Entomofauna carpathica, 2020, 32(2): 19-46

CONTRIBUTION TO THE KNOWLEDGE OF (, ) OF CENTRAL AND NORTHERN LAOS, PART 3.

Karol BUCSEK

Slovak Academy of Sciences, Institute of Zoology, Dúbravská cesta 9, 845 06 Bratislava, Slovak republic; e-mail: [email protected]

BUCSEK, K. 2020. Contribution to the knowledge of Lithosiini (Erebidae, Arctiinae) of central and northern Laos, part 3. Entomofauna carpathica, 32(2): 19-46.

Abstract: This is the third part of study focusing on fauna of Lithosiini (Erebidae, Arctiinae) of central and northern Laos. The studied material consisted of 35 species that were collected in the Nakai (about 500 m), Bolikhamsai (200–300 m) and Huaphanh (1200–1900 m) areas. A new : Volynkinia n. gen. and 11 new species: Aemene cinerea n. sp., Aemene bicornuata n. sp., Aemene lancea n. sp., Aemene spinivalva n. sp., Volynkinia inaequalis n. sp., Halone ivetae n. sp., Stictosia maculata n. sp., Tospitis ogriagra n. sp., Tospitis denisae n. sp., Stictane bayarsaikhanae n. sp., Stictane lao n. sp. are described.

Key words: Erebidae, Lithosiini, new genus, new species, Laos, taxonomy

MATERIAL AND METHODS

The photos of adults were made with NIKON D700 and SONY DSC-WX220. Genitalia were mounted in microscopic slides using Canada balsam, morphological observations were made using and photographed by Digital USB 2,0 microscope camera (Micro Capture Pro). Subsequently the genitalia were demounted,preserved in glycerine in the test tubes and attached to the corresponding specimen. Post-shooting image processing was done using the software Adobe® Photoshop CS5. Holotypes are stored in the Slovak National Museum, Bratislava, paratypes in the collection of Karol Bucsek.

Abbreviations HT – holotype SNM – Slovak National Museum, Bratislava PT – paratype CKB – in coll. Karol Bucsek, Bratislava

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SYSTEMATIC SECTION

Genus Aemene Walker, 1854 Type species: Aemene taprobanis Walker, 1854 (Ceylon=Sri Lanka).

Aemene coniuncta Černý, 2009 (Figs. 1, 63)

Aemene coniuncta Černý, 2009, Dr. Karel Cerny, Bro. Amnuay Pinratana 2009, of Thailand vol.6: 104–105, pl.22: 207, gen. 2: 207 f, v. Aemene somodyi Bucsek, 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula- Malaysia, 65, pl. 12: 157, 157a, Mal207, n. syn. Aemene coniuncta Černý: Volynkin, Černý & Ivanova, 2017, Zootaxa 4306 (3), 448 (list).

Wingspan. Male 18 mm. Type locality. NE Thailand. Distribution in Laos. Huaphanh. Geographical range. Thailand, Laos, Peninsular Malaysia. Note. Aemene somodyi Bucsek is a n. syn. of Aemene coniuncta Černý.

Aemene eudonioides Volynkin & Černý, 2017 (Figs. 2, 64)

Aemene eudonioides Volynkin, Černý & Ivanova 2017, Zootaxa 4306 (3), 445– 446, figs 1, 2, 5, 7.

Wingspan. Male 22 mm. Type locality. N Thailand. Distribution in Laos. Huaphanh. Geographical range. Thailand, Laos.

Aemene cinerea Bucsek n. sp. (Figs. 3, 4, 65)

Holotype. ♂ LAOS, Bolikhamsai, Khoun Ngeun env., 200–300 m, 18°08′48.9″N, 04°27′44.7″E, 29. IV.–1. V. 2019, K. Bucsek leg., SNM with a meseum serial number SZ 10 946. Paratypes. 2 ♀♀ like holotype, CKB.

Wingspan. Male 18 mm, female 20–22 mm.

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Holotype. Male, antennae finely pectinate, head, thorax and abdomen are cinereous, anal margin cream-coloured. Forewing white-gray with irregular black bands in the basal, antemedial, medial and postmedial areas. The black coloration is more pronounced on costa. Discal and antemedial spot black, indistinct. Terminal line with black, regularly intermittent tape. Cilia cream- coloured with black dust. Hindwing light brown-gray, cilia whitish. Male genitalia. Uncus long, nearly straight, with a curved tip. Tuba analis large and conspicuous. Valva wide, only slightly narrowed to the top, distal costal process at the top with a small tip, two larger tips at the side. Discal saccular process with one tip. Aedeagus with a very large field with more or less equally large cornuti and one coarse pointed cornut. Female. Habitus very similar to the male, but forewing a little darker and hindwing gray. Antennae filiform. Female genitalia. Ductus bursae funnel-shaped, a distinctive appendix bursae is derived from the posterior section of the corpus bursae.

Aemene bicornuata Bucsek n. sp. (Figs. 5, 66)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.-13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 947.

Wingspan. Male 20 mm. Holotype. Antennae filiform, cream-coloured head, thorax and abdomen are light brown, anal margin lighter. Forewing light brown with black patterning, black coloration is more pronounced on costa. Discal spot black, distinctive. Hindwing light brown. Male genitalia. Uncus long, conspiciously pubescent, at the end with a curved tip, strongly widened in the middle of the inner side. Tuba analis large and expressive. Valva wide, slightly narrowed to the top, costa markedly serrated at the apex, apex with two distinct tips. Juxta with two sclerotized V-shaped spikes. Aedeagus with two promonent cornuti. Female. Unknown.

Aemene lancea Bucsek n. sp. (Figs. 6, 7, 67, 68)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.-13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 948. Paratype. 1 ♂ like holotype, CKB.

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Wingspan. Male 11–12 mm. Holotype. Male, 12 mm. Antennae filiform, head and thorax white-gray, abdomen light brown, anal margin cream-coloured. Forewing light creamy with dark brown patterning. Basal field with two indistinct dots, antemedial and medial line zigzag, postmedial line zigzag, slightly pronounced. The discal and antemedial spot is indistinct. Hindwing cream-coloured. Male genitalia. Uncus long and straight, with pointed tip. Valva wide, distal costal process (is smaller) and discal saccular process are connected and form a V-shape at the apex. Juxta is massive, creating a large tip that is serrated on the inside. Aedeagus with one large irregular cornut and a field of very small cornuti. Female. Unknown.

