Conditions That Impact Soybean Germination and Emergence

Total Page:16

File Type:pdf, Size:1020Kb

Conditions That Impact Soybean Germination and Emergence Conditions that Impact Soybean Germination and Emergence The sequence of events during germination begins with imbibition (absorbing water) followed by the development of the radicle (primary root) and emergence of the cotyledons (seed leaves). Planting into a moist seedbed with good seed-to-soil contact is important for optimal germination. Dry or flooded soils, cool temperatures, and lack of oxygen in soils are conditions that can hinder soybean germination and emergence. Soybean Germination The cotyledons are a temporary source of stored food for the seedling. Shortly after emergence, they open and turn green Soybean seed germination is referred to as “epigeal” because because of their exposure to light and start making additional food storage structures (cotyledons) are pulled above the soil food through photosynthesis. As the cotyledons open, the surface. In contrast, corn germination is considered “hypogeal” epicotyl is revealed (Figure 3). The epicotyl contains small where the storage structure remains below the surface leaves, buds, and the main growing point. The cotyledons drop following germination. off shortly after the first set of true leaves is formed. After planting, a soybean seed will start to absorb or imbibe water and swell, and as a result, change its moisture content from less than 13% to almost 50% in several hours. Within one or two days under favorable temperatures, the first root (radicle) of the swollen seed emerges through the seed coat (Figure 1) and grows downward to develop the primary root. Lateral roots quickly emerge from the radicle as it elongates and root hairs grow from the radicle and the lateral roots. Root hairs are barely visible and should not be confused with later developing and easily seen branch roots. The root hairs become the main Seed hilum absorbing structures. Five to ten days after planting, the new seedling emerges from below the soil surface. The hypocotyl begins to elongate and Figure 1. (Left) Imbibition of a treated seed causes the radicle forms a hook that pushes through the soil surface, pulling the (red arrow) to emerge through the seed coat (yellow arrow). (Right) cotyledons upward (Figures 2 & 3). The hypocotyl can be easily Seed hilum or scar where the seed was attached to the plant ovary broken if the soil surface is too hard or crusted to push through. within the pod. Seedlings usually die if the hypocotyl is broken. Figure 2. Hooked hypocotyl and cotyledons Figure 3. Cotyledons (yellow arrow) Figure 4. Soybean seedlings with fully expanded breaking through the soil surface. open to expose epicotyl (red arrow). cotyledons (green arrow), unifoliate leaves, and growing point above the soil surface. Conditions that Impact Soybean Germination and Emergence Conditions that Affect Germination and Table 1. Soybean Seedling Structures Emergence Structure Function/Characteristic Moisture. Planting into a moist seedbed with good seed-to-soil contact is necessary as moisture needs to move into the seed for germination to occur. If irrigation is required for adequate Testa (Figure 1) Seed coat soil moisture, it should be applied ahead of soybean planting and not immediately after planting. Planting into dry soil with Seed hilum or scar Location where the seed was attached rainfall or irrigation occurring too soon after can result in soil (Figure 1) to the plant ovary within the pod crusting and poor soybean emergence. Radicle First part of the embryo to penetrate Soil Conditions. Soil crusting can delay or prevent seedling (Figure 1) the seed coat; develops into main root emergence and cause soybean hypocotyls to become swollen or broken when trying to push through the crust. Fields with Tissue between cotyledons and Hypocotyl fine-textured soils, low organic matter, and little surface residue radicle that becomes the stem of the (Figure 2) can be vulnerable to crusting, especially where excessive tillage plant after germination has taken place. Temperature. Soybean seed can begin to germinate when soil Cotyledons First leaves to appear; serve as food temperatures are less than 55° F; however, germination is likely (Figures 2, 3, & 4) storage structures to be slow until soil temperatures warm to the upper 70s. Cold Stem area above the cotyledons that soil temperatures can cause seeds to remain dormant, Epicotyl consists of a stem, two primary increasing their vulnerability to seed and seedling diseases and (Figure 3) feeding by insects and wildlife. When soil temperatures are leaves, and a terminal bud between 70º F and 90º F, seedling emergence should occur in less than a week. Soil temperatures above 95º F can also Sources: cause poor soybean germination and emergence resulting in 1 Soybean as a