naturwissenschaftliche Est de l´Espagne. Terrain crétacé. 1. Néocomien. Mé - Classe, 46 , 126-290. moires de la Société Géologique de France, 4 (fascicle VAŠÍČEK , Z. 1971. Report on the results of paleontological 3), 31-59. research in the Lower Cretaceous of the western part of NIKOLOV , T . 1964. Barremian ammonites from north-eastern Silesian Unit in the Moravo-Silesian Beskydy Moun - Bulgaria. Travaux sur la Géologie de Bulgarie, Série Pa - tains . Sborník Vědeckých Prací Vysoké Školy Báňské v léontologie, 6, 117-141. [ In Bulgarian ] Ostravě, Řada Hornicko-Geologická, 15 (1969), 121- OOSTER , W.A . 1860. Catalogue des Céphalopodes fossiles 143. [ In Czech with English summary ] des Alpes suisses. Couches Crétacées, pp . 98-160. Zur - — 1972. Ammonoidea of the Těšín-Hradiště Formation cher & Furrer ;Zürich. (Lower Cretaceous) in the Moravskoslezské Beskydy Mts. D´O RBIGNY , A.D . 1840-1842. Paléontologie française. Des - Rozpravy Ústředního Ústavu Geologického, 38 , 1-103. cription zoologique et géologique de tous animaux mol - — 1973. Über einige von V. Uhlig (1883) beschriebene lusqes et rayonnés fossiles de France. Terrains crétacés. Unterkreide-Ammoniten . Sborník Geologických Věd, 1. Céphalopodes. 662 p. Arthus-Bertrand ; Paris. Řada P – Paleontologie, 15 , 49-73. EARLY CRETACEOUS AMMONITES FROM THE WESTERN CARPATHIANS 423

— 1979. Die Cephalopoden der schlesischen Unterkreide small heteromorph ammonites from Barremian. und ihre paläogeographische Bedeutung. Aspekte der Palaeontology , 37 , 203-239. Kreide Europas. Beiträge zum 1. Symposium Deutsche VERMEULEN , J . 1996. Le Barremien de Saint-Martin, Escra - Kreide – Bindeglied zwischen Boreal und Tethys (Mün - gnolles (Alpes-Maritimes, France). Riviéra Scientifique ster i. Westfalen, April 1978). International Union of (1996), 53-76. Geological Sciences, Series A, Number 6 , 323-334. E. — 2000. Nouvelles données sur les répartitions stratigra - Schweizerbart´sche Verlagsbuchhandlung ; Stuttgart. phiques, les évolutions et les classifications de trois fa - — 1981 a. Zwei neue faunistische Fundorte der Unterkreide milles d´ammonites du Crétacé inférieur. Géologie Al - in der Silesischen Einheit und Zusammenfassung der Re - pine , 75 (1999), 123-132. vision der von V. Uhlig (1883) beschriebenen Ammoniten. — 2003. Étude stratigraphique et paléontologique de la fa - Sborník Vědeckých Prací Vysoké Školy Báňské v Ostravě, mille des Pulchelliidae (Ammonoidea, Ammonitina, En - Řada Hornicko-Geologická, 25 (1979), 119-134. democerataceae). Géologie Alpine, Mémoire Hors Sé - — 1981 b. Die Clansay-Ammoniten der höchsten Hradiště- rie, 42 (2002), 1-333. Schichten (Silesische Einheit, Tschechoslowakei). — 2005a. Boundaries, ammonite fauna and main subdivisions Sborník Vědeckých Prací Vysoké Školy Báňské v Os - of the stratotype of the Barremian. Géologie Alpine, Sé - travě, Řada Hornicko-Geologická, 25 (1979), 193-204. rie spéciale “Colloques et Excursions”, 7, 147-173. — 1990. Unterkreide-Ammoniten aus neu abgeteuften — 2005b. Sur quatre espèces particulières d´ammonites du Schächten im Gebiet von Frenštát – Trojanovice Barrémien du Sud-est de la France. Annales du Muséum (Äussere Karpaten, Silesische Einheit, ČSFR). Acta d´Histoire Naturelle de Nice, 20 , 1-24. Musei Moraviae (Brno), Scientiae Naturales, 75 , 95-116. — 2006. Nouvelle classification a fondement phylogéné - VAŠÍČEK , Z. & K LAJMON , P . 1998. Contribution to the tique des ammonites hétéromorphes du Crétacé infé - knowledge of some small early Barremian ammonites rieur. Annales du Muséum d´Histoire Naturelle de Nice, from the Silesian Unit (Outer Carpathians, Czech Re - 21 , 137-178. public). Věstník Českého Geologického Ústavu, 73 , VERMEULEN , J. & K LEIN , J. 2006. Endemocerataceae: Pul - 331-342. chelliidae. – Fossilium Catalogus I: Animalia ( Ed . W. VAŠÍČEK , Z. & S KUPIEN , P . 2002. Notes on some rare Bar - RIEGRAF ), pars 141, Lower Cretaceous Ammonites II, pp. remian/Aptian ancyloceratids from the Silesian Unit 187-255. Backhuys Publishers ; Leiden. (Outer Western Carpathians, Czech Republic). Journal WIEDMANN , J . 1966. Stammesgeschichte und System der po - of the Czech Geological Society, 47 , 65-74. sttriadischen Ammonoideen. Neues Jahrbuch für Geo - VAŠÍČEK , Z., S KUPIEN , P. & Š ULGAN , F . 2004. New occur - logie und Paläontologie, Abhandlungen, 127 ,13-81. rences of Lower Cretaceous ammonites in the western WRIGHT , C.W., C ALLOMON , J.H. & H OWARTH , M.K . 1996. part of the Silesian Unit (Barremian – early Aptian, Cretaceous Ammonoidea. – Treatise on Invertebrate Pa - Outer Western Carpathians, Czech Republic). Journal of leontology. Part L, Mollusca 4 Revised. The Geological the Czech Geological Society, 49 , 41-55. Society of America, the University of Kansas: 1-362. VAŠÍČEK , Z. & W IEDMANN , J. 1994. The Leptoceratoidinae: Boulder, Colorado & Lawrence, Kansas.

