Vol. 15(2): 83–93

THE COLLECTION OF TERRESTRIAL (: ) AT THE MUSEUM OF NATURAL HISTORY, WROC£AW UNIVERSITY (POLAND)

ANDRZEJ WIKTOR1 &JOLANTA JURKOWSKA2

Museum of Natural History, Wroc³aw University, Sienkiewicza 21, 50-335 Wroc³aw, Poland (e-mail: [email protected], [email protected]) ABSTRACT: The collection of terrestrial slugs of the Museum of Natural History, Wroc³aw University, comprises 69 descriptive types as well as material representing 172 mainly from the Northern Hemisphere. KEY WORDS: Gastropoda, Pulmonata, terrestrial slugs, type material, Museum of Natural History, Wroc³aw, Po- land

INTRODUCTION

The Museum of Natural History was established in of the Bulgarian materials come from the collection 1814 along with the University of Wroc³aw then com- of the late Prof. Dr. habil. JAROS£AW URBAÑSKI, ing into existence. Its original name was Zoological Poznañ, and were offered as a gift by the family of this Museum, and precisely “Zoologisches Museum” as it researcher after his death. In the last half a century, was German. By the mid-20th century the Museum the Museum’s malacological section gained abundant had housed shells both of land and marine snails, but materials left by Master’s and doctor’s students, and specimens had been lacking. In 1962, with the also specimens collcted occasionally by the employees employment of ANDRZEJ WIKTOR, the Museum’s of this institution. Donations of colleagues, especially malacological collection got enriched with his private A. WIKTOR’s co-authors, as well as of private collectors collection of gastropods, including slugs, which had or museums that the author of the present paper been being built for 10 years then. These specimens identified slugs for, constitute an important part of are labelled as “The collection of Andrzej Wiktor”, the Polish collection. These gifts include a series of and their considerable part is composed of slugs from specimens from the territory of the former USSR the Polish territory. Since A. WIKTOR started to spe- which were presented by Prof. I. M. LIKHAREV from cialise in slugs (until recently named Gastropoda Sankt Petersburg. terrestria nuda), the collection of these invertebrates At present the whole collection of slugs is ordered has been incessantly expanding; the material was and nearly all specimens determined. There are, how- mostly collected personally by the senior author dur- ever, rather few specimens, mainly from Poland, with- ing his field research in Poland, Bulgaria, Croatia, out identification labels, which could be used in fur- Bosnia and Herzegovina, Macedonia, Serbia, Greece, ther research. Spain, the Caucasus, Turkey, the Canary Isl., The majority of the material is kept in jars, the Tadzhikistan, China, Tibet and Papua-New Guinea. specimens preserved in 70–75% ethyl alcohol. They Also other malacologists, particularly Prof. Dr. habil. are arranged according to families and genera, and ADOLF RIEDEL from Warsaw, contributed to the en- within them – alphabetically according to specific largement of the collection, acquiring slugs specially names. Types of all categories are excluded from this for this purpose in different parts of the globe. Some section of the collection and kept separately in an in- 84 Andrzej Wiktor & Jolanta Jurkowska dividual cabinet (along with the type specimens of sions, monographs and larger publications for which other ). A list of types has already partly been the collection concerned constituted voucher mate- published (JADWIGA WIKTOR 1992). At present the rial. collection holds 66 types of slugs. The whole slug collection comprises 520 jars. The ACKNOWLEDGEMENTS total number of specimens is only roughly estimated, and it is about 43,500. On behalf of the Museum of Natural History and The specification below provides detailed data on their own, the authors wish to express their sincere the locality only for the types. In the case of other ma- thanks to all people who have contributed to the en- terial, exclusively for better orientation, the country richment of the Wroc³aw slug collection, and whose or geographical region of origin of the particular names, as collectors, are to be found on the labels. samples are mentioned. The list of references includes only the papers that Wroc³aw, 30th December, 2006 are the sources of descriptions of new species, revi-

LIST OF SLUGS AT THE COLLECTION OF THE MUSEUM OF NATURAL HISTORY, WROC£AW UNIVERSITY (STATE AT THE END OF 2006)

Familia ONCHIDIIDAE Rafinesque, 1815 pictum (Stoliczka, 1873) Onchidiidae (unidentified species) Korea Bahrain, New Guinea Meghimatium rugosum (Chen et Gao, 1982) (juv.) Taiwan Familia VERONICELLIDAE Gray, 1840 ?Meghimatium striatum Hesselt, 1823 (juv.) Laevicaulis alte (Férussac, 1821) India Bahrain (including laboratory-bred specimens), Oahu (Hawaii) hemphilli (Binney, 1885) U.S.A. Phylocaulis sp. Peru carolinianus (Bose, 1802) U.S.A. Veronicella sp. China (unidentified species) Mexico Veronicellus sp. Fiji-Lau Island Familia Godwin-Austen, 1888 (= PARMARIONIDAE Godwin-Austen, Familia RATHOUSIDAE Heude, 1885 1908) Atopus sp. Ariophantidae (unidentified species) Papua-New Guinea, Thailand India

Familia RYTHIDIDAE Pilsbry, 1893 Familia Gray, 1840 Schizoglossa sp. alpinus Pollonera, 1887 (Unidentified) Italy, Slovenia Familia ATHORACOPHORIDAE P. Fischer, Arion anguloi Martin et Gomez, 1988 1883 Paratypes! No. MP 609 Berantevilla, Spain Athoracophorus bitentaculus (Quoy et Gaimard, 1832) Arion anthracius Bourguignat, 1866 Spain Athoracophorus sp. New Zealand Arion ater (Linnaeus, 1758) Spain Familia PHILOMYCIDAE Gray, 1847 Arion baeticus Garrido, Castillejo et Iglesias, 1994 Meghimatium bilineatum (Benson, 1842) Paratypes! No. MP 639 Russia, Korea, India, Taiwan, China Puerto del Madrono, Spain The collection of terrestrial slugs at the Museum of Natural History, Wroc³aw University 85

