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Ent12 1 109 118 Sviridov.P65 Russian Entomol. J. 12 (1): 109118 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2003 Ñîâêè (Lepidoptera, Noctuidae) Âëàäèìèðñêîé îáëàñòè Noctuid Moths (Lepidoptera) of Vladimir Area À.Â. Ñâèðèäîâ*, Ì.Â. Óñêîâ**, Â.È. Ãîðüêàâûé** A.V. Sviridov*, M.V. Uskov**, V.I. Gorkavyi** * Çîîëîãè÷åñêèé ìóçåé Ìîñêîâñêîãî ãîñóäàðñòâåííîãî óíèâåðñèòåòà èì. Ì.Â. Ëîìîíîñîâà, óë. Áîëüøàÿ Íèêèòñêàÿ, 6, Ìîñêâà 125009 Ðîññèÿ. * Zoological Museum of Lomonosov Moscow State University, Bolshaya Nikitskaya Str. 6, Moscow 125009 Russia. ** Êàôåäðà çîîëîãèè, Åñòåñòâåííî-ãåîãðàôè÷åñêèé ôàêóëüòåò, Âëàäèìèðñêèé ãîñóäàðñòâåííûé ïåäàãîãè÷åñêèé óíèâåðñèòåò, ïð. Ñòðîèòåëåé, 11, Âëàäèìèð 600024 Ðîññèÿ. ** Department of Zoology, Natural-Geographic Faculty, Vladimir State Pedagogic University, Stroitelei Str. 11, Vladimir 600024 Russia. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: ñîâêè, Lepidoptera, Noctuidae, Âëàäèìèðñêàÿ îáëàñòü, ôàóíèñòèêà, ñïèñîê. KEY WORDS: Noctuidae, Lepidoptera, Vladimir Area, faunistics, check-list. ÐÅÇÞÌÅ. Äàí áàçîâûé ñïèñîê âèäîâ ñîâîê òåð. Ñ ñåâåðà ê Îïîëüþ ïðèìûêàåò åùå îäèí íåáîëü- (Lepidoptera, Noctuidae) Âëàäèìèðñêîé îáëàñòè. Îí øîé ó÷àñòîê Êëÿçüìèíñêî-Íåðëèíñêîé íèçèíû (Ã). ñîäåðæèò 257 âèäîâ, äàííûå î ìåñòîíàõîæäåíèÿõ è Êëèìàò îáëàñòè óìåðåííî-êîíòèíåíòàëüíûé. Ïî ïî ôåíîëîãèè. Ðàáîòà îñíîâàíà íà ðåçóëüòàòàõ îáðà- ïðèðîäíîìó ðàéîíèðîâàíèþ îáëàñòü âñåöåëî âõî- áîòêè ìíîãîëåòíèõ ñáîðîâ (19932002 ãã.) è ìàòå- äèò â ñîñòàâ çîíû ñìåøàííûõ ëåñîâ, ñîñòàâëÿþùåé ðèàëàõ Çîîëîãè÷åñêîãî ìóçåÿ ÌÃÓ. ñåâåðíóþ ïîëîâèíó ñðåäíåé ïîëîñû Åâðîïåéñêîé Ðîññèè, ïî çîîãåîãðàôè÷åñêîìó â Åâðîïåéñêî- ABSTRACT. The basic check-list of Noctuids Moths Ñèáèðñêóþ ïîäîáëàñòü Ãîëàðêòèêè. (Lepidoptera) of Vladimir Area containing 257 species Ðàñòèòåëüíîñòü îáëàñòè [Âëàäèìèðñêàÿ îáëàñòü, is presented. Data on localities and phenology of each 1971; Êóçíåöîâ è