NEWSLETTER of the MICHIGAN ENTOMOLOGICAL SOCIETY

Volume 41 , Number 1 March, 1996

Opportunities for Ecological Studies at Research Natural Areas

Lucy Ty rrell' and Robert A. Haack 2 , USDA Forest Service, North Central Forest Experiment Station, 1992 Folwell Avenue, St. Paul, Minnesota 55108; 2 USDA Forest Service, North Central Forest Experiment Station, 1' 407 S. Harrison Road, East Lansing, Michigan 48823.

What are Research Natural Areas? Although appropriate research and tional Forest System (Le., the National monitoring are encouraged in RNAs, ac­ Forests) and Forest Service Research (Le., Research Natural Areas (R1':l"As) are tivities that directly or indirectly modify the Experiment Stations). The local Dis­ permanently protected lands within Na­ ecological processes are NOT allowed. The trict Ranger has authority over access and tional Forests that are maintained in their level ofacceptable use differs among RNAs, administration, and the Director of the natural condition. Typically, RNAs are depending on the rarity of taxa, fragility Experiment Station has authority to re­ minimally disturbed natural ecosystems or resilience ofthe ecosystems, the specific view and approve research, monitOring, that exhibit unique ecological features, obj,ectives of the RNA, and cumulative and management activities on RNAs. possess rare plants or animals, or are impacts ofuse. The methods used in each There are two Experiment Stations in the high-quality exampIes ofwidespread eco­ study will be evaluated on a case-by-case Eastern Region of the US Forest Service. systems. RNAs function as biodiversity basis and documented in the permitting The North Central Forest Experiment Sta­ reserves, sites for non-manipulative re­ process. To date, activities that have been tion oversees use of RNAs in lllinois, Indi­ search projects, and areas for lOW-impact approved include limited collecting of ana, Michigan, Minnesota, Missouri, and educational activities. As for research and plantand animal voucher specimens, col­ Wisconsin. For the States of Maine, New monitOring-related studies, RNAs serve as lecting soil samples, and taking incre­ Hampshire, New York, Ohio, Pennsylva­ (1) reference areas for the study of succes­ ment cores. In some of the recently ap­ nia, Vermont, and West Virginia, the sion, (2) baseline areas for measuring proved insect studies on RNAs, collecting Northeastern Forest Experiment Station is long-term ecological changes, and (3) con­ with the use of butterfly nets and pitfall responsible for research use of RNAs. See trol areas for comparing results from ma­ traps hasbeenapproved. Again, all activi­ Tabie 2 for a list of current contacts. nipulative research studies and manage­ ties, and especially those that involve col­ People wishing to conduct studies in ment practices that are conducted nearby lecting, will be evaluatedona case-by-case RNAs must complete an application form but not on the actual RNA. basis, keeping the ecological integrity of and have it approved before initiating the Santa Catalina, the first RNA, was es­ the RNA in mind. When proposed re­ study. The application process allows for tablished in 1927 on the Coronado Na­ search projects are being reViewed, the evaluation of the proposed activities and tional Forest in Arizona. There are now following questions are considered: Is the for record keeping of all activities con­ more than 300 established USDA Forest activity consistent with the management ducted in each RNA. The District Ranger Service RNAs nationwide, and several of the RNA? How valuable will the results may require a Special Use Permit as well. hundred more are being considered. In be? Whatare thepotential environmental If the RNA resides within congressionally the Eastern Region (= Region 9; Figure 1) of effects? Can the activity be conducted else­ designated areas such as Wilderness Ar­ the Forest Service, there are 40 established where instead? Will it negatively impact eas, Wild and Scenic Rivers, or National RNAs andanother95 underconsideration other projects in progress? Are there cu­ Recreation Areas, then the Regional For­ (Table 1). The Bowl RNA, the first RNA in mulative impacts to consider? ester must also approve the study. For the Eastern Region, was established in Unlike most National Forest lands, 1931 on the White Mountain National RNAs are administered jointly by the Na­ CONTINUED ON NEXT PAGE­ Forest in New Hampshire.

MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html will not be conducting activities or collecting samples, do not need a permit, but should contact the local National Forest District Ranger office for approval to enter the RNA.

How to Apply

First, you must obtain and com plete an RNA Use Application Form. These forms are available at Forest Service offices and from the contacts listed in Table 2. The application form may be available soon on the internet homepage of the North Central Forest Experiment Station ( http://www.ncfes.umn.edu/ ). A complete application includes the application form, a brief activi ty proposal orstudyplan, and a topographiC map (or copy) indicating the location ofthe proposed project. For projects in the Eastern Region ofthe Forest Service, com pleted documents should be mailed to: Research Natural Areas Coordinator; USDA Forest Service; North Central Forest ExperimentStation; 1992 Folwell Avenue; St. Paul, MN 55108. The forms can also be sent to the RNA contacts listed in Table 2. In most cases, applications CONTINUED FROM PREVIOUS PAGE- must be approved by the Station Director, after review by the District Ranger and by Forest and Station RNA Coordinators. activities that involve threatened orendangeredspecies, permis­ Because several people a re involved in reviewing each applica­ sion must also be obtained from the appropriate State agency tion, please allow a t least one month to complete the process. and/or the U. S. Fish and Wildlife Service. Before applying, you In the 2-page application form, you will be asked to supply are encouraged to discuss your project with local or regional RNA information on the study objectives; methods; starting and coordinators (Table 2), District Rangers, or Forest Service scien­ ending dates; number ofentry days; number of people involved tists. in the field work; the specific study location; common and scientific names of the plants and animals being studied; antici­ Who Should Apply? pated number ofsamples or specimens to be collected; proposed All researchers and managers, including Forest Service em­ final location of voucher specimens; list of any equipment, ployees, must apply for a permit to conduct studies in an RNA. chemicals, or structures to be used in the proposed activity; Permits are reqUired for all categories of RNAs, e.g., established, anticipated area to be disturbed and the nature of the distur­ candidate, and proposed RNAs. Individuals or small groups who bance; and any potential impacts to the RNA ecosystems as a want to make a short visit (one day or less) to an RNA, and who result of the proposed activities.

Eastern Region National Forests are: USDA Forest Service (1) Chippewa, Eastern Region (2) Superior, (3) Chequamegon-Nicolet, (4) Ottawa, lVIN (5) Hiawatha, (6) Huron-Manistee, (7) Mark Twain, (8) Shawnee, (9) HOOSier, (10) Wayne, (11) Monongahela, (12) Allegheny, (13) Finger Lakes, (14) Green Mountain, and (15) Wh ite Mountain.

• March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) 2 • MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.htrnl 1111 Obligations of RNA Users • A. G. Wheeler, Jr. (Pennsylvania Department of Agriculture) studied the distribution of plant bugs All users of RNAs are reqUired to file a summary report, (Hemiptera: Miridae) on Phlox subulata L. plants that including a copy of all data, and a map indicating the location were growing primarily in shale barrens of Maryland, where the study occurred. If the research lasts more than one year, a yearly update is reqUired. Reports, a copy ofthe data, and Pennsylvania, Virginia, and West Virginia, but also publication reprints should be sent to the Regional RNA Coordi­ including phlox growing on other related rock outcrop nator. In the Eastern Region of the Forest Service, Lucy Tyrrell sites. Buffalo Beats candidate RNA was one of the three now holds this pOSition (see Table 2). Plant, animal, and other Ohio sites. specimens collected during the study must be deposited at the • Donald Snitgen (Northern Michigan University) has location agreed upon in the permit. Researchers are also asked to been involved with a survey of the insect fauna occupy­ provide 1 to 3 color slides showing themselves atworkin the RNA, ing Scott's Marsh candidate RNA, which is a patterned illustrating methods of sampling, or shOwing the organisms fen with prescribed burning management in the Upper being studied. Peninsula of Michigan. Collection sites were located in areas to be burned (for both pre- and post-bum sam­ Examples of Insect Studies at RNAs pling) and in control areas that were not burned. Key aspects of the study involve determining if any rare Several insect-related studies have been conducted at RNAs. insect taxa are present, and studying the effects that Some studies have focused on the life history ofindividual insect prescribed burning have on the insect community spedes. Ollie have invo ved surveys of groups ofinsects such as Odonata and pentatom ids. rne following are examples ofinsect structure. studies that included RNAs as field sites: • Donald Chandler (University of New Hampshire) Opportunities are Numerous compared an old-growth fo~st (The Bowl RNA in New RNAs are extremely valuable resources, but so far they have Hampshire) and a 40-year-old forest for several groups been much underutilized. You should consider using one ormore of Coleoptera (e.g., Cucujidae, Leiodidae, Pselaphidae), RNAs in your upcoming research projects. Also, you may wish to and as a result also published information about new considervolunteering to help monitor or managea nearby RNA. pselaphids from New Hampshire.

Table 1. Established and candidate research natural areas (RNAs) in the Eastern Region of the USDA Forest Service.

