DOCSLIB.ORG

A Taxonomic Revision of the Tragulus Mouse-Deer (Artiodactyla)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
A Taxonomic Revision of the Tragulus Mouse-Deer (Artiodactyla) Blackwell Science, LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082The Lin- nean Society of London, 2004? 2004 1401 63102 Original Article E. MEIJAARD and C. P. GROVESA REVISION OF TRAGULUS Zoological Journal of the Linnean Society, 2004, 140, 63–102. With 23 figures A taxonomic revision of the Tragulus mouse-deer (Artiodactyla) E. MEIJAARD* and C. P. GROVES School of Archaeology and Anthropology, Faculty of Arts, A.D. Hope Building, Australian National University, 0200 ACT, Australia Received October 2002; accepted for publication July 2003 The taxonomy of South-East Asian mouse-deer (Tragulus) is complex, and after some 120 years of considerable tax- onomic revisions of the genus a clear key is still lacking for the determination of species and subspecies. Through craniometrical analysis of 338 skulls of Tragulus and some study of coat coloration patterns we have come to a better understanding of mouse-deer taxonomy. Our results show that there are three species groups: the T. javanicus- group, the T. napu-group, and T. versicolor. Within the T. javanicus-group we recognize three species: T. javanicus (from Java), T. williamsoni (from northern Thailand and possibly southern China), and T. kanchil (from the rest of the range), and within these species we provisionally recognize 16 subspecies. Within the T. napu-group we recognize two species: T. nigricans (from Balabac), and T. napu (from the rest of the range); within these species we provi- sionally recognize eight subspecies. T. versicolor from Nhatrang, south-east Vietnam, is distinct from the two pre- vious groups; it is, however, unclear whether this species is still extant. © 2004 The Linnean Society of London, Zoological Journal of the Linnean Society, 2004, 140, 63–102. ADDITIONAL KEYWORDS: chevrotain – coat colours – craniometrics – island fauna – South-East Asia – taxonomy – Tragulidae. INTRODUCTION Smit-van Dort (1989) studied the morphological dif- ferences between the four conventionally recognized The chevrotains or mouse-deer (Tragulidae) are an species of tragulids, i.e. the African Hyemoschus ancient group of ungulates, with a shared history dat- aquaticus, Moschiola meminna, Tragulus javanicus, ing back to the Miocene. They are considered the and T. napu. She could separate the two Tragulus spe- sister-group of the remaining living Ruminantia cies only by using metrical characters in the skeleton, (Groves & Grubb, 1982). In southern Asia, they are i.e. the greater mouse-deer T. napu has stronger and found in fossil assemblages dated at 18 million years broader feet than the lesser mouse-deer T. javanicus. before present (Myr), although they reached their Skull dimensions of T. napu and T. javanicus, espe- highest diversity with five named and 52 unnamed cially in specimens from small islands, converged to species at around 11.5 Myr (Barry et al., 1991). such an extent that the two species could not be reli- Between 16 and 14 Myr, they were also common in the ably separated using skull characters (also see van area that is now northern Thailand, although species Dort, 1988). Skin patterns do differ between the two diversity may have been low (Ducrocq et al., 1994). species, with T. napu having mottled fur and light After 9 Myr, the tragulid family declined significantly superciliary lines on the head, whereas T. javanicus in diversity in southern Asia (Barry et al., 1991), normally has three strong white lines on the throat which may have been caused by the evolution of more and has uniformly coloured upperparts (Smit-van open vegetation types. Presently there are three gen- Dort, 1989). There is, however, considerable variation era left, of which two, Moschiola and Tragulus, occur within each species in the coloration and striping pat- in the South and South-East Asian region. terns, especially on the many islands where the spe- cies occur, and in several instances it is nearly impossible to determine whether a taxon is more *Corresponding author. E-mail: [email protected] closely related to T. napu or to T. javanicus. In fact, © 2004 The Linnean Society of London, Zoological Journal of the Linnean Society, 2004, 140, 63–102 63 64 E. MEIJAARD and C. P. GROVES Groves (1989) considered the two to be sibling species. We propose instead to designate Gmelin’s Moschus Based on this geographical variation, Chasen (1940), javanicus as type, which, with Pallas’ Tragulus, would the first major reviser of Tragulus, recorded 27 sub- fix the name of the Javan mouse-deer as Tragulus jav- species for T. napu and 26 for T. javanicus (some of anicus (this will be part of our submission to the which he had reallocated from one to the other) for the Commission). Malayan region, and at least ten more subspecies, pri- Opinions as to the identity of javanicus, whether marily from mainland Asia, have been described in dated from Osbeck or Gmelin, have