äð., 2000] ïðåäñòàâëåíà ëåñàìè species are provided. The article is based upon the long- (îêîëî 42 % ïëîùàäè), ñåíîêîñàìè, ïàñòáèùàìè è term (19932002) collectings and materials of the áîëîòàìè. Áîëåå ïîëîâèíû îáëåñåííîé ïëîùàäè Zoological Museum of MSU. çàíÿòî ñîñíîé, ìåíåå 4,5 % åëüþ, 40 % âòîðè÷- íûìè ìåëêîëèñòâåííûìè ïîðîäàìè. Äóáîâûõ ëåñîâ Ââåäåíèå ñîõðàíèëîñü íåìíîãî, îíè â îñíîâíîì ïðîèçðàñòà- þò ïî äîëèíàì ðåê è íà âîäîðàçäåëàõ ïî íåóäîáíûì Âëàäèìèðñêàÿ îáëàñòü [Âëàäèìèðñêàÿ îáëàñòü, äëÿ ñåëüñêîãî õîçÿéñòâà ìåñòàì. 1971; Æóêîâ, Øèìàíþê, 1966; Ôîìèíöåâà, 1991; Ôàóíà ñîâîê Âëàäèìèðñêîé îáëàñòè äî íàñòîÿ- Êóçíåöîâ è äð., 2000] ðàñïîëîæåíà â öåíòðàëüíîé ùåé ðàáîòû íå èññëåäîâàëàñü è ïîòîìó íå íàøëà ÷àñòè Âîñòî÷íî-Åâðîïåéñêîé ðàâíèíû, ãðàíè÷à ñ îòðàæåíèÿ â ëèòåðàòóðå. Èñêëþ÷åíèå ñîñòàâëÿþò Ìîñêîâñêîé, ßðîñëàâñêîé, Èâàíîâñêîé, Íèæåãîðîä- íåìíîãî÷èñëåííûå óïîìèíàíèÿ âðåäíûõ âèäîâ ñî- ñêîé è Ðÿçàíñêîé îáëàñòÿìè. Áîëüøóþ ÷àñòü âîñ- âîê â ïðèêëàäíîé ëèòåðàòóðå (êàïóñòíàÿ è îçèìàÿ òî÷íîé ãðàíèöû îáëàñòè ñîñòàâëÿåò ð. Îêà, ñ þãî- ñîâêè, ñîâêà-ãàììà) [Àíîí., 1900; Ïåòðîâ, 1928; çàïàäà íà ñåâåðî-âîñòîê òåððèòîðèþ îáëàñòè ïåðå- Ñêîðèêîâà, 1929]. Êðîìå òîãî, èìåþòñÿ óïîìèíà- ñåêàåò åå êðóïíûé ïðèòîê ð. Êëÿçüìà. Ïëîùàäü Âëà- íèÿ íåñêîëüêèõ âèäîâ â ðàáîòàõ, íå ïîñâÿùåííûõ äèìèðñêîé îáëàñòè 29 òûñ. êâ. êì. Þæíàÿ ïîëî- ñïåöèàëüíî ôàóíå ñîâîê îáëàñòè.  ñëó÷àå íåîáõî- âèíà îáëàñòè íà çàïàäå çàíÿòà ñåâåðíîé ÷àñòüþ Ìå- äèìîñòè ìû èõ [Kostrowicki, 1961; Î ñîñòîÿíèè..., ù¸ðñêîé íèçìåííîñòè (À ñì. êàðòó ðèñ.1), à íà 1999] ïðèâîäèì, à âèäû, êàêèì-ëèáî îáðàçîì îòìå- âîñòîêå Êîâðîâñêî-Êàñèìîâñêèì ïëàòî (Á). Êðàé- ÷åííûå â ëèòåðàòóðå, íî íàøèìè èññëåäîâàíèÿìè íå íèé ñåâåðî-çàïàä òåððèòîðèè ïðåäñòàâëÿåò ñîáîé îáíàðóæåííûå, ïðèâîäèì â ñïèñêå áåç ïîðÿäêîâûõ âîñòî÷íóþ îêîíå÷íîñòü Êëèíñêî-Äìèòðîâñêîé ãðÿ- íîìåðîâ. Èõ îáíàðóæåíèå íà òåððèòîðèè îáëàñòè â äû (Â), ñåâåðî-âîñòîê çàíÿò íèçèíàìè êëÿçüìèíñêî- äàëüíåéøåì âïîëíå âîçìîæíî.  