Arion circumscriptus Johnston, 1892 Arion subfuscus (Draparnaud, 1805) Poland, Hungary, Roumania, Russia, Bulgaria, Neotype! Lectotype! No. MP 649 (design. Garrido, Germany, South Africa Castillejo et Iglasias, 1995) Montagne Noir, France Arion distinctus Mabille, 1868 Other material: Poland, Latvia, Greece, Slovenia, Poland, Latvia, England, Bulgaria, Hungary, New Macedonia, Herzegovina, Croatia, Montenegro, Zealand, Croatia, Serbia Bulgaria, Hungary, Italy, Spain Arion fagophilus De Winter, 1986 Arion urbiae De Winter, 1986 Spain Paratypes! No. MP 590 (Nilsson, 1823) Urbia, Sierra de Aitzgorri, Spain Poland, Bosnia, Croatia, Hungary, Germany ?Arion vejdovskyi Babor et Kostal, 1894 Arion hessei (Simroth, 1894) Germany Spain Arion wiktori Parejo et Martin, 1990 Arion hispanicus Simroth, 1889 Paratypes! No. MP 622 Topotypes! Valvanera, Spain Portugal Geomalacus anguiformis (Morelet, 1845) Arion hortensis Ferussac, 1819 Spain The Netherlands, England, New Zealand Geomalacus maculosus Allmann, 1843 Arion intermedius (Normand, 1852) (= minimus) Spain Poland, Germany, New Zealand Geomalacus malagensis Wiktor et Norris, 1991 Arion iratii Garrido, Castillejo et Iglesias, 1995 Holotype and paratypes! No. MP 613 Paratypes! No. MP 650 Colmenar, Prov. Malaga, Spain Selva de Irati, Spain Letourneuxia numidica Bourguignat, 1866 Arion isseli (Lessona et Pollonera,1882) Morocco Sardinia Arion lizarrustii Garrido, Castillejo et Iglesias, 1995 Familia OOPELTIDAE Cockerell, 1891 Paratypes! No. MP 651 Oopelta nigripunctata Mich, 1865 Alto de Lizarrustii, Novara, Spain South Africa

Arion lusitanicus Mabille, 1848 Familia ANADENIDAE Pilsbry, 1948 Poland, Slovenia, Bulgaria, England, The Netherlands, Germany, France, Spain, Portugal Anadenus altivagus (Theobald, 1862) Nepal, Pakistan Arion molinae Garrido, Castillejo, Iglesias, 1995 Paratypes! No. MP 652 Anadenus (Neoanadenus) gonggashensis Wiktor, Chen La Molina, Catalonia, Spain et Wu, 2000 Paratypes! No. MP 704 Arion molleri Pollonera, 1889 Sichuan Prov., China Spain Anadenus (Anadenus) nepalensis Wiktor, 2001 Arion ovenii Davies, 1979 Holotype and paratypes! No. MP 817 Great Britain, Ireland (Buncrana) Nepal Arion paularensis Wiktor et Parejo, 1989 Anadenus (Anadenus) yangtzeensis Wiktor, Chen et Holotype! No. MP 592 Wu, 2000 Catas al Paular, Sierra de Guadarama Mts., Spain Paratypes! No. MP 705 Arion rufus (Linnaeus, 1758) Sichuan and Yunnan Prov., China Poland, Spain, the Netherlands, Italy Familia Ellis, 1926 Arion silvaticus Lohmander, 1937 Poland, Bulgaria, Slovenia, Macedonia, Bosnia, aegaeicus Wiktor et Mylonas, 1986 Croatia, Serbia, Montenegro, Herzegovina Holotype and paratypes! No. MP 573 Leukes, Amorgos Isl., Greece Other material: Greece Milax altenai Forcart, 1972 Greece, Turkey 86 Andrzej Wiktor & Jolanta Jurkowska