Aemene spinivalva Bucsek n. sp. (Figs. 8, 69)

Holotype. ♂ LAOS, Khammouane prov., Nakai env., 500 m, 17°34´N, 105°10´E, 25.- 29. IV. 2019, leg. K. Bucsek, SNM with a museum serial number SZ 10 949.

Wingspan. Male 11 mm. Holotype. Antennae filiform, head and thorax creamy, abdomen brown, anal margin in cream-coloured. Forewing creamy with black patterning. In the basal area two indistinct dots, in the antemedial area three dots, the fourth dot on the costa near to the apex. Postmedial line indistinct, in the subterminal area two spots. Antemedial spot small, discal spot distinctive. Terminal line is black, regularly intermittent. Cilia creamy. Hindwing creamy. Male genitalia. Uncus more or less straight, blunt at the termination. Vincular lobe sclerotized, wide and flat, terminated by two opposite spikes, inner spike pointed. Valva markedly sclerotized, distinctly serrated on the costal margin from the base to the apex. Ventral saccular process pronounced, slender, wider at the base. Aedeagus: carinal plate with three distinctive pointed processes of different sizes, apical field with a number of large cornuti. Female. Unknown.

Genus Volynkinia n. gen. Type species: Volynkinia inaequalis Bucsek (C Laos)

Genus is represented by one species. Small , both sexes have antennae filiform, forewing white-gray, with black patterning. Male genitalia: uncus club-shaped, massive, both sides of the tegumen with massive, symmetrical projections. Valvae asymmetric, left one is much more robust. Aedeagus very long and tenuous. Female genitalia: ductus bursae thin, sclerotized, corpus bursae ovoid.

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Etymology. I named this genus after an excellent expert on Arctiinae (Plaearct, Oriental region and Africa) Dr. Anton Valeryevich Volynkin (Barnaul, Russia).

Volynkinia inaequalis Bucsek n. sp. (Figs. 9, 10, 70, 71)

Holotype. ♂ LAOS, Bolikhamsai, Khoun Ngeun env., 200–300 m, 18°08′48.9″N, 04°27′44.7″E, 29. IV.–1. V. 2019, K. Bucsek leg., SNM with a meseum serial number SZ 10 950. Paratype. 1 ♀ like holotype, CKB.

Wingspan. Male 9 mm, female 8 mm. Holotype. Male, antennae filiform, head and thorax white, abdomen brown, anal margin creamy. Forewing white-gray, costa with four black spots from base to apex, antemedial and subterminal line zigzag, feebly distinctive. Discal spot indistinct, cilia white-gray. Hindwing gray. Male genitalia. Uncus massive, club-shaped, with pubescence. Massive conic- shaped growths symmetrical on both sides of tegumen. Valvae asymmetrical, the left is much more robust, both have a large lobular protrusion at the top. Juxta semi-circular, vinculum broad, elongated. Aedeagus very long and thin, without diagnostic characters. Female. Habitus very similar to the male. Female genitalia. Dorsal plate wide at the apex, with sharp protrusions on the sides. Ductus bursae thin, sclerotized, corpus bursae spherical, completely filled with spinules, the sclerotized medial band is stretched down from top.

Genus Halone Walker, 1854 Type species: Halone sobria Walker, 1854 (Australia)

Halone ivetae Bucsek n. sp. (Figs. 11-13, 72, 73)

Holotype. ♂ LAOS, Khammouane prov., Nakai env., 500 m, 17°34´N, 105°10´E, 25.- 29. IV. 2019, leg. K. Bucsek, SNM with a museum serial number SZ 10 951. Paratypes. 2 ♀♀ like holotype, CKB.

Wingspan. Male 12 mm, female 12 mm. Holotype. Male, antennae filiform, head, thorax and abdomen brown. Forewing ocher-brown, antemedial and subterminal area dark brown. Discal spot very indistinct, adjacent to subterminal area, cilia brown. Hindwing brown.

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Male genitalia. Uncus long, thin, with curved tip. Tuba analis pronounced. Valva very wide, narrowly tapered to the top, apex of costa with a blunt tip, a row of thorns at the medial costal angle. Juxta with four discernible protuberances. Vinculum broad, elongated and semi-cirkular. Aedeagus: carinal plate with two large serrate spikes, in apical field two large cornuti in a region of very small cornuti. Female. A little darker than the male, anal margin creamy. Female genitalia. Antevaginal plate with distinctive lobes on both sides. Corpus bursae with a band of distinctive cornuti. Etymology. I named the species after Mrs. Iveta Nováková, the wife of my esteemed friend, Mr. Henrich Novák, Bratislava.

Genus Neoduma Hampson, 1918 Type species: Neoduma ectozona Hampson, 1918 (Philippines)

Neoduma songensis Dubatolov & Bucsek, 2016 (Figs. 14, 74)

Neoduma songensis Dubatolov & Bucsek, 2016, Euroasian Ent. J., 15 (3): 236, figs 17, 36.

Wingspan. Male 10–12 mm. Type locality. N Vietnam. Distribution in Laos. Huaphanh, Nakai. Geographical range. Vietnam, Laos.

Neoduma ectozona Hampson, 1918 (Figs. 15, 75)

Neoduma ectozona Hampson, 1918, Novit. Zool 25: 102. Neoduma ectozona Hampson: Holloway, 2001, Moths of Borneo 7: 442–443, pl. 8, fig. 444. Neoduma ectozona Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 8, pl. 1: 7. Neoduma ectozona Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula – Malaysia, 72, Pl. 13: 175, 175a.

Wingspan. Male 11 mm. Type locality. Philippines – Luzon. Distribution in Laos. Nakai. Geographical range. Philippines, Borneo, Peninsular Malaysia, Thailand, Laos.

