ECOSYSTEM INTRODUCTION • Ecosystem Is the Smallest Structural

Total Page:16

File Type:pdf, Size:1020Kb

ECOSYSTEM INTRODUCTION • Ecosystem Is the Smallest Structural ECOSYSTEM INTRODUCTION • Ecosystem is the smallest structural and functional unit of nature or environment where living organisms interact among themselves and also with the surrounding physical environment. • Ecosystem may be large or small. A single drop of water may be an ecosystem. • Ecosystem may be temporary or permanent. TYPES OF ECOSYSTEM • An ecosystem may be natural or artificial. • Natural ecosystem is divided into two basic categories – 1. Terrestrial ecosystem : E.g., forest, grassland, tree, desert ecosystem. 2. Aquatic ecosystem : Aquatic ecosystem is again of two types: a) Lentic ecosystem : E.g., Stagnant fresh water, lake, pond, swamp. b) Lotic ecosystem : Running freshwater ecosystem. E.g., river. • Artificial ecosystem : These are man made ecosystem. E.g., cropland, gardens etc. • On the basis of size, types of ecosystem are :- a) Mega ecosystem – Ocean/Sea b) Macroecosystem – Forest c) Microecosystem – Pond d) Nanoecosystem – Drop of water COMPONENTS OF ECOSYSTEM An ecosystem is composed of two components – • Biotic component • Abiotic component BIOTIC COMPONENTS • Biotic component is made up of many different inter dependent population, e.g., plants, animals, microbes, etc. • Types of biotic component are – a. Producers b. Consumers & c. Decomposers PRODUCERS • All the autotrophs of an ecosystem are called producers. • The green plants are the main producers. • Producers are generally chlorophyll bearing organisms which produce their own food by the process of photosynthesis, in which producers absorb solar energy and convert it into chemical energy. • So producers are also called transducers or converters. E.g., yellow green algae, algal protist etc. • Energy enters into the ecosystem through the producers. • The solar energy is the only ultimate source of energy in the ecosystem. This energy is available for the remaining living organisms. • Other examples of producers are – a) Chemoautotrophs : Iron bacteria, sulphur bacteria, nitrifying bacteria. b) Phytoplankton : In an aquatic ecosystem, phytoplankton are the major autotrophs. Phytoplanktons may produce as much food as produced by the larger shrubs and trees in unit area. CONSUMERS • All the heterotrophs of the ecosystem are known as consumers. It includes micro-organisms. These directly (on herbivores) or indirectly (on carnivores) depend on the producers for food. • Types of consumer are : macroconsumers and micro-consumers. • Macroconsumers / Phagotrophs or holozoic digest their food inside the body of organism i.e. first ingestion then digestion. • Macro-consumers are of following type : a) Primary consumer : Such living organisms which obtain food directly from producers or plants are known as primary consumers. E.g., Herbivores of ecosystem, cow, grazing cattle, rabbit. They are also known as secondary producers as they synthesize complex materials in the cells by the digestion of food which is obtained from the plant. b) Secondary consumers or primary carnivores: Those animals which feed upon primary consumers and obtain food. Those carnivores which kill and eat the herbivores are called predators. E.g., dog, cat, snake. In aquatic system, whale is a secondary consumer. It is an example of a filter feeder because it feeds on plankton. c) Top (tertiary) consumers : Those animals which kill other animals and eat them, but are not killed & eaten by other animals in nature are called tertiary consumers. E.g., Lion, man, peacock. • Micro-consumers/Decomposers or Scavengers/Osmotrophs are those living organisms which decompose the dead body of producers and consumers. • The main decomposers in ecosystem are bacteria and fungi. • Decomposers play a significant role in mineral cycle. ABIOTIC COMPONENTS • Abiotic components include inorganic substances or minerals; organic substance and atmospheric factor. • Inorganic substances are P, S, N, H, Mg, K, CO2, NO2 etc. These are raw materials for plants. • Organic substances are proteins, carbohydrates, lipids in dead organic substances. • Atmospheric factors are light, temperature, humidity, rain, water, gas etc. PRODUCTIVITY • Productivity of the ecosystem refers to the rate of biomass production i.e. the amount of organic matter accumulated per unit area per unit time. • It is generally expressed in