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By Howard Zimmerman
by Howard Zimmerman DINO_COVERS.indd 4 4/24/08 11:58:35 AM [Intentionally Left Blank] by Howard Zimmerman Consultant: Luis M. Chiappe, Ph.D. Director of the Dinosaur Institute Natural History Museum of Los Angeles County 1629_ArmoredandDangerous_FNL.ind1 1 4/11/08 11:11:17 AM Credits Title Page, © Luis Rey; TOC, © De Agostini Picture Library/Getty Images; 4-5, © John Bindon; 6, © De Agostini Picture Library/The Natural History Museum, London; 7, © Luis Rey; 8, © Luis Rey; 9, © Adam Stuart Smith; 10T, © Luis Rey; 10B, © Colin Keates/Dorling Kindersly; 11, © Phil Wilson; 12L, Courtesy of the Royal Tyrrell Museum, Drumheller, Alberta; 12R, © De Agostini Picture Library/Getty Images; 13, © Phil Wilson; 14-15, © Phil Wilson; 16-17, © De Agostini Picture Library/The Natural History Museum, London; 18T, © 2007 by Karen Carr and Karen Carr Studio; 18B, © photomandan/istockphoto; 19, © Luis Rey; 20, © De Agostini Picture Library/The Natural History Museum, London; 21, © John Bindon; 23TL, © Phil Wilson; 23TR, © Luis Rey; 23BL, © Vladimir Sazonov/Shutterstock; 23BR, © Luis Rey. Publisher: Kenn Goin Editorial Director: Adam Siegel Creative Director: Spencer Brinker Design: Dawn Beard Creative Cover Illustration: Luis Rey Photo Researcher: Omni-Photo Communications, Inc. Library of Congress Cataloging-in-Publication Data Zimmerman, Howard. Armored and dangerous / by Howard Zimmerman. p. cm. — (Dino times trivia) Includes bibliographical references and index. ISBN-13: 978-1-59716-712-3 (library binding) ISBN-10: 1-59716-712-6 (library binding) 1. Ornithischia—Juvenile literature. 2. Dinosaurs—Juvenile literature. I. Title. QE862.O65Z56 2009 567.915—dc22 2008006171 Copyright © 2009 Bearport Publishing Company, Inc. All rights reserved. -
The Princeton Field Guide to Dinosaurs, Second Edition
MASS ESTIMATES - DINOSAURS ETC (largely based on models) taxon k model femur length* model volume ml x specific gravity = model mass g specimen (modeled 1st):kilograms:femur(or other long bone length)usually in decameters kg = femur(or other long bone)length(usually in decameters)3 x k k = model volume in ml x specific gravity(usually for whole model) then divided/model femur(or other long bone)length3 (in most models femur in decameters is 0.5253 = 0.145) In sauropods the neck is assigned a distinct specific gravity; in dinosaurs with large feathers their mass is added separately; in dinosaurs with flight ablity the mass of the fight muscles is calculated separately as a range of possiblities SAUROPODS k femur trunk neck tail total neck x 0.6 rest x0.9 & legs & head super titanosaur femur:~55000-60000:~25:00 Argentinosaurus ~4 PVPH-1:~55000:~24.00 Futalognkosaurus ~3.5-4 MUCPv-323:~25000:19.80 (note:downsize correction since 2nd edition) Dreadnoughtus ~3.8 “ ~520 ~75 50 ~645 0.45+.513=.558 MPM-PV 1156:~26000:19.10 Giraffatitan 3.45 .525 480 75 25 580 .045+.455=.500 HMN MB.R.2181:31500(neck 2800):~20.90 “XV2”:~45000:~23.50 Brachiosaurus ~4.15 " ~590 ~75 ~25 ~700 " +.554=~.600 FMNH P25107:~35000:20.30 Europasaurus ~3.2 “ ~465 ~39 ~23 ~527 .023+.440=~.463 composite:~760:~6.20 Camarasaurus 4.0 " 542 51 55 648 .041+.537=.578 CMNH 11393:14200(neck 1000):15.25 AMNH 5761:~23000:18.00 juv 3.5 " 486 40 55 581 .024+.487=.511 CMNH 11338:640:5.67 Chuanjiesaurus ~4.1 “ ~550 ~105 ~38 ~693 .063+.530=.593 Lfch 1001:~10700:13.75 2 M. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
Rule Booklet