Milax ater (Collinge, 1895) fejervaryi (Wagner, 1929) Algeria Croatia Milax barypus Bourguignat, 1866 ? Tandonia kaleniczenkoi (Clessin, 1883) Israel Crimea Milax caucasicus obscurus (Simroth, 1912) Tandonia kusceri (Wagner, 1931) Azerbaijan (Talysh) Bulgaria, Croatia, Macedonia, Serbia (Draparnaud, 1801) Tandonia macedonica (Rähle, 1974) Canary Islands, Spain, Corsica, Italy, Morocco, Macedonia Algeria, South Africa Tandonia melanica Wiktor, 1986 Milax monstruosus Wiktor, 1986 Holotype and paratypes! No. MP 569 Holotype! No. MP 571 S of Joanina, Therakison, Greece Latakia, Syria Tandonia pageti (Forcart, 1972) Milax nigricans (Schulz, 1836) (= insularis) Turkey Algeria, Spain, Majorca, Corsica, Canary Islands, Tandonia piriniana (Wiktor, 1975) Croatia, Malta, Italy, The Netherlands, Turkey Holotype and paratypes! No. MP 468 Milax (Milax) parvulus Wiktor, 1968 Pirin, Koprivlen, Bulgaria Holotype and paratypes! No. MP 457 Tandonia rara Wiktor, 1996 Stara Planina, Iskrec, Bulgaria Holotype and paratype! No. MP 647 Milax (Tandonia) pinteri Wiktor, 1975 Knin, Croatia (at present =Tandonia pinteri (Wiktor, 1975)) Tandonia reuleauxi (Clessin, 1887) Holotype and paratypes! No. MP 468 Croatia, Montenegro, Bosnia, Herzegovina Rodopi Mts: Èudni Mostove, Bulgaria Tandonia robici (Simroth, 1885) Milax riedeli Wiktor, 1968 Italy, Slovenia, Croatia Holotype! No. MP 572 Iskenderun (=Alexandreta), Turkey (Millet, 1843) Poland, Italy Milax (Subamalia) totevi Wiktor, 1975 (at present = Tandonia totevi (Wiktor, 1975) Tandonia serbica (Wagner, 1931) Holotype and paratypes! No. MP 459 Bulgaria Rodopi Mts, Bulgaria, Kriti Tandonia simrothi (Hesse, 1923) Other material: Bulgaria Italy Milax verrucosus Wiktor, 1969 (Férussac, 1823) Holotype and paratypes! No. MP 460 Greece, Croatia, Serbia, The Netherlands, Spain, Stara Planina Mts, Gabrovo, Bulgaria Italy, Great Britain, Albania, Turkey Other material: Bulgaria Tandonia albanica (Soós, 1924) Familia P. Fischer, 1856 Albania, Macedonia Candaharia(Levanderia) levanderi (Simroth, 1901) (Hazay, 1881) (= koznakovi) Poland, Bulgaria, Italy, Macedonia, Serbia, Hungary, Kirghizia, Tadzhikistan Turkey, Germany, The Czech Republic, Candacharia rutelum (Hutton, 1849) Roumania, New Zealand Afghanistan, Russia (environs of St. Petersburg), Tandonia bosnensis Wiktor, 1986 Pakistan, Kirghizia, Tadzhikistan Holotype and paratypes! No. MP 570 (Cryptella) canariensis, Webb et Berthold, Trebevic Mt., Bosnia and Herzegowina 1833 Tandonia cavicola (Simroth, 1916) Canary Islands (Lanzarote Isl.) Croatia, Herzegovina, Montenegro Parmacella deshayesi Moquin-Tandon, 1848 Tandonia cretica (Simroth, 1885) Morocco, South Africa Greece, Turkey Parmacella festae Gambetta, 1925 Tandonia cristata (Kaleniczenko, 1851) Libya Bulgaria, Crimea, Roumania Parmacella ibera Eichwald, 1841 Russia (Caucasus), Azerbaijan The collection of terrestrial slugs at the Museum of Natural History, Wroc³aw University 87

Parmacella tenerifensis Alonso, Ibanez et Diaz, 1985 horezia Grossu et Lupu, 1967 Paratypes! No. MP 564 Paratypes! No. MP 1003 La Laguna (Tenerife Isl.), Spain Vaideeni – Horezu, Roumania Other material: Bulgaria Parmacella valencienni Web et Vanbenenden, 1836 Spain, Canary Islands Lehmannia janetscheki Forcart, 1966 Italy Parmacella sp. – a subfossil shell Greece Lehmannia jaroslaviae Grossu, 1967 Paratypes! No. MP 664 Familia TESTACELLIDAE Gray, 1840 Jaroslavae – Salatures Arges, Roumania haliotidea Draparnaud, 1801 Other material: Roumania Canary Islands Lehmannia macroflagellata Grossu et Lupu, 1967 Testacella sp. Paratypes! No. MP 1004 Spain, Italy Giumalau, Roumania Other material: Poland Familia Fitzinger, 1833 (Müller, 1774) (= PLUTONIINAE Cockerell, 1893) Poland, Spain, Slovenia, Croatia, Serbia, Bosnia and Plutonia atlantica (Morelet, 1860) Herzegovina, Italy Azores (Bourguignat, 1861) Bulgaria, Poland, Montenegro Familia Lamark, 1801 Lehmannia rupicola Lessona et Pollonera, 1880 Subfamilia Rafinesque, 1815 Germany, Spain Caspilimax keyserlingi (Martens, 1880) (= kollyi) Lehmannia szigethyae Wiktor, 1975 Azerbaijan (Lenkoran – Talysh) Holotype and paratypes! No. MP 456 Galaèica, Macedonia Gigantomilax (Gigantomilax) lederi (Boettger, 1883) Other material: Greece, Albania, Macedonia (= kollyi – forma) Georgia, Azerbaijan, Turkey, Adzhariya Lehmannia valentiana (Férussac, 1823) Poland, Spain, Canary Islands, Majorca, Corsica, Gigantomilax (Monochroma) brunneus (Simroth, 1901) Greece, Morocco, The Netherlands, Israel, China Armenia ? (Limax) albipes Dumont et Mortillet, 1852 Gigantomilax (Monochroma) lenkoranus Simroth, 1912 Austria Azerbaijan ?Limax (Limax) bielzi Siebert, 1873 Gigantomilax (Vitrinoides) cecconi Simroth, 1906 Poland Israel Limax (Limax) cephalonicus Simroth, 1886 Gigantomilax (Vitrinoides) dagestanus (Simroth, 1886) Greece, Macedonia (= florenskii) Turkey Limax (Limax) cinereoniger Wolf, 1803 Poland, Bulgaria, Slovenia, Croatia, Serbia, Bosnia, Gigantomilax (Vitrinoides) eustrictus Heynemann, 1882 Macedonia, Montenegro, Roumania, Italy, Israel Greece Gigantomilax (Vitrinoides) majoricensis (Heynemann, Limax (Limax) conemenosi Boettger, 1882 1863) Bulgaria, Greece, Macedonia, Kosovo, Italy Neotype! No. MP 999 Barranc d'Algendar (Ferreries, Minorca) Limax (Limax) corsicus Moquin-Tandon, 1855 Corsica Isl., Capraia Isl., Sardinia Isl. Gigantomilax (Vitrinoides) monticola armeniacus (Simroth, 1886) Limax (Limax) graecus Simroth, 1889 (= macedonicus) Georgia, Azerbaijan, Turkey Bulgaria, Greece, Macedonia Lehmannia brunneri (Wagner, 1931) (= Gigantomilax Limax (Limax) maximus Linnaeus, 1758 occidentalis) Poland, Spain, Bulgaria, Croatia, Serbia, Bosnia and Bulgaria, Croatia, Serbia, Montenegro, Macedonia, Herzegovina, Montenegro, Roumania, Albania, Greece Greece 88 Andrzej Wiktor & Jolanta Jurkowska