g–2 yr –1 or (kcal m–2) yr–1. • There are two types of productivity- primary and secondary. PRIMARY PRODUCTIVITY • Primary productivity is defined as the amount of biomass or organic matter produced per unit area over a time period by plants during photosynthesis. • Primary productivity depends on the plant species inhabiting a particular area. It also depends on a variety of environmental factors, like, availability of nutrients and photosynthetic capacity of plants. Therefore, it varies in different types of ecosystems. • The annual net primary productivity of the whole biosphere is approximately 170 billion tons (dry weight) of the organic matter. • Productivity of the ocean is only 55 billion tons. • In per unit area, maximum productivity is found in tropical rainforest. • In water, least productive ecosystems are very deep lakes and highly productive ecosystems are coral reefs. • Nitrogen is the limiting factor in ocean and phosphorus is the limiting factor in lake productivity. • Most productive agro-ecosystem is sugarcane and rice ecosystem - 3 - 4 kg/m sq/year. • Primary productivity can be divided into - gross primary productivity (GPP) and net primary productivity (NPP). • Gross primary productivity (G.P.P.) is the total amount of energy fixed (organic food) in an ecosystem (in producers) in unit time including the organic matter used up in respiration during the measurement period. It is also known as total (gross) photosynthesis. A considerable amount of GPP is utilized by plants in respiration. • Net primary productivity (N.P.P.) is the amount of stored organic matter in plant tissues after respiratory utilization. NPP = GPP – R (R = Respiration + Metabolic activities) . or GPP = NPP + R SECONDARY PRODUCTIVITY • Secondary productivity is the rate of formation of new organic matter by consumers. • The food of consumers has been produced by the primary producers & secondary productivity reflects only the utilization of this food for the production of consumer biomass. • Net community productivity or Net productivity is the rate of storage of organic matter not used by the heterotrophs NCP = N.P.P. – HR (HR = Energy used by heterotrophs or consumers) ECOLOGICAL EFFICIENCY (EE) [EXTRA INFORMATION FOR ENTRANCE EXAMINATIONS] The percentage of energy transferred from one trophic level to the next is called ecological efficiency or food chain efficiency. Ecological efficiencies may be expressed as – • Assimilation efficiency (AE) It is the proportion of consumed energy that is assimilated. • Net production efficiency (NPE) It is the percentage of net primary productivity in relation to gross primary productivity. • Photosynthetic efficiency (PE) It is the percentage of incident solar radiation trapped by producers to perform photosynthesis & produce gross primary productivity. Photosynthetic efficiency is 1-5%. It can also be expressed in relation to PAR when it is 2-10%. • Ecological efficiency(EE)/trophic level efficiency (TLE) : It is the percentage of energy converted into biomass by a higher trophic level over the energy of food resources available at the lower trophic level. DECOMPOSITION • Decomposers are responsible for converting complex organic material of dead minerals or plants into simpler organic matter through the process of decomposition and release mineral substances into the soil where these are reused by the producers, so that soil is considered as the best resource of minerals. • The upper layer of the soil is the main site for decomposition process in the ecosystem. • The process of decomposition involves several processes. These processes can be categorised as: A. Fragmentation of detritus B. Leaching C. Catabolism D. Humification E. Mineralization • Detritivores (e.g., earthworm) breakdown detritus into smaller particles (called fragmentation). • By the process of leaching, water soluble inorganic nutrients' go down into the soil horizon and get precipitated as unavailable salts. • In bacteria and fungi, process of decomposition completely takes place outside the body. They carry out catabolism and release extracellular enzymes from their body on dead remains and decompose it into simpler organic substances and then absorb it. So these are called as osmotrophs (absorptive). • In the process of decomposition, some nutrients get tied up with the biomass of microbes and become temporarily unavailable to other organisms. Such incorporation of nutrient in living