Genozoic Mammal Horizons of Western North America

Total Page:16

File Type:pdf, Size:1020Kb

Genozoic Mammal Horizons of Western North America DEPARTMENT OF THE INTERIOR UNITED STATES GEOLOGICAL SURVEY GEORGE OTIS SMITH, DIRECTOR 361 GENOZOIC MAMMAL HORIZONS OF WESTERN NORTH AMERICA BY HENRY FAIRFIELD OSBORN WITH FAUNAL LISTS OF THE TERTIARY MAMMALIA OF THE WEST WILLIAM D1LLER MATTHEW WASHINGTON GOVERNMENT PRINTING OFFICE 1909 CONTENTS. Page. INTRODUCTION ........................................................... 7 Formations and zones................................................. 7 Life zones........................................................ 7 Geologic formations...........................'..................... 7 '. Correlation................................................................. 8 Bibliography........................................................... 9 CHAPTER I. General geologic and climatic history of the Tertiary............ 19 The Mountain Region................................................. 19 The Plains Region..................................................... 20 Resemblances and contrasts between Mountain and Plains regions........ 21 Resemblances..................................................... 21 Contrasts.......................................................... 21 Geologic history of Mountain basin deposits of the Eocene and Oligocene. 24 Geologic history of the Great Plains deposits of the .Oligocene to lower Pleistocene.......................................................... 26 Extent............................................................ 26 History of opinion as to mode of deposition.......................... 26 Summary.......................................................... .28 CHAPTER II. Time correlation of mammal-bearing horizons................... 29 The two grand problems............................................... 29 American correlation............................................... 29 American and Eurasiatic correlation................................. 29 Methods of correlation.............'.................................... 30 Bases............................................................. 30 Sources of error.................................................... 30 Preliminary correlation of the Eocene and Oligocene Mountain deposits... 31 Preliminary correlation of the Oligocene to lower Pleistocene Mountain and Plains deposits...................i.................................. 31 CHAPTER III. Western American Cenozoic horizons......................... 33 Eocene................................................................ 33 I. First faunal phase.............................................. 33 Post-Cretaceous or basal Eocene (e"tage Thane"tien)................ 33 1. Puerco formation; Poiymastodon zone..................... 33 2. Torrejon formation; Pantolambda zone................... 34 II. Second faunal phase............................................ 35 Lower Eocene (6tages Sparnacien, Yprdsien).................... 36 3. Wasatch formation; Coryphodon zone...................... 36 3a. Wasatch of the Bighorn basin .......................... 41 III. Third faunal phase............................................ 42 Lower to middle Eocene (etages Ypr<§sien, Lut^tien infdrieur)..... 43 4. Wind River formation; Lambdotherium and Bathyopsis zones.................................................... 43 4a. Huerfano formation; Lambdotherium and ? Uintatherium zones.................................................... 48 3 91406 4 CONTENTS. CHAPTER III. Western American Cenozoic horizons Continued. ' Page. Eocene Continued. III. Third faunal phase Continued. Middle Eocene (Stages Lute"tien supe"rieur, Bartonien)........... 50 5. Bridger formation; Orohippus and Uintatherium zones..... 50 Middle to upper Eocene (<§tage Bartonien)...................... 53 Later Eocene deposits of Washakie basin; Uintatherium and Eobasileus zones......................................... 53 Upper Eocene (Stages Bartonien in part, Ludien (Ligurien) in part)........................................................ 54 7. Later Eocene deposits of Uinta basin; Uintatherium, Eobasileus, and Diplacodon zones.......................... 54 Oligocene............................................................. 57 IV. Fourth faunal phase. .^....................................\.... .57 Lower Oligocene, White River group of Hayden (etage Sannoisien [Tongrien infe"rieur])........ =................................. 60 8. Chadron formation; Titanotherium zone.................. 60 Middle Oligocene (e"tage Stampien [Tongrien sup6rieur]).......... 62 9. Lower part of Brule clay; Oreodonzone and " Metamynodon sandstones".............................................. 62 Upper Oligocene,. first phase................................... 