State National Forest RNA Ranger District Status1 Acres Major Features

ILLI NOIS Shawnee NF Atwood Ridge Jonesboro E-1990 955 barrens,prairie,dry to mesic forest Barker Bluff El,izabethtown E-1990 60 glade/forest complex Burke Branch Vienna E-1991 206 dry to mesic forest, mesic barrens Cave Hill Elizabethtown E-1990 465 xeric to dry-mesiC forest, barrens, sandstone glades, cliffs Dennison Hollow Elizabethtown E-1989 205 xeric to dry-mesic forest, barrens, sandstone glades, cliffs Grantsburg Swamp Vienna S 751 cypress/tupelo gum swamp LaRue-Pine Hills/Otter Pond Jonesboro E-1991 2,585 dry to mesic forests, swamps, ponds, cliffs Ozark Hilll'rairie Jonesboro E-1991 535 prairie, dry to mesic forest Panther Holl'ow Elizabethtown E-1989 180 dry to mesic forest, deep hollows, cliffs Stoneface Elizabethtown E-1990 176 xeric to dry-mesic forest, loess, prairie, sandstone glades, cliffs, barrens Whoopie Cat Mountain Elizabethtown E-1990 17 dry forest, limestone glade

INDIANA Hoosier NF Bear Hollow/Goblin Rocks Tell City C 275 dry mesic to mesic forest &: cliffs Browning Hill Brownstown C 305 dry to mesic forest Harding Flats Tell City C 65 dry forest &: barrens Hemlock Cliffs Tell City C 1,570 dry to mesic forest &: cliffs Pioneer Mothers Mem. Forest Tell City E-1944 88 old-9rowth mesic forest Rockhouse Hollow Tell City C 170 dry to mesic forest, barrens &: cliffs

March, 1996 NEWSLETTER ofthe MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) MES Homepage on the WWW: http://insects.ummz.lsa.um ich.edu/mes/mes.h tml • 3 State National Forest RNA Ranger District Status1 Acres Major Features

MICHIGAN Hiawatha NF Betchler Tamarack Flats Sault Ste. Marie C 217 tamarack Dukes Munising E-1974 233 rich conifer swamp 18 Mile lakes Rapid River C 370 pine, birch, white cedar, ash, elm, maple Grand Island Munising [-1977 59 northern hardwood forest Horseshoe Bay St. Ignace C 1,500 wet cobble beach lake Sixteen Rapid River C 16 birch, white cedar lake Stella Munising C 80 patterned fen little Waiska Basin Sault Ste. Marie C 305 red maple Nahma Rapid River C 377 birch, white cedar, ash, elm, maple, red oak Oakridge North Sault Ste. Marie C 46 red oak Ogontz Rapid River C 17 birch, white cedar Ogontz lake Plain Rapid River C 277 sand ridge - lake plain complex Pointe Aux Chenes St. Ignace C 4,830 interdunal wetland Ramsey - lost Rapid River C 7 red pine Rock River Canyon Munising C 80-100 moist non-acid cliff, mesic northern forest st. Martin's Point St. Ignace C 520 wet cobble beach, fen-like wetlands Scott's Marsh Munising C 10,000 patterned fen Shingleton Bog Munising C 3,180 patterned fen Sturgeon River Floodplain Rapid River C 649 hardwood floodplain Summerby Swamp St. Ignace C 580 northern fen Upper 18 Mile lake Rapid River C 447 northern fen Upper Sturgeon River Rapid River C 43 white cedar, ash, elm, rnaple Wedens Bay Rapid River C 362 birch, white cedar

Huron-Manistee NFs Bear Swamp Cadillac-Manistee &: C 3,485 intermittent wetland community, rich Baldwin-White Cloud conifer swamp, southern hardwood swamp Big South Baldwin-White Cloud C 82 floodplain forest Brandy Brook Cadi Ilac-M an istee P 1,570 bog, muskeg, poor conifer swamp, rich conifer swamp, hardwood-conifer swamp Boardwalk Mio C 64 rich conifer swamp, dry-mesic forest Fry l ake Baldwin-White Cloud C 60 coastal plain marsh Hayes Tower Huron Shores P 388 high quality pine-oak forest Hopper Swamp Cadillac-Manistee P 597 regularly inundated hardwood swamp Indian lake Baldwin-White Cloud P ? 2 dry sand prairie, oak barrens Knapp Prairie Baldwin-White Cloud P mesic an d dry sand prairie, northem wet meadow, oak barrens little Robinson lake Baldwin-White Cloud P ? coastal plain marsh loon lake Baldwin-White Cloud P ? coastal plain marsh, rare plants Newaygo Prairies Baldwin-White Cloud E-1988 180 dry sand prairie, undisturbed oak-pine Nordhouse Dunes Cadillac-Manistee E-1987 795 sanddunes, jack pine interdunal wetlands, swam ps, hardwood dune forest Pearl lake Baldwin-White Cloud P ? bog, rare plant Pine Island Marsh Baldwin-White Cloud C 14 coastal plain marsh Sischo Prairies Baldwin-White Cloud P 18 dry sand prairie, oak-pine barrens TImmerman lake Cadillac-Manistee P 97 intermittent wetland Toft Lake Baldwin-W hite Cloud P ? bog, rare plant Yonkers Meadow Cadillac-Manistee P 49 intermittent wetland

Ottawa NF McCormick Kenton E-1971 3,675 northem hardwoods and conifers, conifer swamp, lakes Sturgeon River Kenton C 281 deep v-shaped valley with terraces, oxbow lakes, hemlock stand Sytvania Watersmeet C 3,200 Winegar Moraine ecosystem: old-growth hemlock, hemlock-yellow birch, sugar maple, perched-water glacial lakes

• March, 1996 NEWSLETTER ofthe MICHIGAN ENTOMOLOGICAl SOCIElY V41(1) 4 • MrS Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html State National Forest RNA Ranger District Status1 Acres Major Features

MINNESOTA Chippewa NF Battle Point Deer River E-1991 329 sugar maple-basswood forest Clustered Bur Reed M arcell E-1991 79 open and forested bog, shrub swamp, marsh Pine Point Walker E-1932 1,239 red, jack, and white pines Stony Point Black Duck E-1991 404 wet-mesic northern hardwoods Sunken Lake Deer River C 952 forest and lake ecosystems of Bemidji Sand Plain

Superior NF Kawishiwi Pines Kawishiwi P 240 old-growth white pine Keeley Creek Kawishiw i E-1976 640 jack pine, black spruce, sedge meadow Lac La Croix LaCroix E-1942 973 virgin red an d w hite pine Marble Lake Laurentian E-1987 120 northern hardwoods, rare plants Lake Agnes Tofte C 80 northem hardwoods, rare plants Schroeder Tofte E-1973 360 northern hardwoods

MISSOURI Mark TwaIn NF Grasshopper Hollow Salem C 30 prairie fen Hayden Bald Ava-Cassville-Willow Springs C 40 dolomite glade Indian Creek Ava-Cassville-Willow Springs C 300 Ozark stream Sugar Camp/Yellowwood Ava-Cassvill e-Wi lIow Springs C 50 Cladrastis kentukea (yellowwood) site Tupelo Gum !Pond Eleven-Point C 32 sinkhole pond Wolfpen Bluff Ava-Cassville-Willow Springs C 5 Juniperus ashei site

NEW HAMPSHIRE White Mountain NF Alpine Gardens Androscoggin E- 1989 300 alpine tundra The Bowl Saco E-1931 S10 northern hardwood-spruce forest Bowl Expansion Saco C 1,100 northern hardwood-spruce forest Church Pond/Bog Saco C 300 bog, wetland Cone Pond Pemigewasset- Ammonoosuc C 160 pond, spruce forest Gibbs Brook Saco C 1,650 spruce-fir, subalpine Mon roe Flats Pemigewasset- Ammonoosuc C 45 alpine tundra Mountain Pond Saco C 300 northem hardwood forest Nancy Brook Pemigewasset- Ammonoosuc E-1991 1,385 northem hardwood-spruce forest Owls Head Pemigewasset- Ammonoosuc C 70 talus slope Peabody Mountain Androscoggin C 230 northern hardwood-spruce forest

NEW YORK Finger Lakes NF Butcher Hill Shrub Opening Hector C 57 shrublands Hector Oak Woods Hector C 29 mature oak-hickory forests Mill Creek Ravine Hector C 39 ravine forests, rare fish Potomac Creek Woods Hector C 120 hemlock-northern hardwoods ravine Potomac Road Shrub Opening Hector C 54 old fields, cattail marsh Sawmill Ravine Hector C 139 mature ravine forest South of Blueberry Patch Hector C 97 mature oak-hickory Townsend Road Oak Woods Hector C 15 Appalachian oak-hickory forests

OHIO Wayne NF Buffalo Beats Athens C 14 remnant tallgrass prairie Kaiser Hollow Ironton C 800 mixed mesophytic forest dominated by sugar maple, beech, and white oak Reas Run Marietta Unit E-1975 77 successional stages of Virginia pine

March, 1996 NEWSLETTER of the MICHIGAN ENfOMOLOGICAL SOCIETY V41(1) 5 • MES Homepage on the WWW: http://insects.ummz.lsa.umkh.edu/mes/mes.html • State National Forest RNA Ranger District Status1 Acres Major Features

PENNSYLVANIA Allegheny NF Tionesta Sheffield E-1940 2,113 hemlock-beech old growth

VERMONT Green Mountain NF Beaver Meadows/Abbey Pond Middlebury C 470 wetlands, rare plants Blue Ridge Bog Middlebury C 50 high-elevation bog The Cape Middlebury E-1993 290 old growth, rich hardwoods Hi Elevation Ponds Manchester C 918 soft-water ponds, rare plants Mount Horrid Rochester C 105 cliffs, rare elements

WE ST VIRGINIA Monongahela NF 3 Bear Heaven Cheat C 147 red spruce, rare squirrel habitat Bear Rocks Potomac C 66 bog, rare plants Big Run Bog Cheat C 632 bog, rare plants Black Cherry Greenbrier C 120 representative northern hardwoods Cranberry Glades Gauley C 799 bog, rare plants Fisher Spring Run Potomac C 554 bog, rare plants Hawthorn Greenbrier C 13 hawthorn M eadow Creek White Sulphur C 14 shale barren, rare plants M t. Porte Crayon Potomac C 425 red spruce Red Spruce Greenbrier C 60 red spruce Rohrbaugh Plains Potomac C 149 bog, rare plants Stuart Knob Cheat C 400 red spruce, rare squirrel habitat Yellow Poplar Gauley C 143 representative mixed mesophytic Whites Draft White Sulphur C 74 shale barren, rare plants

WISCONSIN Chequamegon-Nlcolet NFs Alvin Creek Eagle River-Florence C 11 9 hardwood ridges, sedge meadow, spring pond an d lake Bose Lake Eagle River-Florence E- 1992 81 hemlock-ha rdwoods, lakeshore Camp 9 Pines Washburn P ? red pine Chequamegon Hardwoods Glidden-Hayward E-1988 80 northern hardwood forest Giant White Pine Grove Eagle River-Florence C 20 old-growth w hite pine, hardwood-pine Grandma Lake Wetland Eagle River-Florence E-1991 495 lake and open bog Ke ntuck lake Swale Eagle River-Florence C 208 beach ridge, bog swale, northern dry-mesic and wet forest McCarthy Lake an d Cedars Glidden- Hayward E-1989 363 lake, streams, w hite cedar swamp McCaslin Mountain Laona-Lakewood E-1992 524 hardwoods, xeric ridges M emorial Grove Hemlocks Park Falls-Medford E-1989 64 hemlock-h a rdwooUs Moquah Washburn E-1936 640 fire cessation (pine-oak barrens) Scott lake- Shelp Lake Eagle River-Florence C 266 softwater lake, open bog, old-growth pine and hemlock South Branch Grove laona-lakewood C 160 river, floodplain Spider Lake Glidden-Hayward E-1989 94 ash swamp Tucker l ake Hemlocks Park Falls-Medford E-1991 158 hemlock-hardwoods, lakeshore Twin Lakes Bog Park Falls-Medford E-1989 38 conifer swamp, bog Waupee Lake laon a-Lakewood C 255 shallow hardwater lake, mixed conifer- hardwood swamp, wet meadows, bog

. Stat s: E-1S xx == Established-date of RNA; C =Candidate RNA in Forest Plan, still under consideration for RNA; = X) sed 0 in Forest Plan; S =Under study for consideration as a candidate RNA. : . = 52 . bE ce:e"...... ·.... ea ased on evaluation of feature boundaries and/or land acquisition. i Sc;: - He_ e:r a ::- c_ g~ - 0: 'onra y r ted i the Forest Plan, is a subdivision of Stuart Knob RNA.

