Alpine Tundra

Total Page:16

File Type:pdf, Size:1020Kb

Load more

Alpine Tundra Where mountaintops rise like islands above a sea of trees lies the world of the alpine tundra. John Muir called it "a land of desolation covered with beautiful light." Yet this light shines on a tapestry of living detail. Tundra lands, too cold for trees, support over 300 kinds of plants, as well as animals from bighorn to butterflies. Alpine Tundra Arctic tundra occurs around the north Reserve because of the beauty and . a world by itself pole. Alpine tundra crowns mountain research value of its alpine wild lands. in the sky. areas above treeline. Alpine tundra is a sensitive indicator of such climatic changes as global warming Enos Mills Rocky Mountain National Park is and acid rain. Over one-third of the recognized worldwide as a Biosphere park is tundra. A Fragile World After Trail Ridge Road opened in 1932, Some areas along Trail Ridge Road are the tundra was not fully protected. signed as Tundra Protection Areas. No People walking on the tundra caused walking off trail is allowed in these areas. much damage. Park scientists initiated In other areas walking on the tundra is an impact study in 1959, fencing off permitted. Walk with care! Step lightly, certain areas. Some areas show little or do not scuff the surface. Step on rocks no sign of recovery, even today, and may whenever possible. Spread out instead take centuries to heal. High winds and of walking in a single line. long winters slow new growth. Snow in July Summer on the tundra is brief, about 40 Average annual precipitation at the frost-free days per year. Temperatures Contintental Divide is 40 inches, of stay below freezing for over five months. which 65% falls as snow. In summer Winter winds exceed 150 mph. Summer frequent brief afternoon thunderstorms days seldom reach 70 F. Snow might fall with lightning, hail, and high winds any day of the year. occur. Watch the clouds; storms can develop within minutes. Alpine Adaptations Strong winds, shallow soil, a short Only a few animals live all year on the growing season, thin air, intense tundra. Pika cut and dry stacks of sunlight, temperature extremes, and plants in summer for their winter larder. limited water challenge life on the Ptarmigans, the only birds that winter tundra. Plants cope by hugging the on the tundra, grow feathers on their ground. Waxy or hairy leaves help hold feet and comb-like teeth on their toes water. Red pigments act as sunscreen. that work like snowshoes. Marmots Roots are extensive; tundra plants have hibernate. Other animals migrate or up to 90% of their bulk below commute: ravens, hawks, coyotes, and ground level. elk. March, 2009 Park information is available at 970-586-1206 or on-line at www.nps.gov/romo EXPERIENCE YOUR AMERICA™ THE ALPINE TUNDRA ECOSYSTEM It may look barren--but look again! While standing in one spot you could touch a meadow, a marsh, a rocky desert, or perhaps all three. On the tundra slight differences in climate (microclimates) result in very different ecosystems. Microclimates provide growing conditions for certain plants by providing slightly warmer temperatures, especially where rocks shelter the plants from wind. FELLFIELDS These "fields of rock," lie on exposed slopes where winter winds blow the snow away. Water and soil are scarce, but lichens— SNOWBED COMMUNITIES crust-like plants that tolerate ALPINE TURFS A distinct community forms where extremes of cold and drought— AND MEADOWS deep snow collects. Late-melting can grow on the rocks. Among Much of Rocky Mountain snow shortens the already brief the rocks pioneer leafy plants National Park's alpine tundra is growing season for plants beneath grow in the shape of rounded KRUMMHOLZ covered with dense turfs of sedges it but insulates them in winter and "cushions," hugging the ground Alpine tundra begins where trees and grasses that support many yields a bonus of water during to avoid wind. As dead leaves cannot survive extremes of cold animals such as elk, marmots, the growing season. Blossoms and soil collect within a cushion and wind, and the short growing and pikas. Rich soils support a of the yellow snow buttercup plant, other plants may sprout in season. At Rocky Mountain colorful diversity of wildflowers (Ranunculus adoneus) often bloom this fertile bed. In time invading National Park this happens whose colors peak in late June or while the plant is in the snow. Look plants may replace the cushion near 11,500 feet (3505 m). The early July. The largest flower on for snake-like casts of soil left by plant. Moss campion (Silene transition is marked by low, the tundra, the alpine sunflower pocket gophers tunneling through acaulis) is a common cushion wind-shaped spruce and fir trees (Rydbergia grandiflora), grows the snow. The gophers eat plant plant in the park. Nearly half of called krummholz (German for only in the Rocky Mountains. It roots and shoots. The soil they Rocky Mountain National Park's "crooked wood"). Many of these stores solar energy for seven to 10 plow makes a seedbed for pioneer alpine plants grow in other tundra short, twisted trees have lived summers before blooming only plant communities: the "gopher areas throughout the northern more than a thousand winters. once. Then the whole plant dies. garden." hemisphere..
Recommended publications
  • Reduced Net Methane Emissions Due to Microbial Methane Oxidation in a Warmer Arctic