Limax (Limax) maximus altenae Grossu et Lupu, 1960 Subfamilia EUMILACINAE Likharev et Wiktor, Roumania 1980 Limax (Limax) polipunctatus Pollonera, 1888 Eumilax brandti (Martens, 1880) Italy Georgia, Dagestan, Abkhazia, Turkey Limax (Limax) punctulatus Sordelli, 1870 Eumilax brandti salamandroides (Simroth, 1901) Bulgaria Caucasus Limax (Limax) seticus Wiktor et Bössneck, 2004 Eumilax intermittens (O. Boettger, 1883) (= reddei) Holotyp and paratypes! No. MP 968 Caucasus (Abkhazia, Karachai-Cherkessia), Iran, Chachour, Prov. Seti, Nepal Azerbaijan, Dzungaria Limax (Limax.) subalpinus Lessona, 1880 Metalimax varius (O. Boettger, 1884) Bulgaria Caucasus (Abkhazia) Limax (Limax) wohlberedti Simroth, 1900 Croatia, Herzegovina, Albania Familia H. Wagner, 1935 Limax () ecarinatus (Boettger, 1881) Subfamilia AGRIOLIMACINAE H. Wagner, 1935 (= maculatus = grossui) (Deroceras) adolphi Wiktor, 1998 Bulgaria, Greece, Ireland, Turkey Holotype and paratypes! No. MP 668 Limax (Limacus) flavus Linnaeus, 1758 Moulis near St. Girons, France Poland, Bulgaria, Greece, Germany, Croatia, Deroceras (Deroceras) agreste (Linnaeus, 1758) Macedonia, Hungary, Albania, Spain, Canary Poland, Sweden, Georgia, Hungary, Spain, Bulgaria Islands, Majorca, Crimea, Turkey, Azerbaijan, Brazil, China, New Zealand, Latvia, South Africa Deroceras (Deroceras) altaicum Simroth, 1886 Russia (Sakhalin), Caucasus (Kabardino-Balkaria), kostali Babor, 1894 Altai Mts, China, Kirghizia, Kazakhstan Herzegovina Deroceras (Deroceras) altimirai Altena, 1969 Malacolimax mrazeki (Simroth, 1904) Spain Bosnia, Herzegovina, Croatia, Montenegro, Macedonia, Slovenia Deroceras (Deroceras) astypalaeensis Wiktor et Mylonas, 1992 (Müller, 1774) Holotype and paratypes! No. MP 630 Poland Dodecanesos Archipelago, Astipalaia Isl., Greece Malacolimax wiktori Alonso et Ibanez, 1889 Deroceras (Deroceras) bakurianum (Simroth, 1912) Paratypes! No. MP 624 Russia (Caucasus – Sochi), Ukraine (Crimea), La Laguna (Tenerife Isl.), Spain Azerbaijan (Baku), Turkey Turcomilax (Michaelisia) natalianus (Michaelis, 1892) Deroceras (Deroceras) barceum (Gambetta, 1889) Dzungaria, Kurdistan, Kirghizia, Kazakhstan Libya (Cyrenaica) Turcolimax (Turcolimax) nanus (Simroth, 1901) Deroceras (Deroceras) berytensis (Bourguignat, 1852) Kazachstan, Kirghizia (Przewalsk), Dzungaria Israel, Turkey Turcomilax (Tanlimax) oli Wiktor, Naggs et Gupta, Deroceras (Deroceras) bistimulatum Wiktor, 2000 1999 Holotype and paratypes! No. MP 629 Holotype and paratypes! No. MP 703 Pilio Mts., Crete, Greece Naina Peak, Himalayas, India Deroceras (Agrolimax) boeoticum Wiktor, 1984 Turcolimax (Tanlimax) turkestanus (Simroth, 1898) (at present = Deroceras (Deroceras)) Kirghizia (Przewalsk) Holotype and paratypes! No. MP 478 Subfamilia BIELZINAE Likharev et Wiktor, Levadhia, Prov. Voicia, Greece 1980 Deroceras (Deroceras) callatis Grossu et Lupu, 1963 Paratypes! No. MP 665 (Bielz, 1851) Mangalia, Comarova Wald, Roumania Poland, Roumania Deroceras (Deroceras) cazioti (Pollonera, 1894) Corsica The collection of terrestrial slugs at the Museum of Natural History, Wroc³aw University 89