63 10. Upper part of Bfule clay; Leptauchenia zone and "Pro- toceras sandstones"...................................... 63 Oregon Cenozoic formations................................. 64 R6sum6 of the Oregon deposits as a whole............... 64 John Day formation.................................... 64 Upper Oligocene, second phase................................. 67 11. Middle part of John Day formation; Diceratherium zone (also upper part of John Day, transitional)................ 67 Upper Oligocene, latest phase.................................. 68 12. Upper part of John Day formation; Promerycochoerus zone.. 68 Miocene.............................................................. 70 IV. Fourth faunal phase Continued ............................... 70 Lower Miocene (Stages Aquitanien, Burdigalien)................ 70 13. Arikaree formation; Promerycochoerus zone............... 70 General features......................................... 70 Westerly section....................................... 73 A. Lower division................................. 73 (a) Gering formation........................... 73 (6) Monroe Creek formation............ ....... 73 (c) Harrison formation, Hatcher................. 73 (d) Harrison formation, Agate Spring quarry.... 74 B. Upper division.................................. 74 (e) Upper part of Harrison formation........... 74 Easterly section....................................... 74 A. Lower part of Rosebud.......................... 75 B. Upper part of Rosebud.....:................... 75 V. Fifth faunal phase........ '.'.".................................... 76 Middle Miocene (Stages Helvetien, Sarmatien, Tortonien)........ 76 Faunal changes........................................... 76 14. Deep River sequence; Ticholeptus zone................. 76 Upper Miocene (e"tage Pontien).................................. 79 15. Ogalalla formation (in part); Procamelus zone........... 79 : - CONTENTS. 5 . CHAPTER III. Western American Cenozoic horizons Continued. Page. * Miocene Continued. £! V. Fifth faunal phase Continued. Last phase of Miocene of first phase 0- Pliocene (6tages Pontien, Messinien).................................................. 80 \ 16. Ogalalla formation (in part); Peraceras zone............. 80 \ 16a. Rattlesnake formation of John Day Valley, Oregon...... 81 i Pliocene............................................................... 81 ' VI. Sixth faunal phase............................................ 82 Middle Pliocene or second phase (e"tage Astien).................. 82 17. Blanco formation; Glyptotherium zone.................... 82 '* Upper Pliocene or lower Pleistocene............................ 83 18. Elephas imperator zone.................................. 83 Pleistocene............................................................ 84 \ VII. Seventh faunal phase.......................................... 84 * Lower Pleistocene (preglacial)................................... 84 ^ 19. Equus zone............................................ 85 i Middle Pleistocene (glacial)..................................... 86 / General characters.......................................... 86 Early phases....................... ...................... 87 I Subsequent phases......................................... 88 > Late phase................................................ 90 i Conclusion. ............................................................. 90 APPENDIX. Faunal lists of the Tertiary Mammalia of the West, by W. D. ? Matthew................................................................ 91 ' INDEX. ................................................................... 121 ILLUSTRATIONS. Page, PLATE I. Map of the United States, showing general Mountain and Great Plains regions, also location of the principal formations, sections, and deposits considered in this paper.........."....................... 7 II. Oligocene and Miocene exposures in South Dakota, northwestern Nebraska, and eastern Wyoming ................................. 60 III. Idealized bird's-eye view of the great Badlands of South Dakota, showing channel and overflow deposits in the Oligocene and lower Miocene....................................................... 64 FIG. 1. Composite section of the Tertiary deposits of the West................ 23 .2. Map of southwestern Wyoming and northern Utah, showing partial areas of the Wasatch, Wind River, Bridger, and Uinta formations.... 37 3. Composite columnar section of the Wasatch formation of Bighorn Basin.. 38 4. Columnar section showing the relations of the typical Wasatch section, including the Knight formation, to the overlying and underlying formations........................................................ 39 5. Columnar section of the Wind River basin, based on the descriptions of Hayden and Loomis.............................................. 44 6.