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Neoduma kuangtungensis (Daniel, 1951) (Figs. 16, 17, 76, 77)

Siccia kuangtungensis Daniel, 1951, Bonn. Zool. Beitr. 2: 303, pl. 1: 10. Siccia kuangtungensis Daniel: Fang, 2000, Fauna Sin. Ins.19, Lep. Arctiidae: 151–152, pl.9: 16. Neoduma kuangtungensis Daniel: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 8, pl. 1: 7, gen. 4: 8f, 8v. Neoduma kuangtungensis Daniel: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 72, Pl. 13: 176, 176a.

Wingspan. Male 12 mm, female 12 mm. Type locality. China – Kuangtung. Distribution in Laos. Huaphanh. Geographical range. China (Kuangtung, Chekiang), Laos, Thailand, Peninsular Malaysia.

Genus Eucyclopera Hampson, 1895 Type species: Eucyclopera plagidisca Hampson, 1895 (Bhutan)

Eucyclopera plagidisca Hampson, 1895 (Fig. 18)

Eucyclopera plagidisca Hampson, 1895, Trans. Ent. Soc., 294. Eucyclopera plagidisca Hampson: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 263, fig. 180. Eucyclopera plagidisca Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 8, pl. 1: 7, gen. 4: 8f, 8v.

Wingspan. Male 12 mm. Type locality. Bhutan. Distribution in Laos. Huaphanh. Geographical range. Bhutan, Laos, Thailand.

Genus Stictosia Hampson, 1900 Type species: Conchylis flexilisana Walker, 1863 (Borneo)

Stictosia flava (van Eecke, 1927) (Figs. 19, 78)

Siccia flava van Eecke, 1927; Zool. Meded. 10: 94. Stictosia flava van Eecke: Holloway, 2001, Moths of Borneo 7: 446, pl. 8, f. 436, 438. Stictosia flava van Eecke: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 73, Pl. 13: 179, 179a, gen. Mal108.

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Wingspan. Male 12 mm. Type locality. Sumatra – Fort de Kock. Distribution in Laos. Nakai. Geographical range. Laos, Peninsular Malaysia, Sumatra, Borneo.

Stictosia maculata Bucsek n. sp. (Figs. 20, 21, 79, 80)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.-13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 952. Paratypes. 2 ♂♂, 1 ♀ like holotype, CKB.

Wingspan. Male 13–14 mm, female 14 mm. Holotype. Male 14 mm, antennae filiform, head ochreous, thorax cream- coloured, abdomen brown, anal margin creamy. Forewing ochreous, discal spot black, medial and postmedial lines bent and running in parallel around the discal spot. Anal margin of the forewing with large dark brown spot, the edges of the spot are not sharply delimited. Hindwing cream. Male genitalia. Uncus folded, conical, pointed. Tuba analis pronounced. Valva along the entire length nearly equally wide, apical lobe of valva prominent, distal saccular process is pointed and bent at right angle inside the valva, medial costal process of irregular shape. Aedeagus with one thin and long cornut in a field of very small cornuti. Female. Habitus similar to the male, but hinwing with brown dust. Female genitalia. Ductus bursae with two fields of cornuti, the lower field partially located in the appendix bursae, corpus bursae with semi-circular row of larger cornuti, near to it, from the outside a distinctive signum.

Genus Tospitis Walker, 1863 Type species: Tospitis nulliferana Walker, 1863 (Borneo)

Tospitis ochreus Bucsek, 2014 (Figs. 22, 81)

Tospitis ochreus Bucsek, 2014, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, Suppl., 21, Pl. 3: 41, 41a, 41b, gen. MalS34.

Wingspan. Female 11 mm. Type locality. Malaysia – Perak. Distribution in Laos. Nakai. Geographical range. Laos, Peninsular Malaysia. 26 Entomofauna carpathica, 2020, 32(2): 19-46

Tospitis ogriagra Bucsek n. sp. (Figs. 23, 82)

Holotype. ♂ LAOS, Khammouane prov., Nakai env., 500 m, 17°34´N, 105°10´E, 25.–29. IV. 2019, leg. K. Bucsek, SNM with a museum serial number SZ 10 953.

Wingspan. Male 10 mm. Holotype. Antennae filiform, head, thorax and abdomen creamy. Forewing light brown, antemedial and postmedial area ochreous. Antemedial area near costa a little darker. Hindwing light brown. Male genitalia. Uncus long, slightly curved, with curved tip. Valva more or less equally wide, distal costal process curved, pointed. Saccular process slightly bent, ending with a blunt tip. Aedeagus conical, with one small field of medium- sized cornuti. Female. Unknown.

Tospitis denisae Bucsek n. sp. (Figs. 24–29, 83, 84)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.–13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 954. Paratypes. 14 ♂♂, 8 ♀♀ like holotype, CKB, 1 ♂, 1 ♀ SNM.

Wingspan. Male 9–10 mm, female 9–10 mm. Holotype. Male 10 mm, antennae filiform, head creamy, thorax and abdomen ochreous. Forewing ochreous, antemedial line brown, slightly pronounced, thickened on the costa. Costal edges of the medial and postmedial areas form dark brown spots. Apex with a small whitish spot. Cilia dark ochreous. Hindwing whitish, apex with light brown dusting. Male genitalia. Uncus long, slightly curved, pointed. Valva with strong pubescence, saccular process long, narrow, slightly bent at the apex. Distal costal process significantly separated from valva, slightly bent, conical with a sharp tip. Aedeagus conical with one large cornut. Female. The same as a male, slightly darker and with a more pronounced patterning. Female genitalia. Ductus bursae very long, in the corpus bursae one horizontal and two vertical bands of cornuti. Etymology. I named the species after Mrs. Denisa Hergovitsová, the wife of my esteemed friend Mr. Roman Hergovits, Bratislava.

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Genus Tortricosia Hampson, 1900 Type species: Tortricosia excisa Hampson, 1900 (Myanmar=Burma)

Tortricosia classeyi Holloway 2001 (Fig. 30)

Tortricosia classeyi Holloway, 2001, Moths of Borneo 7: 407– 408, pl. 6, fig. 335. Tortricosia classeyi Holloway: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 119, pl. 24: 236. Tortricosia classeyi Holloway: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 77–78, Pl. 13: 191, 191a. Tortricosia classeyi Holloway: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 154, Pl. 14: 109.

