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Lepidoptera: Papilionoidea) Q ⇑ Marianne Espeland A,B, , Jason P.W
Molecular Phylogenetics and Evolution 93 (2015) 296–306 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Ancient Neotropical origin and recent recolonisation: Phylogeny, biogeography and diversification of the Riodinidae (Lepidoptera: Papilionoidea) q ⇑ Marianne Espeland a,b, , Jason P.W. Hall c, Philip J. DeVries d, David C. Lees e, Mark Cornwall a, Yu-Feng Hsu f, Li-Wei Wu g, Dana L. Campbell a,h, Gerard Talavera a,i,j, Roger Vila i, Shayla Salzman a, Sophie Ruehr k, David J. Lohman l, Naomi E. Pierce a a Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA b McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Powell Hall, 2315 Hull Road, Gainesville, FL 32611, USA c Department of Systematic Biology-Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560-127, USA d Department of Biological Sciences, University of New Orleans, 2000 Lake Shore Drive, New Orleans, LA 70148, USA e Department of Zoology, University of Cambridge, Cambridge CB2 3EJ, UK f Department of Life Science, National Taiwan Normal University, Taipei, Taiwan g The Experimental Forest, College of Bio-Resources and Agriculture, National Taiwan University, Nantou, Taiwan h Division of Biological Sciences, School of Science, Technology, Engineering & Mathematics, University of Washington Bothell, Box 358500, 18115 Campus Way NE, Bothell, WA 98011-8246, USA i Institut de Biologia Evolutiva (CSIC-UPF), Pg. Marítim de la Barceloneta 37, 08003 Barcelona, Spain j Faculty of Biology & Soil Science, St. -
Title Lorem Ipsum Dolor Sit Amet, Consectetur Adipiscing Elit
Volume 26: 102–108 METAMORPHOSIS www.metamorphosis.org.za ISSN 1018–6490 (PRINT) LEPIDOPTERISTS’ SOCIETY OF AFRICA ISSN 2307–5031 (ONLINE) Classification of the Afrotropical butterflies to generic level Published online: 25 December 2015 Mark C. Williams 183 van der Merwe Street, Rietondale, Pretoria, South Africa. E-mail: [email protected] Copyright © Lepidopterists’ Society of Africa Abstract: This paper applies the findings of phylogenetic studies on butterflies (Papilionoidea) in order to present an up to date classification of the Afrotropical butterflies to genus level. The classification for Afrotropical butterflies is placed within a worldwide context to subtribal level. Taxa that still require interrogation are highlighted. Hopefully this classification will provide a stable context for researchers working on Afrotropical butterflies. Key words: Lepidoptera, Papilionoidea, Afrotropical butterflies, classification. Citation: Williams, M.C. (2015). Classification of the Afrotropical butterflies to generic level. Metamorphosis 26: 102–108. INTRODUCTION Suborder Glossata Fabricius, 1775 (6 infraorders) Infraorder Heteroneura Tillyard, 1918 (34 Natural classifications of biological organisms, based superfamilies) on robust phylogenetic hypotheses, are needed before Clade Obtectomera Minet, 1986 (12 superfamilies) meaningful studies can be conducted in regard to their Superfamily Papilionoidea Latreille, 1802 (7 evolution, biogeography, ecology and conservation. families) Classifications, dating from the time of Linnaeus in the Family Papilionidae Latreille, 1802 (32 genera, 570 mid seventeen hundreds, were based on morphology species) for nearly two hundred and fifty years. Classifications Family Hedylidae Guenée, 1858 (1 genus, 36 species) based on phylogenies derived from an interrogation of Family Hesperiidae Latreille, 1809 (570 genera, 4113 the genome of individual organisms began in the late species) 20th century. -
Superior National Forest