    Reduced Net Methane Emissions Due to Microbial Methane Oxidation in a Warmer Arctic

    LETTERS https://doi.org/10.1038/s41558-020-0734-z Reduced net methane emissions due to microbial methane oxidation in a warmer Arctic Youmi Oh 1, Qianlai Zhuang 1,2,3 ✉ , Licheng Liu1, Lisa R. Welp 1,2, Maggie C. Y. Lau4,9, Tullis C. Onstott4, David Medvigy 5, Lori Bruhwiler6, Edward J. Dlugokencky6, Gustaf Hugelius 7, Ludovica D’Imperio8 and Bo Elberling 8 Methane emissions from organic-rich soils in the Arctic have bacteria (methanotrophs) and the remainder is mostly emitted into been extensively studied due to their potential to increase the atmosphere (Fig. 1a). The methanotrophs in these wet organic the atmospheric methane burden as permafrost thaws1–3. soils may be low-affinity methanotrophs (LAMs) that require However, this methane source might have been overestimated >600 ppm of methane (by moles) for their growth and mainte- without considering high-affinity methanotrophs (HAMs; nance23. But in dry mineral soils, the dominant methanotrophs are methane-oxidizing bacteria) recently identified in Arctic min- high-affinity methanotrophs (HAMs), which can survive and grow 4–7 eral soils . Herein we find that integrating the dynamics of at a level of atmospheric methane abundance ([CH4]atm) of about HAMs and methanogens into a biogeochemistry model8–10 1.8 ppm (Fig. 1b)24. that includes permafrost soil organic carbon dynamics3 leads Quantification of the previously underestimated HAM-driven −1 to the upland methane sink doubling (~5.5 Tg CH4 yr ) north of methane sink is needed to improve our understanding of Arctic 50 °N in simulations from 2000–2016. The increase is equiva- methane budgets.
  • Alpine Tundra

    Alpine Tundra

    National Park Rocky Mountain Colorado National Park Service U.S. Department of the Interior Alpine Tundra WHAT IS ALPINE TUNDRA? Where mountaintops rise Arctic tundra occurs around like islands above a sea of the north pole. Alpine tundra trees lies the world of the crowns mountains that alpine tundra. John Muir reach above treeline. called it "a land of deso­ ...a world by itself lation covered with beautiful Rocky Mountain National in the sky. light." Yet this light shines Park is recognized world­ on a tapestry of living detail. wide as a Biosphere Reserve —Enos Mills Tundra lands too cold for because of the beauty and trees support over 200 kinds research value of its alpine of plants, as well as animals wild lands. Alpine tundra is from bighorn to butterflies. a sensitive indicator of such climatic changes as global Tundra is a Russian word for warming and acid rain. Over "land of no trees." 1/3 of the park is tundra. HOW FRAGILE IS IT? For 25 years after Trail That is why busy stops Ridge Road opened in 1932, along Trail Ridge Road are people had free run on the marked as Tundra Protection tundra. Repeated trampling Areas where no walking off damaged popular places. the trail is allowed. Some of these areas, fenced Elsewhere, walking on the off in 1959 for study, show tundra is permitted. But almost no sign of recovery walk with care! Step lightly, today. High winds and long without scuffing the winters make new growth surface. Step on rocks slow. Trampled places may when you can.
  • The Exchange of Carbon Dioxide Between Wet Arctic Tundra and the Atmosphere at the Lena River Delta, Northern Siberia

    The Exchange of Carbon Dioxide Between Wet Arctic Tundra and the Atmosphere at the Lena River Delta, Northern Siberia