Wingspan. Female 13 mm. Type locality. Borneo – Sabah. Distribution in Laos. Huaphanh. Geographical range. Laos, Cambodia, Thailand, Peninsular Malaysia, Borneo.

Genus Oxacme Hampson, 1894 Type species: Oxacme dissimilis Hampson, 1894 (Sikkim)

Oxacme dissimilis Hampson, 1894 (Figs. 31–33)

Oxacme dissimilis Hampson, 1894, Moths Ind. 2: 94, fig. 209. Oxacme dissimilis Hampson: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 292, fig. 209. Oxacme dissimilis Hampson: Draudt, 1914: in Seitz, Macrolep. World 10: 176, pl. 16f. Oxacme dissimilis Hampson: Fang, 1982, Icon. Het. Sin. 2: 194, fig. 1395. Oxacme dissimilis Hampson: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 179, fig. 104, pl. 5: 15. Oxacme dissimilis Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 116, pl. 24: 232. Oxacme dissimilis Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 81, Pl. 14: 202, 202a, gen. Mal114. Oxacme dissimilis Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 156, Pl. 14: 111.

Wingspan. Male 14–15 mm, female 15 mm. Type locality. India – Sikkim. Distribution in Laos. Huaphanh. Geographical range. India (Sikkim, Khasis), China, Laos, Thailand, Cambodia, Peninsular Malaysia.

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Oxacme marginata Hampson, 1896 (Figs. 34–37, 85–87)

Oxacme marginata Hampson, 1896, Moths Ind. 4: 496. Oxacme marginata Hampson: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 292, pl. 26: 27. Oxacme marginata Hampson: Draudt, 1914: in Seitz, Macrolep. World 10: 176, pl. 16f. Oxacme marginata Hampson: Fang, 1982, Icon. Het. Sin. 2: 194, fig. 1394. Oxacme marginata Hampson: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 179, pl. 5: 14. Oxacme marginata Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 117, pl. 24: 233. Oxacme marginata Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 80, Pl. 14: 201, 201a. Oxacme marginata Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 155, Pl. 14: 110.

Wingspan. Male 15–16 mm, female 15–16 mm. Type locality. India – Meghalaya. Distribution in Laos. Huaphanh. Geographical range. India (Meghalaya, Khasis), Myanmar, China, Laos, Thailand, Cambodia, Peninsular Malaysia. Note. In Laos, there are specimens with conspicuous black dusted forewing.

Genus Orieosia Bucsek, 2012 Type species: Pasteosia orientalis Hampson, 1909 (Singapore)

Orieosia orientalis (Hampson, 1909) (Figs. 38, 88)

Pasteosia orientalis Hampson, 1909, A. M. N. H. (8) iv. p. 354. Pasteosia orientalis Hampson: Hampson 1914, Cat. Lep. Phal. Br. Mus., Suppl. Vol. 1: 539, f. 136. Orieosia orientalis Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 83 - 84, Pl. 14: 208, 208a, gen. Mal117. Orieosia orientalis Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 157, Pl. 14: 112.

Wingspan. Male 16 mm. Type locality. Singapore. Distribution in Laos. Nakai. Geographical range. Singapore, Peninsular Malaysia, Cambodia, Laos.

Genus Padenia Moore, 1882 Type species Cyllene transversa Walker, 1854 (Ceylon = Sri Lanka)

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Padenia acutifascia de Joanis, 1928 (Fig. 39)

Padenia acutifascia de Joanis, 1928, Ann. Soc. Ent. Fr. 97: 260–261. Padenia acutifascia de Joanis: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 185, pl. 5: 23. Padenia acutifascia de Joanis: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 109–110, pl. 24: 219a, 219b. Padenia acutifascia de Joanis: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 161, Pl. 14: 116.

Wingspan. Male 17 mm. Type locality. Vietnam. Distribution in Laos. Huaphanh. Geographical range. Vietnam, China, Laos, Thailand, Cambodia.

Genus Garudinistis Hampson, 1900 Type species: Tospitis eburneana Walker, 1863 (Borneo)

Garudinistis cumcuneusa Bucsek, 2012 (Figs. 40, 41)

“Garudinistis” cumcuneusa Bucsek, 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 93, Pl. 16: 231, gen. Mal135. Garudinistis cumcuneusa Bucsek: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 166, Pl. 15: 121.

Wingspan. Male 12 mm, female 14 mm. Type locality. Malaysia – Pahang. Distribution in Laos. Huaphanh. Geographical range. Laos, Cambodia, Peninsular Malaysia.

Genus Stictane Hampson, 1900 Type species: Pitane fractilinea Snellen, 1880 (Sumatra)

Stictane bayarsaikhanae Bucsek n. sp. (Figs. 42–46, 89–92)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.–13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 955. Paratypes. 3 ♂♂, 4 ♀♀ like holotype, CKB.

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Wingspan. Male 13–14 mm, female 12–14 mm. Holotype. Male 14 mm, antennae finely pectinated, head and tegulae brown, thorax dark brown, abdomen brown, anal margin creamy. Forewing ochreous with black brown patterning, subbasal area dark brown, antemedial area ochreous. Medial area dark brown, clearly delimited, with a significant emargination in the middle of the subbasal side. Discal double-spot black, the upper one is smaller. Postmedial line intermittent, arched, subterminal area bear the apex black brown, cilia gray. Hindwing light gray-brown. Male genitalia. Uncus long and thin, slightly curved. Valva has in the middle of costal margin a pronounced protrusion, the lobe of valva is more or less equaly wide, markedly hairy, rounded at the top. Distal saccular extension forms a sharp apex-facing point. Sacculus pronounced, saccular process S-shaped and pointed. Apical lobe of juxta heart-shaped, juxta distinctive, slightly sclerotized, with two rounded tips, vinculum V-shaped. Aedeagus conical, with one large cornut and a bunch of smaller cornuti. Female. Like a male, a little darker with a more pronounced patterning. Female genitalia. V-shaped antrum, ductus bursae more or less cylindrical, corpus bursae of irregular spherical shape with a band of distinctive spines, between them is placed a large smooth sclerotized plate, around which is an array of spinules. Appendix bursae large, of tubular shape, at the mouth with a field of distinctive thorns. Similar species. Stictane obscura (Inoue) – has a shallower depression on the inside of the medial area. The apical lobe of valva has a conical shape, distal saccular extension is longer and more outwardly curved, with smaller apical saccular extension. Juxta is less pronounced. The female without a field of thorns in the mouth of appendix bursae. Etymology. The new species is named in honour of Dr. Ulziijargal Bayarsaikhan (Incheon, Republic of Korea), excellent expert on Arctiinae of Oriental region.