Dig for fossils, build skeletons, and attract the most visitors to your museum! TM SCAN FOR VIDEO RULES AND MORE! FOSSILCANYON.COM Dinosaurs of North America edimentary rock formations of western North America are famous for the fossilized remains of dinosaurs The rules are simple enough for young players, but and other animals from the Triassic, Jurassic, and serious players can benefit Cretaceous periods of the Mesozoic Era. Your objective from keeping track of the cards that is to dig up fossils, build complete skeletons, and display have appeared, reasoning about them in your museum to attract as many visitors as possible. probabilities and expected returns, and choosing between aggressive Watch your museum’s popularity grow using jigsaw-puzzle and conservative plays. scoring that turns the competition into a race! GAME CONTENTS TM 200,000300,000 160,000 VISITORS VISITORS PER YEAR 140,000 VISITORS PER YEAR 180,000 VISITORS PER YEAR 400,000 VISITORS PER YEAR Dig for fossils, build skeletons, and 340,000 VISITORS PER YEAR RD COLOR ELETONS CA GENUS PERIODDIET SK FOSSIL VISITORSPARTS 360,000 VISITORS PER YEAR PER YEAR attract the most visitors to your museum! VISITORS PER YEAR PER YEAR Tyrannosaurus K C 1 4 500,000 Brachiosaurus J H 1 3 400,000 ON YOUR TURN: TM SCAN FOR VIDEO Triceratops K H 1 3 380,000 RULES AND MORE! Allosaurus J C 2 Dig3 a first360,000 card. If it is a fossil, keep it hidden. FOSSILCANYON.COM Ankylosaurus K H 2 If it3 is an340,000 action card, perform the action. -
R~;: PHYSIOLOGICAL, MIGRATORIAL
....----------- 'r~;: i ! 'r; Pa/eont .. 62(4), 1988, pp. 64~52 Copyright © 1988, The Paleontological Society 0022-3360/88/0062-0640$03.00 PHYSIOLOGICAL, MIGRATORIAL, CLIMATOLOGICAL, GEOPHYSICAL, SURVIVAL, AND EVOLUTIONARY IMPLICATIONS OF CRETACEOUS POLAR DINOSAURS GREGORY S. PAUL 3109 North Calvert Street, Baltimore, Maryland 21218 ABSTRACTT- he presence of Late Cretaceous social dinosaurs in polar regions confronted them with winter conditions of extended dark, coolness, breezes, and precipitation that could best be coped with via an endothermic homeothermic physiology of at least the tenrec level. This is true whether the dinosaurs stayed year round in the polar regime-which in North America extended from Alaska south to Montana-or if they migrated away from polar winters. More reptilian physiologies fail to meet the demands of such winters -in certain key ways, a· point tentatively confirmed by the apparent failure of giant Late Cretaceous phobosuchid crocodilians to dwell north of Montana. Low metabolisms were also insufficient for extended annual migrations away from and towards the poles. It is shown that even high metabolic rate dinosaurs probably remained in their polar habitats year-round. The possibility that dinosaurs had avian-mammalian metabolic systems, and may have borne insulation at least seasonally, severely limits their use as polar paleoclimatic and Earth axial tilt indicators. Polar dinosaurs may have been a center of dinosaur evolution. The possible ability of polar dinosaurs to cope with conditions of cool and dark challenges theories that a gradual temperature decline, or a sudden, meteoritic or volcanic induced collapse in temperature and sunlight, destroyed the dinosaurs. INTRODUCTION America suggests that dinosaurs were regularly crossing, and NCREASINGNUMBERSof remains show that dinosaurs lived living upon, the Bering Land Bridge within a few degrees of the I near the North and South Poles during the Cretaceous. -
A New Caenagnathid Dinosaur from the Upper Cretaceous Wangshi
www.nature.com/scientificreports OPEN A new caenagnathid dinosaur from the Upper Cretaceous Wangshi Group of Shandong, China, with Received: 12 October 2017 Accepted: 7 March 2018 comments on size variation among Published: xx xx xxxx oviraptorosaurs Yilun Yu1, Kebai Wang2, Shuqing Chen2, Corwin Sullivan3,4, Shuo Wang 5,6, Peiye Wang2 & Xing Xu7 The bone-beds of the Upper Cretaceous Wangshi Group in Zhucheng, Shandong, China are rich in fossil remains of the gigantic hadrosaurid Shantungosaurus. Here we report a new oviraptorosaur, Anomalipes zhaoi gen. et sp. nov., based on a recently collected specimen comprising a partial left hindlimb from the Kugou Locality in Zhucheng. This specimen’s systematic position was assessed by three numerical cladistic analyses based