Recommended publications
  • Summary of the Paleontology of the Santa Fe Group (Mio-Pliocene), North-Central New Mexico Barry S
    New Mexico Geological Society Downloaded from: http://nmgs.nmt.edu/publications/guidebooks/30 Summary of the paleontology of the Santa Fe Group (Mio-Pliocene), north-central New Mexico Barry S. Kues and Spencer G. Lucas, 1979, pp. 237-241 in: Santa Fe Country, Ingersoll, R. V. ; Woodward, L. A.; James, H. L.; [eds.], New Mexico Geological Society 30th Annual Fall Field Conference Guidebook, 310 p. This is one of many related papers that were included in the 1979 NMGS Fall Field Conference Guidebook. Annual NMGS Fall Field Conference Guidebooks Every fall since 1950, the New Mexico Geological Society (NMGS) has held an annual Fall Field Conference that explores some region of New Mexico (or surrounding states). Always well attended, these conferences provide a guidebook to participants. Besides detailed road logs, the guidebooks contain many well written, edited, and peer-reviewed geoscience papers. These books have set the national standard for geologic guidebooks and are an essential geologic reference for anyone working in or around New Mexico. Free Downloads NMGS has decided to make peer-reviewed papers from our Fall Field Conference guidebooks available for free download. Non-members will have access to guidebook papers two years after publication. Members have access to all papers. This is in keeping with our mission of promoting interest, research, and cooperation regarding geology in New Mexico. However, guidebook sales represent a significant proportion of our operating budget. Therefore, only research papers are available for download. Road logs, mini-papers, maps, stratigraphic charts, and other selected content are available only in the printed guidebooks. Copyright Information Publications of the New Mexico Geological Society, printed and electronic, are protected by the copyright laws of the United States.
    [Show full text]
  • New Materials of Chalicotherium Brevirostris (Perissodactyla, Chalicotheriidae)
    Geobios 45 (2012) 369–376 Available online at www.sciencedirect.com Original article New materials of Chalicotherium brevirostris (Perissodactyla, Chalicotheriidae) § from the Tunggur Formation, Inner Mongolia Yan Liu *, Zhaoqun Zhang Laboratory of Evolutionary Systematics of Vertebrate, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142, Xizhimenwai Street, PO Box 643, Beijing 100044, PR China A R T I C L E I N F O A B S T R A C T Article history: Chalicotherium brevirostris was named by Colbert based on a skull lacking mandibles from the late Middle Received 23 March 2011 Miocene Tunggur Formation, Tunggur, Inner Mongolia, China. Here we describe new mandibular Accepted 6 October 2011 materials collected from the same area. In contrast to previous expectations, the new mandibular Available online 14 July 2012 materials show a long snout, long diastema, a three lower incisors and a canine. C. brevirostris shows some sexual dimorphism and intraspecific variation in morphologic characters. The new materials differ Keywords: from previously described C. cf. brevirostris from Cixian County (Hebei Province) and the Tsaidam Basin, Tunggur which may represent a different, new species close to C. brevirostris. The diagnosis of C. brevirostris is Middle Miocene revised. Chalicotheriinae Chalicotherium ß 2012 Elsevier Masson SAS. All rights reserved. 1. Abbreviations progress in understanding the Tunggur geology and paleontology (Qiu et al., 1988). The most important result of this expedition so far is
    [Show full text]
  • Carnivora from the Late Miocene Love Bone Bed of Florida
    Bull. Fla. Mus. Nat. Hist. (2005) 45(4): 413-434 413 CARNIVORA FROM THE LATE MIOCENE LOVE BONE BED OF FLORIDA Jon A. Baskin1 Eleven genera and twelve species of Carnivora are known from the late Miocene Love Bone Bed Local Fauna, Alachua County, Florida. Taxa from there described in detail for the first time include the canid cf. Urocyon sp., the hemicyonine ursid cf. Plithocyon sp., and the mustelids Leptarctus webbi n. sp., Hoplictis sp., and ?Sthenictis near ?S. lacota. Postcrania of the nimravid Barbourofelis indicate that it had a subdigitigrade posture and most likely stalked and ambushed its prey in dense cover. The postcranial morphology of Nimravides (Felidae) is most similar to the jaguar, Panthera onca. The carnivorans strongly support a latest Clarendonian age assignment for the Love Bone Bed. Although the Love Bone Bed local fauna does show some evidence of endemism at the species level, it demonstrates that by the late Clarendonian, Florida had become part of the Clarendonian chronofauna of the midcontinent, in contrast to the higher endemism present in the early Miocene and in the later Miocene and Pliocene of Florida. Key Words: Carnivora; Miocene; Clarendonian; Florida; Love Bone Bed; Leptarctus webbi n. sp. INTRODUCTION can Museum of Natural History, New York; F:AM, Frick The Love Bone Bed Local Fauna, Alachua County, fossil mammal collection, part of the AMNH; UF, Florida Florida, has produced the largest and most diverse late Museum of Natural History, University of Florida. Miocene vertebrate fauna known from eastern North All measurements are in millimeters. The follow- America, including 43 species of mammals (Webb et al.