Admirals & Relatives Subfamily Limenitidinae Skippers Family Hesperiidae £ Viceroy Limenitis archippus Spread-wing Skippers Subfamily Pyrginae £ Silver-spotted Skipper Epargyreus clarus £ Dreamy Duskywing Erynnis icelus £ Juvenal’s Duskywing Erynnis juvenalis £ Northern Cloudywing Thorybes pylades Butterflies of the £ White Admiral Limenitis arthemis arthemis Superior Satyrs Subfamily Satyrinae National Forest £ Common Wood-nymph Cercyonis pegala £ Common Ringlet Coenonympha tullia £ Northern Pearly-eye Enodia anthedon Skipperlings Subfamily Heteropterinae £ Arctic Skipper Carterocephalus palaemon £ Mancinus Alpine Erebia disa mancinus R9SS £ Red-disked Alpine Erebia discoidalis R9SS £ Little Wood-satyr Megisto cymela Grass-Skippers Subfamily Hesperiinae £ Pepper & Salt Skipper Amblyscirtes hegon £ Macoun’s Arctic Oeneis macounii £ Common Roadside-Skipper Amblyscirtes vialis £ Jutta Arctic Oeneis jutta (R9SS) £ Least Skipper Ancyloxypha numitor Northern Crescent £ Eyed Brown Satyrodes eurydice £ Dun Skipper Euphyes vestris Phyciodes selenis £ Common Branded Skipper Hesperia comma £ Indian Skipper Hesperia sassacus Monarchs Subfamily Danainae £ Hobomok Skipper Poanes hobomok £ Monarch Danaus plexippus £ Long Dash Polites mystic £ Peck’s Skipper Polites peckius £ Tawny-edged Skipper Polites themistocles £ European Skipper Thymelicus lineola LINKS: http://www.naba.org/ The U.S. Department of Agriculture (USDA) prohibits discrimination http://www.butterfliesandmoths.org/ in all its programs and activities on the basis of race, color, national -
Butterflies (Lepidoptera: Papilionoidea) in a Coastal Plain Area in the State of Paraná, Brazil
62 TROP. LEPID. RES., 26(2): 62-67, 2016 LEVISKI ET AL.: Butterflies in Paraná Butterflies (Lepidoptera: Papilionoidea) in a coastal plain area in the state of Paraná, Brazil Gabriela Lourenço Leviski¹*, Luziany Queiroz-Santos¹, Ricardo Russo Siewert¹, Lucy Mila Garcia Salik¹, Mirna Martins Casagrande¹ and Olaf Hermann Hendrik Mielke¹ ¹ Laboratório de Estudos de Lepidoptera Neotropical, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19.020, 81.531-980, Curitiba, Paraná, Brazil Corresponding author: E-mail: [email protected]٭ Abstract: The coastal plain environments of southern Brazil are neglected and poorly represented in Conservation Units. In view of the importance of sampling these areas, the present study conducted the first butterfly inventory of a coastal area in the state of Paraná. Samples were taken in the Floresta Estadual do Palmito, from February 2014 through January 2015, using insect nets and traps for fruit-feeding butterfly species. A total of 200 species were recorded, in the families Hesperiidae (77), Nymphalidae (73), Riodinidae (20), Lycaenidae (19), Pieridae (7) and Papilionidae (4). Particularly notable records included the rare and vulnerable Pseudotinea hemis (Schaus, 1927), representing the lowest elevation record for this species, and Temenis huebneri korallion Fruhstorfer, 1912, a new record for Paraná. These results reinforce the need to direct sampling efforts to poorly inventoried areas, to increase knowledge of the distribution and occurrence patterns of butterflies in Brazil. Key words: Atlantic Forest, Biodiversity, conservation, inventory, species richness. INTRODUCTION the importance of inventories to knowledge of the fauna and its conservation, the present study inventoried the species of Faunal inventories are important for providing knowledge butterflies of the Floresta Estadual do Palmito. -
Catalogue of the Type Specimens of Lepidoptera Rhopalocera in the Hill Museum