fluctuated other publications. The determination of this plethora between authors, from the large (napu group) to the of subspecies has been based primarily on variation in small (kanchil group) and back. Raffles (1822), the skin patterns, but when, during preliminary research, first to mention the existence of two (or even three) we investigated skins of Tragulus we realized that the species of South-East Asian mouse-deer, called the characteristics of the subspecies reported by Chasen larger species Moschus javanicus Pallas and named (1940) and others were not always consistent, and the smaller M. kanchil, mentioning in addition a there appeared to be considerable morphological over- potential third species, ‘the Pelandok’, based on anec- lap between subspecies. Also, some taxa that Chasen dotal information. His specimens were from the had assigned to subspecies appeared to be distinct Bengkulu district of Sumatra. His descriptions of the enough to qualify as species. We therefore decided to two named species are accurate, and recognizably focus on a craniometrical analysis, because the taxon- those of the two that occur today sympatrically on omists who worked on the genus had paid relatively Sumatra. Hamilton-Smith (1827) named Raffles’ little attention to skull form. Pelandok Moschus pelandoc, although his rather In this research we investigated skulls of Tragulus confused description suggests that he thought his to obtain a more transparent and consistent taxonomy specimen to be the same as Raffles’ ‘Napu’. Miller for the genus. In addition we studied some skins, but (1903b) finally rejected all descriptions of because many type specimens especially of small M. javanicus, T. javanicus, and T. pelandoc, and island taxa are located in the Smithsonian Museum, instead proposed a new species, T. focalinus, to desig- which we did not visit, we were unable to assess the nate the species from Java. validity of all taxa. We intend to address the variation Gray (1836) recognized Moschus javanicus Gmelin between these island taxa, and also variation within (with M. napu F.Cuvier a synonym), Moschus kanchil M. meminna in future publications. Raffles, and two new species: Moschus fulviventer (which he thought is probably Raffles’ ‘pelandok’) and Moschus stanleyanus. After these initial descriptions TAXONOMIC HISTORY OF THE TRAGULIDAE several intermediate revisions appeared. In a later The earliest scientific name generally considered publication, Gray (1843) dropped the genus Moschus, applicable to a tragulid is Cervus javanicus Osbeck included it in the Bovidae, and referred the South- (1757), described from Nieu Bay, Ujung Kulon, Java. East Asian mouse-deer to Tragulus; he also introduced We are not convinced that this name actually applies the name Tragulus affinis from Malacca. Blyth (1858) to a tragulid at all; he may have been describing a suggested that four (if not five) species of Tragulus juvenile cervid, perhaps Muntiacus muntjak. As this were distinguishable. Milne-Edwards (1864) wrote a will be the subject of a future application to the Inter- long review of the Tragulidae, recognizing Tragulus national Commission on Zoological Nomenclature, we javanicus, T. napu, T. kanchil and T. stanleyanus will not argue the case here. (as well as South Asian T. meminna). Gray (1872) Pallas (1777) gave what we consider to be the first changed his species listing yet again and recognized description of a specimen of a mouse-deer that he had T. stanleyanus, T. javanicus (of which he maintained obtained from Java; he did not name it, but Gmelin that T. napu of Milne-Edwards was the adult, and his (1788) considered it a new species of musk-deer (Mos- T. javanicus the young), T. fulviventer (= Tragulus chus), and on the basis of Pallas’ description named it kanchil Milne-Edwards, 1864), and Tragulus pyg- Moschus javanicus (see Appendix 1). Pallas’ descrip- maeus. One year later, Gray (1873) dropped the last tion not only clearly refers to a mouse-deer, but his species again. mention of a grey nape confirms that the specimen The late 19th and early 20th centuries were a time was from Java, as no other mouse-deer species has of prolixity in the taxonomy of Tragulus, when zoolo- this characteristic (e.g. Miller, 1903a). Pallas also cited gists such as Miller, Lyon, Lydekker, Kloss, Sody, Bon- Brisson’s (1762) generic name Tragulus, which had hote and Chasen described numerous species and been substituted for Linnaeus’ Moschus; as Gentry subspecies. They differed in their opinions on how spe- (1994) showed, the type of this genus is pygmaea, but cies and subspecies should be defined. For instance, she proposed, following Ellerman & Morrison-Scott Miller (1911) referred his new taxa to subspecies when (1951), to designate Cervus javanicus Osbeck as type. there was some overlap in skin characteristics © 2004 The Linnean Society of London, Zoological Journal of the Linnean Society, 2004, 140, 63–102 A REVISION OF TRAGULUS 65 between the specimens from two geographical groups, paraocciptal processes (PPW) (not shown in Fig.