õîäå ðàáîòû íàä ãî ëåâîáåðåæüÿ Êëÿçüìèíñêî-Íåðëèíñêîé (Ã) è âûÿâëåíèåì âèäîâîãî ñîñòàâà ñîâîê Âëàäèìèðñêîé Ëóõñêîé (íà êðàéíåì ñåâåðî-âîñòîêå) (Ä).  ñðåäíåé îáëàñòè áûë îïóáëèêîâàí ðÿä ïðåäâàðèòåëüíûõ ñïèñ- ÷àñòè ñåâåðà îáëàñòè, ïðåäñòàâëÿþùåé ñîáîé ìåæ- êîâ îáíàðóæåííûõ íàìè âèäîâ [Ñâèðèäîâ, 1998; äóðå÷üå ëåâûõ ïðèòîêîâ Êëÿçüìû, Ïåêøè è Íåðëè, Óñêîâ è äð., 2000, 2001; Ãîðüêàâûé, Óñêîâ, 2002] è ðàñïîëîæåíà âûñîêàÿ ðàâíèíà Âëàäèìèðñêîå ñòàòüè ñ ïðåäâàðèòåëüíûì àíàëèçîì ôàóíèñòè÷åñ- Îïîëüå (Å), èìåþùàÿ óâàëèñòî-âîëíèñòûé õàðàê- êèõ äàííûõ, ãäå òàêæå óïîìèíàëèñü îòäåëüíûå âèäû 110 à 41 00' â.ä. Þð.-Ïîëüñê. Îìóòñêîå Ñóçä. Ñåðã. Ñà. à Ìñòåðà Êøàðà Ìåò. Å Ìèøí. Ä À.Â. Ñâèðèäîâ, Ì.Â. Óñêîâ, Â.È. Ãîðüêàâûé  Ñóð. Áîã. Ñíîâ. Ãîðîõîâåö Íåâ. ßã. Äàâ. Âëàä. Ñòàâð. Åð. Þð. Ðóê. Êîìì. Ïàòð. Ðàòüê. Øïàë. Èê. Êàä. Ïåðí. Âåð. Ìèòüê. Á Ëàê. Óñò. Ðûæ. Ñóä. Îñòð. Êîñòð. 56 00' ñ.ø. Ãíåçä. Îìóòèùè Ïåò. Àáá. Áîæ. Ñîá. Ãîë. Êàøì. Ëèê. Òèõ. 56 00' ñ.ø. Êîí. ×åð. Ãîðîäèùè Òèì. Áðûê. Ìóðîìöåâî Íîâ. Ìàðê. Êð. Ãîðá. Áîë. Íåêë. À Ìîø. Êðàñí. Ýõî Èâ. Ãóñü-Õðóñò. Êîíñò. Ìóðîì Ãðèã. Çàê. Çëîá. 41 00' â.ä. Ñîâêè (Lepidoptera, Noctuidae) Âëàäèìèðñêîé îáëàñòè 111 [Óñêîâ, 2001, 2002]. Âñå ýòè äàííûå ïîëíîñòüþ ó÷òå- ôàóíå ñîâîê çàäà÷à, òðåáóþùàÿ ñáîðîâ íà ñâåò íû â íàñòîÿùåé áàçîâîé ñòàòüå ïî ôàóíå ñîâîê Âëàäè- èëè íåñðàâíåííî áîëüøèõ óñèëèé, ÷òî â óñëîâèÿõ ìèðñêîé îáëàñòè áåç ñïåöèàëüíûõ óïîìèíàíèé. òîðôÿíûõ áîëîò íå âñåãäà óäàåòñÿ.  òî æå âðåìÿ íàìè îñòàâëåíà áåç âíèìàíèÿ Ìàòåðèàë, ïîñëóæèâøèé îñíîâîé íàøåé ñòàòüè, ðàáîòà ëþáèòåëÿ Ì.Í. Ñàãîâñêîãî [1914], ãäå äëÿ áûë ñîáðàí Ì.Â. Óñêîâûì â ðÿäå ìåñò Âëàäèìèðñ- öåëîãî ðÿäà âèäîâ ñîâîê ïðèâîäèòñÿ ïåðå÷åíü ãó- êîé îáë. â 19932002 ãã., îñîáåííî çíà÷èòåëüíûå ïî áåðíèé, â êîòîðûõ ýòè âèäû îòìå÷àþòñÿ àâòîðîì, ÷èñëó âèäîâ è ýêçåìïëÿðîâ ñáîðû áûëè ïðîâåäåíû âêëþ÷àÿ è Âëàäèìèðñêóþ. Ýòî ñâÿçàíî ñ òåì, ÷òî â èì â Ñóäîãîäñêîì ð-íå (â ïåðâóþ î÷åðåäü â ñ. Áðû- îòíîøåíèè äàííûõ Ñàãîâñêîãî èç-çà îòñóòñòâèÿ â êèíî è îêðåñòíîñòÿõ). Êðîìå