Deroceras (Plathystimulus) cycladicum Wiktor et Deroceras (Deroceras) kasium Rähle, 1993 Mylonas, 1981 Greece (at present = Deroceras (Deroceras)) Deroceras (Deroceras) keaensis Altena, 1973 Holotype and paratypes! No. MP 466 Greece Cyclades, Sikinos, Greece Deroceras (Deroceras) korthionensis Winter et Butot, Deroceras (Deroceras) chrysorrovatensis Rähle, 1984 1985 Cyprus Greece Deroceras (Deroceras) dallai Giusti, 1970 Deroceras (Deroceras) kythirensis Wiktor, Vardino- Italy (Elba Isl.) yannis et Mylonas, 1994 Deroceras (Deroceras) damascensis (Germai, 1911) Holotype and paratypes! No. MP 634 (? = berytensis) Kythira Isl., Greece Israel Deroceras (Agriolimax) laconicum Wiktor, 1984 Deroceras (Deroceras) dewinteri Maassen, 2000 (at present = Deroceras (Deroceras)) Paratype! No. MP 839 Holotype and paratypes! No. MP 476 Crete, Greece Peloponnesus, Greece Deroceras (Deroceras) ercinae De Winter, 1985 Deroceras (Deroceras) laeve (Müller, 1774) Spain Poland, Spain, Bulgaria, Sweden, Labrador, Israel, Turkey, Tadzhikistan, China, Taiwan, India, Deroceras (Deroceras) gavdosensis Wiktor, Vardino- Yakutia, Pakistan, Greece, Macedonia, Croatia, yannis et Mylonas, 1994 Slovenia, New Guinea, Costa Rica Holotype and paratypes! No. MP 632 Gavdos Gavdos Isl., Greece Deroceras (Deroceras) lasithionensis lasithionensis Wiktor, Vardinoyannis et Mylonas, 1994 Derocers (Deroceras) gerasensis Rodriguez, Castillejo Holotype and paratypes! No. MP 635 et Outeiro, 1989 Crete, Nomos Lasithiou, Greece (at present = Furcopenis gerasensis) Other material: Greece Paratypes! No. MP 623 Serra de Geres, Portugal, Spain Deroceras (Deroceras) lasithionensis polylobatum Wiktor, Vardinoyannis et Mylonas, 1994 Deroceras (Deroceras) giustianum Wiktor, 1998 Holotype and paratypes! No. MP 636 Holotype and paratypes! No. MP 669 Crete, Nomos Lasithiou, Greece Sicily, Italy Other material: Greece Deroceras (Deroceras) golcheri Altena, 1962 Deroceras (Deroceras) libanoticum (Pollonera, 1909) Malta, Corsica Israel Deroceras (Deroceras) gorgonium Wiktor, Deroceras (Deroceras) lombricoides (Morelet, 1845) Vardinoyannis et Mylonas, 1994 Spain Holotype and paratypes! No. MP 633 Deroceras (Deroceras) lothari Giusti, 1971 Nomos Rethymnis, Crete, Greece Bulgaria, Italy, Hungary, Croatia, Slovenia, Austria Deroceras (Plathystimulus) hispaniensis Castillejo et Deroceras (Deroceras) malkini Wiktor, 1984 Wiktor, 1983 Holotype and paratypes! No. MP 475 (at present = Deroceras (Deroceras)) Hios, Thymiana, Greece Holotype and paratypes! No. MP 473 Other material: Greece Galizia, Vidan, Santiago, Spain Other material: Spain Deroceras (Deroceras) maasseni Wiktor, 1996 Paratype! No. MP 648 Deroceras (Deroceras) johanae De Winter et Butot, 1985 Valley of the Tara River, Montenegro Greece Other material: Serbia Deroceras (Deroceras) juranum Wütrich, 1993 Deroceras (Plathystimulus) melinum Wiktor et (= rodnae) Mylonas, 1981 Topotypes! (at present = Deroceras (Deroceras)) Switzerland Holotype and paratypes! No. MP 464 Deroceras (Plathystimulus) karnaniensis Wiktor, 1984 Milos Isl., Greece (at present = Deroceras (Deroceras)) Holotype and paratypes! No. MP 477 Karnania, Greece 90 Andrzej Wiktor & Jolanta Jurkowska

Deroceras (Deroceras) minoicum Wiktor, Deroceras (Deroceras) reticulatum (Müller, 1774) Vardinoyannis et Mylonas, 1994 Poland, Bulgaria, Germany, Spain, Canary Islands, Holotype and paratypes! No. MP 637 Slovenia, Croatia, Montenegro, Macedonia, Crete, Nomos Rethymnis, Greece Karachai-Cherkessia (Caucasus), Italy, New Other material: Greece (Crete) Zealand, South Africa Deroceras (Deroceras) neuteboomi Forcart, 1972 Deroceras (Deroceras) ridelianum Wiktor, 1983 Greece Holotype and paratypes! No. MP 470 Skikda, Algeria Deroceras (Deroceras) nitidum (Morelet, 1845) Spain Deroceras (Deroceras) rodnae Grossu et Lupu, 1965 Poland, Slovakia (Mala Fatra), Roumania Deroceras (Deroceras) nyphoni De Winter et Butot, (Dobrudja), Croatia, Germany, Spain, Hungary, 1986 Austria, Italy Paratypes! No. MP 607 Sporades, Skiantos, Greece Deroceras (Deroceras) romanicum Grossu et Lupu, Other material: Greece 1959 (= sturanyi) Hungary Deroceras (Deroceras) oertzeni (Simroth, 1889) Greece Deroceras (Deroceras) samium Rähle, 1983 Greece Deroceras (Deroceras) osseticum (Simroth, 1901) Batumi, Turkey Deroceras (Deroceras) schleschi Grossu et Lupu, 1961 Specimen sent by Grossu as paratype, the label does Deroceras (Deroceras) pageti Grossu, 1972 not agree with the list of paratypes – see Arch. Bulgaria, Greece Mol. 1961, 90: 24 Deroceras (Deroceras) panormitanum (Lessona et Deroceras (Plathystimulus) seriphium Wiktor et Pollonera, 1882) (= caruanae) Mylonas, 1981 Spain, The Netherlands, Greece, Bulgaria, Slovakia, (at present = Deroceras (Deroceras)) South Africa, Austria, Poland, Germany, Italy, Holotype and paratypes! No. MP 467 Juan Fernandez, New Zealand Serifos, Kentarchos, Greece Deroceras (Plathystimulus) parium Wiktor et Mylonas, Other material: Greece 1981 ?Deroceras (Deroceras) steindachneri Wagner, 1931 (at present = Deroceras (Deroceras)) (= berytensis) Holotype and paratypes! No. MP 465 Turkey Paros Isl., Greece Deroceras (Deroceras) sturanyi (Simroth, 1894) Deroceras (Agrolimax) parnasium Wiktor, 1984 Poland, Gotland, Bulgaria, Macedonia, Croatia, (at present = Deroceras (Deroceras)) Bosnia, Montenegro, Serbia, Slovenia, Hungary, Holotype and paratypes! No. MP 474 Austria Parnas, Arakora, Greece Other material: Greece Deroceras (Deroceras) subagreste (Simroth, 1892) Abkhazia Deroceras (Deroceras) planarioides Simroth, 1910 (= rivanus) Deroceras (Deroceras) thersites (Simroth, 1886) Italy Bulgaria Deroceras (Deroceras) praecox Wiktor, 1966 Deroceras (Deroceras) turcicum (Simroth, 1901) Holotype and paratypes! No. MP 461 (= wiktori = forcarti) Biskupia Kopa Massif, Opawskie Mts., Poland Poland, Greece, Turkey, Crimea, Bulgaria, Croatia, Other material: Poland Montenegro, Bosnia, Macedonia, Serbia, Herzegovina, Slovenia, The Czech Republic, Deroceras (Plathystimulus) pseudopanormitanum Albania, Turkey Wiktor, 1984 (at present = Deroceras (Deroceras)) Deroceras (Krynickillus) urbanskii Wiktor, 1971 (at present = Krynickillus urbañskii) Holotype and paratypes! No. MP 521 Tymfi, N of Joannina, Greece Holotype and paratypes! No. MP 455 Other material: Greece Stara Planina, Bulgaria Other material: Bulgaria Deroceras (Deroceras) rethimnonensis De Winter et Deroceras (Deroceras) vascoana De Winter, 1986 Butot, 1986 Greece, Turkey Paratypes! No. MP 608 Navarra, Alto de Lizzarusti, Spain The collection of terrestrial slugs at the Museum of Natural History, Wroc³aw University 91