Load more

Recommended publications
  • Hippotragus Equinus – Roan Antelope
    Hippotragus equinus – Roan Antelope authorities as there may be no significant genetic differences between the two. Many of the Roan Antelope in South Africa are H. e. cottoni or equinus x cottoni (especially on private properties). Assessment Rationale This charismatic antelope exists at low density within the assessment region, occurring in savannah woodlands and grasslands. Currently (2013–2014), there are an observed 333 individuals (210–233 mature) existing on nine formally protected areas within the natural distribution range. Adding privately protected subpopulations and an Cliff & Suretha Dorse estimated 0.8–5% of individuals on wildlife ranches that may be considered wild and free-roaming, yields a total mature population of 218–294 individuals. Most private Regional Red List status (2016) Endangered subpopulations are intensively bred and/or kept in camps C2a(i)+D*†‡ to exclude predators and to facilitate healthcare. Field National Red List status (2004) Vulnerable D1 surveys are required to identify potentially eligible subpopulations that can be included in this assessment. Reasons for change Non-genuine: While there was an historical crash in Kruger National Park New information (KNP) of 90% between 1986 and 1993, the subpopulation Global Red List status (2008) Least Concern has since stabilised at c. 50 individuals. Overall, over the past three generations (1990–2015), based on available TOPS listing (NEMBA) Vulnerable data for nine formally protected areas, there has been a CITES listing None net population reduction of c. 23%, which indicates an ongoing decline but not as severe as the historical Endemic Edge of Range reduction. Further long-term data are needed to more *Watch-list Data †Watch-list Threat ‡Conservation Dependent accurately estimate the national population trend.
    [Show full text]
  • Connochaetes Gnou – Black Wildebeest
    Connochaetes gnou – Black Wildebeest Blue Wildebeest (C. taurinus) (Grobler et al. 2005 and ongoing work at the University of the Free State and the National Zoological Gardens), which is most likely due to the historic bottlenecks experienced by C. gnou in the late 1800s. The evolution of a distinct southern endemic Black Wildebeest in the Pleistocene was associated with, and possibly driven by, a shift towards a more specialised kind of territorial breeding behaviour, which can only function in open habitat. Thus, the evolution of the Black Wildebeest was directly associated with the emergence of Highveld-type open grasslands in the central interior of South Africa (Ackermann et al. 2010). Andre Botha Assessment Rationale Regional Red List status (2016) Least Concern*† This is an endemic species occurring in open grasslands in the central interior of the assessment region. There are National Red List status (2004) Least Concern at least an estimated 16,260 individuals (counts Reasons for change No change conducted between 2012 and 2015) on protected areas across the Free State, Gauteng, North West, Northern Global Red List status (2008) Least Concern Cape, Eastern Cape, Mpumalanga and KwaZulu-Natal TOPS listing (NEMBA) (2007) Protected (KZN) provinces (mostly within the natural distribution range). This yields a total mature population size of 9,765– CITES listing None 11,382 (using a 60–70% mature population structure). This Endemic Yes is an underestimate as there are many more subpopulations on wildlife ranches for which comprehensive data are *Watch-list Threat †Conservation Dependent unavailable. Most subpopulations in protected areas are stable or increasing.