    Biogeosciences, 4, 869–890, 2007 www.biogeosciences.net/4/869/2007/ Biogeosciences © Author(s) 2007. This work is licensed under a Creative Commons License. The exchange of carbon dioxide between wet arctic tundra and the atmosphere at the Lena River Delta, Northern Siberia L. Kutzbach1,*, C. Wille1,*, and E.-M. Pfeiffer2 1Alfred Wegener Institute for Polar and Marine Research, Research Unit Potsdam, Telegrafenberg A43, 14473 Potsdam, Germany 2University of Hamburg, Institute of Soil Science, Allende-Platz 2, 20146 Hamburg, Germany *now at Ernst Moritz Arndt University Greifswald, Institute for Botany and Landscape Ecology, Grimmer Straße 88, 17487 Greifswald, Germany Received: 23 May 2007 – Published in Biogeosciences Discuss.: 25 June 2007 Revised: 1 October 2007 – Accepted: 8 October 2007 – Published: 18 October 2007 Abstract. The exchange fluxes of carbon dioxide between piration continued at substantial rates during autumn when wet arctic polygonal tundra and the atmosphere were inves- photosynthesis had ceased and the soils were still largely un- tigated by the micrometeorological eddy covariance method. frozen. The temporal variability of the ecosystem respiration The investigation site was situated in the centre of the Lena during summer was best explained by an exponential func- River Delta in Northern Siberia (72◦220 N, 126◦300 E). The tion with surface temperature, and not soil temperature, as study region is characterized by a polar and distinctly con- the independent variable. This was explained by the ma- tinental climate, very cold and ice-rich permafrost and its jor role of the plant respiration within the CO2 balance of position at the interface between the Eurasian continent and the tundra ecosystem.
  • Compilation of U.S. Tundra Biome Journal and Symposium Publications

    Compilation of U.S. Tundra Biome Journal and Symposium Publications

    COMPILATION OF U.S • TUNDRA BIOME JOURNAL .Alm SYMPOSIUM PUBLICATIONS* {April 1975) 1. Journal Papers la. Published +Alexander, V., M. Billington and D.M. Schell (1974) The influence of abiotic factors on nitrogen fixation rates in the Barrow, Alaska, arctic tundra. Report Kevo Subarctic Research Station 11, vel. 11, p. 3-11. (+)Alexander, V. and D.M. Schell (1973) Seasonal and spatial variation of nitrogen fixation in the Barrow, Alaska, tundra. Arctic and Alnine Research, vel. 5, no. 2, p. 77-88. (+)Allessio, M.L. and L.L. Tieszen (1974) Effect of leaf age on trans­ location rate and distribution of Cl4 photoassimilate in Dupontia fischeri at Barrow, Alaska. Arctic and Alnine Research, vel. 7, no. 1, p. 3-12. (+)Barsdate, R.J., R.T. Prentki and T. Fenchel (1974) The phosphorus cycle of model ecosystems. Significance for decomposer food chains and effect of bacterial grazers. Oikos, vel. 25, p. 239-251. 0 • (+)Batzli, G.O., N.C. Stenseth and B.M. Fitzgerald (1974) Growth and survi­ val of suckling brown lemmings (Lemmus trimucronatus). Journal of Mammalogy, vel. 55, p. 828-831. (+}Billings,. W.D. (1973) Arctic and alpine vegetations: Similarities, dii'­ ferences and susceptibility to disturbance. Bioscience, vel. 23, p. 697-704. (+)Braun, C.E., R.K. Schmidt, Jr. and G.E. Rogers (1973) Census of Colorado white-tailed ptarmigan with tape-recorded calls. Journal of Wildlife Management, vel. 37, no. 1, p. 90-93. (+)Buechler, D.G. and R.D. Dillon (1974) Phosphorus regeneration studies in freshwater ciliates. Journal of Protozoology, vol. 21, no. 2,· P• 339-343.
  • From Tropics to Tundra: Global Convergence in Plant Functioning