• March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAl SOCIETY V41(1) 6 MES Homepage on the "WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html .11• Table 2. RNA contacts in the USDA Forest Service Eastern Region (R9), North Central Forest Experiment Station (NC), and Northeastern Forest Experiment Station (NE).

Area of Juristiction Contact location' Office Phone Office FAX

Regional Adm inistration: R9/NC/NE RNA Coordinator Lucy Tyrrell NCFES-St. Paul, MN (612) 649-5031 61 2) 649-5256

National Forest Allegheny Brad Nelson SO-Warren, PA (814) 72 3-5150 (814) 726-1465 Chequamegon-Nicolet Linda Parker SO-Park Falls, WI (715) 762·5169 ( 71 5) 762-5179 Chippewa Dave Shad is SO-Cass Lake, MN (21 8) 315-8600 (218) 335·8637 Green Mtn & Finger Lakes Diane Burbank SO-Rutland, VT (802) 747·6741 (802) 747-6766 Hiawatha Jan Schultz Marquette, MI (906) 228·8491 (906) 228-8972 Hoosier Steve Olson DO-Tell City, IN (81 2) 547-7051 (81 2) 547-6144 Huron-Manistee Al ix Cleveland SO-Cadillac, MI (61 6) 775-5023 (616) 775-5551 Mark Twain Lynda Richards SO-Rolla, MO (31 4) 364-4621 (314) 341-7475 Monongahela Tom DeMeo SO-Elkins, \NIl (304) 636-1800 (304) 636·1875 Ottawa Karen Nash SO-Ironwood, MI (906) 932-1 330 (906) 932-0122 Shawnee Beth Shimp SO-Harrisburg, IL (618) 253-7114 (618) 253-1 060 Superior Tom Williams SO-Duluth, MN (218) 720-5426 (218) 720-5600 Wayne Rosemarie Boyle DO-Ironton, OH (614) 532-3223 (614) 532-S615 White Mountah Steve Fay SO -Laconia, NH (603) 528-8721 (603) 528-8783

State(s) MN Gary Brand NCFES -St. Paul (612) 649·5170 (61 2) 649-5285 MI (Upper Peninsula), WI Tom Crow NCFES - Rhinelander (715) 362-7474 (715) 362-781 6 MI (Lower Peninsula) Debbie Carr NCFES - East lansing (517) 355-7740 (517) 355-5121 IL John Dwyer NCF ES - Chicago (847) 866-9311 (847) 866·9506 IN . Dale Weigel NCF ES - Bedford (812) 275-5987 (812) 279-3423 MO Bill Dijak NCF ES - Col umbia (573) 875-5341 (573) 882-1977 ME John Brissette N EF ES -Orono (207) 866-7259 (207) 866-7262 NH Pete Garrett N EF ES - Durham (603) 868-7600 (603) 868·7604 NY,VT Gary Wade N EF ES - Burlington (802) 951·6771 (802) 951 -6368 OH Chip Scott NEF ES - Delaware (614) 368·0 101 (61 4) 368-0152 \NIl Rose- Marie M uzika NEF ES - Morgantown (304) 285-161 1 (304) 285-1505 PA Chris Nowak NEFES -Warren (81 4) 563-1 040 (814) 563-1048

1 NCFES =North Central Forest Experiment Station; NEFES =Northeastern Forest Experiment Station; DO =District Office; RO =Regional Office; SO =Supervisors Office;.

The 1995 NABA -Xerces Fourth of July Butterfly Count

The 21st annual NABA-Xerces 4th of each count added up to 2,946 (a 40% atala) adults, as well as over 11,000 a ta la July Butterfly Count, sponsored by the increase from 1994), althoughsome people larvae on coontie (Zamia spp.), a member North American Butterfly Association, was are tallied more than once, since they of the cycad family. held in the summer of 1995. Participants participated in two or more counts. TO GH THE 1995 COUNT REPORT, conducted a one-day census of all butter­ The greatest species diversity for a single please specify the year ofthe countdesired flies observed at sites within their count count occurred in Mexico, with 132 spe· and send a check or money order payable area, a IS-mile diametercircle. Most counts cies. The highest species diversity for 1995 (in US dollars) to NABA for S6/report (for were held in late June and July. counts north of Mexico took place in NABA members) or SlO/report (for In 1995, 296 butterfly countswere held, Ramsey Canyon, AZ: 102 species. For the non-members) to: 4th of July Butterfly a 16.5% increase from 1994. The 1995 second year in a row the Fox River-Seney, Count; 909 Birch Street; Baraboo, WI counts occurred in45 states (same as 1994) MI count tallied the most individuals: 53913; USA. in the US, 5 Canadian provinces (6 in 20,289 (17,191 in 1994) at a rate of 966 TO GH INFORMATION ON 1996 BUT­ 1994), and 2 states in Mexico. In 1995, individuals per party-hour. TERFLYCOUNTS in your area, send a self­ there were 263 counts in the US, 29 in A continental record high ofEuropean addressed stamped business envelope to Canada, and 4 in Mexico. California had skippers were tallied in 1995: 17,504 indi­ NABA; 4 Delaware Road; Morristown, NJ the most 1995 counts, with 19, followed by viduals (13,660 in 1994). In its first year 07960. New York with 18 and Wisconsin with 17. held, the Palm Beach County North, FL The number of people participating in count tallied 393 Florida atala (Eumaeus

March, 1996 NEWSLETTER ofthe MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) • MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html • 7 Notices Tomicus piniperda Continues to Spread Editor's Note (Notices will be run forayearor4 numbers of the Newsletter unless notified to drop them. Since its initial discovery near Cleveland, Ohio in 1992, the pine shoot beetle, Members desiring longer runs should notity Tomicus piniperda (Coleoptera: Scolytidae), has continued to spread in the United States newsletter editor, Robert Haack, USDA Forest and Canada. By the end of 1995, Tomicuswas found in 147 counties in eight US states Service, Nisbet Bldg., Rm. 220, 1407 S. and in 13 counties in Harrison Rd., East Lansing, MJ 48823.) Ontario. So far in 1996, Tomicus-Infested Counties (December 1995) Tomicus has been found 1oN"'~~';:::!"\ Summer Field Classes in Maine. Ground in one new county in the beetles & tiger beetles, June 23-29; Forest US, that being Hancock entomology, July 7-13; Mayflies, August County, Ohio. The final 4-10. For more information, contact Eagle distribution for 1995 is Hill Field Research Station; P.O. Box 9; shown below.Asa histori­ Steuben, ME 04680-0009; Phone: 207 -546­ cal note, Tomicus was 2821; FAX 207-546-3042; e-mail: found in 42 US counties at [email protected]. the close of 1992, 92 in 1993, and 118 in 1994. EntoPrint: Label Making Software. A Similarly, for Canada, new product from Entomation to prepare Tomicuswas found in ocounties at the close of1992, 7 in 1993, and 10in 1994. For more insect labels for the Apple Macintosh. Print information see: Haack, R.A. and R.K. Lawrence. 1994. Geographic distribution of thousands of multiple-line labels in min­ Tomicus piniperda in North America: 1992-1994. Newsletter of the Michigan Entomo­ utes. You can serialize yourlabels and use logical Society 39(4): 14-15. any font. EntoPrint requires a Macintosh II orbetter, System 7.0, and a laser printer for best results. Cost $29 plus $2 shipping. Volunteers Needed to Help Contact: Entomation, 2742 Beacon Hill, Ann Arbor, MI 48104-6502. Phone: 313­ Survey for Karner Blue Butterflies 971 -6033; Email: [email protected]. Wanted: Michigan Orthoptera Records. The Baldwin-White Cloud Ranger District ofthe Huron-Manistee National Forest in Contact: Roger Bland, Department of Michigan will be conducting surveys for the federally endangered Kamerblue butterfly Biology, Central Michigan University, on 1500 to 3000 a cres of National Forest land in 1996. They would greatly appreciate Mt. Pleasant, MI 48859. Telephone: 517­ assistance from MES members in this effort ifyou have one or more open days. The main 774-3455; FAX 517-774-3462. survey period would occur during the first-generation Kamer blue flight period: late May through June. Staff from the National Forest would offer a training session in May Research Opportunities for qualified to interested parties tha t would include information on (a) identification of the Kamer investigators at the Huron Mountains in blue and other rare insects, (b) identification ofnectaring plants, (c) thesurvey protocol, the upper peninsula of Michigan. Submit and (d) how to complete the survey forms. Generally, volunteers would have to cover project proposal and resume (and budget their own costs for transportation, food, and lodging. However, some lodging may be iffinancial support is requested) by March available at times. 1 of eachyear to: Dr. David Gosling, Direc­ Ifyou'd like more information, please contact Joe Kelly or Tom Walter: voice 616­ tor ofResearch; Huron Mountain Wildlife 745-4631; Fax 616-745-2345; USDA Forest Service, P.O. Drawer D, 650 N. Michigan Foundation; 69063 Wallowa Road; White Avenue, Baldwin, Michigan 49304. P!geon. ~ I 49099-9745. Phone: 616-651­ 4 : ~ · FA': ; 616-65 -3679.