    [Show full text]
  • MALE GENITAL ORGANS and ACCESSORY GLANDS of the LESSER MOUSE DEER, TRAGULUS Fa VAN/CUS
    MALE GENITAL ORGANS AND ACCESSORY GLANDS OF THE LESSER MOUSE DEER, TRAGULUS fA VAN/CUS M. K. VIDYADARAN, R. S. K. SHARMA, S. SUMITA, I. ZULKIFLI, AND A. RAZEEM-MAZLAN Faculty of Biomedical and Health Science, Universiti Putra Malaysia, 43400 UPM Serdang, Selangor, Malaysia (MKV), Faculty of Veterinary Medicine and Animal Sciences, Universiti Putra Malaysia, 43400 UPM Serdang, Selangor, Malaysia (RSKS, SS, /Z), Downloaded from https://academic.oup.com/jmammal/article/80/1/199/844673 by guest on 01 October 2021 Department of Wildlife and National Parks, Zoo Melaka, 75450 Melaka, Malaysia (ARM) Gross anatomical features of the male genital organs and accessory genital glands of the lesser mouse deer (Tragulus javanicus) are described. The long fibroelastic penis lacks a prominent glans and is coiled at its free end to form two and one-half turns. Near the tight coils of the penis, on the right ventrolateral aspect, lies a V-shaped ventral process. The scrotum is prominent, unpigmented, and devoid of hair and is attached close to the body, high in the perineal region. The ovoid, obliquely oriented testes carry a large cauda and caput epididymis. Accessory genital glands consist of paired, lobulated, club-shaped vesic­ ular glands, and a pair of ovoid bulbourethral glands. A well-defined prostate gland was not observed on the surface of the pelvic urethra. Many features of the male genital organs of T. javanicus are pleisomorphic, being retained from suiod ancestors of the Artiodactyla. Key words: Tragulus javanicus, male genital organs, accessory genital glands, reproduc­ tion, anatomy, Malaysia The lesser mouse deer (Tragulus javan­ gulidae, and Bovidae (Webb and Taylor, icus), although a ruminant, possesses cer­ 1980).
    [Show full text]
  • Perkembangan Pendidikan Di Kabupaten Natuna Pasca Pemekaran (2006-2013)
    1 PERKEMBANGAN PENDIDIKAN DI KABUPATEN NATUNA PASCA PEMEKARAN (2006-2013) Musdar Halifah *, Tugiman **, Marwoto Saiman E-mail : [email protected] Telp/No. HP: 085272621116 Program Studi Pendidikan Sejarah Fakultas Keguruan dan Ilmu Pendidikan Universitas Riau Abstrak : Natuna district education office has the arduous task in advancing education in the District Natuna after the split, after the division of the education department autonomously take care of the quality of education and the development of education in its own country, because after the expansion of education in the Natuna has not reached the desired target by the community and local government. Achievement of the National Examination in the 2010-2011 school year. Department of Education District Natuna need accurate measures and hard work. By carrying out the qualifications of teachers in the District Natuna and will provide tutoring outside of school to students. The purpose of the study was to (1) determine the expansion history of the area in Riau Islands District Natuna Regency, especially in the years 2006-2013. (2) determine the development of education in post-Proliferation Natuna Regency 2006-2013. (3) determine the barriers faced by the District Education Office Natuna 2006-2013. This type of research is descriptive qualitative approach that aims to provide a systematic overview of the state of the ongoing research on the object. Where research is Natuna. When the study is planned for 4 months. Data collection techniques in this study included three observation
    [Show full text]
  • Animals of Africa
    Silver 49 Bronze 26 Gold 59 Copper 17 Animals of Africa _______________________________________________Diamond 80 PYGMY ANTELOPES Klipspringer Common oribi Haggard oribi Gold 59 Bronze 26 Silver 49 Copper 17 Bronze 26 Silver 49 Gold 61 Copper 17 Diamond 80 Diamond 80 Steenbok 1 234 5 _______________________________________________ _______________________________________________ Cape grysbok BIG CATS LECHWE, KOB, PUKU Sharpe grysbok African lion 1 2 2 2 Common lechwe Livingstone suni African leopard***** Kafue Flats lechwe East African suni African cheetah***** _______________________________________________ Red lechwe Royal antelope SMALL CATS & AFRICAN CIVET Black lechwe Bates pygmy antelope Serval Nile lechwe 1 1 2 2 4 _______________________________________________ Caracal 2 White-eared kob DIK-DIKS African wild cat Uganda kob Salt dik-dik African golden cat CentralAfrican kob Harar dik-dik 1 2 2 African civet _______________________________________________ Western kob (Buffon) Guenther dik-dik HYENAS Puku Kirk dik-dik Spotted hyena 1 1 1 _______________________________________________ Damara dik-dik REEDBUCKS & RHEBOK Brown hyena Phillips dik-dik Common reedbuck _______________________________________________ _______________________________________________African striped hyena Eastern bohor reedbuck BUSH DUIKERS THICK-SKINNED GAME Abyssinian bohor reedbuck Southern bush duiker _______________________________________________African elephant 1 1 1 Sudan bohor reedbuck Angolan bush duiker (closed) 1 122 2 Black rhinoceros** *** Nigerian