Manuscript submitted: 24 th July 2007 Revised version accepted: 15 th June 2008 ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 1

PLATE 1

1 – Phyllopachyceras vergonsense DELANOY & JOLY ; B13808. Heavily deformed original shell with corroded ventral area. Spoil heap M5, Malenovice. Late Bar - remian. 2 – Macroscaphites cf. abchasiensis M. K AKABADZE ; B13810. A small part of shell coiled evolutely and its transition to the shaft. Spoil heap M1, Malenovice. Late Barremian – earliest Aptian. 3, 4 – Melchiorites blayaci (KILIAN ); 3 – B13812, 4 – B13811; On the phragmocone in both specimens the original shell is partly removed; body chambers with origi - nal shells. In 4 the body chamber is dissolved, incomplete on the circumference . Spoil heap T5, Tichá. Early Barremian. 5 – ?Valdedorsella hoheneggeri sp. nov.; B13813. Original shell preserved imper - fectly mainly in the area of the body chamber. Spoil heap KZ2, Kozlovice – Žáry ; late Late Barremian. 6-8 – Haplobrancoceras subquadratum AVRAM ; 6 – B13816, juvenile shell best pre - served ; 7 – B13817, enlarged deformed original shell with growth lines preserved on the last whorl, × 2; 8 –B13818, deformed shell, in the area of the aperture, with three incomplete ribs . Early and Late Barremian, spoil heap T1, Tichá and spoil heap M5, Malenovice. 9, 10 – Kotetishvilia cf. compressissima (D´O RBIGNY ); B13814; 9 – × 1, 10 – × 2. External mould of juvenile specimen. Outcrop T9 near the municipality of Tichá ; Early Barremian. 11, 12 – Kotetishvilia ex gr. sauvageaui (HERMITE ); B13815; 11 – × 1, 12 – × 2. External mould with the remains of the original shell. Spoil heap T3, Tichá ; Late Bar - remian.

All figures natural size unless otherwise stated ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 1

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12 11

10 ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 2

PLATE 2

1 – ?Heteroceras ex gr. emerici (D´O RBIGNY ); B13822. Fragment of deformed shaft . Spoil heap M5, Malenovice ; Late Barremian. 2 – Heteroceras aff. couleti DELANOY ; B1313821 . Small fragment of the shaft. Spoil heap KZ2, Kozlovice – Žáry ; late Late Barremian. 3 – ?Heteroceras sp. ; B13823 . Straight shafts and flexus. Outcrop in the Bílý potok stream near the municipality of Čeladná ; late Late Barremian. 4 – ?Audouliceras sp. ; B13819 . A fragment of the coiled part of the shell. Spoil heap M1, Malenovice ; Late Barremian or earliest Aptian. 5 – Lyticoceras cf. jauberti (ASTIER ); B13820 . A hook-like fragment of the adult shell with characteristically diverging shafts. Location as for fig. 4.

All figures natural size ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 2

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4

3 2 5 ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 3

PLATE 3

1 – Duyeina aff. glemmbachensis (IMMEL ); B13825. Incomplete proversum, retroversum and flexus. Spoil heap K7 between Kozlovice and Tichá ; Barremian. 2 – Pachyhemihoplites sp. ; B13824. Fragments of a penultimate and ultimate whorls. Spoil heap M5, Malenovice ; Late Barremian. 3 – Paradeshayesites semenovi (BOGDANOVA ); B13826. Strongly deformed original shell. Outcrop on the right bank of Ostravice River near the municipality of Ostravice ; Early Aptian.

All figures natural size

Photos taken by K. Mezihoráková (Ostrava). All specimens were coated with ammonium chloride before photography. Specimens are deposited in the Ostrava Museum in Ostrava under inventory numbers which are referred to in the text ACTA GEOLOGICA POLONICA, VOL. 58 Z. VAŠÍČEK , PL. 3

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3