Stictane lao Bucsek, n. sp. (Figs. 47–49, 93–95)

Holotype. ♂ LAOS, Huaphanne prov., Mt. Phu Pane, 1200–1900 m, Ban Saluei v. env., 3.–13.V.2019, 20°12´N, 103°59´E, leg. K. Bucsek, SNM with a museum serial number SZ 10 956. Paratypes. 1 ♂, 2 ♀♀ like holotype, CKB.

Wingspan. Male 13–14 mm, female 12–13 mm. Holotype. Male 14 mm, antennae finely pectinated, head and thorax creamy, abdomen brown, anal margim creamy. Forewing light ochreous with black- brown patterning, three dots on the outer edge of the subbbasal area, antemedial area light ochreous. Medial area dark brown, clearly demarcated and straight on the inside, scattered on the outside. Discal double dot black, the 31 Entomofauna carpathica, 2020, 32(2): 19-46 upper one is smaller. Postmedial line formed by dots, arched, subterminal area at apex black brown, cilia creamy. Hindwing whitish. Male genitalia. Uncus long and thin, slightly curved. Valva from the middle to the top with strong pubescence. Costal margin with slight blunt protrusion in the middle, lobe of valva evenly wide along its entire length, on apex bent (wider than in Stictane bayarsaikhanae). Distal saccular extension forms a sharp tip bent at a 45° angle, above which is a smaller apical saccular extension. Sacculus pronounced, saccular process bent in a shape of S and pointed (it is shorter than in Stictane bayarsaikhanae). Apical lobe of juxta heart-shaped, juxta is rounded at the top. Aedeagus conical, with one large cornut and a bunch of smaller cornuti. Female. Like a male, a little darker and with a more pronounced patterning. Female genitalia. Ductus bursae broad, with a field of spinules and with a single row of cornuti extending into corpus bursae. Corpus bursae egg-shaped with a band of distinctive U-shaped cornuti and heavily filled by spinules. Appendix bursae large and round, irregular in shape. Similar species. Stictane obscura (Inoue) – has a shallower depression on the inside of the medial area, valva with conical apex, apical lobe of juxta is narrower, and vinculum is wider. The female with long cylindrical appendix bursae and without spinules in ductus bursae.

Stictane munda (de Joanis, 1928) (Figs. 50–52, 96, 97)

Manoba munda de Joanis, 1928, Ann. Soc. Ent. Fr. 97: 258, pl. 2, f. 1. Stictane munda de Joanis: Bayarsaikhan, Bae, 2015, Zootaxa 3981 (2): 246. Stictane munda de Joanis: Bayarsaikhan, Bae, 2019, Zootaxa 4686 (2): 256, f. 12. Stictane munda de Joanis: Bayarsaikhan, Bae, 2021, Journ. Asia-Pacific Biodiv., s. 3, figs. 1A-C, 3A-B, 4B, in press.

Wingspan. Male 10 mm, female 10 mm. Type locality. Vietnam. Distribution in Laos. Nakai. Geographical range. Vietnam, Laos. Note. Individuals from Laos are smaller than ones from Vietnam, medial area on subbasal side is nearly straight, without emargination.

Stictane pectenicorniculum Bucsek, 2012 (Fig. 53)

Stictane pectenicorniculum Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 83 - 84, Pl. 14: 208, 208a, gen. Mal117. 32 Entomofauna carpathica, 2020, 32(2): 19-46

Stictane pectenicorniculum Bucsek: Bayarsaikhan, Bae, 2015, Zootaxa 3981 (2): 245, Figs. 6, 13. Stictane pectenicorniculum Bucsek: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 052, Pl. 1: 7. Stictane pectenicorniculum Bucsek: Bayarsaikhan et al., 2019, Zootaxa 4686 (2): 254 (list).

Wingspan. Male 13 mm. Type locality. Malaysia – Pahang. Distribution in Laos. Nakai. Geographical range. Peninsular Malaysia, Thailand, Cambodia, Laos.

Genus Cyana Walker, 1854 Type species: Cyana detrita Walker, 1854 (Bangladesh)

Cyana bellissima (Moore, 1878) (Fig. 54)

Bizone bellissima Moore, 1878, Proc.Zool. Soc. Lond. 1878: 27, pl. 3: 13. Cyana bellissima Moore: Hampson, 1894, Moths Ind. 2: 61. Chionaema bellissima Moore: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 318, 319-320, fig. 229. Chionaema bellissima Moore: Seitz, 1910, Macrolep. World 2: 62: pl. 12 b. Cyana bellissima Moore: Fang, 1992, Sinozoologia 9: 259. Cyana bellissima Moore: Kishida, 1993, Tinea 13 (Suppl. 3): 38, pl. 41: 1, 2. Cyana bellissima Moore: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 171–172, pl. 5: 3. Cyana bellissima Moore: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 56, pl. 12: 112a, 112b. Cyana bellissima Moore: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 75. Cyana bellissima Moore: Singh et al., 2020, Zootaxa 4738 (1): 44–45, Figs 118, 119, 214–218, 275.

Wingspan. Male 44 mm. Type locality. NW Himalaya. Distribution in Laos. Huaphanh. Geographical range. Tibet, India, Myanmar, Nepal, China, Vietnam, Laos, Thailand.