crop. Modern corn and soybean production. MCSP, http://www.mcsp- reduced stands. pubs.com. Other sources: Hoeft, R.G., Nafziger, E.D., Johnson, R.R., and Aldrich, S.A. 2000. Modern corn and soybean production. First edition. MCSP Publications. Champaign, IL. Pedersen, P. Oxygen. Because seed respiration increases during the 2007. Soybean growth stages. Soybean growth & development. PM 1945. Soybean germination process, saturated, flooded, and compacted soils Extension and Research Program. Iowa State University. http://extension.agron.iastate.edu. Pedersen, P. Soybean planting date. Iowa State University. can reduce germination and emergence due to the lack of http://extension.agron.iastate.edu oxygen. Soil pore spaces filled with water reduce the amount Web sources verified: 4/16/15 of oxygen available for seed respiration. Compacted soil reduces the availability of water and oxygen required for For additional agronomic information, please contact your local seed representative. germination, root and plant growth, and nutrient uptake. Developed in partnership with Technology, Development, & Agronomy by Monsanto. Individual results may vary, and performance may vary from location to location and from year to year. This result may not be an indicator of results you may obtain as local growing, soil and weather conditions may vary. Growers should evaluate data from multiple locations and years whenever possible. ©2015 Monsanto Company. 140513060107 051314SMK .
Recommended publications
  • Plant Physiology
    PLANT PHYSIOLOGY Vince Ördög Created by XMLmind XSL-FO Converter. PLANT PHYSIOLOGY Vince Ördög Publication date 2011 Created by XMLmind XSL-FO Converter. Table of Contents Cover .................................................................................................................................................. v 1. Preface ............................................................................................................................................ 1 2. Water and nutrients in plant ............................................................................................................ 2 1. Water balance of plant .......................................................................................................... 2 1.1. Water potential ......................................................................................................... 3 1.2. Absorption by roots .................................................................................................. 6 1.3. Transport through the xylem .................................................................................... 8 1.4. Transpiration ............................................................................................................. 9 1.5. Plant water status .................................................................................................... 11 1.6. Influence of extreme water supply .......................................................................... 12 2. Nutrient supply of plant .....................................................................................................
    [Show full text]
  • What Are Soybeans?
    candy, cakes, cheeses, peanut butter, animal feeds, candles, paint, body lotions, biodiesel, furniture soybeans USES: What are soybeans? Soybeans are small round seeds, each with a tiny hilum (small brown spot). They are made up of three basic parts. Each soybean has a seed coat (outside cover that protects the seed), VOCABULARY cotyledon (the first leaf or pair of leaves within the embryo that stores food), and the embryo (part of a seed that develops into Cultivar: a variety of plant that has been created or a new plant, including the stem, leaves and roots). Soybeans, selected intentionally and maintained through cultivation. like most legumes, perform nitrogen fixation. Modern soybean Embryo: part of a seed that develops into a new plant, cultivars generally reach a height of around 1 m (3.3 ft), and including the stem, leaves and roots. take 80–120 days from sowing to harvesting. Exports: products or items that the U.S. sells and sends to other countries. Exports include raw products like whole soybeans or processed products like soybean oil or Leaflets soybean meal. Fertilizer: any substance used to fertilize the soil, especially a commercial or chemical manure. Hilum: the scar on a seed marking the point of attachment to its seed vessel (the brown spot). Leaflets: sub-part of leaf blade. All but the first node of soybean plants produce leaves with three leaflets. Legume: plants that perform nitrogen fixation and whose fruit is a seed pod. Beans, peas, clover and alfalfa are all legumes. Nitrogen Fixation: the conversion of atmospheric nitrogen Leaf into a nitrogen compound by certain bacteria, such as Stem rhizobium in the root nodules of legumes.