òîãî , â ñáîðàõ ïðèíè- åãî ðàáîòå óêàçàíèé íà ãåîãðàôè÷åñêèå òî÷êè ñåé- ìàë àêòèâíîå ó÷àñòèå öåëûé ðÿä âëàäèìèðñêèõ ýí- ÷àñ íåâîçìîæíî äîñòîâåðíî óòâåðæäàòü, îòíîñÿòñÿ òîìîëîãîâ, êðàåâåäîâ è ýíòóçèàñòîâ. Ñðåäè íèõ ñòó- ëè îíè ê òåððèòîðèÿì íûíåøíèõ Âëàäèìèðñêîé èëè äåíò-ýíòîìîëîã Âëàäèìèðñêîãî ïåäàãîãè÷åñêîãî Èâàíîâñêîé îáëàñòåé. Òàê æå ñòîèò îòíîñèòüñÿ è óíèâåðñèòåòà Â.È. Ãîðüêàâûé, ñáîðû êîòîðîãî ñó- äðóãèì óêàçàíèÿì íà Âëàäèìèðñêóþ ãóáåðíèþ áåç ùåñòâåííî äîïîëíÿþò ñïèñîê âèäîâ. Âñåì ïðèíÿâ- óòî÷íåíèÿ ýòèêåòî÷íûõ äàííûõ. Åùå óïîìÿíåì, ÷òî øèì ó÷àñòèå â ñîáèðàíèè ìàòåðèàëà, êîòîðûé ïî- ìû ðàñïîëàãàåì ñïèñêîì âèäîâ èç ðóêîïèñè áûâøå- ìîã ðàñøèðèòü ïðåäñòàâëåíèå î ðàñïðîñòðàíåíèè ãî ñòóäåíòà êàô. ýíòîìîëîãèè ÌÃÓ Ã.Ñ. Åð¸ìêèíà âèäîâ â îáëàñòè àâòîðû ñåðäå÷íî ïðèçíàòåëüíû. Ýòî ïî íî÷íûì áàáî÷êàì Ïåòóøèíñêîãî ðàéîíà.  ñïèñ- ñáîðû 19992002 ãã. èçâåñòíîãî çîîëîãà ïðîôåññîðà êå íàçâàíî 79 âèäîâ ñîâîê.  îñíîâíîì ýòî áàíàëü- Ã.À.Âåñ¸ëêèíà, Ì.Ñ. Ãóñüêîâîé, Î.È. Èâàíîâîé, À.Ñ. íûå âèäû, çà èñêëþ÷åíèåì ïÿòè âñå îíè áûëè îáíà- Êîðíåâà, À.À. Êîâçóíà, È.Â. Ìàëüöåâà, Â.Å. Ìèõëè- ðóæåíû íàìè âî Âëàäèìèðñêîé îáëàñòè.  ïðàâèëü- íà, Ä.Â. Ìîðåâà, Ð.Å. Íå÷àåâà, À.Â. Ïàâëîâà, Ì.Â. íîñòè îïðåäåëåíèÿ ÷åòûðåõ èç ïÿòè ìû íå óâåðåíû, Ôîòèìñêîãî, Ì.Þ. ×èêèð¸âà è Ò.Å. Ùèïóíîâîé. â íåïðàâèëüíîñòè îïðåäåëåíèÿ îäíîãî óâåðåíû.  ðàáîòå ó÷òåíû âñå ìàòåðèàëû ôîíäîâ Íàó÷íî- Ìàòåðèàë ðóêîïèñè íå ïðîâåðÿëñÿ ñïåöèàëèñòàìè èññëåäîâàòåëüñêîãî çîîëîãè÷åñêîãî ìóçåÿ ÌÃÓ (íà- ïî ñîâêàì, à íûíå, ïî ñëîâàì åå àâòîðà, íå ìîæåò êîëîòûå è íàõîäÿùèåñÿ íà âàòå). Îíè íåìíîãî÷èñ- áûòü ïðîâåðåí, ïîñêîëüêó óòðà÷åí. Êðîìå òîãî, ìà- ëåííû, îò åäèíè÷íûõ íàêîëîòûõ ýêçåìïëÿðîâ äî òåðèàë ïî íåêîòîðûì âèäàì âîâñå íå êîëëåêòèðî- äâóõ òðåõ ñòîïîê âàòíûõ ìàòðàñèêîâ (ñáîðû Ë.Ê. âàëñÿ, à ëèøü "çàïèñûâàëñÿ ñ ýêðàíà". Ïîýòîìó äàí- Ñåëèâàíîâà è Â.