microbes (bacteria & fungi) is called nutrient immobilisation. • Humification and mineralization occur during decomposition in the soil. Humification leads to formation of humus which is highly resistant to microbial action & undergoes extremely slow decomposition. Humus serves as a reservoir of nutrients. • Mineralization results in the release of inorganic substances (e.g., CO2, H2O) + ++ ++ + and variable nutrients (NH4 , Ca , Mg , K etc.) in the soil. • Decomposition is largely an oxygen requiring process. Factors affecting decomposition – a. Temperature and soil moisture are important and regulate decomposition through their effects on the activates of soil microbes. b. Warm and moist environment favour decomposition whereas low temperature and anaerobiosis inhibit decomposition. Fig. : Diagrammatic representation
Recommended publications
  • Environmental Science (Ce-101) Unit- I
    Lecture Notes of Environmental Science ENVIRONMENTAL SCIENCE (CE-101) UNIT- I Introduction to Environmental Science: The science of Environment studies is a multi-disciplinary science because it comprises various branches of studies like chemistry, physics, medical science, life science, agriculture, public health, sanitary engineering etc. It is the science of physical phenomena in the environment. It studies of the sources, reactions, transport, effect and fate of physical a biological species in the air, water and soil and the effect of from human activity upon these. Environment – The word environment comes from the greek word “environner” meaning surroundings around us. Scope of Environmental studies: The environment consists of four segments as under: 1. Atmosphere: The atmosphere implies the protective blanket of gases, surrounding the earth: It sustains life on the earth. It saves it from the hostile environment of outer space. It absorbs most of the cosmic rays from outer space and a major portion of the electromagnetic radiation from the sun. It transmits only here ultraviolet, visible, near infrared radiation (300 to 2500 nm) and radio waves. (0.14 to 40 m) while filtering out tissue-damaging ultraviolet waves below about 300 nm. The atmosphere is composed of nitrogen and oxygen. Besides, argon, carbon dioxide, and trace gases. 2. Hydrosphere: The Hydrosphere comprises all types of water resources oceans, seas, lakes, rivers, streams, reservoir, polar icecaps, glaciers, and groundwater. Nature 97% of the earth’s water supply is in the oceans, Prepared by: Er. Arshad Abbas Deptt. of Civil Engg. KMCUAF University Lucknow About 2% of the water resources are locked in the polar icecaps and glaciers.
    [Show full text]
  • The Bodwad Sarvajanik Co-Op.Education Society Ltd., Bodwad Arts, Commerce and Science College, Bodawd Question Bank Class :-S.Y
    The Bodwad Sarvajanik Co-Op.Education Society Ltd., Bodwad Arts, Commerce and Science College, Bodawd Question Bank Class :-S.Y.B.Sc SEM:- IV Subject: - BOTANY- 402 Plant Ecology 1. The science which deals with relationship between organisms and their environment is called a) Morphology b) Palynology c) Taxonomy d) Ecology 2. The meaning of Greek word Oikas a) Nature b) Environment c) House d) Temple 3. The term ecology coined by a) Odum b) Tansley c) Haeckel d) None 4. Autecology deals with the study of a) Ecology of individual species b) Ecology of many species c) Ecology of community d) All of these 5. Synecology deals with the study of a) Ecology of individual species b) Ecology of many species c) Ecology of community d) All of these 6. The branch of ecology which deals with the study of the organisms and geological environments of past is called a) Cytoecology b) Palecology c) Synecology d) Autoecology 7. Ecology deals with the study of a) Living beings b) Living and non living components c) Reciprocal relationship between living and non living components d) Biotic and Abiotic components 8. Phylloclade is modified a) Root b) Leaf c) Stem d) Bud 9. Cuscuta is a) Parasite b) Epiphyte c) Symbiont d) Lichen 10. Mycorrhiza is example of a) Symbiotic relationship b) Parasitic relationship c) Saprophytic relationship d) Negative interaction 11. Edaphic ecological factors are concerned with a) Rainfall b) Light c) Competition d) Soil 12. The soil is said to be physiologically dry when a) Temperature band light available to plants is insufficient b) There is abundance of water in soil c) Soil water is with high concentration of salts d) Both b and c 13.