Deroceras (Deroceras) zilchi Grossu, 1969 Megalopelte simrothi Lindholm, 1914 Bulgaria Turkey, Adzharia Deroceras (Liolytopelte) bureschi (Wagner, 1934) Toxolimax hoplites Simroth, 1899 Bulgaria Turkey

Deroceras (Liolytopelte) caspium (Simroth, 1901) Subfamilia MESOLIMACINAE Hausdorf, 1998 Armenia Mesolimax brauni Pollonera, 1888 Deroceras (Liolytopelte) caucasicum (Simroth, 1901) Greece, Turkey Armenia, Georgia, Kabardino-Balkaria (Caucasus), Tadzhikistan Mesolimax escherichi Simroth, 1899 Turkey Deroceras (Liolytopelte) moldavicum (Grossu et Lupu, 1961) Familia BOETTGERILLIDAE Wiktor et Poland Likharev, 1980 Deroceras (Liolytopelte) trabzonensis Wiktor, 1994 pallens Simroth, 1912 Holotype and paratypes! No. MP 612 Poland, Turkey Trabzon, Turkey Boettgerilla vermiformis Wiktor, 1959 Furcopenis circularis Castillejo et Moscato, 1987 Lectotype and paralectotypes! No. MP 462 Paratype! No. MP 610 Miêdzygórze, Poland Montefurado, Lugo, Spain See: WIKTOR 1960 Furcopenis darioi Castillejo et Wiktor, 1983 Holotype and paratypes! No. MP 471 Familia PAPILLODERMATIDAE Wiktor, Martin Bosque los Cabaninos los Ancares, Spain et Castillejo, 1999 (= PAPILLODERMIDAE) Other material: Spain Papilloderma altonagai Wiktor, Martin et Castillejo, Furcopenis gallaeciensis Castillejo et Wiktor, 1983 1990 Holotype and paratypes! No. MP 472 Holotype and paratypes! No. MP 599 Galicia, Spain Asturias, Covandonga, Spain Other material: Spain Familia Hesse, 1882 Lytopelte maculata (Koch et Heynemann, 1874) Tadzhikistan, Ferganski Hrebiet Hyrcanolestes velitaris (Martens, 1880) (= bicolor) Azerbaijan Krynickillus melanocephalus Kaleniczenko, 1851 Georgia, Latvia, Germany, Karachai-Cherkessia Selenochlamys pallida Boettger, 1883 (Caucasus), Kabardino-Balkaria (Caucasus) Turkey Krynickillus urbanskii (Wiktor, 1971) Trigonochlamys imitatrix (Martens, 1880) See: Deroceras (Krynickillus) urbanskii Turkey

REFERENCES

ALONSO R., IBANEZ M., DIAZ J. A. 1985. A new slug from Ca- (Gastropoda: Pulmonata: Arionidae). Malak. Abh. 17: nary Islands (Pulmonata: Parmacellidae). Arch. Moll. 37–45. 116: 57–65. GARRIDO C., CASTILLEJO J., IGLESIAS J. 1995. The Arion sub- CASTILLEJO J., MASCATO R. 1987. Morphology and anatomy fuscus complex in the eastern part of the Iberian Penin- of new species of Furcopenis (Gastropoda, Pulmonata, sula, with redescription of Arion subfuscus (Draparnaud, Agriolimacidae). Monitore zool. ital. (N.S.) 21: 33–40. 1805) (Gastropoda, Pulmonata, Agrolimacidae). Arch. CASTILLEJO J., WIKTOR A. 1983. Furcopenis gen. n. with its two Moll. 124: 103–118. new species and a new Deroceras species from Spain GROSSU A. 1969. Beschreibung einiger neuer Deroceras Arten (Gastropoda, Pulmonata, Agrolimacidae). Malak. Abh. (Gastropoda, Limacidae). Arch. Moll. 99: 157–170. 9: 1–16. GROSSU A., LUPU D. 1961a. Revizuirea speciilor genului DRAPARNAUD J. P. R. 1805. Histoire naturelle des mollusques Milax Gray, 1855 existente in Romînia. An. Univ. C. I. trrestres et fluviatiles de la France. Paris. Parhon, Ser. ªtiinþi. Nat. (Biol.), 28: 127–148. GARRIDO C., CASTILLEJO J., IGLESIAS J. 1994. Description of GROSSU A., LUPU D. 1961b. Zwei neue Arten der Gattung Arion baeticus spec. nov. from Iberian Peninsula Deroceras Rafinesque, 1820. Arch. Moll. 90: 21–25. 92 Andrzej Wiktor & Jolanta Jurkowska