Original from and digitized by National University of Singapore Libraries Original from and digitized by National University of Singapore Libraries Original from and digitized by National University of Singapore Libraries Original from and digitized by National University of Singapore Libraries CATALOGUE OF THE Type Specimens of Lepidoptera Rhopalocera IN THE HILL MUSEUM BY A. G. GABRIEL, F.E.S. Issued June, 1932 LONDON JOHN BALE, SONS & DANIELSSON, LTD. 83-91, GBEAT TITCHFIELD STEEET, OXEOED STEEET, W. 1 1932 Price 20/- Original from and digitized by National University of Singapore Libraries Unfortunately Mr. Joicey did not live to see the publication of this Catalogue. It will however remain, together with the four completed volumes of the " Bulletin of the Hill Museum," as a lasting memorial to to the magnificent collection of Lepidoptera amassed by Mr. Joicey, and to the work carried out at the Hill Museum under his auspices. G. Talbot. Original from and digitized by National University of Singapore Libraries CATALOGUE OF THE TYPE SPECIMENS OF LEPIDOPTERA RHOPALOCERA IN THE HILL MUSEUM. By A. G. GABRIEL, F.E.S. INTRODUCTION BY G. TALBOT. It is important to know exactly where type specimens are to be found. The British Museum set an example by publishing catalogues of some of their Rhopalocera types, and we hope this will be continued. Mr. Gabriel, who was responsible for that work, has been asked by Mr. Joicey to prepare a catalogue for the Hill Museum. The original description of almost every name in this catalogue has been examined for the correct reference, and where the sex or habitat was wrongly quoted, the necessary correction has been made. -
A Distinctive New Species of Cloud Forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru
WILLMOTT ET AL.: New species of Erichthodes TROP. LEPID. RES., 28(1): 39-45, 2018 39 A distinctive new species of cloud forest Euptychiina (Lepidoptera: Nymphalidae: Satyrinae) from Ecuador and Peru Keith R. Willmott1, Gerardo Lamas2, James Radford3, Mario A. Marín4, Shinichi Nakahara1, Marianne Espeland5, Lei Xiao1, and Jason P. W. Hall6 1. McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL, USA: [email protected] 2. Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru. 3. Cambridge, UK. 4. Departamento de Biologia Animal and Museu de Zoologia, Instituto de Biologia, Universidade Estadual de Campinas, Rua Monteiro Lobato, 255 - Cidade Universitária Zeferino Vaz - Barão Geraldo, 13083-862, Campinas, São Paulo, Brazil. 5. Arthropoda Department, Zoological Research Museum Alexander Koenig, Adenauer Allee 160, 53113 Bonn, Germany. 6. Department of Entomology, National Museum of Natural History, Smithsonian Institution, Washington, DC, USA Date of issue online: 13 July 2018 Zoobank Registered: urn:lsid:zoobank.org:pub:F4A0F8EB-600F-4973-9D52-DDA7E27C3EF8 Electronic copies (ISSN 2575-9256) in PDF format at: http://journals.fcla.edu/troplep; https://zenodo.org; archived by the Institutional Repository at the University of Florida (IR@UF), http://ufdc.ufl.edu/ufir;DOI : 10.5281/zenodo.1309677 © The author(s). This is an open access article distributed under the Creative Commons license CC BY-NC 4.0 (https://creativecommons.org/ licenses/by-nc/4.0/). Abstract: A new species of Euptychiina, Erichthodes eremita Lamas, Willmott & Radford, n. sp., is described and illustrated. DNA sequence data suggest that the new species is sister to a species currently placed in Erichthodes Forster, 1964, although ongoing revision of the generic taxonomy of the subtribe might result in the reclassification of both of these species in future. -
The Radiation of Satyrini Butterflies (Nymphalidae: Satyrinae): A