Cyana dudgeoni Hampson, 1895 (Figs. 55, 56)

Cyana dudgeoni Hampson, 1895, Trans. Ent. Soc.: 293. Chionaema dudgeoni Hampson: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 321, pl. 27: 5. Chionaema dudgeoni Hampson: Draudt, 1914: in Seitz, Macrolep. World 10: 174, pl. 16 l. Cyana dudgeoni Hampson: Tams, 1924, Journ. Nat. Hist. Soc. Siam VI (3): 229-234. Cyana dudgeoni Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 44, pl. 8: 89a, 89b. Cyana dudgeoni Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 103, Pl. 17: 254, 254a. 33 Entomofauna carpathica, 2020, 32(2): 19-46

Cyana dudgeoni Hampson: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 74. Cyana dudgeoni Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 183, Pl. 17: 138. Cyana dudgeoni Hampson: Bayarsaikhan, Bae, 2016, Zootaxa 4114 (4): 456-457, Pl. 2: 15a, 15b; 4: 15; 5: 15. Cyana dudgeoni Hampson: Singh et al., 2020, Zootaxa 4738 (1): 18 - 19, Figs 39–41, 178, 251.

Wingspan. Male 19–20 mm, female 25 mm. Type locality. India – Sikkim. Distribution in Laos. Nakai. Geographical range. NE India, Nepal, China, Vietnam, Laos, Thailand, Cambodia, Peninsular Malaysia.

Cyana obliquilineata (Hampson, 1900) (Fig. 57)

Chionaema obliquilineata Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 299, pl. 26: 24 Chionaema obliquilineata Hampson: Draudt, 1914: in Seitz, Macrolep. World 10: 169, pl. 16g. Chionaema obliquilineata Hampson: Kaleka, 2002, Zoos′ Print J. 17 (10): 894. Chionaema obliquilineata Hampson: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 44, pl. 8: 88. Chionaema obliquilineata Hampson: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 76. Chionaema obliquilineata Hampson: Bayarsaikhan et al. 2016, Biod. of Cambodia: tiger moths, 181, Pl. 17: 136. Chionaema obliquilineata Hampson: Bayarsaikhan, Bae, 2016, Zootaxa 4114 (4): 455-456, Pl. 2: 13; 4: 13. Chionaema obliquilineata Hampson: Singh et al., 2020, Zootaxa 4738 (1): 15–16, Figs 33–36, 176, 249.

Wingspan. Male 23 mm. Type locality. India – Sikkim. Distribution in Laos. Nakai. Geographical range. NE India, Nepal, China, Laos, Thailand, Cambodia.

Cyana quadrinotata (Walker, 1869) (Fig. 58)

Bizone quadrinotata Walker, 1869, Cat. Lep. Het.: 90. Bizone pallens Butler, 1877, Trans. Ent. Soc. Lond. 1877: 338 Cyana alborosea Hampson, 1896, Fauna Br. India (Moths) 4: 492. Chionaema alborosea quadrinotata Walker: Draudt, 1914: in Seitz, Macrolep. World 10: 168. Cyana quadrinotata Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 45, pl. 8: 91a, 91b. Cyana alborosea Hampson: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 103, Pl. 17: 254, 254a. Cyana quadrinotata Walker: Kirti, Singh, 2015, Arctiid Moths of India, vol. 1: 76. Cyana quadrinotata Walker: Singh et al. 2020, Zootaxa 4738 (1): 24– 25, Figs 58–62, 188, 189, 256. 34 Entomofauna carpathica, 2020, 32(2): 19-46

Wingspan. Male 20 mm. Type locality. Not stated. Distribution in Laos. Huaphanh. Geographical range. NE India, Nepal, Bhutan, Bnagladesh, Myanmar, S China, Laos, Vietnam, Thailand, Cambodia, Malay Peninsula, Java.

Cyana centripuncta Černy, 2016 (Fig. 59)

Cyana centripuncta Černy, 2016, Entomofauna 37 (26): 425, pl. 1: 11–15. Cyana centripuncta Černy: Bayarsaikhan, Bae, 2016, Zootaxa 4114 (4): Pl. 2: 16a, 16b; 4: 16; 5: 16.

Wingspan. Male 18 mm. Type locality. S Vietnam. Distribution in Laos. Nakai. Geographical range. Laos, Vietnam, Cambodia.

Cyana interrogationis (Poujade, 1886) (Fig. 60)

Bizone interrogationis Poujade, 1886, Bull. Soc. Ent. France (6) 6: 125. Chionaema interrogationis Poujade: Hampson, 1900, Cat. Lep. Ph. Br. Mus. 2: 320, pl. 27, f. 4. Chionaema interrogationis Poujade: Seitz, 1910, Macrolep. World 2: 62, pl. 12 c. Chionaema interrogationis Poujade: Daniel, 1953, Bonn. zool. Beitr. 3 (3–4): 310. Chionaema interrogationis Poujade: Fang, 1982, Icon. Het. Sin. 2: 204, fig. 1502. Cyana interrogationis Poujade: Fang, 1992, Sinozoologia 9: 259. Cyana interrogationis Poujade: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 172–173, pl. 5: 4. Chionaema interrogationis Poujade: Dubatolov, Kishida & Wang, 2012, Tinea 22 (1): 47, f. 37.

Wingspan. Male 28 mm. Type locality. China. Distribution in Laos. Huaphanh. Geographical range. China, Laos, Vietnam, Thailand.

Cyana costifimbria (Walker, 1862) (Fig. 61)

Bizone costifimbria Walker, 1862, Journ. Lin. Soc. Zool. 6: 121. Chionaema costifimbria Walker: Hampson, 1900, Cat. Lep. Phal. Br. Mus. 2: 307, fig. 223. Chionaema rhadota Swinhoe, 1907, Ann. Mag. Nat. Hist. (7), 20: 76. Chionaema costifimbria Walker: Draudt, 1914: in Seitz, Macrolep. World 10: 170. 35 Entomofauna carpathica, 2020, 32(2): 19-46

Chionaema javanica ab. niasensis Strand, 1922, Lep. Cat. 26: 685. Chionaema rhadota borneensis Talbot, 1926, Sarawak Mus. J., 3 (2): 135. Cyana pectinata Roepke, 1946, Tijd. Ent. 87: 31. Doliche costifimbria Walker: Černý, 1993, Nacht. Ent. Ver. Apollo, Suppl. 12: 47. Cyana costifimbria Walker: Fang, 2000, Fauna Sin., Ins. 19, Lep., Arctiidae: 163, pl. 4: 17. Cyana costifimbria Walker: Holloway, 2001, Moths of Borneo 7: 336, pl. 1, figs 131. Cyana costifimbria Walker: Cerny, Pinratana 2009, Moths of Thailand, vol. 6: 47, pl. 9: 95a, 95b. Cyana costifimbria Walker: Lourens, 2011, Nacht. Ent. Ver. Apollo, NF 32 (1/2): 84, 95 (list). Cyana costifimbria Walker: Bucsek 2012, Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula-Malaysia, 104–105, Pl. 18: 259, 259a, 259b. Cyana costifimbria Walker: Bayarsaikhan, Bae, 2016, Zootaxa 4114 (4): 451, Pl. 1: 3a, 3b; 3: 3; 5: 3.

