The Phasmid Study Group CHAIRMAN: Judith Marshall
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Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2018-07-01 Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Life Sciences Commons BYU ScholarsArchive Citation Pacheco, Yelena Marlese, "Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea" (2018). Theses and Dissertations. 7444. https://scholarsarchive.byu.edu/etd/7444 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco A thesis submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Master of Science Michael F. Whiting, Chair Sven Bradler Seth M. Bybee Steven D. Leavitt Department of Biology Brigham Young University Copyright © 2018 Yelena Marlese Pacheco All Rights Reserved ABSTRACT Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Department of Biology, BYU Master of Science Phasmatodea exhibit a variety of cryptic ecomorphs associated with various microhabitats. Multiple ecomorphs are present in the stick insect fauna from Papua New Guinea, including the tree lobster, spiny, and long slender forms. While ecomorphs have long been recognized in phasmids, there has yet to be an attempt to objectively define and study the evolution of these ecomorphs. -
Phasmid Studies 1 (June & December 1992)
PSG 118, Aretaon asperrimus (Redtenbacher) Paul Jennings, 14, Grenfell Avenue, Sunnybill, Derby, DE3 7JZ. U.K•• Taxonomic and distribution notes by P .E. Bragg. lliustration of male by E. Newman. Key words Phasmida, Aretaon asperrimus, Breeding, Rearing. Classification This species was originally described as Obrimus asperrimus by Redtenbacher in 1906. In 1938 Rehn & Rehn established the new genus Aretaon (1938: 419), with asperrimus as the type species. I have examined the type specimens of this species and A. muscosus Redtenbacher, which are all in Vienna, and I found that the type specimens of A. asperrimus are all adults while those of A. muscosus are all nymphs. A. muscosus is distinguished by having more prominent spines, particularly on the front tibiae and the top of the abdomen. However, having reared A. asperrimus it is clear that nymphs of this genus are very spiny and these spines in particular are reduced when the insect becomes adult. It is therefore quite likely that A. asperrimus and A. muscosus are the same species. This possibility was considered and rejected by Gunther (1935: 123) but as he had not reared them he would not have known that the spines are reduced when the insects become adult. The species in culture is clearly A. asperrimus, however it has smaller spines than the type specimens so clearly the species is variable. The type specimens of A. muscosus are much more spiny than those in culture, I cannot therefore be certain that A. asperrimus and A. muscosus are the same species. As there is a strong possibility that they are the same species, I am listing the published references and distribution records for both species. -
Insecta: Phasmatodea) and Their Phylogeny
insects Article Three Complete Mitochondrial Genomes of Orestes guangxiensis, Peruphasma schultei, and Phryganistria guangxiensis (Insecta: Phasmatodea) and Their Phylogeny Ke-Ke Xu 1, Qing-Ping Chen 1, Sam Pedro Galilee Ayivi 1 , Jia-Yin Guan 1, Kenneth B. Storey 2, Dan-Na Yu 1,3 and Jia-Yong Zhang 1,3,* 1 College of Chemistry and Life Science, Zhejiang Normal University, Jinhua 321004, China; [email protected] (K.-K.X.); [email protected] (Q.-P.C.); [email protected] (S.P.G.A.); [email protected] (J.-Y.G.); [email protected] (D.-N.Y.) 2 Department of Biology, Carleton University, Ottawa, ON K1S 5B6, Canada; [email protected] 3 Key Lab of Wildlife Biotechnology, Conservation and Utilization of Zhejiang Province, Zhejiang Normal University, Jinhua 321004, China * Correspondence: [email protected] or [email protected] Simple Summary: Twenty-seven complete mitochondrial genomes of Phasmatodea have been published in the NCBI. To shed light on the intra-ordinal and inter-ordinal relationships among Phas- matodea, more mitochondrial genomes of stick insects are used to explore mitogenome structures and clarify the disputes regarding the phylogenetic relationships among Phasmatodea. We sequence and annotate the first acquired complete mitochondrial genome from the family Pseudophasmati- dae (Peruphasma schultei), the first reported mitochondrial genome from the genus Phryganistria Citation: Xu, K.-K.; Chen, Q.-P.; Ayivi, of Phasmatidae (P. guangxiensis), and the complete mitochondrial genome of Orestes guangxiensis S.P.G.; Guan, J.-Y.; Storey, K.B.; Yu, belonging to the family Heteropterygidae. We analyze the gene composition and the structure D.-N.; Zhang, J.-Y. -
PROCEEDINGS of the WORKSHOP on TRADE and CONSERVATION of PANGOLINS NATIVE to SOUTH and SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S
PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S. Pantel and S.Y. Chin Wildlife Reserves Singapore Group PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 JUNE –2JULY 2008, SINGAPORE ZOO EDITED BY S. PANTEL AND S. Y. CHIN 1 Published by TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia © 2009 TRAFFIC Southeast Asia All rights reserved. All material appearing in these proceedings is copyrighted and may be reproduced with permission. Any reproduction, in full or in part, of this publication must credit TRAFFIC Southeast Asia as the copyright owner. The views of the authors expressed in these proceedings do not necessarily reflect those of the TRAFFIC Network, WWF or IUCN. The designations of geographical entities in this publication, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of TRAFFIC or its supporting organizations concerning the legal status of any country, territory, or area, or its authorities, or concerning the delimitation of its frontiers or boundaries. The TRAFFIC symbol copyright and Registered Trademark ownership is held by WWF. TRAFFIC is a joint programme of WWF and IUCN. Layout by Sandrine Pantel, TRAFFIC Southeast Asia Suggested citation: Sandrine Pantel and Chin Sing Yun (ed.). 2009. Proceedings of the Workshop on Trade and Conservation of Pangolins Native to South and Southeast Asia, 30 June-2 July -
Phasmid Studies
ISSN 0966-0011 PHASMID STUDIES. volume 13, numbers 1 & 2. September 2005. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 13, numbers I & 2. Contents Phasmids from Sabah Robert Bradburne I A reassessment of some Bornean Lonchodinae and Aschiphasmalidae, with some lectotype designations, new synonyms, and the description of (WO new species P.E. Bragg ................ ........ .. 11 Hap/opus Burmeisler, 1838, replacement name for Aplopus Gray, 1835 (Phasmalodea). Oliver Zornpro ... .. 30 A new species of the genus Baculofraclum. the first record of the genus from Borneo. P.E. Bragg .. ............................. .. 31 Reviews and Abstracts Phasmid Abstracts 38 Cover illustration: Female Parafoxopsis kQrySll!.~ Gilmher, 1932 by r.E. Bragg. Br.dburn, R. (2005) Phasmid Studies, 13(1&2): \-10. - Phasmids from Sabah Robert Bradburne, 26 Royal Avenue, Tonbridge, Kent, TN9 208, UK. Abstract This paper describes a trip (Q six locations in Sabah, Borneo, during October 2003. A 101:11 of around 20 species of stick insects were found al four of these locations, including an undescribed species found at 3300m on Moum Kinabalu. The most commonly encountered species in the lowland forest were Lonchodes spp., Haaniella spp., and Asceles margarilatus. Key words Phasmida, Borneo, Sabah, Sukau, Kinabalu, Danum Valley, Haaniella, Asceles, Prosemoria, Necroscia, Presbistus, Carausius, PhellQcephoms, Dinophasmo. Introduction In October 2003 I travelled to Sabah in North Borneo to spend [WO weeks searching for the wildlife of the region. Our group stayed in six locations, four of which yielded many species of phasmid. The rainy season had started early and therefore it frequently rained all afternoon, and often into the night. -
Phasmida (Stick and Leaf Insects)
● Phasmida (Stick and leaf insects) Class Insecta Order Phasmida Number of families 8 Photo: A leaf insect (Phyllium bioculatum) in Japan. (Photo by ©Ron Austing/Photo Researchers, Inc. Reproduced by permission.) Evolution and systematics Anareolatae. The Timematodea has only one family, the The oldest fossil specimens of Phasmida date to the Tri- Timematidae (1 genus, 21 species). These small stick insects assic period—as long ago as 225 million years. Relatively few are not typical phasmids, having the ability to jump, unlike fossil species have been found, and they include doubtful almost all other species in the order. It is questionable whether records. Occasionally a puzzle to entomologists, the Phasmida they are indeed phasmids, and phylogenetic research is not (whose name derives from a Greek word meaning “appari- conclusive. Studies relating to phylogeny are scarce and lim- tion”) comprise stick and leaf insects, generally accepted as ited in scope. The eggs of each phasmid are distinctive and orthopteroid insects. Other alternatives have been proposed, are important in classification of these insects. however. There are about 3,000 species of phasmids, although in this understudied order this number probably includes about 30% as yet unidentified synonyms (repeated descrip- Physical characteristics tions). Numerous species still await formal description. Stick insects range in length from Timema cristinae at 0.46 in (11.6 mm) to Phobaeticus kirbyi at 12.9 in (328 mm), or 21.5 Extant species usually are divided into eight families, in (546 mm) with legs outstretched. Numerous phasmid “gi- though some researchers cite just two, based on a reluctance ants” easily rank as the world’s longest insects. -
Phasmid Study Group Newsletter, 28