    [Show full text]
  • A Scoping Review of Viral Diseases in African Ungulates
    veterinary sciences Review A Scoping Review of Viral Diseases in African Ungulates Hendrik Swanepoel 1,2, Jan Crafford 1 and Melvyn Quan 1,* 1 Vectors and Vector-Borne Diseases Research Programme, Department of Veterinary Tropical Disease, Faculty of Veterinary Science, University of Pretoria, Pretoria 0110, South Africa; [email protected] (H.S.); [email protected] (J.C.) 2 Department of Biomedical Sciences, Institute of Tropical Medicine, 2000 Antwerp, Belgium * Correspondence: [email protected]; Tel.: +27-12-529-8142 Abstract: (1) Background: Viral diseases are important as they can cause significant clinical disease in both wild and domestic animals, as well as in humans. They also make up a large proportion of emerging infectious diseases. (2) Methods: A scoping review of peer-reviewed publications was performed and based on the guidelines set out in the Preferred Reporting Items for Systematic Reviews and Meta-Analyses (PRISMA) extension for scoping reviews. (3) Results: The final set of publications consisted of 145 publications. Thirty-two viruses were identified in the publications and 50 African ungulates were reported/diagnosed with viral infections. Eighteen countries had viruses diagnosed in wild ungulates reported in the literature. (4) Conclusions: A comprehensive review identified several areas where little information was available and recommendations were made. It is recommended that governments and research institutions offer more funding to investigate and report viral diseases of greater clinical and zoonotic significance. A further recommendation is for appropriate One Health approaches to be adopted for investigating, controlling, managing and preventing diseases. Diseases which may threaten the conservation of certain wildlife species also require focused attention.
    [Show full text]
  • Wild Ways Well and Deer
    Wild Ways Well and Deer Today’s Wild Ways Well task is to go for a walk in your local greenspace and keep an eye out for deer… Remember to follow the guidelines on Social Distancing, stay 2m apart from other people and only walk in your local area – and remember to wash your hands! You’ll Be Active by carefully walking outdoors (observing social distancing) keeping your mind busy and occupying your time looking for signs of these elusive mammals. Deer are quite common, even in urban areas, but spotting them can be difficult. We can Connect with deer by opening up our senses and empathising with the way they live their lives. Deer have many of the same needs as us – how do they find food, water and shelter in Cumbernauld? How do their senses compare to ours? Do they see the world in the same way we do? We can Keep Learning, there are hundreds of web pages, book and tv programmes dedicated to deer. Deer have been part of human culture for thousands of years, we can learn what our ancestors thought of them and how we can live alongside them today. Although they are secretive and hard to see Deer area actually all around us, and are vital to the ecosystem we all share but we rarely Take Notice and look very closely at them. It’s amazing how much we miss out in nature when we just walk through without paying attention to what is around us. We can Give by giving ourselves a break from the drama of the current events and focusing on the little things around us that give us pleasure and by sharing these with others, in person or online.
    [Show full text]
  • Cervid Mixed-Species Table That Was Included in the 2014 Cervid RC