    [Show full text]
  • When Corridors Work: Insights from a Microecosystem
    ecological modelling 202 (2007) 441–453 available at www.sciencedirect.com journal homepage: www.elsevier.com/locate/ecolmodel When corridors work: Insights from a microecosystem Martin Hoyle ∗ School of Biology, University of Nottingham, Nottingham NG7 2RD, UK article info abstract Article history: Evidence for a beneficial effect of corridors on species richness and abundance in habitat Received 14 February 2006 patches is mixed. Even in a single microecosystem of microarthropods living in moss patches Received in revised form connected by a moss corridor, experiments have had different results (positive and neutral). 2 November 2006 This paper attempts to provide an explanatory framework for understanding these results. I Accepted 7 November 2006 developed a stochastic individual-based model of the moss—microarthropod microecosys- Published on line 12 December 2006 tem. Some of the movement parameter values were estimated from two manipulation experiments. Assuming mortality independent of the season, and assuming the corridors Keywords: merely increase migration rates between patches, only a very weak beneficial effect of corri- Conservation dors was possible in simulations. Incorporating a seasonal pattern to mortality caused some Habitat fragmentation simulated populations to die out, which were then occasionally rescued by migrants from Individual-based model the adjacent patch. Corridors were slightly beneficial if there was little or no immigration Metapopulation from the surrounding matrix. In contrast, corridors were very beneficial in simulations that Microarthropod incorporated lower emigration to the matrix when a corridor was present, even for moder- ate levels of immigration from the matrix. Thus corridors may reduce the chance of species extinction in patches even when the lifespan of the individuals is long relative to the time- scale in question.
    [Show full text]
  • Information to Users
    INFORMATION TO USERS This manuscript has been reproduced from the microfilm master. UMI films the text directly from the original or copy submitted. Thus, some thesis and dissertation copies are in typewriter face, while others may be from any type of computer printer. The quality of this reproduction is dependent upon the quality of the copy submitted. Broken or indistinct print, colored or poor quality illustrations and photographs, print bleedthrough, substandard margins, and improper alignment can adverselyaffect reproduction. In the unlikely event that the author did not send UMI a complete manuscript and there are missing pages, these will be noted. Also, if unauthorized copyright material had to be removed, a note will indicate the deletion. Oversize materials (e.g., maps, drawings, charts) are reproduced by sectioning the original, beginning ,H the upper left-hand corner and continuing from left to right in equal sections with small overlaps. Each original is also photographed in one exposure and is included in reduced form at the back of the book. Photographs included in the original manuscript have been reproduced xerographically in this copy. Higher quality 6" x 9" black and white photographic prints are available for any photographs or illustrations appearing in this copy for an additional charge. Contact UMI directly to order. University Microfilms International A Beil & Howell Information Company 300 North Zeeb Road. Ann Arbor. M148106-1346 USA 313, 761-4700 800.521-0600 ~_..,------ Order Number 9215041 Stability in closed ecological systems: An examination of material and energetic parameters Shaffer, Jonathon Andrew, Ph.D. University of Hawaii, 1991 Copyright @1991 by Shaffer, Jonathon Andrew.
