Eur. J. Entomol. 111(2): 299–302, 2014 doi: 10.14411/eje.2014.034 ISSN 1210-5759 (print), 1802-8829 (online) First fossil tooth-necked fungus beetle (Coleoptera: Derodontidae): Juropeltastica sinica gen. n. sp. n. from the Middle Jurassic of China CHENYANG CAI 1, 2, John F. LawrENCE 3, AdAm Ślipiński 3 and diying HUANG 1* 1 state key laboratory of palaeobiology and stratigraphy, Nanjing institute of Geology and palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China; e-mails:
[email protected],
[email protected] 2 Graduate School, University of Chinese Academy of Sciences, 19A Yuquanlu, Beijing 100049, China 3Australian National Insect Collection, CSIRO Ecosystem Sciences, GPO Box 1700, Canberra ACT 2601, Australia; e-mails:
[email protected],
[email protected] Key words. Coleoptera, Derodontidae, Juropeltastica gen. n., fossil, Daohugou beds, Middle Jurassic, China Abstract. The first fossil tooth-necked fungus beetle, Juropeltastica sinica gen. n. sp. n., is described and illustrated based on a single impression fossil from the Middle Jurassic Daohugou beds (ca. 165 Ma) of northeastern China. it represents the first definitive fossil be- longing to the extant family Derodontidae. Juropeltastica is placed in Derodontidae based on its overall body shape and size, head with complex systems of tubercles and grooves, pronotum with dentate lateral carinae, open mesocoxal cavities bordered by mesepimeron and metanepisternum, excavate metacoxae, and 5-segmented abdomen. The occurrence of a reliable derodontid fossil from 165 mil- lion years ago places Derodontidae among the small but growing number of beetle families of known Middle Jurassic age, which is important in the dating of phylogenetic trees.