Philippine Carahidae (Col.) Collected by the Noona Dan Expedition
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Entomologiske Meddelelser 35 (1967) (Noona Dan Papers No. 47.) Philippine Carahidae (Col.) collected by the Noona Dan Expedition. By C. J. Louwerens (Hilversum). I have to thank Dr Borge Petersen (Universitetets Zoologiske Museum, Kobenhavn), who was kind enough as to submit to me for study the Carabidae collected by the Danish Noona Dan Ex pedition to the Philippines and the Bismarck Islands in 1961 and 1962 (Petersen 1966). I am much obliged too to Prof. P. J. Darlington Jr (Cambridge, U.S.A.), Ing. Jedlicka Arnost (Praha), Mr. R. D. Pope (British Museum, London), Dr Shun-ichi Ueno (Tokyo) and Prof. S. L. Straneo (Milano) for their kind help and advice, which they all readily gave me, when asked for. This paper contains the descriptions of two new species from the Philippines and an enumeration of all Philippine species col lected (except two species described by Straneo 1967), with a few notes as to geographical distribution, etc. A second paper will deal with the Carabidae brought home from the Bismarck Islands. The arrangement of the species examined is alphabetical. It is to be understood, that all specimens are collected in the second half of 1961, most of them caught by Mercury light or in Malaise traps. Subfam. Scaritinae. Clivina vulgivaga Boh. (Eugenie's Resa, Zool. Inst. IV. Col. 1861, p. 9). BALABAC: Dalawan Bay, 1 sp., 8. X. The small species is, so far I know, restricted to the Philippines. Enl. Medd. 35 196 C. J. Louwerens Subfam. Bembidiinae. Armatocillenus yokohamae Bates (Tr. Ent. Soc. Lond. 1883, p. 268). PALAWAN: Brooke's Point, Uring Uring, 36 sp. 14. VIII. BALABAC: Dalawan Bay, 21 sp. 4. X and 2 sp. 5. X. Through the kindness of Dr Shun-ichi Ueno I was able to examine 4 typical specimens of yokohamae of Japan. The Phi lippine specimens (Fig. 1) are extremely like yokohamae in size Fig. 1. Armatocillenus yokohamae Bates, of Palawan, left side. as well in color, but most of them also bear an ill defined apical spot on the elytra. They differ only slightly in the shape of the pronotum. In yokohamae the pronotal anterior angles are more advanced and the sides of the pronotum a little more contracted behind. In my opinion the differences are of minor importance. Also in Japan. Tachys fumi.gatus Motch. (Bull. Soc. Imp. Nat. Mosc. 1851, p. 509). PALAWAN: Brooke's Point, Uring Uring, 1 sp. 14. VIII. - TA WI TA WI: Tarawakan, 1 sp. 15. XI; Lapid Lapid, 1 sp. 19. XL Readily recognizable by the deeply emarginate labrum and clypeus and by the long, deep frontal furrows. A variable species with an enormous distribution extending throughout South East Asia including the Malay Region and the Philippines. Also in Japan, Formosa, New Guinea and Africa. Tachys impressi.pennis Motch. (Et. Ent. 1859, p. 39). PALAWAN: Brooke's Point, Uring Uring, 31 sp. 14. VIII and 3 sp. 21. VIII. - TAWI TAWI: Tarawakan, 1 sp. 21. X, 5 sp. 11.-15. XL Common throughout South East Asia with the exception of Japan. Also in Australia. Entomologiske 1\,f eddelelser 35 (1967) 197 Tachys klugi Nietn. (Ann. Mag. Nat. Hist. 1859, p. 423). PALAWAN: Brooke's Point, Uring Uring, 2 sp. 14.VIII. BALABAC: Dalawan Bay, 1 sp. 4. X.