    [Show full text]
  • New Postcranial Specimens of the Anthracotheriidae (Mammalia; Artiodactyla) from the Paleogene of Fayum Depression, Egypt
    International Journal of Scientific Engineering and Applied Science (IJSEAS) - Volume-1, Issue-8,November 2015 ISSN: 2395-3470 www.ijseas.com New postcranial specimens of the Anthracotheriidae (Mammalia; Artiodactyla) from the Paleogene of Fayum Depression, Egypt 1 2 Afifi H. Sileem , Abdel Galil A Hewaidy 1 Vertebrate paleontology section, Cairo Geological Museum, Cairo, Egypt, [email protected] 2Department of Geology, Faculty of Science, Al-Azhar University, Egypt, <[email protected]> Abstract: The fossiliferous deposits exposed north of Birket Qarun in the Fayum Depression, northeast Egypt, have produced a remarkable collection of fossil mammals from localities that range in age from earliest late Eocene (~37 Ma) to latest early Oligocene (~29 Ma). Anthracotheriidae are among the most common mammals that are preserved in these deposits. Here we describe a new fossil specimens of the Anthracotheriidae (Mammalia, Artiodactyla) discovered in the Jebel Qatrani Formation of Fayum. The specimens consist of a seven astragalus, which is referable to Bothriogenys sp. from the formation. The specimens Bothriogenys sp. show a higher degree of size variation and some feature suggest that the anthracothere are not closely related to Hippopotamus. Key word: anthracothere, Bothriogenys; astragalus; Fayum; Early Oligocene. 376 International Journal of Scientific Engineering and Applied Science (IJSEAS) - Volume-1, Issue-8,November 2015 ISSN: 2395-3470 www.ijseas.com Introduction: The fossiliferous sedimentary deposits exposed north of Birket (lake) Qarun in the Fayum Depression (Fig.1), northeast Egypt, have produced a remarkable collection of a wide variety of fish, amphibian, reptile, bird and mammal taxa (e.g. Andrews, 1906; Simons and Rasmussen, 1990; Murray et al.