    Appendix III. Cervid Mixed Species Attempts (Successful) Species Birds Ungulates Small Mammals Alces alces Trumpeter Swans Moose Axis axis Saurus Crane, Stanley Crane, Turkey, Sandhill Crane Sambar, Nilgai, Mouflon, Indian Rhino, Przewalski Horse, Sable, Gemsbok, Addax, Fallow Deer, Waterbuck, Persian Spotted Deer Goitered Gazelle, Reeves Muntjac, Blackbuck, Whitetailed deer Axis calamianensis Pronghorn, Bighorned Sheep Calamian Deer Axis kuhili Kuhl’s or Bawean Deer Axis porcinus Saurus Crane Sika, Sambar, Pere David's Deer, Wisent, Waterbuffalo, Muntjac Hog Deer Capreolus capreolus Western Roe Deer Cervus albirostris Urial, Markhor, Fallow Deer, MacNeil's Deer, Barbary Deer, Bactrian Wapiti, Wisent, Banteng, Sambar, Pere White-lipped Deer David's Deer, Sika Cervus alfredi Philipine Spotted Deer Cervus duvauceli Saurus Crane Mouflon, Goitered Gazelle, Axis Deer, Indian Rhino, Indian Muntjac, Sika, Nilgai, Sambar Barasingha Cervus elaphus Turkey, Roadrunner Sand Gazelle, Fallow Deer, White-lipped Deer, Axis Deer, Sika, Scimitar-horned Oryx, Addra Gazelle, Ankole, Red Deer or Elk Dromedary Camel, Bison, Pronghorn, Giraffe, Grant's Zebra, Wildebeest, Addax, Blesbok, Bontebok Cervus eldii Urial, Markhor, Sambar, Sika, Wisent, Waterbuffalo Burmese Brow-antlered Deer Cervus nippon Saurus Crane, Pheasant Mouflon, Urial, Markhor, Hog Deer, Sambar, Barasingha, Nilgai, Wisent, Pere David's Deer Sika 52 Cervus unicolor Mouflon, Urial, Markhor, Barasingha, Nilgai, Rusa, Sika, Indian Rhino Sambar Dama dama Rhea Llama, Tapirs European Fallow Deer
    [Show full text]
  • Bangka Tin, and the Collapse of the State Power
    GSTF Journal of Law and Social Sciences (JLSS) DOI 10.7603/s40741-016-0001-9 DOI: 10.5176/2251-2853_5.1.190 PrintI SSN: 2251-2853,E-periodical: 2251-2861 ; Volume 5, Issue 1; 2016 pp 1-7 © The Author(s) 2016. This article is published with open access by the GSTF. BANGKA TIN, AND THE COLLAPSE OF THE STATE POWER Ibrahim Department of Political Science Faculty of Politics and Social Sciences, University of Bangka Belitung Republic of Indonesia [email protected] Abstract - Bangka Belitung Islands is a region with the most Bangka Island has been indeed the biggest tin producer victorious tin route in the world. This tin wealth spans from in South East Asia and now it even becomes the only area Singkep to Belitung islands. Since Malaysia and Thailand producing tin in this region since Malaysia and Thailand closed their production and followed by Singkep on the late closed their production in 1990’s (Sujitno, 2007:5-7; of 90’s, Bangka Belitung islands have become the only Erman, 2010:3). region producing tin in Indonesia and South East Asia. Interestingly, since reformation rolling, tin that initially under full control of government has turned to be free Tin has been such a problematic matter, not only because commodity without clear management. Tin has entered the of its high price as an un-replaceable industrial whirlpool playing in all arenas, i.e. politics, law, ecology, component, but tin in Bangka Island also dealt with such social, up to the very complicated economy domain. How complicated management with very long management can the state loss its control over this nonrenewable journey.
    [Show full text]
  • This Regulation Shall Enter Into Force on the Day Of
    19 . 10 . 88 Official Journal of the European Communities No L 285 / 1 I (Acts whose publication is obligatory) COMMISSION REGULATION ( EEC ) No 3188 / 88 of 17 October 1988 amending Council Regulation ( EEC ) No 3626 / 82 on the implementation in the Community of the Convention on international trade in endangered species of wild fauna und flora THE COMMISSION OF THE EUROPEAN COMMUNITIES Whereas the measures provided for in this Regulation are in accordance with the opinion of the Committee on the Convention on International Trade in Endangered Species of Wild Fauna and Flora , Having regard to the Treaty establishing the European Economic Community , HAS ADOPTED THIS REGULATION : Having regard to Council Regulation ( EEC ) No 3626 / 82 Article 1 of 3 December 1982 on the implementation in the Corrimunity of the Convention on international trade in Appendix III of Annex A to Regulation ( EEC ) No 3626 / 82 endangered species of wild fauna and flora ( J ), as last is hereby replaced by the Annex to this Regulation . amended by Regulation ( EEC ) No 869 / 88 ( 2 ), and in particular Article 4 , thereof, Article 2 Whereas alterations were made to Appendix III to the This Regulation shall enter into force on the day of its publication in the Official Journal of the European Convention ; whereas Appendix III of Annex A to Communities . Regulation ( EEC ) No 3626 / 82 should now be amended to incorporate the amendments accepted by the Member States parties to the abovementioned Convention ; It shall apply from 21 September 1988 . This Regulation shall be binding in its entirety and directly applicable in all Member States .