    [Show full text]
  • International Union for the Protection of New Varieties of Plants
    E TG/81/7(proj.1) ORIGINAL: English DATE: 2018-04-05 INTERNATIONAL UNION FOR THE PROTECTION OF NEW VARIETIES OF PLANTS Geneva DRAFT * COMMON SUNFLOWER UPOV Code(s): HLNTS_ANN Helianthus annuus L. GUIDELINES FOR THE CONDUCT OF TESTS FOR DISTINCTNESS, UNIFORMITY AND STABILITY prepared by experts from Hungary to be considered by the Technical Working Party for Agricultural Crops at its forty-seventh session, to be held in Naivasha, Kenya, from 2018-05-21 to 2018-05-25 Disclaimer: this document does not represent UPOV policies or guidance Alternative names:* Botanical name English French German Spanish Helianthus annuus L. Common Sunflower Soleil, Tournesol Sonnenblume Girasol The purpose of these guidelines (“Test Guidelines”) is to elaborate the principles contained in the General Introduction (document TG/1/3), and its associated TGP documents, into detailed practical guidance for the harmonized examination of distinctness, uniformity and stability (DUS) and, in particular, to identify appropriate characteristics for the examination of DUS and production of harmonized variety descriptions. ASSOCIATED DOCUMENTS These Test Guidelines should be read in conjunction with the General Introduction and its associated TGP documents. * These names were correct at the time of the introduction of these Test Guidelines but may be revised or updated. [Readers are advised to consult the UPOV Code, which can be found on the UPOV Website (www.upov.int), for the latest information.] TG/81/7(proj.1) Common Sunflower, 2018-04-05 2 TABLE OF CONTENTS PAGE 1.
    [Show full text]
  • Effects of Root Pruning on Germinated Pecan Seedlings
    PROPAGATION AND TISSUE CULTURE HORTSCIENCE 50(10):1549–1552. 2015. if pruning the roots shortly after germination stimulates lateral root formation. Similarly, little work has been attempted to consider Effects of Root Pruning on Germinated the effects of taproot pruning in young pecan seedlings back to different lengths on root Pecan Seedlings regeneration. The objective of this study 1 was to determine if root pruning at different Rui Zhang, Fang-Ren Peng , Pan Yan, Fan Cao, periods after seed germination affects pecan and Zhuang-Zhuang Liu seedling root and shoot growth and to evaluate College of Forestry, Nanjing Forestry University, 159 Longpan Road, different degrees of taproot pruning on root Nanjing 210037, China initiation. Dong-Liang Le Materials and Methods College of Forestry, Nanjing Forestry University, 159 Longpan Road, Nanjing 210037, China; and Nanjing Green Universe Pecan Science and The test was established at Lvzhou Pecan Research Station, Nanjing, China (lat. Technology Co. Ltd., 38 Muxuyuan Street, Nanjing 210007, China 32.05°N, long. 118.77°E). Open-pollinated Peng-Peng Tan seeds of the Chinese selection ‘Shaoxing’ were used as seed stock to produce seedlings. College of Forestry, Nanjing Forestry University, 159 Longpan Road, ‘Shaoxing’ had a small nut of very good Nanjing 210037, China quality with 50% percentage fill. The seeds averaged 30.4 mm in length, 20.9 mm in Additional index words. Carya illinoinensis, taproot, lateral root, shoot growth width, and 5.4 g in weight. Seeds were Abstract. Root systems of pecan trees are usually dominated by a single taproot with few collected on 7 Nov.
    [Show full text]
  • Germination Structure of Seeds Quote
    Germination Structure of Seeds Quote: Thomas Fuller's Gnomologia, 1732: "The greatest Oaks have been little Acorns." Seeds are found in a staggering array of shapes and sizes, but the process by which seeds germinate is similar in all species. Abies koreana Acer griseum Wisteria floribunda 'Korean Fir' by Roger Culos. CC BY-SA. 'Paperbark Maple' by Roger Culos. CC 'Japanese Wisteria' by Roger BY-SA. Culos. CC BY-SA. Alsomitra macrocarpa Macrozamia communis Strelitzia reginae 'Alsomitra macrocarpa seed' by Scott 'BurrawangSeeds' by AYArktos. CC BY- 'Paradiesvogelblumensamen' by Zona. CC BY. SA. Sebastian Stabinger. CC BY-SA. Taraxicum officinale Stephanotis floribunda Phleum pratense 'Achane of Taraxacum sect. 'Stephanotis seed' by L. Marie"/Lenore 'Timoteegras vruchten Phleum Ruderalia' by Didier Edman, Sunnyvale, CA. CC BY. pratense' by Rasbak. CC BY-SA. Descouens. CC BY-SA. Dicotyledon seeds testa epicotyl plumule hypocotyl cotyledon radicle 'Aesculus hippocastanum seed section' by Boronian. CC BY. plumule epicotyl hypocotyl testa hilum radicle cotyledon micropyle endosperm Monocotyledon seeds endosperm epicotyl testa hypocotyl cotyledon radicle Parts of a seed Testa The seed coat. A protective layer which is tough and hard and it protects the seed from attack by insects, fungi and bacteria. Cotyledon Dicotyledons have 2 cotyledons Monocotyledons have 1 cotyledon A cotyledon is an embryonic leaf. It is the first leaf to appear when a seedling grows. They often contain reserves of food which the developing seedling can use to grow. Epicotyl The section of stem between the cotyledon(s) and the plumule. In a seedling it is the section of stem between the cotyledons and the first true leaves.