    [Show full text]
  • Three-Toed Browsing Horse Anchitherium (Equidae) from the Miocene of Panama
    J. Paleonl., 83(3), 2009, pp. 489-492 Copyright © 2009, The Paleontological Society 0022-3360/09/0083-489S03.00 THREE-TOED BROWSING HORSE ANCHITHERIUM (EQUIDAE) FROM THE MIOCENE OF PANAMA BRUCE J. MACFADDEN Florida Museum of Natural History, University of Florida, Gainesville FL 32611, <[email protected]> INTRODUCTION (CRNHT/APL); L, left; M, upper molar; R upper premolar; R, DURING THE Cenozoic, the New World tropics supported a rich right; TRN, greatest transverse width. biodiversity of mammals. However, because of the dense SYSTEMATIC PALEONTOLOGY vegetative ground cover, today relatively little is known about extinct mammals from this region (MacFadden, 2006a). In an Class MAMMALIA Linnaeus, 1758 exception to this generalization, fossil vertebrates have been col- Order PERISSODACTYLA Owen, 1848 lected since the second half of the twentieth century from Neo- Family EQUIDAE Gray, 1821 gene exposures along the Panama Canal. Whitmore and Stewart Genus ANCHITHERIUM Meyer, 1844 (1965) briefly reported on the extinct land mammals collected ANCHITHERIUM CLARENCI Simpson, 1932 from the Miocene Cucaracha Formation that crops out in the Gail- Figures 1, 2, Table 1 lard Cut along the southern reaches of the Canal. MacFadden Referred specimen.—UF 236937, partial palate (maxilla) with (2006b) formally described this assemblage, referred to as the L P1-M3, R P1-P3, and small fragment of anterointernal part of Gaillard Cut Local Fauna (L.E, e.g., Tedford et al., 2004), which P4 (Fig. 1). Collected by Aldo Rincon of the Smithsonian Tropical consists of at least 10 species of carnivores, artiodactyls (also see Research Institute, Republic of Panama, on 15 May 2008.
    [Show full text]
  • The Carnivora (Mammalia) from the Middle Miocene Locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina)
    Palaeobiodiversity and Palaeoenvironments https://doi.org/10.1007/s12549-018-0353-0 ORIGINAL PAPER The Carnivora (Mammalia) from the middle Miocene locality of Gračanica (Bugojno Basin, Gornji Vakuf, Bosnia and Herzegovina) Katharina Bastl1,2 & Doris Nagel2 & Michael Morlo3 & Ursula B. Göhlich4 Received: 23 March 2018 /Revised: 4 June 2018 /Accepted: 18 September 2018 # The Author(s) 2018 Abstract The Carnivora (Mammalia) yielded in the coal mine Gračanica in Bosnia and Herzegovina are composed of the caniform families Amphicyonidae (Amphicyon giganteus), Ursidae (Hemicyon goeriachensis, Ursavus brevirhinus) and Mustelidae (indet.) and the feliform family Percrocutidae (Percrocuta miocenica). The site is of middle Miocene age and the biostratigraphical interpretation based on molluscs indicates Langhium, correlating Mammal Zone MN 5. The carnivore faunal assemblage suggests a possible assignement to MN 6 defined by the late occurrence of A. giganteus and the early occurrence of H. goeriachensis and P. miocenica. Despite the scarcity of remains belonging to the order Carnivora, the fossils suggest a diverse fauna including omnivores, mesocarnivores and hypercarnivores of a meat/bone diet as well as Carnivora of small (Mustelidae indet.) to large size (A. giganteus). Faunal similarities can be found with Prebreza (Serbia), Mordoğan, Çandır, Paşalar and Inönü (all Turkey), which are of comparable age. The absence of Felidae is worthy of remark, but could be explained by the general scarcity of carnivoran fossils. Gračanica records the most eastern European occurrence of H. goeriachensis and the first occurrence of A. giganteus outside central Europe except for Namibia (Africa). The Gračanica Carnivora fauna is mostly composed of European elements. Keywords Amphicyon . Hemicyon .