Wingspan. Male 26 mm. Type locality. Borneo – Sarawak. Distribution in Laos. Nakai. Geographical range. Palawan, Tawi Tawi, Borneo, Sumatra, Java, Nias, Peninsular Malaysia, Cambodia, Thailand, Vietnam, Laos, China.

Genus Nephelomilta Hampson, 1900 Type species: Lyclene suffusa Hampson, 1891 (Nilgiris)

Nephelomilta sumatrana effractoida Volynkin & Černý, 2018 (Fig. 62)

Nephelomilta sumatrana effractoida Volynkin & Černý, 2018, Zootaxa 4472 (3): 408– 409, Figs.13–16, 89, 90, 129, 130.

Wingspan. Male 23 mm. Type locality. India – Darjeeling. Distribution in Laos. Nakai. Geographical range. North and North-East India, Thailand, Laos, Vietnam.

ACKNOWLEDGEMENTS

Thanks to Mr. Henrich Novák, Bratislava for translating the manuscript into English and Dr. Ladislav Roller, Slovak Academy of Sciences, Bratislava for English correction. I also express my thanks to Dr. Ľubomír Vidlička, Slovak Academy of Sciences, Bratislava for technical help during editing of the manuscript.

36 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 1-17. Adults. 1. Aemene coniuncta, ♂. 2. Aemene eudonioides, ♂. 3-4. Aemene cinerea n. sp., 3. ♂, HT, 4. ♀, PT. 5. Aemene bicornuata n. sp., ♂, HT. 6-7. Aemene lancea n. sp., 6. ♂, HT, 7. ♂, PT. 8. Aemene spinivalva n. sp., ♂, HT. 9-10. Volynkinia inaequalis n. sp., 9. ♂, HT, 10. ♀, PT. 11-13. Halone ivetae n. sp., 11. ♂, HT, 12-13. ♀♀, PT. 14. Neoduma songensis, ♂. 15. Neoduma ectozona, ♂. 16-17. Neoduma kuangtungensis, 16. ♂, 17. ♀. 37 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 18-37. Adults. 18. Eucyclopera plagidisca, ♂. 19. Stictosia flava, ♂. 20-21. Stictosia maculata n. sp., 20. ♂, HT, 21. ♀, PT. 22. Tospitis ochreus, ♀. 23. Tospitis ogriagra n. sp., ♂, HT. 24-29. Tospitis denisae n. sp., 24. ♂, HT, 25. ♂, PT, 26-29. ♀♀, PT. 30. Tortricosia classeyi, ♀. 31-33. Oxacme dissimilis, 31-32. ♂♂, 33. ♀. 34-37. Oxacme marginata, 34. ♂, 35. ♀, 36. ♂, dark form, 37. ♀, dark form. 38 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 38-53. Adults. 38. Orieosia orientalis, ♂. 39. Padenia acutifascia, ♂. 40-41. Garudinistis cumcuneusa, 40. ♂, 41. ♀. 42-46. Stictane bayarsaikhanae n. sp., 42. ♂, HT, 43. ♂, PT, 44-46. ♀♀, PT. 47-49. Stictane lao, n. sp., 47. ♂, HT, 48. ♂, PT, 49. ♀, PT. 50-52. Stictane munda, 50-51. ♂♂, 52. ♀. 53. Stictane pectenicorniculum, ♂. 39 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 54-62. Adults. 54. Cyana bellissima, ♂. 55-56. Cyana dudgeoni, 55. ♂, 56. ♀. 57. Cyana obliquilineata, ♂. 58. Cyana quadrinotata, ♂. 59. Cyana centripuncta, ♂. 60. Cyana interrogationis, ♂. 61. Cyana costifimbria, ♂. 62. Nephelomilta sumatrana effractoida, ♂. 40 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 63-70. Genitalia. 63. Aemene coniuncta, ♂. 64. Aemene eudonioides, ♂. 65. Aemene cinerea n. sp., ♂, HT. 66. Aemene bicornuata n. sp., ♂, HT. 67-68. Aemene lancea n. sp., 67. ♂, HT, 68. ♂, PT. 69. Aemene spinivalva n. sp., ♂, HT. 70. Volynkinia inaequalis n. sp., ♂, HT. 41 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 71-79. Genitalia. 71. Volynkinia inaequalis n. sp., ♀, PT. 72-73. Halone ivetae n. sp., 72. ♂, HT, 73. ♀, PT. 74. Neoduma songensis, ♂. 75. Neoduma ectozona, ♂. 76-77. Neoduma kuangtungensis, 76. ♂, 77. ♀. 78. Stictosia flava, ♂. 79. Stictosia maculata n. sp., ♂, HT. 42 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 80-88. Genitalia. 80. Stictosia maculata n. sp., ♀, PT. 81. Tospitis ochreus, ♀. 82. Tospitis ogriagra n. sp., ♂, HT. 83-84. Tospitis denisae n. sp., 83. ♂, HT, 84. ♀, PT. 85-87. Oxacme marginata, 85. ♂, 86. ♀, 87. ♂, dark form. 88. Orieosia orientalis, ♂. 43 Entomofauna carpathica, 2020, 32(2): 19-46

Figures 89-97. Genitalia. 89-92. Stictane bayarsaikhanae n. sp., 89. ♂, HT, 90. ♂, PT, 91-92. ♀♀, PT. 93-95. Stictane lao n. sp., 93. ♂, HT, 94. ♂, PT, 95. ♀, PT. 96-97. Stictane munda, 96. ♂, 97. ♀. 44 Entomofauna carpathica, 2020, 32(2): 19-46