    [Show full text]
  • Soybean Growth and Development & Management Information Fo Replant Decisions
    MAGRCORE Metadata, citation and similar papers at core.ac.uk GOVSProvided by University of Minnesota Digital Conservancy MN 2500 AGFO-5701 MINNESOTA EXTENSION SERVICE I:,:, I UNIVERSITY OF MINNESOTA AGRICULTURE Soybean Growth and Development & Management Information fo UNIVf:R&TYoF MINNESOTA J DOCUMENTS ~ Replant Decisions JUL ·2s 1991 ,1 ·I shoot apex emerging decomposing radicle · - seed coat primary root L.L. Hardman J.L. Gunsolus Extension Agronomist-Crops Extension Agronomist-Weed Control Dept. of Agronomy & Dept. of Agronomy & Plant Genetics Plant Genetics Univ. of MN, St. Paul Univ. of MN, St. Paul SOYBEAN GROWTH AND DEVELOPMENT stored food is removed the cotyledons tum yellow and shrivel before dropping off the plant. Loss of one or more of the cotyledons before the food reserves are fully utilized can slow An understanding of how a soybean plant develops can help you early plant growth or result in death of the plant if photosynthetic make important management decisions. This section discusses leaf tissue is not formed quickly. various seed and plant parts, explains how the soybean plant develops utilizing standardized vegetative and reproductive stage descriptions which are used by the National Crop Insurance Services (NCIS), and describes some of the important factors which affect growth and development of the soybean plant. A soybean seed consists of a miniature plant attached to two nutrient storage reservoirs (cotyledons or seed halves) surrounded by a protective outer wrapper or seed coat called a testa (Figure 1). The cotyledons contain about 40% protein, 20% oil, and about 35% carbohydrate. These materials provide nutrients to the developing embryo during the germination process.
    [Show full text]
  • Drupe. Fruit with a Hard Endocarp (Figs. 67 and 71-73); E.G., and Sterculiaceae (Helicteres Guazumaefolia, Sterculia)
    Fig. 71. Fig. 72. Fig. 73. Drupe. Fruit with a hard endocarp (figs. 67 and 71-73); e.g., and Sterculiaceae (Helicteres guazumaefolia, Sterculia). Anacardiaceae (Spondias purpurea, S. mombin, Mangifera indi- Desmopsis bibracteata (Annonaceae) has aggregate follicles ca, Tapirira), Caryocaraceae (Caryocar costaricense), Chrysobal- with constrictions between successive seeds, similar to those anaceae (Licania), Euphorbiaceae (Hyeronima), Malpighiaceae found in loments. (Byrsonima crispa), Olacaceae (Minquartia guianensis), Sapin- daceae (Meliccocus bijugatus), and Verbenaceae (Vitex cooperi). Samaracetum. Aggregate of samaras (fig. 74); e.g., Aceraceae (Acer pseudoplatanus), Magnoliaceae (Liriodendron tulipifera Hesperidium. Septicidal berry with a thick pericarp (fig. 67). L.), Sapindaceae (Thouinidium dodecandrum), and Tiliaceae Most of the fruit is derived from glandular trichomes. It is (Goethalsia meiantha). typical of the Rutaceae (Citrus). Multiple Fruits Aggregate Fruits Multiple fruits are found along a single axis and are usually coalescent. The most common types follow: Several types of aggregate fruits exist (fig. 74): Bibacca. Double fused berry; e.g., Lonicera. Achenacetum. Cluster of achenia; e.g., the strawberry (Fra- garia vesca). Sorosis. Fruits usually coalescent on a central axis; they derive from the ovaries of several flowers; e.g., Moraceae (Artocarpus Baccacetum or etaerio. Aggregate of berries; e.g., Annonaceae altilis). (Asimina triloba, Cananga odorata, Uvaria). The berries can be aggregate and syncarpic as in Annona reticulata, A. muricata, Syconium. Syncarp with many achenia in the inner wall of a A. pittieri and other species. hollow receptacle (fig. 74); e.g., Ficus. Drupacetum. Aggregate of druplets; e.g., Bursera simaruba THE GYMNOSPERM FRUIT (Burseraceae). Fertilization stimulates the growth of young gynostrobiles Folliacetum. Aggregate of follicles; e.g., Annonaceae which in species such as Pinus are more than 1 year old.