on recently published theropod phylogenetic datasets, with the inclusion of several new characters. Anomalipes zhaoi difers from other known caenagnathids in having a unique combination of features: femoral head anteroposteriorly narrow and with signifcant posterior orientation; accessory trochanter low and confuent with lesser trochanter; lateral ridge present on femoral lateral surface; weak fourth trochanter present; metatarsal III with triangular proximal articular surface, prominent anterior fange near proximal end, highly asymmetrical hemicondyles, and longitudinal groove on distal articular surface; and ungual of pedal digit II with lateral collateral groove deeper and more dorsally located than medial groove. The holotype of Anomalipes zhaoi is smaller than is typical for Caenagnathidae but larger than is typical for the other major oviraptorosaurian subclade, Oviraptoridae. Size comparisons among oviraptorisaurians show that the Caenagnathidae vary much more widely in size than the Oviraptoridae. Oviraptorosauria is a clade of maniraptoran theropod dinosaurs characterized by a short, high skull, long neck and short tail. -
State of the Palaeoart
Palaeontologia Electronica http://palaeo-electronica.org State of the Palaeoart Mark P. Witton, Darren Naish, and John Conway The discipline of palaeoart, a branch of natural history art dedicated to the recon- struction of extinct life, is an established and important component of palaeontological science and outreach. For more than 200 years, palaeoartistry has worked closely with palaeontological science and has always been integral to the enduring popularity of prehistoric animals with the public. Indeed, the perceived value or success of such products as popular books, movies, documentaries, and museum installations can often be linked to the quality and panache of its palaeoart more than anything else. For all its significance, the palaeoart industry ment part of this dialogue in the published is often poorly treated by the academic, media and literature, in turn bringing the issues concerned to educational industries associated with it. Many wider attention. We argue that palaeoartistry is standard practises associated with palaeoart pro- both scientifically and culturally significant, and that duction are ethically and legally problematic, stifle improved working practises are required by those its scientific and cultural growth, and have a nega- involved in its production. We hope that our views tive impact on the financial viability of its creators. inspire discussion and changes sorely needed to These issues create a climate that obscures the improve the economy, quality and reputation of the many positive contributions made by palaeoartists palaeoart industry and its contributors. to science and education, while promoting and The historic, scientific and economic funding derivative, inaccurate, and sometimes exe- significance of palaeoart crable artwork. -
New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia
Bull. Natn. Sci. Mus., Tokyo, Ser. C, 30, pp. 95–130, December 22, 2004 New Oviraptorid Dinosaur (Dinosauria: Oviraptorosauria) from the Nemegt Formation of Southwestern Mongolia Junchang Lü1, Yukimitsu Tomida2, Yoichi Azuma3, Zhiming Dong4 and Yuong-Nam Lee5 1 Institute of Geology, Chinese Academy of Geological Sciences, Beijing 100037, China 2 National Science Museum, 3–23–1 Hyakunincho, Shinjukuku, Tokyo 169–0073, Japan 3 Fukui Prefectural Dinosaur Museum, 51–11 Terao, Muroko, Katsuyama 911–8601, Japan 4 Institute of Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 5 Korea Institute of Geoscience and Mineral Resources, Geology & Geoinformation Division, 30 Gajeong-dong, Yuseong-gu, Daejeon 305–350, South Korea Abstract Nemegtia barsboldi gen. et sp. nov. here described is a new oviraptorid dinosaur from the Late Cretaceous (mid-Maastrichtian) Nemegt Formation of southwestern