REFERENCES

BAYARSAIKHAN, U. & BAE, Y. S. 2015. Three new species of Stictane Hampson, 1900 (Erebidae, Arctiinae) from Cambodia, with Checklist of Stictane, Zootaxa 3981 (2): 241–252. BAYARSAIKHAN, U., & BAE, Y. S. 2016. A review of the genus Cyana Walker, 1854 (, Erebidae, Arctiinae) from Cambodia, with description of new species. Zootaxa 4114 (4): 447–463. BAYARSAIKHAN, U. & BAE, Y.S. 2021. The genus Stictane Hampson, 1900 (Lepidoptera: Erebidae: Arctiinae) in Vietnam. Journal of Asia-Pacific Biodiversity, in press. BAYARSAIKHAN, U., BAE, Y.S. & KIMSUN, CH. 2016. Biodiversity of Cambodia: tiger moths (Insecta, Lepidoptera). National Institute of Biological Resources, Incheon, 303 pp. BUCSEK, K. 2012. Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula – Malaysia. Institute of Zoology, Bratislava, 170 pp+51 pls. BUCSEK, K. 2014. Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula – Malaysia, Supplementum. Institute of Zoology, Bratislava, 170 pp+51 pls. CERNY, K. & PINRATANA, A. 2009. Moths of Thailand. Vol. 6. Brothers of Saint Gabriel in Thailand, Thailand, 227 pp + 52 pls. ČERNÝ K. 2016. A Contribution to the Knowledge of the Genus Cyana Walker in the South East Asia (LEPIDOPTERA: Erebidae, Arctiinae, Lithosiinae). Entomofauna 37 (26): 421–436. DANIEL, F. 1954. Beiträge zur Kenntnis der Arctiidae Ostasiens unter besonderer Berücksichtigung der Ausbeuten von Dr. h. c. H. Höne aus diesem Gebiet (Lep. Het.). III. Teil: Lithosiinae. Bonner zoologische Beiträge 5 (1-2): 89–138, Taf. III. DUBATOLOV, V.V. & BUCSEK, K. 2013. New species of lichen˗moths from South˗East Asia (Lepidoptera, , Lithosiini). Tinea 22 (4): 279–291. DUBATOLOV, V.V. & BUCSEK K. 2016. New lichen-moth taxa (Lepidoptera, Arctiidae, Lithosiinae) from Vietnam. Euroasian Entomological Journal [Евразиатский энтомологический журнал] 15 (3): 228–238. DUBATOLOV, V.V., KISHIDA, Y. & WANG, M. 2012. New records of lichen-moth from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae). Tinea 22 (1): 25–52. EECKE VAN, R. 1930. De Heterocera van Sumatra. Zoologische Mededeelingen 1925–1929; republished by E. J. Brill in one volume 1930, pp. 67–218. FANG, C. 2000. Lepidoptera. Arctiidae. Fauna Sinica (Insecta) 19. Science Press, Beijing, 590 pp. (In Chinese). HAMPSON, G.F. 1891. Illustrations of typical specimens of Lepidoptera Heterocera in the collection of the British Museum, 8. The Lepidoptera Heterocera of the Nilgiri District. British Museum (Natural History), London, 144 pp + 18 pls.

45 Entomofauna carpathica, 2020, 32(2): 19-46

HAMPSON, G.F. 1898. The moths of India. Supplementary paper to the volumes in „The fauna of British India“. Journal of the Bombay Natural History Society 11: 438–462. HAMPSON, G. F. 1900. Catalogue of the Lepidoptera Phalaenae in the British Museum, 2: Catalogue of the Arctiadae (Nolinae, Lithosianae) in thecollection of the British Museum. Taylor and Francis Ltd., London, 590 pp. HOLLOWAY, J.D. 2001. The Moths of Borneo, Vol. 7. Malayan Nature Society, Kuala Lumpur, 207 pp. JOANNIS, J. de, 1928. Lepidopteres Heteroceres du Tonkin. Annales de la Société entomologique de France 97: 241-368, pl. 1–2. MOORE, F. 1878. A revision of certain genera of European and Asiatic Lithosiidae, with characters of new genera and species. Proceedings of the Zoological Society of London 1878: 3–37. SEITZ, A. 1914. 2. Subfamilie: Lithosiinae, Flechtenbärchen. In: SEITZ, A. (Ed.) Die Gros- Schmetterlinge der Erde. 10. Spinner und Schwarmer des indoautralischen Gebiets. Fritz Lehmann Verlag, Stuttgart, pp. 118–223. SINGH, N. & KIRTI, J.S. 2015. Arctiid Moths of India, Vol. 1. Nature Books India, New Delhi, 205 pp. SINGH, N. & KIRTI, J.S. 2016. Arctiid Moths of India, Vol. 2. Nature Books India, New Delhi, 214 pp. SINGH, N., VOLYNKIN, A.V., KIRTI, J.S., DATTA, H.S. & IVANOVA M.S. 2020. A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa 4738 (1): 1–93. STRAND, E. 1922. Arctiidae: Subfam. Lithosiinae. In: WAGNER, H. (Ed.) Lepidopterorum Catalogus 15-16 (26): 499–899, Junk, Berlin & The Hague. VOLYNKIN, A.V., ČERNÝ, K. & IVANOVA, M. 2017. Description of a new Aemene Walker, 1854 from North Thailand and Myanmar, with a checklist of the genus (Lepidoptera, Erebidae, Arctiinae). Zootaxa 4306 (3): 444-450. VOLYNKIN A.V., ČERNÝ K. 2018. Revision of the genus Nephelomilta Hampson, 1900, with descriptions of twelve new species and two new subspecies (Lepidoptera, Erebidae, Arctiinae). Zootaxa 4472 (3): 401–451. WALKER, F. 1858 [1857]. Catalogue of Lepidoptera Heterocera. Third Series. List of the Specimens of Lepidopterous in the Collection of the British Museum, Part 12. – Noctuidae: 765-982. WALKER, F. 1862. Catalogue of the Heterocerous Lepidopterous Insects collected at Sarawak, in Borneo, by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the Procedings of the Linnean Society (Zoology) 6: 82–145, 171–198.

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