    [Show full text]
  • Difference Between Radicle and Plumule Key Difference
    Difference Between Radicle and Plumule www.differencebetween.com Key Difference - Radicle vs Plumule All seeds contain embryos. Seed embryo creates a new plant after germination. Therefore the embryo is protected inside the seed by a hard covering called testa. The seed begins to grow when the right conditions are found such as moisture, warmth, sunlight, nutrient-rich soil etc. There are two main components of a plant; shoot and root. Stem, leaves and roots develop from different parts of the seed embryo. The radicle is the first part that emerges from the seed during the germination through the micropyle (seed pore). It makes the roots of the new plant. Plumule emerges after the radicle and makes the stem of the new seedling. Cotyledons form the first leaves of the seedling. The key difference between radicle and plumule is that radicle is the root forming part of the seed embryo while plume is the stem forming part of the seed embryo. The cotyledons of the seed embryo hold the radicle and the plume. What is Radicle? The radicle is the embryonic root of the plant. It is a part of the seedling that comes first from the seed during the germination. It comes out from the seed through the micropyle. Radicle grows downwards into the soil. A root cap protects the tip of the radicle. It absorbs water and nutrients and supplies to the leaves for starting photosynthesis. Embryonic stem or the hypocotyl is found above the radicle. Figure 01: Radicle Radicle emerges from the seed as a short white structure.
    [Show full text]
  • Early Soybean Growth and Development
    EARLY SOYBEAN GROWTH AND DEVELOPMENT A soybean seed has two distinct parts: the cotyledons and the embryo. The two cotyledons are the main food storage structure, which supply food during emergence and for the seven to ten days after emergence through the V1 growth stage. The embryo itself is comprised of three parts. 1. The radicle is the first root and it’s the first part of the embryo to penetrate the seed coat. Lateral roots form from the radicle, and tiny root hairs then develop on the lateral roots. Root hairs are the main absorbing surface of the entire root system. A soybean’s root system branches and rebranches within the first four to five weeks after planting, with the bulk of root mass persisting in the top six inches of soil. 2. The hypocotyl is the stem below the cotyledon. It begins to elongate after the radicle and forms an arch, which is pushed upward. As the arch breaks the soil surface, it pulls the cotyledons and epicotyl up. The uppermost cells of the hypocotyl stop growing as cells on the underside continue to straighten the arch. This process lifts the cotyledons into an upright position. 3. The epicotyl is comprised of the small leaves, stem and the apical growing point. The epicotyl is protected between the cotyledons until after emergence. The loss of a cotyledon is not lethal to the seedling, but damage to the epicotyl is. The hypocotyl arch is a large structure that is relative to seed size, utilizing considerable seed energy to push through the soil.