GROSSU A., LUPU D. 1962a. Contributions a la connaissance WIKTOR A. 1982. Contributions to the knowledge of the du Genre Lehmannia Heynemann (Gastropoda, Lima- slugs of Yugoslavia (Arionidae, Milacidae, Limacidae, cidae). Journ. Conchyl. CVI, fasc. 4: 119–128. Agriolimacidae – Gastropoda, Pulmonata). Ann. Zool. GROSSU A., LUPU D. 1962b. Zur Kenntnis der Gattung Leh- 36: 465–489. mannia Heynemann (Limacidae) und deren grosse Va- WIKTOR A. 1983a. The slugs of Bulgaria (Arionidae, riabilität, nebst Beschreibung neuer Arten. Arch. Moll. Milacidae, Limacidae, Agriolimacidae – Gastropoda, 91: 191–201. ). Ann. Zool. 37: 71–206. GROSSU A., LUPU D. 1963. Varibilité du genre Deroceras (fam. WIKTOR A. 1983b. Some data on slugs of Morocco and Alge- Limacidae, Gasteropodes terrestres) et description ria with description of new Deroceras species (Gastro- d’une nouvelle espéce. Travaux du Museum d’Histoire poda, Pulmonata). Malak. Abh. 8: 155–166. Naturelle “Grigoei Antipa” 4: 193–200. WIKTOR A. 1983c. Parmacellidae of the Mediterranean Area LESSONA M., POLLONERA C. 1882. Monografia dei Limacidi and Canary Islands (Gastropoda, Pulmonata). Malak. Italiani. Torino. Abh. 9: 81–96. MAASEN W. J. M. 2000. Deroceras dewinteri spec. nov. from is- WIKTOR A. 1984. Six Deroceras species from Greece, new for land of Crete, Greece (Gastropoda, Pulmonata, Agrio- science (Gastropoda, Pulmonata: Agriolimacidae). limacidae). Basteria 64: 125–126. Malak. Abh. 9: 151–164. MARTIN R., GOMEZ B. J. 1988. A new slug from the Iberian WIKTOR A. 1986. New for science Milacidae (, peninsula: Arion anguloi n. sp. (Pulmonata: Arionidae). Pulmonata). Malak. Abh. 11: 155–165. Arch. Moll. 118: 167–174. WIKTOR A. 1987a. Spermatophores in Milacidae and their PARJO C., MARTIN R. 1990. Arion wiktori sp. n. from Iberian significance for classification (Gastropoda, Pulmonata). Peninsula (Gastropoda, Pulmonata: Arionidae). Malak. Malak. Abh. 12: 85–100. Abh. 15: 25–35. WIKTOR A. 1987b. Milacidae (Gastropoda, Pulmonata) – sys- RODRIGUEZ T., CASTILLEJO J., OUTEIRO A. 1989. Morphol- tematic monograph. Ann. Zool. 41: 153–319. ogy and anatomy of new Iberian species: Deroceras WIKTOR A. 1994. Contribution to the knowledge of the slugs gerasensis (Gastropoda: Pulmonata: Agriolimacidae). of Turkey (Gastropoda terrestria nuda). Arch. Moll. 123: The Veliger 32: 95–100. 1–47. SIMROTH H. 1886. Weitere Mitteilungen über palaearktische WIKTOR A. 1996. The slugs of the former Yugoslavia Nacktschnecken. Malak. Jahrb. 13: 16–34. (Gastropoda terrestria nuda – Arionidae, Milacidae, URBAÑSKI J., WIKTOR A. 1968. Beiträge zur Kenntnis Bulga- Limacidae, Agriolimacidae). Ann. Zool. 46: 1–110. rischer Nacktschnecken (Moll., Pulm.) (Systematische, WIKTOR A. 1998. Two new for science Deroceras species from Zoogeoraphische und Ökologische studien über die France and the Italian island of Sicily (Gastropoda: Molluken der Balkan-Halbinsel. VIII). Bull. Soc. Amis Pulmonata: Agriolimacidae). Malak. Abh. 19: 13–19. Sci. Lett. Poznañ Ser. D 8: 47–95. WIKTOR A. 2000. Agriolimacidae (Gastropoda: Pulmonata) WIKTOR A. 1960. Description of Boettgerilla vermiformis Wik- – a systematic monograph. Ann. Zool. 49: 347–590. tor (Mollusca, Pulmonata). Bull. Soc. Amis Sci. Lett. WIKTOR A. 2001. A review of Anadenidae (Gastropoda: Poznañ. Ser. D 1: 151–156. Pulmonata) with a description of a new species. Folia WIKTOR A. 1961. Materialien zur Kenntnis der Gattung Boett- Malacol. 9: 3–26. gerilla Simroth, 1910 (Gastropoda, Limacidae). Ann. WIKTOR A., BÖSSNECK U. 2004. Limax (Limax) seticus n. sp. Zool. 19: 125–145. from high mountains in Nepal (Gastropoda: Pulmonata: WIKTOR A. 1966. Eine neue Nacktschnecken-Art (Gastropo- Limacidae). Folia Malacol. 12: 183–187. da, Limacidae) aus Polen. Ann. Zool. 23: 449–457. WIKTOR A., CASTILLEJO J. 1987. Deroceras lombricoides WIKTOR A. 1968. Eine Nacktschnecke Milax (Milax) parvulus (Morelet, 1845) and Deroceras malzani (Simroth, 1885) Wiktor, 1968 aus Bulgarien (Mollusca, Pulmonata) Neu- with their synonymy (Gastropoda, Pulmonata, Agrioli- beschreibng. Bull. Acad. Pol. Sci., Sér. sci. biol. 16: maidae). Malak. Abh. 12: 13–19. 419–422. WIKTOR A., CHEN DE-NIU,WU MING. 2000. Stylommato- WIKTOR A. 1969. Milax verrucosus sp. n., eine neue Nackt- phoran slugs of China (Gastropoda: Pulmonata) – schnecke aus Bulgarien (Mollusca, Pulmonata). Bull. Prodromus. Folia Malacol. 8: 3–35. Acad. Pol. Sci., Sér. sci. biol. 17: 47–49. WIKTOR A., MARTIN R., CASTILEJO J. 1990. A new slug family WIKTOR A. 1971. Die von der Niederlandischen Biologi- Papillodermidae with description of a new and schen Expedition in die Türkei in 1959 gesammelten species from Spain. Malak. Abh. 15: 1–18. Nacktschnecken (Milacidae und Limacidae, Pulmonata) WIKTOR A., MIENIS H. K. 2006. Redescription of Deroceras mit Beschreibung einer neuen Deroceras-Art aus dem Bal- libanoticum (Pollonera, 1909) (Mollusca: Pulmonata: kan-Gebiet und Türkei. Zool. Meded. 45: 261–280. Agriolimacidae). Folia Malacol. 14: 31–34. WIKTOR A. 1973. Die Nacktschnecken Polens, Arionidae, WIKTOR A., MILANI N. 1995. Contribution to the knowledge Milacidae, Limacidae (Gastropoda, Stylommatophora). of two scarcely known Alpine slugs, Tandonia simrothi Monogr. Fauny Polski 1. PWN, Warszawa–Kraków. (Hesse, 1923) and Deroceras planarioides (Simroth, 1910) WIKTOR A. 1975. New slug species (Pulmonata: Milacidae (Gastropoda: Pulmonata: Milacidae et Agriolimacidae). and Limacidae) from the Balkan Peninsula. Ann. Zool. Malak. Abh. 17: 151–160. 33: 77–91. The collection of terrestrial slugs at the Museum of Natural History, Wroc³aw University 93