Mongolia. It differs from other oviraptorids in the skull having a well-developed crest, the anterior margin of which is nearly vertical, and the dorsal margin of the skull and the anterior margin of the crest form nearly 90°; the nasal process of the premaxilla being less exposed on the dorsal surface of the skull than those in other known oviraptorids; the length of the frontal being approximately one fourth that of the parietal along the midline of the skull. Phylogenetic analysis shows that Nemegtia barsboldi is more closely related to Citipati osmolskae than to any other oviraptorosaurs. Key words : Nemegt Basin, Mongolia, Nemegt Formation, Late Cretaceous, Oviraptorosauria, Nemegtia. dae, and Caudipterygidae (Barsbold, 1976; Stern- Introduction berg, 1940; Currie, 2000; Clark et al., 2001; Ji et Oviraptorosaurs are generally regarded as non- al., 1998; Zhou and Wang, 2000; Zhou et al., avian theropod dinosaurs (Osborn, 1924; Bars- 2000). -
The Systematic Position of the Enigmatic Thyreophoran Dinosaur Paranthodon Africanus, and the Use of Basal Exemplifiers in Phyl
1 The systematic position of the enigmatic thyreophoran dinosaur Paranthodon africanus, 2 and the use of basal exemplifiers in phylogenetic analysis 3 4 Thomas J. Raven1,2 ,3 and Susannah C. R. Maidment2 ,3 5 61Department of Earth Science & Engineering, Imperial College London, UK 72School of Environment & Technology, University of Brighton, UK 8 3Department of Earth Sciences, Natural History Museum, London, UK 9 10Corresponding author: Thomas J. Raven 11 12Email address: [email protected] 13 14 15 16 17 18 19 20 21ABSTRACT 22 23The first African dinosaur to be discovered, Paranthodon africanus was found in 1845 in the 24Lower Cretaceous of South Africa. Taxonomically assigned to numerous groups since discovery, 25in 1981 it was described as a stegosaur, a group of armoured ornithischian dinosaurs 26characterised by bizarre plates and spines extending from the neck to the tail. This assignment 27that has been subsequently accepted. The type material consists of a premaxilla, maxilla, a nasal, 28and a vertebra, and contains no synapomorphies of Stegosauria. Several features of the maxilla 29and dentition are reminiscent of Ankylosauria, the sister-taxon to Stegosauria, and the premaxilla 30appears superficially similar to that of some ornithopods. The vertebral material has never been 31described, and since the last description of the specimen, there have been numerous discoveries 32of thyreophoran material potentially pertinent to establishing the taxonomic assignment of the 33specimen. An investigation of the taxonomic and systematic position of Paranthodon is therefore 34warranted. This study provides a detailed re-description, including the first description of the 35vertebra. Numerous phylogenetic analyses demonstrate that the systematic position of 36Paranthodon is highly labile and subject to change depending on which exemplifier for the clade 37Stegosauria is used. -
Days & Hours for Social Distance Walking Visitor Guidelines Lynden
53 22 D 4 21 8 48 9 38 NORTH 41 3 C 33 34 E 32 46 47 24 45 26 28 14 52 37 12 25 11 19 7 36 20 10 35 2 PARKING 40 39 50 6 5 51 15 17 27 1 44 13 30 18 G 29 16 43 23 PARKING F GARDEN 31 EXIT ENTRANCE BROWN DEER ROAD Lynden Sculpture Garden Visitor Guidelines NO CLIMBING ON SCULPTURE 2145 W. Brown Deer Rd. Do not climb on the sculptures. They are works of art, just as you would find in an indoor art Milwaukee, WI 53217 museum, and are subject to the same issues of deterioration – and they endure the vagaries of our harsh climate. Many of the works have already spent nearly half a century outdoors 414-446-8794 and are quite fragile. Please be gentle with our art. LAKES & POND There is no wading, swimming or fishing allowed in the lakes or pond. Please do not throw For virtual tours of the anything into these bodies of water. VEGETATION & WILDLIFE sculpture collection and Please do not pick our flowers, fruits, or grasses, or climb the trees. We want every visitor to be able to enjoy the same views you have experienced. Protect our wildlife: do not feed, temporary installations, chase or touch fish, ducks, geese, frogs, turtles or other wildlife. visit: lynden.tours WEATHER All visitors must come inside immediately if there is any sign of lightning. PETS Pets are not allowed in the Lynden Sculpture Garden except on designated dog days. -