    From Tropics to Tundra: Global Convergence in Plant Functioning

    Proc. Natl. Acad. Sci. USA Vol. 94, pp. 13730–13734, December 1997 Ecology From tropics to tundra: Global convergence in plant functioning PETER B. REICH*†,MICHAEL B. WALTERS‡, AND DAVID S. ELLSWORTH§ *Department of Forest Resources, University of Minnesota, St. Paul, MN 55108; ‡Faculty of Natural Resources and Environmental Studies, University of Northern British Columbia, Prince George, British Columbia, Canada V2N 4Z9; and §Department of Applied Science, Brookhaven National Laboratory, Upton, NY 11973 Communicated by Harold Alfred Mooney, Stanford University, Stanford, CA, September 12, 1997 (received for review May 21, 1997) ABSTRACT Despite striking differences in climate, soils, species, light-saturated net photosynthetic capacity (Amax) and and evolutionary history among diverse biomes ranging from leaf diffusive conductance were measured in the field (7–9) tropical and temperate forests to alpine tundra and desert, we under ambient CO2 concentrations and SLA (projected leaf found similar interspecific relationships among leaf structure area per dry mass), dark respiration (at 25°C), and N concen- and function and plant growth in all biomes. Our results thus tration of those leaves were assessed, on both mass and area demonstrate convergent evolution and global generality in bases. Average leaf life-span (month 5 30.4 days) was assessed plant functioning, despite the enormous diversity of plant for each species in the field, from long term monitoring of species and biomes. For 280 plant species from two global data numerous leaves andyor phenological observation. Field data sets, we found that potential carbon gain (photosynthesis) and on leaf, plant, and ecosystem characteristics also were com- carbon loss (respiration) increase in similar proportion with piled from the literature for species from several continents decreasing leaf life-span, increasing leaf nitrogen concentra- and an equally broad array of biomes.
  • Carbon Dioxide Sources from Alaska Driven by Increasing Early Winter Respiration from Arctic Tundra

    Carbon Dioxide Sources from Alaska Driven by Increasing Early Winter Respiration from Arctic Tundra

    Carbon dioxide sources from Alaska driven by increasing early winter respiration from Arctic tundra Róisín Commanea,b,1, Jakob Lindaasb, Joshua Benmerguia, Kristina A. Luusc, Rachel Y.-W. Changd, Bruce C. Daubea,b, Eugénie S. Euskirchene, John M. Hendersonf, Anna Kariong, John B. Millerh, Scot M. Milleri, Nicholas C. Parazooj,k, James T. Randersonl, Colm Sweeneyg,m, Pieter Tansm, Kirk Thoningm, Sander Veraverbekel,n, Charles E. Millerk, and Steven C. Wofsya,b aHarvard John A. Paulson School of Engineering and Applied Sciences, Cambridge, MA 02138; bDepartment of Earth and Planetary Sciences, Harvard University, Cambridge, MA 02138; cCenter for Applied Data Analytics, Dublin Institute of Technology, Dublin 2, Ireland; dDepartment of Physics and Atmospheric Science, Dalhousie University, Halifax, NS, Canada, B3H 4R2; eInstitute of Arctic Biology, University of Alaska Fairbanks, Fairbanks, AK 99775; fAtmospheric and Environmental Research Inc., Lexington, MA 02421; gCooperative Institute of Research in Environmental Sciences, University of Colorado Boulder, Boulder, CO 80309; hGlobal Monitoring Division, National Oceanic and Atmospheric Administration, Boulder, CO 80305; iCarnegie Institution for Science, Stanford, CA 94305; jJoint Institute for Regional Earth System Science and Engineering, University of California, Los Angeles, CA 90095; kJet Propulsion Laboratory, California Institute of Technology, Pasadena, CA 91109; lDepartment of Earth System Science, University of California, Irvine, CA 92697; mEarth Science Research Laboratory, National Oceanic and Atmospheric Administration, Boulder, CO 80305; and nFaculty of Earth and Life Sciences, Vrije Universiteit, 1081 HV Amsterdam, The Netherlands Edited by William H. Schlesinger, Cary Institute of Ecosystem Studies, Millbrook, NY, and approved March 31, 2017 (received for review November 8, 2016) High-latitude ecosystems have the capacity to release large amounts of arctic and boreal landscapes.
  • Alpine Tundra of the Krkonoše and Carpathian Mountains