- TAWI TAWI: Tarawa kan, 1 sp. 14. XI. Range: throughout South East Asia including Japan. Tachys klugi Nietn. var. sulcatopunctatus Putz. (Ann. Mus. Civ. Stor. Nat. Gen. 1875, p. 742). PALAWAN: Brooke's Point, Uring Uring, 1 sp. 14. VIII. BALABAC: Dalawan Bay, 1 sp. 4. X. Also in Sumatra, Java, Celebes and Andonare Isl. Differs from the typical form in the unspotted elytra. Tachys quadrillum Schaum (Berl. Ent. Zeit. 1860, p. 201). PALA WAN: Brooke's Point, Uring Uring, 135 sp. 14.-23. VIII. A common and variable species occuring from India, Ceylon and southern China to New Guinea. Tachys singularis Andr. (Ann. Mus. Civ. Stor. Nat. Gen. 1925, p. 393). PALAWAN: Brooke's Point, Uring Uring, 9 sp. 14. VIII. TAWI TAWI: Tarawakan, 1 sp. 14. XI. Also in Celebes and the Bismarck Archipelago (Noona Dan Expedition). This species is very much like yunnax Darl., originally describ ed from Dominican Republic by P. J. Darlington Jr (1939) and also occuring in New Guinea and the Aru Isl. (Darlington, 1962). Through the kindness of Prof. Darlington I was able to compare the specimens of the Philippines and the Bismarck Isl. with 2 paratypes of yunnax from Sanchez, Dominican Republic. After careful examination I could not find sufficient differences to re cognize two species. Nevertheless it is possible, that yunnax and singularis are different. According to Darlington (1962) yunnax is slightly larger than singularis, with more prominent eyes and relatively slightly wider prothoracic base. Tachys umbrosus Motch. (Bull. Soc. Imp. Nat. Mosc. 1851, p. 507). MINDANAO: Sapamoro, Curuan district, 3 sp. 20. XII. 13* 198 C. J. Louwerens Throughout South East Asia except Japan. In India it is con fined to the Hymalayans. Also in New Guinea and the Solomon Isl. Limnastis pilosus Bates (Ann. Mus. Civ. Stor. Nat. Gen. 1851, p. 296). TAWI TAWI: Tarawakan, 2 sp. 20. X, 10 sp. 13.-15. XI; Lapid Lapid, 3 sp. 19. and 21. XI. Common and widely distributed in India, Formosa, Tonkin, the Greater Sunda Isl., the Philippines, the Moluccas, New Guinea, eastern Australia and New Britain. Subfam. Trechinae. Perileptus sp. The expedition collected two specimens of the same species. According to Dr Shun-ichi Ueno, to whom I sent them for examin ation, they are very close to a new species that will be described in his forthcoming revision of the vVest Pacific species of Perilep tini, and may probably be the same species. Subfam. Pterostichinae. Besides the two new species, described by Prof. Straneo (1967), the following species are collected: Lesticus sp. near gregori Kuntz. (Ent. Rundsch. 1911, p. 175). PALAWAN: Mantalingajan, Tagembung, 1150 m, 1 sp. 17. IX. Identified by Prof. Straneo. L. gregori is only known from the Philippines. Trigonotoma palavanica Tchitch. (Hor. Soc. Ent. Ros. 1900, p. 184). BALABAC: Dalawan Bay, 6 sp. 5.-9. X. Confined to the Philippine Isl. Subfam. Anchomeninae. Euplenes viridis Andr. (Treubia 1933, p. 282). PALAWAN: Mantalingajan, Pinigisan, 600 m, 6 sp. 5.-9. IX. - BALABAC: Dalawan Bay, 5 sp. 7.