far only a few ofyou have sent dress to me. Please continue to do 8 • 1111 42nd Annual MES Meeting

The Forty-Second Annual Meeting of Unfortunately, housing will not be Registration will be $15 per person. the Michigan Entomological Society will available at KBS at the time ofthe annual This will cover facility fees, coffee/teal be held on Friday, 17 May 1996, at the meeting. There is one hotel nearby: The snacks, and lunch. Please pay in advance. Education Centerofthe Kellogg Biological Gull Lake Inn (616-731-4131). State that You NEED TO REGISTER even if you are Station (KBS) on Gull Lake, near Hickory you are attending the MES meetingatKBS, NOT presenting a talk because we need to Comers, Michigan. KBS is located a short and perhaps they will give you a discount. give KBS a headcountfor lunch. For guests distance to the northwest of Battle Creek Call now to reserve a room! There are who are NOT attending the meeting, but (see maps). severalotherhotelsnearby in Battle Creek want to eat lunch at KBS, the cost will be On-site registration begins at 8:30 am, and other surrounding towns. $5. We need to confirm the number of and the first talk begins at 9 am. lunches with KBS by Tuesday, 16 May. So please register as soon as you can with Cathy Bristow, tell us how many are com­ ing in your party, and pay in advance. Make checks out to the: Michigan Entomological Society. If you wish to present an oral paper, complete that portion of the registration form and mail, fax, or e-mail it to Cathy Bristow at MSU by 30 April. We hope to see you at KBS on 17 May. For further information contact the program orga­ nizer, Cathy Bristow, Department of Entomology, Michigan State University, East Lansing, Michigan 48824. Phone: 517-355-9542; FAX: 517-353-4354; e-mail ([email protected]).

~ l if From T:8 Hickory Corns,,. I Oirection6 to KI35 III

Michigan State University's The above map shows the W. K. Kellogg Biological Station l\+~"-II-,,- location of KBS in relation to Kalamazoo and Battle Creek. The bottom map is a detailed representation of the KBS Gull lake area.

From I·U t.1ea ~f 40. Sprlnlcl. Rd. to M-Ij (Gu" Rd.)

March, 1996 NEWSLETTER ofthe MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) • • MIS Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html 9 Survey Improves Understanding of Michigan Lepidoptera

Robert D. Kriegel, Mogens C. Nielsen, Ron Priest, and Fred W. Stehr Michigan Lepidoptera Survey steering committee members

As biolo~ists , m ost ofus appreciate the In 1995, Michigan Lepidoptera SUIVey Erynnis persius, and Incisalia irus. However, importance of preseIVing species diver­ received a grant from the Michigan De­ by far the la rgest increase occurred in the sity. In recent years we have witnessed partment of Agriculture to sUIVey listed category ofspecial concern species (Table grOwing public support for conseIVa tion Lepidoptera species (endangered, threat­ 3). Here, MLS documented 46 county a nd the preseIVation of our natural heri­ ened, andspecialconcern) in insect collec­ records for 14 species not previously re­ tage. Internationa l, federal, and state tions at Michiga n State University (MSU), corded by the Natural Features Inventory. agencies have enacted legislation to pro­ University of Michigan, and several pri­ The presentstudy collected data from over tect biodiversity. In Michigan, 87 species vate collections. Michigan Department of 50 specimens for each of 11 special con· of insects are "listed" by the Department Agriculture administers a voluntary pro­ cern species and 90+ specimens for each of of Natural Resources as endangered, gram with counties and the USDA Forest five species: Boloria freija, Boloria frigga, threatened, or of special concern (Anony­ Selvice to suppress gypsy moth popula. Calephelis muticum, Oeneis macounii, and mous 1994). The majority of these, 54 tions using Bacillus thuringiensis var. Phyciodes batesii. Data collected by the species, are Lepidoptera. Efforts to protect kurstaki. Federal andstate regula tions pro­ Michigan Lepidoptera SUIVey were also threatened andendangeredspecies should hibit pesticide a pplications that impact used in the 1995 biennial review of be guidedby a thorough understanding of listed species. The agencyfunded this study Michigan's state-listed insects (Haack an organism's biology and habitat re­ to improve knowledge about the distribu­ 1994). quirements. Unfortunately, relatively little tion, flight times, and likely periods of Currently, the sUIVey is focusing on is known about the distribution and biol­ lQIVal occurrence for Michigan's listed developing a database ofMichigan' s listed ogy of many invertebrates, particularly moths and butterflies. Additional infor­ Lepidoptera. Later, this database will be , insects. A group of Michigan Lepidopter­ mation a bout the distribution and biol­ expanded to include all Michigan Lepi­ ists is trying to improve this situation by ogy of these Lepidoptera would provide a doptera. SU IVey participants are concen­ embarking on a long-term, cooperative stronger decision-making baSis to deter· trating field studies in critical habitats (Le. . effort to expand our knowledge of Michi­ mine whether a n area could be treated coa staldunes, remna nt tall·grass prairies, gan moths and butterflies. with pesticides without impacting listed fens, and h eath bogs) and seldom col­ The Michigan Lep idoptera Survey species. lected areas of the state such as Berrien, (MLS), a project of the Michigan Entomo­ Prior to this study, Michiga n Natural Gogebic, Monroe, Ontanogon, Presque Isle logical Society, is composed of individuals Features Inventory (MNFI) had collected counties and Great Lakes islands. The interested in studying Lepidoptera (Priest informationfor 39 ofthe listed Lepidoptera project continues to gather data on listed 1994). The long term mission ofMLS is to species (Anonymous 1995). The present Lepidoptera from other institutional and document the identity, distribUtion, and MLS study documented records for 2,608 private collections. Project participants biology of Michigan's diverse lepidopteran specimens covering a1154 species. In addi­ believe there are still over a dozen such fauna. The project's essentia l tasks are: (1) tion, county records for Michigan listed collections to be sUIVeyed both within to develop a computerized database of Lepidoptera increased from 138 counties Michigan and outside of the state. Project Miciligan Lepidoptera records; (2) to con­ in the MNFI database to 242 counties in leaders hope to collect records from 2,500 duct field investigations to improve our the MLS database. Five new county records more specimens during 1996. understanding of Lepidopterabiologyand were documented for three endangered All specimens included in the Michi· biodiversity, particularly in critical habi­ species (Table 1). The study also discov­ gan Lepidoptera Survey da tabase oflisted tats and poorly suzveyed areas ofthe state; ered a specimen of the regal fritillary, species were collected in accordance with and (3) to distribute information about Speyeria idalia, housed in the MSU collec· state and federal regulations. All endan­ W chigan Lepidoptera through publica­ tion, tha t was captured in 1984. Previ­ gered or threatened species were collected ::o1"..s and ed 'ca 'onal outreach projects. ously, the most recent documented cap­ prior to listing or under appropriately is­ T!".ls S'~ey is a ong·:em: pro'ect thatwill ture of this species in Michigan was from sued permits. Datagatheredinthis project :~'~t::!:€ !":e.:pO!'.d coope:a :ion ofmany 1978. Substantial new information was is kept strictly confidential. Only informa­ p€'C p:e w!:: a..t: se:: '~s: - ~. :e:es:eri :r. . also collected for several threatened spe· tion pertaining to species distribution, andi

March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAl SOCIETY V41(1) 10 • MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html II tions of endangered or threatened species are only reported to References appropriately authorized agencies. Names of collectors or pri­ Anonymous. 1994. Michigan's spedal animals. Endangered, vate collectionsarespecifically not included inproject reports. All threatened, sped al concern, and probably extirpated. Michi­ spedes detenninations are verified by knowledgeable lepidop­ gan Department of Natural Resources. June 1, 1994. llpp. terists. Anonymous. 1995. Threatened, endangered, and special con ­ Individualsinterestedinsubmitting spedmen data or becom­ cern insects of Michigan. Michigan Natural Features inven­ ing involved in the Michigan Lepidoptera Survey shouldcontact tory database, Michigan Department of Natural Resources. Fred Stehr (51 7-353-8739; e-mail [email protected]) or Mogens Lansing, MI. (liMo" ) C. Nielsen (51 7 -353-7294) atthe DepartmentofEntomol­ Haack, R. A. 1994. A review of Michigan's state-listed insects: ogy; Michigan State University; East Lansing, MI 48824. Agen­ your assistance is requested. Newsletter of the Michigan des interested in funding future efforts of the Michigan Lepi­ Entomological Society. 39(4): 10-12. doptera Survey should contact Fred Stehra t MSU or Ron Priest a t Priest, R. 1994. Michigan Lepidoptera survey: a proposal. News­ 5464 fo Pass; East Lansing, Michigan 48823 or phone evenings letter of the Michigan Entomological Society. 39(2&3): 17. at 517 -349-7407.

Table 3. Summary data for Michigan listed special concern spedes of Lepidoptera.