Index of Subjects
Cambridge University Press 978-1-108-47594-5 — Dinosaurs 4th Edition Index More Information INDEX OF SUBJECTS – – – A Zuul, 275 276 Barrett, Paul, 98, 335 336, 406, 446 447 Ankylosauridae Barrick, R, 383–384 acetabulum, 71, 487 characteristics of, 271–273 basal dinosauromorph, 101 acromial process, 271, 273, 487 cladogram of, 281 basal Iguanodontia, 337 actual diversity, 398 defined, 488 basal Ornithopoda, 336 adenosine diphosphate, see ADP evolution of, 279 Bates, K. T., 236, 360 adenosine triphosphate, see ATP ankylosaurids, 275–276 beak, 489 ADP (adenosine diphosphate), 390, 487 anpsids, 76 belief systems, 474, 489 advanced characters, 55, 487 antediluvian period, 422, 488 Bell, P. R., 162 aerobic metabolism, 391, 487 anterior position, 488 bennettitaleans, 403, 489 – age determination (dinosaur), 354 357 antorbital fenestra, 80, 488 benthic organisms, 464, 489 Age of Dinosaurs, 204, 404–405 Arbour, Victoria, 277 Benton, M. J., 2, 104, 144, 395, 402–403, akinetic movement, see kinetic movement Archibald, J. D., 467, 469 444–445, 477–478 Alexander, R. M., 361 Archosauria, 80, 88–90, 203, 488 Berman, D. S, 236–237 allometry, 351, 487 Archosauromorpha, 79–81, 488 Beurien, Karl, 435 altricial offspring, 230, 487 archosauromorphs, 401 bidirectional respiration, 350 Alvarez, Luis, 455 archosaurs, 203, 401 Big Al, 142 – Alvarez, Walter, 442, 454 455, 481 artifacts, 395 biogeography, 313, 489 Alvarezsauridae, 487 Asaro, Frank, 455 biomass, 415, 489 – alvarezsaurs, 168 169 ascending process of the astragalus, 488 biosphere, 2 alveolus/alveoli, -
A New Middle Jurassic Diplodocoid Suggests an Earlier Dispersal and Diversification of Sauropod Dinosaurs
ARTICLE DOI: 10.1038/s41467-018-05128-1 OPEN A new Middle Jurassic diplodocoid suggests an earlier dispersal and diversification of sauropod dinosaurs Xing Xu1, Paul Upchurch2, Philip D. Mannion 3, Paul M. Barrett 4, Omar R. Regalado-Fernandez 2, Jinyou Mo5, Jinfu Ma6 & Hongan Liu7 1234567890():,; The fragmentation of the supercontinent Pangaea has been suggested to have had a profound impact on Mesozoic terrestrial vertebrate distributions. One current paradigm is that geo- graphic isolation produced an endemic biota in East Asia during the Jurassic, while simul- taneously preventing diplodocoid sauropod dinosaurs and several other tetrapod groups from reaching this region. Here we report the discovery of the earliest diplodocoid, and the first from East Asia, to our knowledge, based on fossil material comprising multiple individuals and most parts of the skeleton of an early Middle Jurassic dicraeosaurid. The new discovery challenges conventional biogeographical ideas, and suggests that dispersal into East Asia occurred much earlier than expected. Moreover, the age of this new taxon indicates that many advanced sauropod lineages originated at least 15 million years earlier than previously realised, achieving a global distribution while Pangaea was still a coherent landmass. 1 Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology & Paleoanthropology, Chinese Academy of Sciences, 100044 Beijing, China. 2 Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK. 3 Department of Earth Science and Engineering, Imperial College London, South Kensington Campus, London SW7 2AZ, UK. 4 Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW7 5BD, UK. 5 Natural History Museum of Guangxi, 530012 Nanning, Guangxi, China.