    [Show full text]
  • ENVIRONMENTAL SYSTEMS and SOCIETIES SL IB Academy Environmental Systems and Societies Study Guide
    STUDY GUIDE ENVIRONMENTAL SYSTEMS AND SOCIETIES SL www.ib.academy IB Academy Environmental systems and societies Study Guide Available on learn.ib.academy Author: Laurence Gibbons Design Typesetting This work may be shared digitally and in printed form, but it may not be changed and then redistributed in any form. Copyright © 2020, IB Academy Version: ESS.2.1.200320 This work is published under the Creative Commons BY-NC-ND 4.0 International License. To view a copy of this license, visit creativecommons.org/licenses/by-nc-nd/4.0 This work may not used for commercial purposes other than by IB Academy, or parties directly licenced by IB Academy. If you acquired this guide by paying for it, or if you have received this guide as part of a paid service or product, directly or indirectly, we kindly ask that you contact us immediately. Laan van Puntenburg 2a ib.academy 3511ER, Utrecht [email protected] The Netherlands +31 (0) 30 4300 430 0 Welcome to the IB Academy guide book for IB Environmental Systems and Society Standard Level. This guide contains all the theory you should know for your final exam. To achieve top marks this theory should be complimented with case studies. Although not covered in this booklet, we provide some in our online podcast series. The guide starts with an explanation of systems and models which are the foundations for the whole course. We will then look at systems in the natural world before turning our attention to humans and their impact. Throughout the guide there are helpful hints from the former IB students who now teach with IB Academy.
    [Show full text]
  • A Keystone Predator Controls Bacterial Diversity in the Pitcher-Plant (Sarracenia Purpurea) Microecosystem
    Environmental Microbiology (2008) 10(9), 2257–2266 doi:10.1111/j.1462-2920.2008.01648.x A keystone predator controls bacterial diversity in the pitcher-plant (Sarracenia purpurea) microecosystem Celeste N. Peterson,1,2* Stephanie Day,3,4† Introduction Benjamin E. Wolfe,1 Aaron M. Ellison,4 Bacterial assemblages may be structured by many Roberto Kolter2 and Anne Pringle1 forces at both local and regional scales. They are inte- 1Department of Organismic and Evolutionary Biology, grated into complex food webs and their composition, in Harvard University, Cambridge, MA 02138, USA. terms of species diversity and cell abundance, may be 2Department of Microbiology and Molecular Genetics, controlled by a combination of ‘top-down’ factors, such Harvard Medical School, Boston, MA 02115, USA. as grazing by predators, and ‘bottom-up’ factors, such 3Department of Biology, Howard University, Washington, as nutrient availability or other environmental param- DC 20059, USA. eters (Moran and Scheidler, 2002). Simple and well- 4Harvard Forest, Harvard University, Petersham, characterized natural model ecosystems can be used to MA 01366, USA. determine how each of these forces function and inter- act with each other at different scales. However, there is Summary a dearth of such systems available for field studies. That makes the model ecosystem of the carnivorous pitcher- The community of organisms inhabiting the water- plant Sarracenia purpurea (Sarraceniaceae) particularly filled leaves of the carnivorous pitcher-plant Sarrace- valuable. nia purpurea includes arthropods, protozoa and The food web that forms in the water-filled leaves of bacteria, and serves as a model system for studies of S. purpurea (Sarraceniaceae) has been used as a model food web dynamics.
    [Show full text]
  • Molecular Musings in Microbial Ecology and Evolution Rebecca J Case* and Yan Boucher*
    Case and Boucher Biology Direct 2011, 6:58 http://www.biology-direct.com/content/6/1/58 REVIEW Open Access Molecular musings in microbial ecology and evolution Rebecca J Case* and Yan Boucher* Abstract: A few major discoveries have influenced how ecologists and evolutionists study microbes. Here, in the format of an interview, we answer questions that directly relate to how these discoveries are perceived in these two branches of microbiology, and how they have impacted on both scientific thinking and methodology. The first question is “What has been the influence of the ‘Universal Tree of Life’ based on molecular markers?” For evolutionists, the tree was a tool to understand the past of known (cultured) organisms, mapping the invention of various physiologies on the evolutionary history of microbes. For ecologists the tree was a guide to discover the current diversity of unknown (uncultured) organisms, without much knowledge of their physiology. The second question we ask is “What was the impact of discovering frequent lateral gene transfer among microbes?” In evolutionary microbiology, frequent lateral gene transfer (LGT) made a simple description of relationships between organisms impossible, and for microbial ecologists, functions could not be easily linked to specific genotypes. Both fields initially resisted LGT, but methods or topics of inquiry were eventually changed in one to incorporate LGT in its theoretical models (evolution) and in the other to achieve its goals despite that phenomenon (ecology). The third and last question we ask is “What are the implications of the unexpected extent of diversity?” The variation in the extent of diversity between organisms invalidated the universality of species definitions based on molecular criteria, a major obstacle to the adaptation of models developed for the study of macroscopic eukaryotes to evolutionary microbiology.