MNFI MichIgan Lepidoptera Table 1. Summary data for Michigan's state-listed # Survey endangered species of Lepidoptera. Species Name Counties # Counties # Specimens Acronicta falcula 0 2 2 MNFP Michigan Lepidoptera Basi/odes pepita 1 1 18 # Survey Boloria freija 0 6 97 Species Name Counties # Counties # Specimens Boloria frigga 1 5 210 Brachionyncha borealis 3 3 72 Catocala ames tris 21 Calephelis muticum 10 11 149 Catocala duldola Lycaeides melissa samuelis 7 8 190 1 2 29 Catocala il/ecta 0 3 31 Neonympha mitchellii 8 10 160 Catocala robinsoni 0 1 3 Schinia indiana 2 1 Chlosyne gorgone cariota 0 1 1 Schinia lucens 1 8 f ades imperialis pini 0 5 23 Speyeria idalia 12 14 60 Erebia discoidalis 4 3 30 Erora laeta 2 2 5 35 440 Total 31 Erynnis baptisiae 1 5 51 fuchloe ausonides 0 1 19 1. Michigan Natural Features Inventory (MNFI) data for all f uxoa aurulenta 2 5 58 three tables exerpted from Anonymous 1995. Rxsenia favonius ontario 0 1 1 Hemi/euca maia 2 6 Sl Heterocampa subrotata 0 4 28 Table 2. Summary data for Michigan's state-listed Heteropad7a rileyana 0 2 15 Hyalophora columbia 1 14 73 threatened species of Lepidoptera. Indsalis henrid 6 8 41 Merolon che dolli 3 2 24 MNFI Michigan Lepidoptera Meropleon ambifusca 2 4 41 # Survey Oeneis macoun;i 0 1 92 Species Name Counties # Counties # Specimens Oncocnemis piffardi 1 1 1 ~ Pachypo/ia atricomis 2 1 2 Atrytonopsis hianna 4 8 65 Papaipema aweme 1 1 1 Battus phi/enor 2 5 52 Papaipema beeriana 5 5 S5 Papaipema cerina 2 3 23 0 11 150 Erynnis persius Papaipema maritima 3 3 6 f uphyes dukes; 3 4 31 Papaipema sdata 6 6 18 Hesperia ottoe 5 4 70 Papaipema spedosissima 5 6 35 Indsalia irus 3 7 140 Phydodes batesii 0 16 100 Polygonia gradlis 2 Lycaeides idas nabokovi 4 3 68 0 2 Proserpinus flavofasdata 0 1 1 Oarisma powesheik 6 4 77 Pygarctia spraguei 4 6 32 Papaipema si/phii 6 3 9 Pyrgus wyandot . 4 6 47 Total 33 49 662 Spartiniphagainops 2 3 19 Total 74 158 1506

March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAL SOCIETY V41 (1) • 11 ••• MIS Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes!mes.html • Prairie Insects Studied in Southern Michigan by The Nature Conservancy

Christopher A. Clampitt, Gary A. Dunn, Tara Fletcher, and Jon Bedick Michigan Fiel d Office, The Nature Conservancy; 2840 East Grand Rive r Ave.; East Lansing, MI 48823. (517) 332-1741; e-mail: [email protected]

For years, The Nature Conservancy has used prescribed burn­ ber. Three study blocks (one per preserve) were burned in April ing as a tool to restore and maintain prairies and other native 1995 as part of our regular management. grasslands. While the positive response ofmost prairie plants to During this project, over 15,000 specimens were collected and frequent fires hasbeen well-documented, over the pastdecade or pinned. Soft-bodied invertebrates, nymphs, and some duplicate so there has been a growing realization that prairie insect specimens were stored in alcohol. Several hundred additional populations could be harmed by fire. specimens were collected on other grassland sites in southern To reduce the likelihood that prescribed burns will adversely Michigan: Grand Beach Marsh (site G, Berrien County - coastal affect the native insects on our preserves, we have divided each plain marsh); Ives Road Fen (site I, Lenawee County -prairie fen); prairie into several units that are burned in rotation over a Ore-Ida Prairie (site 0, Newaygo County - sand prairie); Tama­ multi-year cycle. However, this substantially increases ourman­ rackSwamp(site T, Cass County -wet meadow); Thompson Road agement costs. Given the ecological and evolutionary history of Prairie (site P, Cass County -mesic prairie); andThree Rivers State prairiespecies, landscape fragmentation by humans, and inten­ Game Area (site R, St. Joseph County - wet meadow). More than sive fires in the past, it is possible that fire-sensitive, half of the specimens have been identified to genus or species. prairie-dependent insects do not currently inhabit these pre­ Roughly halfofthe remaining specimens havebeen identified to serves. If so, less-costly management techniques may be appro­ family. The rest, mostly Diptera, are identified only to order. priate. Once we have completed our analyses, these specimens will be In 1994, we initiated a study of the direct effects ofprescribed deposited in a public institution for study by the entomological burning on insects inhabiting prairies and wet prairies in south­ com munity. west Michigan. For this study, two comparable study blocks (15 Because of the growing interest in prairie insects on the part x 76 m) were established on each of four preserves: Bakertown of entomologists and natural area managers in the region, we Fen (site B, Berrien County - wet prairie/prairie fen); Dayton Wet present our current list ofprairie insect taxa here. We recognize Prairie (site D, Berrien County-wetprairie); Klumbis Road Prairie that we may not have used the latest taxonomic revisions for (site K, Cass County - mesic prairie); and Lawton Prairie (site L, various grou ps, and that identification ofsomespecies is difficult Yan Buren County - mesic prairie). During 1994 and 1995, the forthenon-specialist. Therefore, we welcome commentsfrom the invertebrates in these study blocks were sampled with a sweep entomological community on current nomenclature and on the net on a bi-weekly schedule between mid-June and early Octo- ecological significance of the species on this list. Our specimens can also be made available for review by interested experts.

Order, family, genus, species, and site information for Ischnura verticalis (Say) B,D,I,K Nehalennia gracilis Morse B insects collected in various prairies in southwest Michigan. Cordulegastridae Cordulegaster diastatops Seyls DERMAPTERA (27 specimens) Site Cordullldae Forficulidae Doracordulia libera (Seyls) T,R Doru aculeatum (Scudder) D ,L Tetragoneuria cynosura (Say) L Forficu!a auricularia L. D,K ,L Lestidae Lestes unguiculatus Hagen L Ubellulidae Erythemis simplicicollis (Say) T,R T Leuco"hinia hudsonica (Selys) L,T Libel/ula luduosa Burmeister I,K B,I ,T Libel/ula pulche/la Drury B,K,L Pachydiplax longipennis (Burmeister) T B.I Plathemis lydia Drury B,D 5ymepetrum rubicundulum (Say) K,L 5ympetrom semidnctum (Say) I

• March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAL SOCIETY Y41(1) 12 • MES Homepage on the WWW: http://insects.ummz.1sa.umich.edu/mes/mes.html PLECOPTIRA (8 specimens) Prosapia ignipectus Perlidae undetermined spp. undetermined spp. B,D Cicadellidae Agallia constricta Van Duzee B,D,K,l ORTHOPTERA (400 specimens) Agallia novella (Say) B,D,K,l ----- Acrididae Agallia spp. B,D,K,l Arphia sulphurea Fabricius l Aphrodes albifrons (l.) B,D,K,l Chorthippus curtipennis (Harris) B,l,M Aphrodes bicincta (Schrank) K,l Dissosteira carolina (l.) L Aphrodes costata (Panzer) B,K,l Melanoplus bivittatus Say B,D Athysanus argentarius Metcalf B,D,I,K,l,O,P Melanoplus confusus Scudder K,l Aulacizes irrorata (Fabricius) D,K,l Melanoplus differentialis (Thomas) B,I,K,l Balclutha impicta (Van Duzee) K,l M elanoplus femurrubrum femurrubrum (DeGeer) B,D,K, l ,M Balclutha punctata (Thumb) L Melanoplus viridipes viridipes Scudder B,D,K,l Chloanthanus acutus (Say) B,D,I,K,L Pseudopomala brachyptera (Scudder) B,l Chloanthanus bicolor (DeLong) B,D,K,L Stethophyma Iineatum Scudder R Chloanthanus frontalis (Van Duzee) B,D,K,l undetermined spp. B,D,K,l Chlorotettix unicolor (Fitch) B,D,K,l Gryllidae Gcadula punctifrons (Fall) D Allonemobius allardi Alexander & Thomas l Cicadula variata (Fall) B,D Nemobius affinis Beutenmuller K,l Gminius hartii (Ball) K Oecanthus fultoni T.). Walker I Commel/us comma B Oecanthus nigricornis F. Walker B,D,K Cuerna Iimbatus (Say) l Oecanthus quadripunctatus Beutenmuller D,K Deltocephalus balli (Osborn) B, D,K,l undetermined spp. B Deltocephalus configuratus (Uhler) l Mantidae Deltocephal us flavicosta (Stal) Mantis religiosa (l.) B,D,I,K,L Deltocephalus misellus (Ball) L Phasmatidae Deltocephalus reflexus (0. & B.) K Diapheromera femorata Say K,l Deltocephalus sayi (Fitch) K, L Tetrigfdae Deltocephalus stylata Boheman B, D,K Nomotettix cristatus (Scudder) D,L Deltocephalus spp. B, l Tetrix ornata Say B,D,K Dicraneura celtidis (Osborn) B, l Tetrix subulata (L.) D Doratura stylata (Boheman) K,l Tettigidea lateralis I Draeculocephala angulifera (Walker) B, D,K,l Tettigoniidae Draeculacephala antica l Amblycorypha oblongifolia Scudder B Draeculacephala constricta P Amblycorypha rotundifolia Scudder K Draeculacephala spp. L Conocephalus fasciatus (DeGeer) D,K,l Draeculocephala novemboracensis (Fitch) B,D, K Conocephalus strictus (Scudder) B Empoasca alboneura (Gill) B,K, l Neoconocephalus ensiger Harris l Empoasca faboe (Harris) B, D,K,l Orchelimum gladiator Bru nner B Empoasca f1avescens (Fabricius) B,D,K,l Orchelimum nigripes Scudder B, D,K,l Empoasca spp. B, l Orchelimum vulgare Harris D Erasmoneura obliqua (Say) B,l Scudderia furcata Brunner B,D Euscelis bicolor (Van Duzee) K,l Scudderia pistil/ata Bru n ner B, D,l,T Exitanus exitiosus Scudderia texensis Saussure & Pictet B,D Exitianus spp. °I undetermi ned spp. B,L Fitchana vitellina (Fitch) L Graphalocepha/a coccinea (Forster) B,D,K HOMOPTERA (4,900 spedmens) Gypona ma/anota Spanberg B,K,L Acanaloniidae Gypona octolineata (Say) B, D,K,l Acanalonia bivittata Say B,D,I,K,L,T,P Gypona unicolor Stal K Acanalonia conica Say B Gypona spp. l Achllidae Hecalus Iineatus (Uhler) B Catonia cinctifrons (F itch) B Helochara communis Fitch B,K,l Catonia lunata Metcalf K,l Idiocerus duzeei Provancher B Aphidldae Idiocerus pallidus Fabricius B undetermined spp. B, D,K,l Idiocerus raphus G Cercopidae Idiocerus rotundens Delong & Caldwell K Aphrophora quadrinotata Say B,D Idiocerus snowi G. & B. K Gastoptera hyperici Gibson B Jassus olitorius Say B, D,K,l Gastoptera obtusa (Say) l Macropsis bifasciata (Van Duzee) D,K Clastoptera proteus Fitch B,D,K,l,P Macropsis viridis (Fitch) D Lepyronia angulifera I Macroste/es variata (Fallen) B,D Lepyronia gibbosa Ball l M esamia nigridorsus Ball K,L Lepyronia quadrangufaris Say B,D,I,T Neko/la hieroglyphica Say B,D,K, l Neophi/aenus lineatus L. B Oncopis spp. B, D,K,L Phi/aeneus killa Ophiola uhleri (Ball) B,D Phi/aenus spumarius (L.) B,D,I,K,L,P° Parabolocratus viridis Uhler B,D,K Phi/aronia canadensis (Walley) B,K Paraphlepsius spp. B,D,K,l