WIKTOR A., MYLONAS M. 1981. New Deroceras species from poda: Pulmonata: Limacidae) from Balearic Islands. In: Cyclades Islands (Greece) (Gastropoda, Pulmonata, Die Land- und Süsswassermollusken der Balearischen Agriolimacidae). Malak. Abh. 7: 183–193. Inseln (BECKMANN K. H. ed.), pp. 187–197, WIKTOR A., MYLONAS M. 1986. Milax aegaeicus n. sp. from ConchBooks, Hackenheim. Cyclades (Gastropoda, Pulmonata, Milacidae). Malak. WIKTOR A., VARDINOYANNIS K., MYLONAS M. 1994. Slugs of Abh. 11: 147–153. the Greek Southern Aegean Islands (Gastropoda terres- WIKTOR A., MYLONAS M. 1992. Deroceras astypalaeensis n. sp. tria nuda: Milacidae, Agriolimacidae et Limacidae). from the Greek island Astipalaia (Dodecanesos) (Gastro- Malak. Abh. 17: 1–36. poda, Pulmonata: Agriolimacidae). Malak. Abh. 16: 1–6. WIKTOR J. 1992. Type specimens of invertebrates (excluding WIKTOR A., NAGGS F., GUPTA P. K. 1999. Turcomilax (Tauli- insects) in the W³adys³aw Rydzewski Museum of Natural max) oli sp. n. from the Kumaun Himalaya, India History Wroc³aw University. Acta Uniw. Wratislaviensis 23 (Gastropoda: Pulmonata: Limacidae). Malak. Abh. 19: (Prace zoologiczne) 1124: 137–155. 225–231. WINTER A. J. DE 1986. Little known and new south-west Eu- WIKTOR A., NORRIS A. 1991. Geomalacus malagensis sp. n. and ropean slugs (Pulmonata: Agriolimacidae, Arionidae). some remarks concerning generic level systematics of Zool. Meded. 60: 135–158. Arionidae (Gastropoda: Pulmonata). J. Conch. 34: WINTER A. J. DE,BUTOT L. J. M. 1986. Three new Deroceras 19–29. species from Greece (Gastropoda, Pulmonata, Agrio- WIKTOR A., PAREJO C. 1989a. Geomalacus (Arrudia) angui- limacidae). Basteria 50: 35–44. formis (Morelet, 1845) its morphology and distribution WÜTRICH M. 1993. Deroceras (Plathystimulus) juranum n. sp. (Gastropoda, Pulmonata: Arionidae). Malak. Abh. 14: eine endemische Nacktschnecke aus dem Schweizer-Jura. 17–25. Arch. Moll. 122: 123–131. WIKTOR A., PAREJO C. 1989b. Arion (Kobeltia) paulariensis sp. n. from Central Spain (Gastropoda, Pulmonata: Received: February 10th, 2007 Arionidae). Malak. Abh. 14: 27–33. Accepted: May 15th, 2007 WIKTOR A., QUINTANA J., BECKMANN K. H. 2007. Redescri- ption of Limax majoricensis (Heynemann 1863) (Gastro-