-13. X. Also in Java, Celebes and Amboina Isl. In all specimens examined the elytra are bluish green with the Entomologiske M edde/elser 35 (1967) 199 underside and the legs much darker than in the typical form. Nevertheless I think they belong all to viridis. Dicranoncus quadriens Motch. (Et. Ent. 1859, p. 32). PALAWAN: Mantalingajan, Pinigisan, 600 m, 3 sp. 1.-9. IX. Commonly met with from Assam, India, Burma, Andaman Isl. to the Malay Peninsula and Archipelago. Also in Indo China. Colpodes obscuritarsis Chaud. (Ann. Soc. Ent. Fr. 1878, p. 375). PALAWAN: Brooke's Point, Uring Uring, 2 sp. 19. and 20. VIII. Also in Rangoon, Java, Sumatra and Borneo. Colpodes salsus J edl. (Sb. ent. odd. Nar. Mus. V Praze 1934, p. 190). TAWI TAWI: Tarawakan, 1 sp. 23. X. So far as I know only known from the Philippine Isl. Colpodes saphyrinus Chaud. (Ann. Soc. Ent. Fr. 1878, p. 366). PALAWAN: Mantalingajan, Pinigisan, 600 m, 1 sp. 17. IX. Also in Penang, Laos, Sumatra, Java, Borneo, Celebes, Ban guey and Tonda. The var. sloanei Maindron (Bull. Soc. Ent. Fr. 1908, p. 185) in Celebes, Amboina Isl. and New Guinea. (Darling ton, 1952). Colpodes sp. BALABAC: Dalawan Bay, 1 sp. 13. X. Notagonum (Colpodes) anceps Jedl. (Sb. ent. odd. Nar. Mus. V Praze 1934, p. 197). PALAWAN: Mantalingajan, Tagemburg, 1150 m, 1 sp. 17. IX. Only known from the Philippines. Notagonum (Colpodes) pereus Jedl. (Sb. ent. odd. Nar. Mus. V Praze 1934, p. 185). PALAWAN: Mantalingajan, Tagemburg, 1150 m, 2 sp. 19. and 20.X. Confined to the Philippines. Subfam. Harpalinae. Platymetopus flavilabris F. (Suppl. Ent. Syst. 1798, p. 59). PALAWAN: Brooke's Point, Uring Uring, 1 sp. 21. VIII. - BALABAC: Dalawan Bay, 1 sp. 4. X. 200 C. J. Louwerens A variable species. In the specimens examined the femora and tibiae are dark. In one or more of its diverse forms it ranges from China and Japan through India and Ceylon and through the whole of the Malay Archipelago. Oxycentropsis grandis Emd. (Arb. m. t. Ent. B. D. 1937, p. 116). PALA WAN: Brooke's Point, Uring Uring, 1 sp. 14. VIII. - BALABAC: Dalawan Bay, 2 sp. 5.-8. X. Also in Java. The 3 specimens of the Philippines are a little smaller than the Javan form, the basal striole longer, basal foveae and sides of pronotum more densely and more finely punctate, the punctures not very distinct. Stenolophus smaragdulus F. (Suppl. Ent. Syst. 1798, p. 60). P ALA WAN: Mantalingajan, Pinigisan, 600 m, 1 sp. 3. IX. A variable species both in size and the shape of the pronotum. Undoubtedly the commonest Carabid beetle found in the East. It abundantly occurs throughout the whole of South East Asia from Japan in the North to Queensland in the South. Stenolophus smaragdulus F. var. quinquepustulatus Wied. (Zool. Mag. 1823, p. 58). PALAWAN: Brooke's Point, Uring Uring, 3 sp. 22. VIII. - BALABAC: Dalawan Bay, 15 sp. 4.-7. X.- TAWI TAWI: Tara wakan, 2 sp. 14. XL - MINDANAO: Sapamoro, Curuan district, 1 sp. 15. XII. As variable and common as the typical form and with the same enormous distribution. Stenolophus sp. P ALA WAN: Mantalingajan, Pinigisan, 600 m, 1 sp. 14. VIII. Trichotichnus sp PALAWAN: Mantalingajan, Pinigisan, 600 m, 1 sp. 21. IX. TAWI TAWI: Tarawakan, 2 sp. 20.-21. X. Carbanus straneoi Louw.