March, 1996 NEWSLElTER of the MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) • II MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html • 13 Penthimia american us Fitch K,L Megalostomus quinquespinosus (Say) K,L Scaphoideus spp. B,D,K,L Protenor be/fragei Haglund B,D,L Thamnotettix clitellarius Say K Anthocoridae Thamnotettix kennicotti Uhler K,L Orius insidiosus Say 8,D,K,L Thamnotettix melanogaster (Provancher) B,D,K 8erytidae Thamnotettix nigrifrons (Forbes) B,D,K,L Neides muticus (Say) K,L undetermined spp. B,D,I,K,L Coreidae Cicadidae Catorhintha mendica Stal K Tibicen canicularis (Harris) L Chariesterus antennator (Fabricius) K,L Cixiidae f uthochtha galea tor (Fabricius) D,K,L Cixius basalis Van Duzee K,L Cydnidae Myndus fulvus Osborn B Sehiris cinctus (Palisot) B,K Oliarus vicarius Walker B,L Gerridae Delphacidae Gerrfs remigis Oelphacodes foveata (Van Duzee) B, D,K,L Lygaeidae Oelphacodes puella (Van Duzee) B,D,K,L C)'l1lI.G angustatus Stal B,I Oelph acodes spp. B,D,K,L Cymus coriocipef1nis (Stal) B Pentagramma vittatifrons (Uhler) B IschfKXiemus falic s (Say) B,D Pissonotus delicatus (Van Duzee) B Isthmogecxoris piceus Say L Pissonotus marginatus (Van Duzee) Lygaeus kalmii Sta l K Stobaera tricarinata (S ay) B Meacoryphus bicrucis (Say) B Derbidae Myodocha serripes Olivier K Amalopota fitchi (Van Duzee) B,D,K,L Nys ius niger Baker B,K Cedusa vulgaris (Fitch) B,D,K,L Neortholomus scolopax (Say) B, D,K,L Oictyopharldae Oedancala dorsalis (Say) B, D Phylloscelis atra Germar B Perigenes constrictus (Say) B, D, K,L Phylloscelis pallescens (Germar) B Phlegyas abbreviatus (Uhler) K,L Scolops sulcipes (Say) B,D,K,L,P Miridae Stobaera tricarinata Say B,D Ceratocapsus pumilis (Uhler) B,K,L Flatidae Collaria meilleurii Provancher B,D,K, L Anomenis septentrionaJis (Spinola) B,D,G,I,K Criocoris saliens (Reuter) D Ormenis pruinosa (Say) B,D,K,L Hyaliodes vitripennis (Say) K,L Metcalfia pruinosa I Jlnacora stallii Reuter K undetermined spp. B,D,L Leptopterna do/obrata (L.) D,K,L Issidae Lopidea media (Say) B,D,L Bruchomorpha dorsata (Fitch) L Lygocoris communis Knight K Bruchomorpha oculata (Newman) K,L Lygocoris vibumi Knight L Bruchomorpha tristis Stal L Lygus atrifJavus Knight 0 undetermined spp. L Lygus lineolaris (Palisot) B,D,I,K,L Membracidae Megaloceroea recticomis (Geoff.) B,K,L AcutaJis semicrema (Say) B,D Metriiorrhynchomiris dislocatus (Say) B,D,K,L Acutalis tartarea (Say) B,D,T Monalocoris american us Wagner & Slater B,D Antianthe arcuatus (Emmons) L Neurocolpus nubi/us (Say) K,L Archasia galeata Say K,L Orthops scutellatus (Uhler) B,K Campylencia latipes Say B,D,K,L,T Phytocoris tibialis Reuter B,D Ceresia bisonia Kopp & Yonke B,D,K,L,T Plagiognathus obscurus Uhler B,D,K,L Ceresa diceros (Say) D Poecilocapsus lineatus (Fabricius) B,D,K Ceresa inemis Fabricius K,L Polymerus venaticus (Uhler) B,D,K Ceresa lutea Walker I,L pycnoderes quadrimaculatus Guerin-Meneville B,D,K,L Ceresa spp. B,D,L Sericophanes heidemanni Poppius L Cyrtolobus inemis (Emmons) L Sixeonotus insignis Reuter B,K Entylia concava (Say) B,D,L Slaterocoris spp. B,D,K Entylia reticulata (Van Duzee) D Stenotus binotatus Fabricius L Micrutalis calva (Say) B,D,K,L Trigonotylus coelestialium (Kirkaldy) B,D,K,L Ophioderma evelyna Woodruff L undetermined spp. B,D,K,L Smilia camelus (Fabricius) L Nabldae Strictocepha/a spp. I Oolichonabis propinqus Xantholobus muticus (Fabricius) K Nabicula subcoleoptrata Kirby B,I,K,L determined spp. D,L Nabis ameriocoferus Carayon B,D,K,L Pentatomldae B,D,K,L Acrosternum hi/are (Say) D,K Acrosternum pennsylvanicum (Gmelin) B,K,L B,L Apateticus cynicus (Say) L Brochymena quadripustulata (Fabricius) K 5) Coenus delius (Say) K,L Cosmopepla bimaculata (Thomas) B,D,I,K,L 8,O,K,L Dendrocoris humeralis (Uhler) L Euschistus tristigmus (Say) 8,D,I,K,L

• March, 1996 NEWSLETTER of the MlCHIGAN ENTOMOLOGICAL SOCIETY V41(1) 14 • MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html •• Euschistus variolarius (Palisot) B,I,K,L Bruchidae Holocostethus limbolarius (Stal) K undetermined spp. B,I,K,L Mormidea lugens (Fabricius) K,L Buprestidae Podisus maculiventris (Say) D,K,L Acmacodera pulchella Hbst. I Podisus placidus Uhler K Agrilus arcuatus (Say) Thyanta accerra McAtee K,L Agrilus otiosus Say ) I,K,~ undetermined spp. D,I,K,L Agrilus politus (Say) B,K,L Phymatldae Agrilus ruficollis (Fabricius) K Phymata americana Melin B,D,K Anthaxia spp. B,D,K,L Phymata erosa I Brachys aero sus Melsheimer K Phymata pennsylvanica Handlirsch B,D,K Cantharidae undetermined spp. D Cantharis bilineatus Say I Reduviidae Gantharis excavatus LeConte B,I Fitchia aptera Stal L Cantharis lineola Fabricius I Pselliopus cinctus (Fabricius) D,L Cantharis rectis Melsheimer B,D Sinea diadema (Fabricius) B,D,K Gantharis spp. B,D Zelus exsanguis (Stal) L Chauliognathus marginatus Fabricius K,L undetermined spp. B,L Chauliognathus pennsylvanica DeGeer B,D,I,K Rhopalldae Podabrus modestus (Say) B Hypogeocoris piceus Say B,K Podabrus rugulosus (Say) B,I Niesthrea louisianica (Sailer) K Podabrus spp. D,L Scutelleridae undetermined spp. K Eurygaster altemata (Say) M Carabidae Thyreocorldae Agonum thoyeri Dejean B Corimelaena lateralis (Fabricius) B,I,K,L Calathus ingratus Dejean L Corimelaena pulicaria (Germar) D,K,L Calleida punctata LeConte B,D,I,K, Galgupha loboprostethia (Sailer) K,L Harpalus puncticeps Stephens K undetermined spp. I Lebia ornata Say D Tingidae Lebia spp. I Corythuca arcuata (Say) L Lebia viridis Say D Corythuca ciliata (Say) B,D,K,L Leptotrachelus dorsalis Fabricius B Corythuca marmorata L Platynus decorus I,T PhysatocheiJa spp. B Platynus hypo/ithos Say K Undetermined Family Synuchus impunctatus Say L undetermined spp. B,D,I,K,L Cerambyddae Batyleoma suturale (Say) K PSOCOPTERA (13 specimens) Dectes sayi Dillon & Dillon D,T Pseudocaedllidae Hippopsis lemniscata (Fabricius) D,I undet ermined spp. L Megacyllene robiniae Forst. I Psocidae Oberea basalis LeConte K undetermined spp. B,K,L Oberea ruficollis (Fabricius) L Undetermined Family Oberea tripunctata (Swederus) D,L undet ermined spp. B,L Psyrassa unicolor (Randall) IK Romaleum rufulum (Haldeman) L THYSANOPTERA (7 specimens) Straglina luteicornis (Fabricius) K Undetermined Family Tetraopes tetrophthalmus (Forster) K undetermined spp. B,K,L Chrysomelidae Anomoea Jatic/avia (Forster) K,L NEUROPTERA (17 specimens) Anoplitis inaequalis Weber Chrysopidae Babia quadriguttata (Olivier) K,L Chrysopa oculata Say B, D,K,L Blepharius rhois Forster K Hemerobiidae Cal/igrapha bidenticola Brown K undetermined spp. K Cal/igrapha multipunctata (Say) L Mantlspldae Cal/igrapha spp. I Oimaciella brunnea Chaetocnema denticulata (Illiger) B Chaetocnema minuta Melsheimer B,D,K,L COLEOPTERA (2,200 specimens) Chalepus dorsalis Thunberg A1leculldae Chelymorpha sp,p. K Isomira spp. K,L Chlamisus gibbosa Fabricius K,L undetermined spp. L Criocerus duodecimpunctata (L.) L Anobildae Cryptocephala mutabilis Melsheimer K Caenocara spp. B,L Cryptocephala notatus Fabricius K,L Anthicidae Cryptocephalus venustus Fabricius K Notoxus monodus Fabricius B,K,L Cryrtocehalus spp. K,L Sapintus spp. B,D,K,L Deloyala guttata (Olivier) B undetermined spp. B Diabrotica longicomis barberi Sm ith & Lawr. B,L Anthribidae Diabrotica undecimpunctata howardi Barber D,K,L Eurymyeter fasciatus Diabrotica virgifera LeConte B,D,K