ISSN 0268-3806 Chairman: Mrs Judith Marshall Department of Entomology British Museum (Natural History) Cromwell Road, London SW7 5BD Membership: Paul Brock (Phone 0753-79447) "Papillon", 40 Thorndike Road Slough, Berks SL2 1SR NEWSLETTER NO. 28 September 1986 Once again thanks to Peter Curry (No. 91) for opening up the Centre for Life Studies for the Group's summer meeting. Some 30 members attended (including three from abroad), about half of whom gave away some 20 different species - thanks also to Tim Branney (No. 239) for coping with the surplus. Paul Brock (No. 26) showed his new video, which included shots of some "British" sticks in their natural habitats in this country. Three members showed the very latest species 91, 92 and 93~ The Committee discussed a proposal from Peter Curry to build up, in conjunc tion with the British Museum (Natural History), an up-to-date reference collection of all phasmid species within the Group, showing eggs, nymphs and adults and including morphological variations. As part of this, Tony James (No. 1) offered to collect a similar file of photographs, and members are invited to send these (preferably in colour) to him for inclusion. (Peter recommends Kodachrome 64· or 25 for goo d colour rendering.) COMPUTER LIST OF MEMBERS' NAMES AND ADDRESSES - Members are asked to note that their names and addresses are held in a computer by David Robinson (No. 29) for generating address labels (for posting Newsletters, etc), in case anyone wishes to object , This list is registered under the recent Data Protection Act. and its release for other purposes without the consent of the individuals is prohibited. -
The Culture of Bornean Phasmids. P.E
The culture of Bornean phasmids. P.E. Bragg, 51 Longfield Lane, Dkeston, Derbyshire, DE7 4DX, UK. Key words Phasmida, Culture potential, Borneo, Taxonomic assessment, Polyphagy, Monophagy Introduction At the PSG meeting at the Natural History Museum, London, on 22Dd 1995, I gave an illustrated lecture on the species of Bornean phasmids which have been reared in captivity. The talk stressed that the "culture potential" i.e. the likelihood of establishing a culture, of a given species appears to depend upon the flight capability of the species. There is a close correlation between culture potential and taxonomic position; this is not unexpected as flight capability is generally closely linked to the taxonomic position of phasmids. This article summarises the lecture. Since December 1987 I have visited Borneo on 7 occasions, spending a total of about six months on the island. Borneo is the third largest island in the world and is composed of four political units: Brunei (an independent country), Kalimantan (part of Indonesia), Sarawak (a State of Malaysia) and Sabah (also part of Malaysia). Most of my collecting has been done in Sarawak, although I spent five weeks in Kalimantan, three in Brunei, and two in Sabah. My first visit to Borneb was a two week holiday with my wife and Lee Yong Tsui in December 1987 and January 1988, during this time we stayed with Dr Lee's family in Sarawak. Although not a collecting trip, seven species were collected over two nights and five of these were bred in the UK. All subsequent trips have been made specifically to collect phasmids and, whenever possible, to attempt to rear them in the UK. -
Belalong Forest: Anarchoides Iyralus (A Male) Was Taken on Bukit Belalong
B A T R E 0 p I L C A A L L R A I By 0 N F Earl of Cranbrook 0 N R David SEdwards E G~ With other contributions by George Argent Charles M Francis Penroose Saleha Martin Barker Freddy Gathome-Hardy Axel Dalberg Poulsen Roger A Beaver David T lanes Klaus Riede Peter F Becker Kamariah Abu Salim Gaden S Robinson Peter S Bellwood Chris J Kofron Sheila M Ross Elizabeth L Bennett Ruth Levy David Roubik Webber E Booth A Q Malik Samhan Nyawa Philip E Bragg C1iveF Mann Martin Sands Dielrich Burkhardl Nick Mawdsley Haji Serudin Tinggal loe K Charles lason, Lord Medway Gavin C Smith Salish C Choy Tom C Mitchell Rowland Snazell Mark A Cook Jonathan Moran Brian M Spooner Ind rane iI Das D Neil Morgan Mary C Slockdale Martin Drake Ingrid de la Motte Peter Wilkie John Dransfield Mark Mulligan Yanilla Willcox Soejatmi Drullsfield Nico Nieser John TWills Alan P Dykes Alben G Orr Yong Nget Yung Peter P Eaton Colin A Pendry Twenty-nine species of Phasmida have been collected in the Belalong forest: Anarchoides Iyralus (a male) was taken on Bukit Belalong . In the daily cycle of changing sounds in the abdomen, and stabbing or pinching Belalong, many noises are produced by movements of the powerfully spined hind insects, their chief purposes to attract po legs. Two species of leaf insects (Phyllium) tential mates and/or advertise territory. recorded from Borneo produce sounds by Orthoptera: Many Orthoptera produce rubbing together the thickened third seg noises, using a great variety of sound ments of their antennae. -