    [Show full text]
  • Paleocene Mammalian Faunas of the Bison Basin in South-Central Wyoming
    SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 131, NUMBER 6 Cftarlesi ©, anb JHarp ^aux SKHalcott 3Res!earcf) Jf unb PALEOCENE MAMMALIAN FAUNAS OF THE BISON BASIN IN SOUTH-CENTRAL WYOMING (With 16 Plates) By C. LEWIS GAZIN Curator, Division of Vertebrate Paleontology United States National Museum Smithsonian Institution (Publication 4229) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION FEBRUARY 28, 1956 THE LORD BALTIMORE PRESS, INC. BALTIMORE, MD., U. S. A. CONTENTS Page Introduction i Acknowledgments 2 History of investigation 2 Occurrence and preservation of material 3 The Bison basin faunas 4 Environment and relationships between the Bison basin faunas 7 Age and correlation of the faunas lo Systematic description of vertebrate remains I2 Reptilia 12 Sauria 12 Anguidae 12 Mammalia 12 Multitubcrculata 12 Ptilodontidae 12 Marsupialia 14 Didelphidae 14 Insectivora 15 Leptictidae 15 Pantolestidae 17 Primates 19 Plesiadapidae 19 Carnivora 25 Arctocyonidae 25 Mesonychidae 35 Miacidae 35 Condylarthra 36 Hyopsodontidae 36 Phenacodontidae 42 Pantodonta 47 Coryphodontidae 47 References 51 Explanation of plates 54 ILLUSTRATIONS Plates (All plates following page 58.) 1. Mullituberculates and insectivores from the Bison basin Paleocene. 2. Primates and marsupials from the Bison basin Paleocene. 3. Pronothodcctes from the Bison basin Paleocene. 4. Plcsiadapis from the Bison basin Paleocene. 5. Triccntcs and Chriacus from the Bison basin Paleocene. 6. Thryptacodon from the Bison basin Paleocene. 7. Clacnodon from the Bison basin Paleocene. 8. Litomylus and Protoselcnc? from the Bison basin Paleocene. 9. Haplaletes and Gidleyina from the Bison basin Paleocene. 10. Phcnacodusl from the Bison basin Paleocene. 11. Condylarths and Titanoidcs from the Bison basin Paleocene. 12. Caenolambda from the Bison basin Paleocene.
    [Show full text]
  • Unit-V Evolution of Horse
    UNIT-V EVOLUTION OF HORSE Horses (Equus) are odd-toed hooped mammals belong- ing to the order Perissodactyla. Horse evolution is a straight line evolution and is a suitable example for orthogenesis. It started from Eocene period. The entire evolutionary sequence of horse history is recorded in North America. " Place of Origin The place of origin of horse is North America. From here, horses migrated to Europe and Asia. By the end of Pleis- tocene period, horses became extinct in the motherland (N. America). The horses now living in N. America are the de- scendants of migrants from other continents. Time of Origin The horse evolution started some 58 million years ago, m the beginning of Eocene period of Coenozoic era. The modem horse Equus originated in Pleistocene period about 2 million years ago. Evolutionary Trends The fossils of horses that lived in different periods, show that the body parts exhibited progressive changes towards a particular direction. These directional changes are called evo- lutionary trends. The evolutionary trends of horse evolution are summarized below: 1. Increase in size. 2. Increase in the length of limbs. 3. Increase in the length of the neck. 4. Increase in the length of preorbital region (face). 5. Increase in the length and size of III digit. 6. Increase in the size and complexity of brain. 7. Molarization of premolars. Olfactory bulb Hyracotherium Mesohippus Equus Fig.: Evolution of brain in horse. 8. Development of high crowns in premolars and molars. 9. Change of plantigrade gait to unguligrade gait. 10. Formation of diastema. 11. Disappearance of lateral digits.