È. Öâåòêîâà), íî äîïîëíÿþò êàðòè- íûå ðóêîïèñè áîëüøåé ÷àñòüþ íå ó÷èòûâàëèñü íàìè. íó ðàñïðîñòðàíåíèÿ âèäîâ ïî ðàéîíàì, îòêóäà èìå- Ìû ó÷ëè ëèøü äåâÿòü ýêçåìïëÿðîâ, à èìåííî òå, äëÿ þòñÿ ëèøü ôðàãìåíòàðíûå äîñòóïíûå ïðîâåðêå äàí- îïðåäåëåíèÿ êîòîðûõ àâòîð ðóêîïèñè îáðàòèëñÿ ê íûå. Ýòî ñáîðû Å.Ì. Àíòîíîâîé (Ñîáèíñêèé ð-í, À.Â. Ñâèðèäîâó.  ñëó÷àå îòñóòñòâèÿ áîëåå òî÷íûõ 19511952, 1958 ãã.), Â.Ì. Åôàíîâà (Ïåòóøèíñêèé äàííûõ îá ýòèõ íàõîäêàõ â Ïåòóøèíñêîì ðàéîíå, ð-í, 1992 ã.), À.Ë. Êàëóöêîãî (Ïåòóøèíñêèé ð-í, ìû äàåì ññûëêó íà ðàéîí. Çàìåòèì, ÷òî ìàòåðèàëû 1984 ã.), È.Â.Êóçíåöîâà (Ñîáèíêà, 1987 ã.), Â. Îðëî- ñáîðîâ Ã.Ñ. Åðåìêèíà ÷àñòè÷íî ñîäåðæàòñÿ â èíòå- âà (Ïåòóøèíñêèé ð-í, 1965 ã.), À.Ã. Ïîíîìàðåíêî ðåñíîé êíèãå ïî ñîçîáèîëîãèè Ä.Ì. Î÷àãîâà è äð. (Ñîáèíñêèé ð-í, 1993 ã.), Ë.Ê. Ñåëèâàíîâà (Ãóñü- [2000] íà ïðèìåðå òîðôÿíûõ áîëîò Ïåòóøèíñêîãî Õðóñòàëüíûé ð-í, 1995 ã.), Ä.Þ. Òèøå÷êèíà (Ñóç- ðàéîíà. Îäíàêî èíôîðìàöèè î ñîâêàõ â íåé íåò. Ýòî äàëü, 1980 ã.), Â.È. Öâåòêîâà (Ãóñü-Õðóñòàëüíûé ð- è ïîíÿòíî: ñáîðû áóëàâîóñûõ ÷åøóåêðûëûõ è ïÿäå- í, 1983 ã.). Òàêæå ó÷òåíû ñâåäåíèÿ èç ñïèñêîâ ñîâîê, íèö âîçìîæíû è äíåì, à ñîñòàâèòü âïå÷àòëåíèå î îïðåäåëÿâøèõñÿ ðàíåå À.Â. Ñâèðèäîâûì, êîòîðûå Ðèñ.1. Êàðòà ìåñò ñáîðà ìàòåðèàëà è ïðèðîäíîå ðàéîíèðîâàíèå Âëàäèìèðñêîé îáëàñòè. Fig. 1. Map of localities and natural spacing of Vladimir Area. Ïðèðîäíûå ðàéîíû îáëàñòè: À Ñåâåðíàÿ ÷àñòü Ìåù¸ðñêîé íèçìåííîñòè, Á Êîâðîâñêî-Êàñèìîâñêîå ïëàòî,  Âîñòî÷íàÿ îêîíå÷íîñòü Êëèíñêî-Äìèòðîâñêîé ãðÿäû, à Êëÿçüìèíñêî-Íåðëèíñêàÿ íèçèíà, Ä Ëóõñêàÿ íèçèíà, Å ðàâíèíà Âëàäèìèðñêîå Îïîëüå. Áóêâåííûìè àááðåâèàòóðàìè íà êàðòå îáîçíà÷åíû ãåîãðàôè÷åñêèå òî÷êè, èç êîòîðûõ ïðîèñõîäèò ìàòåðèàë, ðåæå îíè íàçâàíû öåëûì íàçâàíèåì òî÷êè. Àëôàâèòíûé ñïèñîê ìåñòîíàõîæäåíèé ñ óêàçàíèåì àááðåâèàòóð ñëåäóþùèé: Àááàêóìîâñêîå Àáá., Áîãîëþáîâî Áîã., Áîæåíîâêà Áîæ., Áîëîãî Áîë., Áðûêèíî Áðûê. (çäåñü æå Êîùååâî, Ëóíüêîâî), Âåðèãèíî Âåð., Âëàäèìèð Âëàä., Ãíåçäèíî Ãíåçä., Ãîëîâèíî Ãîë., Ãîðîäèùè, Ãîðîõîâåö, Ãðèãîðüåâî Ãðèã., Ãóñü-Õðóñòàëüíûé Ãóñü-Õðóñò., Äàâûäîâî Äàâ., Åðîñîâî
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