    [Show full text]
  • Studi Geologi Awal Untuk Calon Tapak PLTN Di Pulau Singkep Dan Lingga, Kepulauan Riau
    Jurnal Pengembangan Energi Nuklir Vol. 21, No. 1, (2019) 35-44 Jurnal Pengembangan Energi Nuklir Laman Jurnal: jurnal.batan.go.id/index.php/jpen Studi Geologi Awal untuk Calon Tapak PLTN di Pulau Singkep dan Lingga, Kepulauan Riau Frederikus Dian Indrastomo*1, Heri Syaeful1, Kurnia Anzhar2, June Mellawati3 1Pusat Teknologi Bahan Galian Nuklir, BATAN, Jalan Lebak Bulus raya No. 9 Ps. Jumat, Jakarta, Indonesia 2Pusat Kajian Sistem Energi Nuklir, BATAN, Jalan Kuningan Barat, Mampang Prapatan, Jakarta, Indonesia. 3Pusat Teknologi Keselamatan dan Metrologi Radiasi, BATAN, Jalan Lebak Bulus raya No.49 Ps. Jumat, Jakarta, Indonesia INFORMASI ARTIKEL ABSTRAK Riwayat Artikel: STUDI GEOLOGI AWAL UNTUK CALON TAPAK PLTN DI PULAU SINGKEP DAN LINGGA, Diterima: KEPULAUAN RIAU. Pulau Singkep dan Lingga adalah bagian dari Kepulauan Riau, seperti Pulau 27 Juni 2019 Batam, Bintan, Karimun, terletak pada daerah strategis yang dapat menjadi lokasi Diterima dalam bentuk revisi: dikembangkannya perdagangan dan perindustrian. Guna memenuhi kebutuhan teknologi dan 01 Juli 2019 listriknya, PLTN merupakan salah satu alternatif pilihan. Berdasarkan hal ini telah dilakukan studi Disetujui: geologi awal guna mengetahui keberadaan daerah interes untuk lokasi PLTN di Kepulauan Riau. 15 Juli 2019 Tujuan penelitian untuk mengetahui kelayakan Pulau Singkep dan Lingga dari aspek geologi sebagai daerah interes untuk ditindaklanjuti sebagai calon tapak PLTN. Metodologi penelitian meliputi studi geologi regional melalui pengumpulan data sekunder, dan survei lapangan untuk verifikasi data sekunder. Verifikasi data lapangan meliputi pengamatan batuan untuk menentukan jenis dan karakteristiknya, pengukuran struktur geologi untuk mengetahui potensi sesar dan Kata kunci: gempa, pendataan sumber air panas untuk mengetahui aktivitas magmatisme. Pengamatan Lingga dilakukan di wilayah pesisir Pulau Singkep dan Lingga.
    [Show full text]
  • Annu Al Report 2019
    ANNUAL REPORT 2019 ANNUAL 1 ANNUAL REPORT 2019 Building on the initiatives of previous years, Telkomsel continued to expand and to enrich its digital business to shape the future through internal collaboration, synergies, and partnerships within the digital ecosystem at large. Telkomsel continued to expand and to enrich its digital business At the same time, Telkomsel strove to improve customer experience and satisfaction as key drivers of long-term success. (in billion rupiah) (in million) DIGITAL BUSINESS DATA USERS REVENUE 58,237 110.3 23.1% 3.5% DATA 50,550 LTE USERS 88.3 22.3% (in million) 61.3% DIGITAL SERVICES 7,687 29.0% 2019 63.9% DIGITAL 2018 BUSINESS 53.0% CONTRIBUTION 2 PT TELEKOMUNIKASI SELULAR IMPROVED MOMENTUM Telkomsel has successfully delivered growth and revenue from data supported by solid digital products and services offerings, as shown by TOTAL BTS improved momentum in 2019. 212,235 (in gigabyte) 12.2% CONSUMPTION/ 2019 DATA USER 3G/4G BTS 54.7% 5.2 161,938 16.7% 2018 3.4 (in terabyte) PAYLOAD 6,715,227 53.6% 3 ANNUAL REPORT 2019 Highlights of the Year 6 Key Performance Company 8 Financial Highlights at a Glance 9 Operational Highlights 10 2019 Event Highlights 52 Telkomsel in Brief 18 Awards & Accolades 53 Share Ownership History 23 ISO Certification 54 Organization Structure 54 Key Products & Services 56 Milestones Business Review Remarks from 60 Vision and Mission the Management 61 Corporate Strategy in Brief 62 Transformation Program 65 Marketing 26 Remarks from the President Commissioner 72 Digital Services 30
    [Show full text]