    Alpine Tundra of the Krkonoše and Carpathian Mountains

    ALPINE TUNDRA OF THE KRKONOŠE AND CARPATHIAN MOUNTAINS Report on study tour funded by Alfred Toepfer Natural Heritage Scholarship 2016 by Dr. Tymur Bedernichek, M.M. Gryshko National Botanical Garden (Ukraine) Text: Tymur Bedernichek Photos: most of the photos were taken by Tymur Bedernichek; except for those listed below: fig. 6 credit to the British Museum fig. 7 & 13 credit to Robert Erik fig 10 & 11 credit to censor.net.ua fig. 12 credit to vorobus.com fig. 14 credit to podstudnicnou.cz fig. 16 credit to Roman Cherepanyn Sponsorship: this report is the result of study tour funded by Alfred Toepfer Foundation and EUROPARC Federation © EUROPARC Federation, 2017 © Alfred Toepfer Foundation, 2017 2 Acknowledgements I am extremely grateful to EUROPARC Federation and Alfred Toepfer Foundation for the Scholarship. It was not only a very timely support, but also a key that opened many doors. It entailed many pleasant and useful contacts and lucky coincidences, and in the end, I managed to do much more than I had planned. Several people significantly helped me during the study tour: Prof. Natalia Zaimenko, Dr. Blanka Skočdopolová, Dr. Irena Kholová and Prof. Zofia Rączkowska. They were my colleagues, friends and sometimes, even guardian angels. Without their support, I probably would not have been able to complete the project. I would also like to thank Erika Stanciu. Six years ago, she made a presentation at RAPPAM workshop in Ukraine, which I attended with other PhD students. She told us about various funding opportunities for nature conservationists, including Alfred Toepfer Natural Heritage Scholarship, and emphasized how important it is to prepare a strong application.
  • A Conservation Vision for Maine Using Ecological Systems

    A Conservation Vision for Maine Using Ecological Systems

    A Conservation Vision for Maine Using Ecological Systems Hemlock forest Shrub swamp Cedar swamp Northern hardwood forest Justin Schlawin and Andy Cutko Maine Natural Areas Program Maine Department of Agriculture, Conservation and Forestry February 2014 A Conservation Vision for Maine Using Ecological Systems Contents Summary ....................................................................................................................................................... 3 Introduction: ................................................................................................................................................. 4 Maine’s Conservation Lands: ........................................................................................................................ 4 Ecological Sections: ....................................................................................................................................... 5 Ecological Systems GIS Layer: ....................................................................................................................... 7 Consolidating Comparable Ecological Systems: ........................................................................................ 7 Requirements for Representation ................................................................................................................ 8 Isolating ‘quality’ patches for each ecological system type .................................................................... 12 Aggregation ............................................................................................................................................
  • New Hampshire Wildlife Action Plan Appendix B Habitats -1 Appendix B: Habitats

    New Hampshire Wildlife Action Plan Appendix B Habitats -1 Appendix B: Habitats

    Appendix B: Habitats Appendix B: Habitat Profiles Alpine ............................................................................................................................................................ 2 Appalachian Oak Pine Forest ........................................................................................................................ 9 Caves and Mines ......................................................................................................................................... 19 Grasslands ................................................................................................................................................... 24 Hemlock Hardwood Pine Forest ................................................................................................................. 34 High Elevation Spruce‐Fir Forest ................................................................................................................. 45 Lowland Spruce‐Fir Forest .......................................................................................................................... 53 Northern Hardwood‐Conifer Forest ........................................................................................................... 62 Pine Barrens ................................................................................................................................................ 72 Rocky Ridge, Cliff, and Talus ......................................................................................................................
  • WILD Colorado: Crossroads of Biodiversity a Message from the Director

    WILD Colorado: Crossroads of Biodiversity a Message from the Director

    WILD Colorado: Crossroads of Biodiversity A Message from the Director July 1, 2003 For Wildlife – For People Dear Educator, Colorado is a unique and special place. With its vast prairies, high mountains, deep canyons and numerous river headwaters, Colorado is truly a crossroad of biodiversity that provides a rich environment for abundant and diverse species of wildlife. Our rich wildlife heritage is a source of pride for our citizens and can be an incredibly powerful teaching tool in the classroom. To help teachers and students learn about Colorado’s ecosystems and its wildlife, the Division of Wildlife has prepared a set of ecosystem posters and this education guide. Together they will provide an overview of the biodiversity of our state as it applies to the eight major ecosystems of Colorado. This project was funded in part by a Wildlife Conservation and Restoration Program grant. Sincerely, Russell George, Director Colorado Division of Wildlife Table of Contents Introduction . 2 Activity: Which Niche? . 6 Grasslands Poster . 8 Grasslands . 10 Sage Shrublands Poster . 14 Sagebrush Shrublands . 16 Montane Shrublands Poster . 20 Montane Shrublands . 22 Piñon-Juniper Woodland Poster . 26 Piñon-Juniper Woodland . 28 Montane Forests Poster . 32 Montane Forests . 34 Subalpine Forests Poster . 38 Subalpine Forests . 40 Alpine Tundra Poster . 44 Treeline and Alpine Tundra . 46 Activity: The Edge of Home . 52 Riparian Poster . 54 Aquatic Ecosystems, Riparian Areas, and Wetlands . 56 Activity: Wetland Metaphors . 63 Glossary . 66 References and Field Identification Manuals . 68 This book was written by Wendy Hanophy and Harv Teitelbaum with illustrations by Marjorie Leggitt and paintings by Paul Gray.
  • Canopy Structure and Phenology of Alpine Tundra Vegetation