    [Show full text]
  • Perissodactyla: Tapirus) Hints at Subtle Variations in Locomotor Ecology
    JOURNAL OF MORPHOLOGY 277:1469–1485 (2016) A Three-Dimensional Morphometric Analysis of Upper Forelimb Morphology in the Enigmatic Tapir (Perissodactyla: Tapirus) Hints at Subtle Variations in Locomotor Ecology Jamie A. MacLaren1* and Sandra Nauwelaerts1,2 1Department of Biology, Universiteit Antwerpen, Building D, Campus Drie Eiken, Universiteitsplein, Wilrijk, Antwerp 2610, Belgium 2Centre for Research and Conservation, Koninklijke Maatschappij Voor Dierkunde (KMDA), Koningin Astridplein 26, Antwerp 2018, Belgium ABSTRACT Forelimb morphology is an indicator for order Perissodactyla (odd-toed ungulates). Modern terrestrial locomotor ecology. The limb morphology of the tapirs are widely accepted to belong to a single enigmatic tapir (Perissodactyla: Tapirus) has often been genus (Tapirus), containing four extant species compared to that of basal perissodactyls, despite the lack (Hulbert, 1973; Ruiz-Garcıa et al., 1985) and sev- of quantitative studies comparing forelimb variation in eral regional subspecies (Padilla and Dowler, 1965; modern tapirs. Here, we present a quantitative assess- ment of tapir upper forelimb osteology using three- Wilson and Reeder, 2005): the Baird’s tapir (T. dimensional geometric morphometrics to test whether bairdii), lowland tapir (T. terrestris), mountain the four modern tapir species are monomorphic in their tapir (T. pinchaque), and the Malayan tapir (T. forelimb skeleton. The shape of the upper forelimb bones indicus). Extant tapirs primarily inhabit tropical across four species (T. indicus; T. bairdii; T. terrestris; T. rainforest, with some populations also occupying pinchaque) was investigated. Bones were laser scanned wet grassland and chaparral biomes (Padilla and to capture surface morphology and 3D landmark analysis Dowler, 1965; Padilla et al., 1996). was used to quantify shape.
    [Show full text]
  • Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology
    Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology by Paul David Polly B.A. (University of Texas at Austin) 1987 A dissertation submitted in partial satisfaction of the requirements for the degree of Doctor of Philosophy in Paleontology in the GRADUATE DIVISION of the UNIVERSITY of CALIFORNIA at BERKELEY Committee in charge: Professor William A. Clemens, Chair Professor Kevin Padian Professor James L. Patton Professor F. Clark Howell 1993 Hyaenodontidae (Creodonta, Mammalia) and the Position of Systematics in Evolutionary Biology © 1993 by Paul David Polly To P. Reid Hamilton, in memory. iii TABLE OF CONTENTS Introduction ix Acknowledgments xi Chapter One--Revolution and Evolution in Taxonomy: Mammalian Classification Before and After Darwin 1 Introduction 2 The Beginning of Modern Taxonomy: Linnaeus and his Predecessors 5 Cuvier's Classification 10 Owen's Classification 18 Post-Darwinian Taxonomy: Revolution and Evolution in Classification 24 Kovalevskii's Classification 25 Huxley's Classification 28 Cope's Classification 33 Early 20th Century Taxonomy 42 Simpson and the Evolutionary Synthesis 46 A Box Model of Classification 48 The Content of Simpson's 1945 Classification 50 Conclusion 52 Acknowledgments 56 Bibliography 56 Figures 69 Chapter Two: Hyaenodontidae (Creodonta, Mammalia) from the Early Eocene Four Mile Fauna and Their Biostratigraphic Implications 78 Abstract 79 Introduction 79 Materials and Methods 80 iv Systematic Paleontology 80 The Four Mile Fauna and Wasatchian Biostratigraphic Zonation 84 Conclusion 86 Acknowledgments 86 Bibliography 86 Figures 87 Chapter Three: A New Genus Eurotherium (Creodonta, Mammalia) in Reference to Taxonomic Problems with Some Eocene Hyaenodontids from Eurasia (With B. Lange-Badré) 89 Résumé 90 Abstract 90 Version française abrégéé 90 Introduction 93 Acknowledgments 96 Bibliography 96 Table 3.1: Original and Current Usages of Genera and Species 99 Table 3.2: Species Currently Included in Genera Discussed in Text 101 Chapter Four: The skeleton of Gazinocyon vulpeculus n.