March, 1996 NEWSLEITER ofthe MICHIGAN ENTOMOLOGICAl SOCIETY V41 (1) • IIlIa MIS Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html • 15 . chus aviatus Fabricius D Elateridae Dis ycha caroliniana (Fabricius) Ctenicera spp. D O'sonycha pennsylvanica (Illiger) D Melanotus communis I Disonycha xanthomelaena (Dalman) D undetermined spp. L Donacia subtilis Kunze Helodldae Exema canadensis Pierce B Cyphon spp. B,D,K,L GalerucelJa americana (Fabricius) B,D,L Scirtes spp. B,K,L Longitarsus melanurus Melsheimer I undetermined spp. B,D Longitarsus testaceus (Melsheimer) B,D,K,L Lam pyridae Metronia bicolor Fabricius B ElJychnia corrusculus (L.) B,L Odonata dorsalis Thumb. I Lucidota atra (Fabricius) B,D,I,L Oedionychus circumdata (Randall) D Lucidota punctata I Oulema melanopus (L.) K,L Photinus pyralis (L.) I,K Phaedon viridis (Melsheimer) K,L Photuris consanguineus LeConte B,I,K,L Trirhabda canadensis I Photuris pennsylvanicus DeGeer B,D Trirhabda virgata LeConte B,D,IK,L Pyr0pY9a decipiens (Harris) B,D,I,L Zygogramma suturalis K Languriidae Undetermined spp. B,D,I,K,L,M Aeropteroxys graci/is Newman B,D,I,K Cicindelidae Aeropteroxys spp, K Gcindela punctulata Oliv. Lathridiidae Cleridae Melanopthalmus spp. K,L Enoclerus rosmarus (Say) B,D Leptinidae Isohydnocera curtipennis (Newman) B,D,K,L undetermined spp. L Isohydnocera tabida (LeConte) B,K,L Lycldae PhylJobaenus humeralis difficilis (LeConte) K,L Calopteron disereptans (Newman) PhylJobaenus humeralis humeralis (Melsheimer) K,L Calopteron reticula tum Fabricius PhylJobaenus pallipes (Say) K,L Meloidae Coccinellldae fpicauata fabricii (LeConte) L Anatis quindecimpunctata (Olivier) L Epicauta pennsylvanicus DeGeer B,I,K,L Brachyacantha felina (Fabricius) K,L Epicauta pestifera Werner K,L Brachyacantha ursina Fabricius L Melyridae CoccinelJa novemnotata Herbst B,I,K Attalus spp K,L CoccinelJa trifasciata L. L Colops spp. K Coleomegilla fuscilabris Mulsant B,D,I,K undetermined spp. L Cyc/oneda sanguinea (L.) K,L Mordellidae Epi/achna varivestria Muscat I Anaspis rufa Say L Harmonia axiridis novemduodecimguttata Falderman K MordelJa atra (Melsheimer) K,L Harmonia axiridis frigida Mulsant M ordelJa marginata Melsheimer D,K,L Hippodamia parenthesis (Say) K,L MordelJa melaena Germar D,K,L Hyperaspis proba (Say) L Mordellistena andreae LeConte D Hyperaspis signata Olivier L Mordellistena aspersa I Hyperaspis undulata (Say) B,K Mordellistena marginalis (Say) D PsylJobora vigintimaculata (Say) B Mordellistena pubescens (Say) B,D,I,K SubcoccinelJa vigintiquatuorpunctata L. K Mordellistena pustulata (Melsheimer) B,D,K,L undetermined spp. K Mordellistena spp. B,D,K,L Cucujidae Mordellistena trifasciata (Say) B,D,K Telephanus velox Haldeman B,K undetermined spp. B,L Curculionidae Nitidulidae Apion spp B,D,K,L Carpophi/us sayi Parsons K Attelabus nigripes LeConte K Colopterus spp. K Brachybamus electus Germar B,L Prometopia spp. K Conotrechalus anaglypticus (Say) D,I,K undetermined spp. B,K Conotrechalus juglandis LeConte K Pedilidae Cyrtepistomus castaneus (Roel.) I,K,L Pedi/us spp. D fugnamptus collaris (Fabricius) K,L Phalacridae Hypera postica (Gyllenhal) K Olibrus spp. B,D,K,L _ixus concavus Say B,D Phalaerus spp. B,D,K,L spp. I Stilbus spp. B,D,K,L ~- ~ -ncr!. lMTtus (L.) K,L undetermined spp. D,K,L - .) K Scarabaeidae s Sa K Mocrodactylus subspinosus (Fabricius) K,L B,K,L Onthaphagus pennsylvanicus Harris B 6,D.K,L Popillia j aponica Newman B,D,I,K,l StrigodemlO arboricokI (Fabricius) K :.IS pige ab' ·... 5) K L

L

March, 1996 . ;I\ 'SLITTER ofthe. OGICAL socrrn' '4 16 • MES Homepage on the WWW: ht1p:IIin.sff:s.u rr: mL:.sa.t:.l:!:.!ch.ed IDES rr.es.htm: •• Staphylinidae Pyrgotidae Oxyporus spp. D,K,L Pyrgota undata Wiedemann K Tenebrionidae Rhagionidae Meracantha contracta (Beauvois) Rhagio spp. B,D,K,L Undetermined family Sarcophagidae undetermined spp. B,K,L Camptopyga spp. Sciarldae MECOPTERA (20 specimens) undetermined spp. B,D,K,L Panorpidae Sciomyzidae Panorpa spp. D,K,L Antichaeta melanosoma Melander K Antichaeta spp. K DIPTERA (3,800 specimens) Dictya spp. B,D,L Asilidae Euthycera arcuata (Loew) B,D,K Efferia spp. K,L Limnia boscii Robineau-Desvoidy B,D Leptogaster spp. L Limnia spp. B,D,K,L undetermined spp. I,K,L Pherbellia seticoxa Steyskal K Bibionidae Pherbellia spp. B,D undetermined spp. L Poedlographa decora (Loew) B,D Bombyliidae Sepedon fuscipennis Loew L Exoprosopa spp. Sepedon spp. B,D,K Geron spp. Tetanocera valida Loew D Calliphoridae Tetanocera spp. B,D,I,K,L Phaenicia serricata (Meigen) B,K Trypetoptera canadensis Macquart B,K,L undetermined spp. B,K,L Sepsidae Chironomldae undetermined spp. B,D,K,L undetermined spp. L StraUomyidae Chloropidae Stratiomys meigenii Wiedemann B,L undetermined spp. B,D,K,L Syrphidae Conopidae . Allograpta obliqua Say Thecophora abbreviata (Loew) D Baccha costata Say L Culicidae Baccha spp. L Psorophora sp p. D Eristalis arbostorum (L.) K undetermined spp. B,D,K,L Mesograpta marginata Say B,K,L Diopsidae Metasyrphus americanus (Wiedemann) D Sphyracepha/us brevicornis Say B,D,K Microdon fu/gens Dollchopodidae Platycheirus erraticus (Zetterstedt) B,K un determined spp. B,D,K,L P/atycheirus immarginatus (Zetterstedt) B,K Drosophilldae Toxomerus geminatus (Say) B,D,K,L un determined sp p. B, D,K,L undetemined spp. B,D,K,L Lauxanlidae Tabanidae Ho moneura spp. (d. bispina (loew» L Chrysops spp. B,D,K,L Lonchopteridae Hybomitra spp. B,D Lonchoptera spp. B,D,K,L Tabanus atratus Fabr. I Micropezidae Tachinidae undetermined spp. B, D,K un determined spp. B,D,K,L Muscidae Tephrytldae un determined spp. B,D,K,L Euaresta bella (Loew) B,D,K,L Mycetophagidae Eurosta cribrata (Van der Wulp) B undeterm ined spp. K,L Euaresta festiva (Loew) D Otltidae Euleia fratria (Loew) D Callopistromyia annulipes Macquart D Orellia pa/posa (Loew) L Chaetopsis massy/a Walker D,K Paraxyna albiceps (Loew) D Chaetopsis spp. Paraxyna piccio/a (Bigot) B (anea Weideman or fulvifrons Macquart) B, D,K Procecidochares atra (Loew) D Chaetopsis spp. B,D,L Trupanea actinobola Loew B,D,L De/phinia spp. B,B,L Trupanea spp. B Herina rufipes Van der Wulp D undetermined spp. B,D,K,L Tritoxa f1exa Weidemann B,D TIpulidae Tritoxa incurva Loew B undetermined spp. B,D,K,L Pipunculldae Xytophagidae Pipuncu/a spp. B,D,K,L Xylophagus spp. K Tomosvaryella spp. B,L Platystomatldae HYMENOPTERA (1 ,900 specimens) Rivella brevifasciata Johnson K,L Apldae: Aplnae Rivella viridulans Robineau-Desvoidy B,D,K,L Apis mellifera L. D,I,K Rivella spp. B,K Bambus impatiens I Psilidae Bambus spp. B,D,K Chyliza notata Loew K Pyrobombus vagans I Loxacera cylindrica Say K Apldae: Xylocoplnae Psilla spp. B,D,K,L Ceratina spp. B,D,K,L March, 1996 NEWSLETTER of the MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) •• MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html • 17 Bethylldae Vespidae: Polistinae undetermined spp. D,K,L Polistes fuscatus (Fabricius) B,I,K Braconidae Vespidae: Vespinae Chelonus spp. L Vespula mawlata Rogus terminalis Cresson I Vespula maculifrons (Buysson) B,K undetermined spp. K Vespidae: undetermined family Ceraphronidae Psithyrus ashtoni L undetermined spp. B,D,L Chalcididae LEPIDOPTERA (183 specimens) undetermined spp. B,D,K,L Arctiidae Chrysididae Haploa lecontei Guerin-Meneyille B undetermined spp. K Ctenuchidae Colletidae Gsseps fulvicollis Hbn. Hy/aeus spp. B,K,L Ctenucha virginica Charpentier D Cynipidae Geometridae undetermined spp. B,K,L Erastria co/oraria (Fabricius) L Diaprildae Euchlaena serrata Drury K undetermined spp. B,D,K,L Xanthotype I.Irticaria Drury D,I Encrytidae Hesperiidae undetermined spp. L Epargyreus clarus Cramer I, Eulophidae Erynnis juvenalis Fabricius I undetermined spp . B,D,K,L Poanes hobomok Harris L Eupetmidae Polites mystic Edwards B undetermined spp. B,D,K,L Thorybes pylades Bell L Eurytomidae Thymelius lineola B,K,L undetermined spp. D,K,L Wallengrenia egeremet Scudder I Formicidae Lycaenidae Aphaenogaster rudis (Emery) B,D,K,L Ce/astrina ladon B,K Camponotus ferrugineus (Fabricius) B Everes comyntas Godart I Camponotus ligniiperde I Noduidae Camponotus pennsylvanicus DeGeer B Fettia jaculifera Gn. Crematogaster lineolata (Say) B,D,K,L Nymphalidae Formica exsectoides Forel B,K,L Boloria bellona Thunberg K Formica fusca L. B,D,K,L Euphydryas phaeton Drury B,D, Formica sanguinea Limenitis archippus Cramer B,D,I Lasius alienus (Foerster) B,D,K,L Phycoides tharos Drury D,I,K,R . Prenolepus imparis (Say) K,L Polygonia interrogatonis Fabricius D Tetramorium caespitum (L.) B,D,K,L Speyeria aphrodite Fabricius I undetermined spp. B,D,K,L Speyeria cybele Fabricius D,I,K,R Gasteruptiidae Vanessa atalanta (L.) B,K undetermined spp. K Vanessa virginiensis Drury B,K Halictidae Papl\ionidae Augochlora spp. Papilio troilus L. L undetermined spp. B,D,K,L Pieridae Ichneumonldae Colias eurytheme Boisduyal I undetermined spp. B,D,K,L Colias philodice Godart I,K Leucospidae Pieris rapae L. I Leocospis affinis LeConte B,K Pyralidae Pteromalldae Alypia odomaculata Strom D Perilampus hyalinus Say B,K,L Satyridae undetermined spp. B,D,K,L Cercyonis pegala nephele Fabricius I Proctotrupidae Euptychia cymela Cramer I undetermined spp. B,D,K,L Lethe eurydice Johannson B Scelionidae Sphingidae undetermined spp. D,K Amphion floridensis I Siricidae Eumorpha pandorus Hubner K Tremex columbia (L.) K Undetermined Family Sphecidae undetermined spp. B,D,K,L " america a Spinola