    [Show full text]
  • Glossary of Landscape and Vegetation Ecology for Alaska
    U. S. Department of the Interior BLM-Alaska Technical Report to Bureau of Land Management BLM/AK/TR-84/1 O December' 1984 reprinted October.·2001 Alaska State Office 222 West 7th Avenue, #13 Anchorage, Alaska 99513 Glossary of Landscape and Vegetation Ecology for Alaska Herman W. Gabriel and Stephen S. Talbot The Authors HERMAN w. GABRIEL is an ecologist with the USDI Bureau of Land Management, Alaska State Office in Anchorage, Alaskao He holds a B.S. degree from Virginia Polytechnic Institute and a Ph.D from the University of Montanao From 1956 to 1961 he was a forest inventory specialist with the USDA Forest Service, Intermountain Regiono In 1966-67 he served as an inventory expert with UN-FAO in Ecuador. Dra Gabriel moved to Alaska in 1971 where his interest in the description and classification of vegetation has continued. STEPHEN Sa TALBOT was, when work began on this glossary, an ecologist with the USDI Bureau of Land Management, Alaska State Office. He holds a B.A. degree from Bates College, an M.Ao from the University of Massachusetts, and a Ph.D from the University of Alberta. His experience with northern vegetation includes three years as a research scientist with the Canadian Forestry Service in the Northwest Territories before moving to Alaska in 1978 as a botanist with the U.S. Army Corps of Engineers. or. Talbot is now a general biologist with the USDI Fish and Wildlife Service, Refuge Division, Anchorage, where he is conducting baseline studies of the vegetation of national wildlife refuges. ' . Glossary of Landscape and Vegetation Ecology for Alaska Herman W.
    [Show full text]
  • Xerosere Faculty Name: Dr Pinky Prasad Email: [email protected]
    Course: B.Sc. Botany Semester: IV Paper Code: BOT CC409 Paper Name: Plant Ecology and Phytogeography Topic: Xerosere Faculty Name: Dr Pinky Prasad Email: [email protected] XEROSERE: Xerosere is a plant succession that is limited by water availability. A xerosere may be lithosere initiating on rock and psammosere initiating on sand. LITHOSERE Bare rocks Bare rocks are eroded by rain water and wind loaded with soil particles. The rain water combines with atmospheric carbon dioxide that corrodes the surface of the rocks and produce crevices. Water enters these crevices, freezes and expands to further increase the crevices. The wind loaded with soil particles deposit soil particles on the rock and its crevices. All these processes lead to formation of a little soil at the surface of these bare rocks. Algal and fungal spores reach these rocks by air from the surrounding areas. These spores grow and form symbiotic association, the lichen, which act as pioneer species of bare rocks. There are six steps of xerosere. They are as follows: 1) Crustose lichen stage. 2) Foliage and fruticose lichen stage. 3) Moss stage. 4) Herbaceous stage 5) Shrub stage. 6) Climax forest stage. 1) Crustose lichen stage A bare rock consists of a very thin film of soil and there is no place for rooting plants to colonize. The plant succession starts with crustose lichens like Lecanora, Lecidia, Rhizocarpon lichen which adhere to the thin film of soil on the surface of rock and absorb moisture from atmosphere. These lichens produce lichenic acids which corrode the rock and their thalli collect windblown soil particles among them that help in formation of a thin layer of soil.