    [Show full text]
  • A New Species of the Rhinoceros Alicornops from the Middle Miocene of the Linxia Basin, Gansu, China
    A NEW SPECIES OF THE RHINOCEROS ALICORNOPS FROM THE MIDDLE MIOCENE OF THE LINXIA BASIN, GANSU, CHINA by TAO DENG . A new species of the genus Alicornops, A. laogouense, from Laogou in Hezheng County, Linxia Basin, China, is described. It is mid-sized in the subfamily Aceratheriinae, but is the largest known species of It represents the first discovery of the genus in Asia. The Middle Miocene fauna bearing A. laogouense is lteI1npOlranleOllS with the Dingjiaergou fauna of Tongxin, Ningxia, China, and its age corresponds to MN 6 in Europe. discovery of A. laogouense in the Linxia Basin implies that Alicornops dispersed from Europe to Asia. During 6 times it was relatively widespread throughout Eurasia. Alicornops laogouense lived in open woodland rich in and rivers. WORDS: Linxia Basin, China, Middle Miocene, Rhinocerotidae, Aceratheriinae. HE Late Cenozoic deposits in the Linxia Basin, Gansu, China, are thick and well exposed. Recently, a m""'''''''<> 11' <In fauna consisting of abundant fossils was discovered in this basin in Middle Miocene deposits Laogou in Hezheng County (Text-fig. 1). The deposits consist of greyish yellow, gravelly, generally .dated sands and occasionally hard sandstones. The sands are mostly medium grained and contain fine gravels. The thickness of the deposits is approximately 58 m. The overlying deposits are grey, rusty yellow sandstones and fine gravels. The underlying beds are alternating thin-bedded light and light orange clays. Judging by the percentage of specimens recovered, proboscideans were dominant and rhinocerotids secondary in this fauna. The known fossils include Amphicyon tairumensis, Anchitheriwn gobiensis, herium sp., Hemicyon teilhardi, Hispanotheriwn matritense, Kubanochoerus gigas, Listriodon liensis, Palaeotragus tungurensis, Platybelodon grangeri, Pliopithecus sp., Zygolophodon sp., and rhrnoc:en)s described herein.
    [Show full text]
  • United States
    DEPARTMENT OF THE INTERIOR BULLETIN OF THE UNITED STATES ISTo. 146 WASHINGTON GOVERNMENT Pit IN TING OFFICE 189C UNITED STATES GEOLOGICAL SURVEY CHAKLES D. WALCOTT, DI11ECTOK BIBLIOGRAPHY AND INDEX NORTH AMEEICAN GEOLOGY, PALEONTOLOGY, PETEOLOGT, AND MINERALOGY THE YEA.R 1895 FEED BOUGHTON WEEKS WASHINGTON Cr O V E U N M K N T P K 1 N T I N G OFFICE 1890 CONTENTS. Page. Letter of trail smittal...... ....................... .......................... 7 Introduction.............'................................................... 9 List of publications examined............................................... 11 Classified key to tlio index .......................................... ........ 15 Bibliography ............................................................... 21 Index....................................................................... 89 LETTER OF TRANSMITTAL DEPARTMENT OF THE INTEEIOE, UNITED STATES GEOLOGICAL SURVEY, DIVISION OF GEOLOGY, Washington, D. 0., June 23, 1896. SIR: I have the honor to transmit herewith the manuscript of a Bibliography and Index of North American Geology, Paleontology, Petrology, and Mineralogy for the year 1895, and to request that it be published as a bulletin of the Survey. Very respectfully, F. B. WEEKS. Hon. CHARLES D. WALCOTT, Director United States Geological Survey. 1 BIBLIOGRAPHY AND INDEX OF NORTH AMERICAN GEOLOGY, PALEONTOLOGY, PETROLOGY, AND MINER­ ALOGY FOR THE YEAR 1895. By FRED BOUGHTON WEEKS. INTRODUCTION. The present work comprises a record of publications on North Ameri­ can geology, paleontology, petrology, and mineralogy for the year 1895. It is planned on the same lines as the previous bulletins (Nos. 130 and 135), excepting that abstracts appearing in regular periodicals have been omitted in this volume. Bibliography. The bibliography consists of full titles of separate papers, classified by authors, an abbreviated reference to the publica­ tion in which the paper is printed, and a brief summary of the con­ tents, each paper being numbered for index reference.
    [Show full text]