* Sites: B = Bakertown Fen, Berrien County; D = Dayton Wet Prairie, Berrien County; G = Grand Beach Marsh, Berrien County; B,D,K,l I = Iyes Road Fen, Lenawee County; K= - Klumbis Road Prairie, Cass County; L = Lawton Prairie, Van Buren County; 0 = Ore-Ida raine, Ne'Naygo County; P = Thompson Road Prairie, Cass Co my; R = ree R' ers State Game Area, St. Joseph County ; ~a T = - a ra~ Swam , Cass County.

• March, 1996 NEWSLETTER of the " flCHlGAl " [1 ;rOMOLOGlCAL SOCIETY V41(1) 18 • MES Homepage on the WWW: http://insects.ummzJsa.umich.edu/mes/mes.html II • Biological Control in Glasshouses at the Kew Gardens Director, Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, UK ..

Each year up to a third of the world's crops are lost as a result • Scale insects are controlled using the paraSitic wasp, of damage caused by pests and diseases. Although this damage Metaphycus helvolus, Which lays its eggs in the pest's canhe greatly reduced byusing chemical pesticides to control the nymphal stage. harmful pests, other methods, less detrimental to the environ­ • Thrips are sprayed with the pathogenic fungus, Verticil­ ment, are becoming increasingly popular. Biological control is lium lecanii, which permeates the body of the pest. one such methodand operates byha rnessing the activities ofone • Aphids are attacked by the predatory midge Aphidoletes living organism to control another living orga nism - the pest. aphidiomyza and the paraSitic wasp Aphldius matricariae The control organisms may be predators which eat various which lays its eggs in adult a phids. Naturally occurring stages ofthe pest; parasi tes Iiving on orin the pestandeventually ladybirds, lacewings and hover flies also control aphids. killing it; or pathogen s, such as bacteria, fungi and viruses, Plants such as Convolmollus tricolor are sometimes which cause disease in the pest. grown in the nursery glasshouse to attract these Currently biological control is widely used against various beneficial insects insects, particularly glasshouse pests including aphids, white­ • Carnation tortrix moth caterpilla r (Cacoedmorph flies, spider mites, mealybugs, and scale insects. It can also be pronubana) is controlled using the bacterium Bacillus employedto control a range of weeds, plantandanimal diseases, thuringiensis, which produces protein crystals toxic to snails a nd other organisms. Biological control is often integrated some species of bu tterflies and moths. with other control methods such as non-persistent pesticides, • The two-spotted spider mite (fetrarhynchus urticae) is resistant plant varieties and culturpl techniques. controlled by introducing the predatory South Ameri­ Biological pest con trol was first introduced a t Kew during the can mite Phytoseiulus persimilis. 1970's in a few small areas such as the WaterliIy House. Then, in • Vine weevil (Otiorhynchus sulcatus) is a ttacked by 1987, an intensive program of biological pest control was estab­ Heterorhabitis nematodes. lished in the research glasshouse where plants are raised for • Sciarid flies are controlled by the nematode Dteinemema biochemical studies. As some of these plants are being investi­ bibionis. gated as sources of chemicals with anti-insect properties, the results ofthe experiments would be misleading ifthe plants were Other biologica l control agents are tested regularly a t Kew. contaminatedwith chemicals pesticides. Biological control meth­ For example, Chinese painted quail have been introduced into ods are now used in both the nursery and display glasshouses. the n ursery glasshouses to find out whether they would control Biological control requires careful monitoring ofpest infesta­ cockroaches. So far, the quail have proved to give effident weed tions so that the introduction of the control agent can be timed control in the glasshouses. for maximum effectiveness. Careful checking of tempera ture, humidity and light levels is also necessary to ensure tha t condi­ Integrated Pest Management at Kew tions are suitable for the pest control agent. Levels of the control agents are also monitored so that they can be reintroduced early Biological control is just one of a variety of pest control methods integrated throughoutthe glasshouses to protect Kew's enough to maintain adequate control. Here are some examples of pest problems at the Kew Gardens and the biological control diverse plant collections from pests and diseases. The aim is to maintain pest populations at levels below those causing unac­ agents used to suppress them. ceptable levels of damage. Other methods being used include traps, cultural techniques, • Whitefly (Trialeuroides vaporariorum) are controlled using and compatible pesticides. For example, sticky traps are used in the paraSitic wasp (Encarsia formosa) . nursery glasshouses to control whiteflies and are also an essen­ • Some species of mealybugs including Pseudococcus tial method for monitoring infestations. Tape placed on benches obscurus are controlled with the larvae and adults of the is used to trap thrips. Spraying water over plant foliage helps to Australian predatory ladybird beetle, Cryptolaemus reduce the spread of spider mites. montrouzieri, which will also feed on scale insects. The For more information consult: Upson, T.M. 1991. Biological citrus mealybug (Planococcus citri) is controlled with the control at the Royal Botanic Gardens, Kew. Professional Horticul­ parasite Leptomastix dactylopii. ture, 5: 10-14

March, 1996 NEWSLETTER ofthe MICHIGAN ENTOMOLOGICAL SOCIETY V41(1) MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html 19 Open House at the MSU Butterfly House

The Botany Greenhouse and Butterfly 355-0229. Last year's Open House drew end as well. Any help will be greatly ap­ House Will be having an Open House from hundreds of people and I anticipate an­ preciated. 8:00am to 5:00pm on Saturday April 13 other record turnout. The Natural Science and Sunday April 14 on the Michigan Open House is the following weekend and John MU99, Manager State University campus. The Butterfly will presumablydrawanotherlarge crowd. MSU Botany Greenhouse House is in the greenhouse complex, just Volunteers are also needed for that week- and Butterfly House northwest ofthe Natural Science Building on the MSU campus. For an area map, please look at page 16 of the December 1995 issue of the MES Newsletter. We Will be highlighting butterfly gar­ Update on the MES Homepage dening, wetlands conservation and possi­ bly herbs. Volunteers are needed to help During the past three months (6 December 1995 to 26 February 1996), interpret the Butterfly House displays. the MES Homepage on the World Wide Web has been "hit" (accessed) by Also I'm looking to put together a display about 800 users from at least 40 countries worldwide. Check out our on insect collecting techniques With inter­ Society's Homepage if you haven'talready. It includes information on the pretive signs. If anyone has materials SOCiety, the governing board, how to join, and MES publications. In or ideas, please e-mail me at addition, all of the MES Entomology Notes can also be read and down­ [email protected] or call (517) loaded. The address of the MES Homepage is

http://lnsects.ummz.lsa.umich .edu /mes/m es.html

and it Will be printed from now on in all issues of the MES Newsletter (bottom of every page) and in our Society's journal The Great Lakes Entomologist. Please send comments and suggestions to Mark O'Brien (e-mail: [email protected]).

MICHIGAN ENTOMOLOGIC AL SOCIETY NONPROFIT ORG. U. S. POSTAG E

DE PART MENT OF EMTO MOLOGY PAID MICHIG AN STATE UNIVERSITY EAST LANSING, MI EAST LANSING, MI CHIGAN 48 82 3 PERMIT NO. 14

ADDR ESS CO RRECT ION REQ UESTE D

MES Homepage on the WWW: http://insects.ummz.lsa.umich.edu/mes/mes.html