    [Show full text]
  • Linking the Development and Functioning of a Carnivorous Pitcher Plant’S Microbial Digestive Community
    The ISME Journal (2017) 11, 2439–2451 © 2017 International Society for Microbial Ecology All rights reserved 1751-7362/17 www.nature.com/ismej ORIGINAL ARTICLE Linking the development and functioning of a carnivorous pitcher plant’s microbial digestive community David W Armitage1,2 1Department of Integrative Biology, University of California Berkeley, Berkeley, CA, USA and 2Department of Biological Sciences, University of Notre Dame, Notre Dame, IN, USA Ecosystem development theory predicts that successional turnover in community composition can influence ecosystem functioning. However, tests of this theory in natural systems are made difficult by a lack of replicable and tractable model systems. Using the microbial digestive associates of a carnivorous pitcher plant, I tested hypotheses linking host age-driven microbial community development to host functioning. Monitoring the yearlong development of independent microbial digestive communities in two pitcher plant populations revealed a number of trends in community succession matching theoretical predictions. These included mid-successional peaks in bacterial diversity and metabolic substrate use, predictable and parallel successional trajectories among microbial communities, and convergence giving way to divergence in community composition and carbon substrate use. Bacterial composition, biomass, and diversity positively influenced the rate of prey decomposition, which was in turn positively associated with a host leaf’s nitrogen uptake efficiency. Overall digestive performance was
    [Show full text]
  • Biodiversity and Ecosystem Functioning in Soil
    Article Lijbert Brussaard et al. Biodiversity and Ecosystem Functioning in Soil ognition of the pivotal role of the world's biota as the life-sup- We review the current knowledge on biodiversity in soils, port system for planet Earth, has revived interest in soil its role in ecosystem processes, its importance for human biodiversity as an asset to conserve, to understand and to man- purposes, and its resilience against stress and disturbance. age wisely in terms of its contribution to ecosystem services. The number of existing species is vastly higher than the The objective of this paper is to review the knowledge on the number described, even in the macroscopically visible diversity of soil biota and its role in ecosystem functioning, and taxa, and biogeographical syntheses are largely lacking. to identify key areas for future research. A major effort in taxonomy and the training of a new Although the diversity of soil organisms is worth conserving generation of systematists is imperative. This effort has to and studying in its own right, their functional roles offer a use- be focussed on the groups of soil organisms that, to the ful framework for making this effort more meaningful. We will best of our knowledge, play key roles in ecosystem functioning. To identify such groups, spheres of influence first define functional roles in a utilitarian way as ecosystem (SOI) of soil biota-such as the root biota, the shredders services. We will then have a closer look at what we mean by of organic matter and the soil bioturbators-are recognized biodiversity in soil, emphasizing spheres of influence (2; 'bio- that presumably control ecosystem processes, for logical systems of regulation' in 3) of the biota in soil and vari- example, through interactions with plants.
    [Show full text]
  • Microbial Experimental Systems in Ecology
    Microbial Experimental Systems in Ecology CHRISTINE M. JESSUP, SAMANTHA E. FORDE AND BRENDAN J.M. BOHANNAN I. Summary . .............................................. 273 II. Introduction. .......................................... 274 III. The History of Microbial Experimental Systems in Ecology . ...... 275 A. G.F. Gause and His Predecessors . ...................... 275 B. Studies Since Gause . .................................. 276 IV. Strengths and Limitations of Microbial Experimental Systems ...... 277 A. Strengths of Microbial Experimental Systems . .............. 277 B. Limitations of Microbial Experimental Systems . .......... 281 V. The Role of Microbial Experimental Systems in Ecology: Consensus and Controversy. .............................. 283 A. The Role of Microbial Experimental Systems . .............. 283 B. Controversies Surrounding Experimental Systems and Microbial Experimental Systems . ...................... 285 VI. Recent Studies of Microbial Experimental Systems . .............. 289 A. The Ecological Causes of Diversity . ...................... 290 B. The Ecological Consequences of Diversity.................. 296 C. The Response of Diversity to Environmental Change . ...... 298 D. Directions for Future Research .......................... 299 VII. Conclusions . .......................................... 300 Acknowledgments. .......................................... 300 References................................................... 300 I. SUMMARY The use of microbial experimental systems has traditionally been limited
    [Show full text]