    Canopy Structure and Phenology of Alpine Tundra Vegetation

    Utah State University DigitalCommons@USU All Graduate Theses and Dissertations Graduate Studies 5-1972 Canopy Structure and Phenology of Alpine Tundra Vegetation Marcee Fareed Follow this and additional works at: https://digitalcommons.usu.edu/etd Part of the Plant Sciences Commons Recommended Citation Fareed, Marcee, "Canopy Structure and Phenology of Alpine Tundra Vegetation" (1972). All Graduate Theses and Dissertations. 6314. https://digitalcommons.usu.edu/etd/6314 This Thesis is brought to you for free and open access by the Graduate Studies at DigitalCommons@USU. It has been accepted for inclusion in All Graduate Theses and Dissertations by an authorized administrator of DigitalCommons@USU. For more information, please contact [email protected]. ,, rn.ea . ,1 ,.I,. "'' CANOPYSTRUCTURE AND PHENOLOGY OF ALPINE TUNDRA VEGETATION by Marcee Fareed A thesis submitted in partial fulfillment · of the requirements for the degree of MASTEROF SCIENCE in Plant Ecology Approved: MaJor Professor Committeeman ·committeeman Dean of Graduate Studies UTAHSTATE UNIVERSITY Logan, Utah 1972 ii ACKNOWLEDGEMENTS Grateful acknowledgement is made to Dr. Martyn Caldwell for his suggestions on the fieldwork and the preparatton of the thesis. Appreciation is also extended to Thomas Shoemaker and Diane and Roger Hanson for their assistance in gathering the data, and to Professor Arthur Holmgren for helping in the identification of plant species in the voucher collection. Dr. Larry Tieszen is responsible for developing the computer programs used in the analysis of the data. This study was supported by the Tundra Biome of the International Biological Program. ~~ Marcee Fareed iii TABLE OF CONTENTS Page Acknowledgements ii List of Tables .. iv Li st of Figures v Abstract vi Introduction .
  • Alpine Ecosystems

    Alpine Ecosystems

    TWENTY-NINE Alpine Ecosystems PHILIP W. RUNDEL and CONSTANCE I. MILLAR Introduction Alpine ecosystems comprise some of the most intriguing hab­ writing about the alpine meadows of the Sierra Nevada, felt itats of the world for the stark beauty of their landscapes and his words were inadequate to describe “the exquisite beauty for the extremes of the physical environment that their resi­ of these mountain carpets as they lie smoothly outspread in dent biota must survive. These habitats lie above the upper the savage wilderness” (Muir 1894). limit of tree growth but seasonally present spectacular flo­ ral shows of low-growing herbaceous perennial plants. Glob­ ally, alpine ecosystems cover only about 3% of the world’s Defining Alpine Ecosystems land area (Körner 2003). Their biomass is low compared to shrublands and woodlands, giving these ecosystems only a Alpine ecosystems are classically defined as those communi­ minor role in global biogeochemical cycling. Moreover, spe­ ties occurring above the elevation of treeline. However, defin­ cies diversity and local endemism of alpine ecosystems is rela­ ing the characteristics that unambiguously characterize an tively low. However, alpine areas are critical regions for influ­ alpine ecosystem is problematic. Defining alpine ecosystems encing hydrologic flow to lowland areas from snowmelt. based on presence of alpine-like communities of herbaceous The alpine ecosystems of California present a special perennials is common but subject to interpretation because case among alpine regions of the world. Unlike most alpine such communities may occur well below treeline, while other regions, including the American Rocky Mountains and the areas well above treeline may support dense shrub or matted European Alps (where most research on alpine ecology has tree cover.