    [Show full text]
  • Agricultural Marketing Service, USDA § 201.56–10
    Agricultural Marketing Service, USDA § 201.56–10 (A) One or more essential structures and become thin, leaf-like, and photo- impaired as a result of decay from pri- synthetic. mary infection. (3) Shoot system: The hypocotyl (B) Albino. elongates carrying the cotyledons above the soil surface. The epicotyl [59 FR 64504, Dec. 14, 1994] usually does not show any development § 201.56–8 Flax family, Linaceae. within the test period. Areas of yel- lowish pigmentation may develop on Kind of seed: Flax. the hypocotyl in cotton. (a) General description. (4) Root system: A primary root, with (1) Germination habit: Epigeal dicot. secondary roots usually developing (Due to the mucilaginous nature of the within the test period. Areas of yel- seed coat, seedlings germinated on lowish pigmentation may develop on blotters may adhere to the blotter and the root in cotton. appear to be negatively geotropic.) (b) Abnormal seedling description. (2) Food reserves: Cotyledons which (1) Cotyledons: expand and become photosynthetic. (i) Less than half of the original cot- (3) Shoot system: The hypocotyl yledon tissue remaining attached. elongates carrying the cotyledons (ii) Less than half of the original cot- above the soil surface. The epicotyl yledon tissue free of necrosis or decay. usually does not show any development (Remove any attached seed coats at within the test period. the end of the test period for evalua- (4) Root system: A primary root, with tion of cotyledons.) secondary roots usually developing (2) Epicotyl: within the test period. (i) Missing. (May be assumed to be (b) Abnormal seedling description. present if both cotyledons are intact.) (1) Cotyledons: (ii) [Reserved] (i) Less than half of the original cot- (3) Hypocotyl: yledon tissue remaining attached.
    [Show full text]
  • Growing a Sunflower from Seed
    Growing a sunflower from seed First, make a pot for your sunflower seeds from newspaper. Fill the pot ¾ full with potting oil. Plant two seeds 1/4” deep in the center of your pot. When you get your pot home: Place your pot on a plastic plate or tray in a sunny window. Water your seeds very gently so you don’t wash the seeds or the soil out of the pot. Keep the soil damp but not wet. When the newspaper pot turns grey, you have watered enough. When the newspaper pot has turned white it is time to water again. Your plant will not need additional food (fertilizer) while it is growing in the pot. When your plants sprout, one will probably be bigger and stronger (dominant)- cut the weaker plant stem with scissors at the soil line. When the remaining plant is approximately 5”-6” tall and has two sets of true leaves, it is ready to be planted outside. The newspaper does not have to be removed when planting your plant as the newspaper will breakdown (decompose). You can plant your sunflower plant in a large container or in a prepared site like a garden. Be sure to plant your plant at the same soil line as it is growing in the pot. Enjoy! Germination of the Sunflower Seed After the seed is wetted and put in a warm dark place: First the fruit wall splits and radicle emerges Then the hypocotyl extends down and forms root hairs Plant development seed coat lateral cotyledons roots cotyledons .
    [Show full text]
  • Federal Aviation Agency
    FEDERAL REGISTER VOLUME 30 • NUMBER 117 Friday, June 18,1965 • Washington, D.C. Pages 7863-7938 Agencies in this issue— Agricultural Research Service Area Redevelopment Administration Atomic Energy Commission Civil Aeronautics Board Civil Service Commission Consumer and Marketing Service Federal Aviation Agency Federal Communications Commission Federal Maritime Commission Federal Trade Commission Food and Drug Administration Interstate Commerce Commission Land Management Bureau Securities and Exchange Commission Detailed list of Contents appears inside. Subscriptions Now Being Accepted S L I P L A W S 89th Congress, 1st Session 1965 Separate prints of Public Laws, published immediately after enactment, with marginal annotations and legislative history references. Subscription Price: $12.00 per Session Published by Office of the Federal Register, National Archives and Records Service, General Services Administration Order from Superintendent of Documents, U.S. Government Printing Office, Washington, D.C.,' 20402 Published dally, Tuesday through Saturday (no publication on Sundays, Monday», or FEDERAL®REGISTER on the day after an official Federal holiday), by the Office of the Federal Register, National Area Code 202 Phone 963-3261 A1®1117®8 a n d R ecords Service, G eneral Services A d m in istra tio n (m ail address Nations _ . , _ _ Archives Building, Washington, D.C. 20408), p u r s u a n t to the authority contained in tne Federal Register Act, approved July 26, 1935 (49 Stat. 500, as amended; 44 U.S.C., ch. 8B), under regulations prescribed by the Admin­ istrative Committee of the Federal Register, approved by the President (1 CFR Ch. I). Distribution is made only by the Superintendent or Documents, Government Printing Office, Washington, D.C.
    [Show full text]