The Phasmid Study Group Newsletter No
The Phasmid Study Group Newsletter No. 122 June 2010 ISSN 0268-3806 Male Brasidas foveolatus with spermatophores. PSG SUMMER MEETING, Saturday, 10th JULY 2010.........................................................................2 News, Information & Updates ......................................................................................................................3 The Committee..........................................................................................................................................3 Diary Dates................................................................................................................................................3 Articles, Reviews & Submissions.................................................................................................................3 Book Review: Big Bugs Life~size...........................................................................................................3 Book Announcement:................................................................................................................................4 Mantids of the Euro-Mediterranean Area .................................................................................................4 Culture survey 2010 ..................................................................................................................................5 Book Announcement: Silent Summer: The State of Wildlife in Britain and Ireland ..............................9 Spermatophores from PSG 301 Brasidas -
Evolution of Flight Morphology in Stick Insects
bioRxiv preprint doi: https://doi.org/10.1101/774067; this version posted October 23, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 2 A tale of winglets: evolution of flight morphology in stick insects 3 4 Yu Zeng1,2,†, Conner O’Malley1, Sonal Singhal1,3, Faszly Rahim4,5, 5 Sehoon Park1, Xin Chen6,7, Robert Dudley1,8 6 7 1Department of Integrative Biology, University of California, Berkeley, CA 92870, 8 USA 9 2Schmid College of Science and Technology, Chapman University, Orange, CA 10 92866, USA 11 3 Department of Biology, CSU Dominguez Hills, Carson, CA 90747 USA 12 4Islamic Science Institute, Universiti Sains Islam Malaysia, 71800 Bandar Baru 13 Nilai, Negeri Sembilan, Malaysia 14 5Centre for Insect Systematics, Universiti Kebangsaan Malaysia, 43600 Bangi, 15 Selangor, Malaysia 16 6Department of Biology, The College of Staten Island, The City University of New 17 York, NY 10314, USA 18 7Department of Biology, The Graduate School and University Center, The City 19 University of New York, NY 10016, USA 20 8Smithsonian Tropical Research Institute, Balboa, 21 Republic of Panama 22 23 †Corresponding author: [email protected] 24 25 1 bioRxiv preprint doi: https://doi.org/10.1101/774067; this version posted October 23, 2019. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. -
Insect Egg Size and Shape Evolve with Ecology but Not Developmental Rate Samuel H
ARTICLE https://doi.org/10.1038/s41586-019-1302-4 Insect egg size and shape evolve with ecology but not developmental rate Samuel H. Church1,4*, Seth Donoughe1,3,4, Bruno A. S. de Medeiros1 & Cassandra G. Extavour1,2* Over the course of evolution, organism size has diversified markedly. Changes in size are thought to have occurred because of developmental, morphological and/or ecological pressures. To perform phylogenetic tests of the potential effects of these pressures, here we generated a dataset of more than ten thousand descriptions of insect eggs, and combined these with genetic and life-history datasets. We show that, across eight orders of magnitude of variation in egg volume, the relationship between size and shape itself evolves, such that previously predicted global patterns of scaling do not adequately explain the diversity in egg shapes. We show that egg size is not correlated with developmental rate and that, for many insects, egg size is not correlated with adult body size. Instead, we find that the evolution of parasitoidism and aquatic oviposition help to explain the diversification in the size and shape of insect eggs. Our study suggests that where eggs are laid, rather than universal allometric constants, underlies the evolution of insect egg size and shape. Size is a fundamental factor in many biological processes. The size of an 526 families and every currently described extant hexapod order24 organism may affect interactions both with other organisms and with (Fig. 1a and Supplementary Fig. 1). We combined this dataset with the environment1,2, it scales with features of morphology and physi- backbone hexapod phylogenies25,26 that we enriched to include taxa ology3, and larger animals often have higher fitness4.