Zoological Journal of the Linnean Society, 2011, 161, 64–87. With 8 figures The radiation of Satyrini butterflies (Nymphalidae: Satyrinae): a challenge for phylogenetic methods CARLOS PEÑA1,2*, SÖREN NYLIN1 and NIKLAS WAHLBERG1,3 1Department of Zoology, Stockholm University, 106 91 Stockholm, Sweden 2Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Av. Arenales 1256, Apartado 14-0434, Lima-14, Peru 3Laboratory of Genetics, Department of Biology, University of Turku, 20014 Turku, Finland Received 24 February 2009; accepted for publication 1 September 2009 We have inferred the most comprehensive phylogenetic hypothesis to date of butterflies in the tribe Satyrini. In order to obtain a hypothesis of relationships, we used maximum parsimony and model-based methods with 4435 bp of DNA sequences from mitochondrial and nuclear genes for 179 taxa (130 genera and eight out-groups). We estimated dates of origin and diversification for major clades, and performed a biogeographic analysis using a dispersal–vicariance framework, in order to infer a scenario of the biogeographical history of the group. We found long-branch taxa that affected the accuracy of all three methods. Moreover, different methods produced incongruent phylogenies. We found that Satyrini appeared around 42 Mya in either the Neotropical or the Eastern Palaearctic, Oriental, and/or Indo-Australian regions, and underwent a quick radiation between 32 and 24 Mya, during which time most of its component subtribes originated. Several factors might have been important for the diversification of Satyrini: the ability to feed on grasses; early habitat shift into open, non-forest habitats; and geographic bridges, which permitted dispersal over marine barriers, enabling the geographic expansions of ancestors to new environ- ments that provided opportunities for geographic differentiation, and diversification. -
Hypandrium As a Key Character in Resolving Species-Level Taxonomy on the Example of Perisama Oppelii (Latreille) (Lepidoptera: Nymphalidae, Biblidinae)
Zootaxa 3990 (1): 032–040 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2015 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3990.1.2 http://zoobank.org/urn:lsid:zoobank.org:pub:BA4D723A-DBCD-4DBF-91A9-8C6620621565 Hypandrium as a key character in resolving species-level taxonomy on the example of Perisama oppelii (Latreille) (Lepidoptera: Nymphalidae, Biblidinae) ANNA ZUBEK1, JADWIGA LORENC-BRUDECKA & TOMASZ W. PYRCZ Zoological Museum of the Jagiellonian University, Gronostajowa 5, 30-387 Kraków, Poland 1Corresponding author. E-mail: [email protected] Abstract A detailed comparative study of the subspecies of Perisama oppelii (Latreille, [1809]) is carried out. Ten out of eleven subspecies are illustrated, including the females of P. o. bleuzeni Attal & Crosson du Cormier, 1996 and P. o. cristal Attal & Crosson du Cormier, 1996 for the first time. Male genitalia of P. o. amalia Oberthür, 1916, P. o. bleuzeni, P. o. cristal, P. o. erebina Oberthür, 1916, P. o. oppelii (Latreille, [1809]), P. o. viridinota Butler, 1873, and P. o. xanthica (Hewitson, 1868), as well as female genitalia of P. o. bleuzeni, P. o. cristal and P. o. oppelii are also presented for the first time. Based on the characters of hypandrium, a separate specific status is proposed for P. bleuzeni stat. n. Taxonomical value of the hypandrium is evaluated. The co-evolution of male and female genitalia is discussed. Key words: systematics, Perisama bleuzeni Attal & Crosson du Cormier n. stat., male genitalia, female genitalia, adult morphology Introduction Biblidinae is a diverse and widespread group of butterflies, whose taxonomical status of sub-family is well supported by both, morphological and molecular data (Brower 2000; Wahlberg et al. -
A New Perisama (Nymphalidae: Callicorini) from Bolivia
10 JOURNAL OF THE LEPIDOPTERISTS' SOCIETY The specimen might have been sent by Walsingham to Rebel from London to Vienna, because Rebel was dealing also with the former collective genus "Lita auct.," to which this species belonged. LITERATURE CITED KEIFER, H. H., 1936. California Microlepidoptera. X. Mo. Bull. Calif. Dept. Agric.,25: 349-359. POVOLNY, D., 1967. Genitalia of wmc Nearctic and Neotropic members of the tribe Gnorimoschemini (Lep., GeL). Acta ent. Mus. Nat. Pragae, 37: 51-126. A NEW PERISAMA (NYMPHALIDAE: CALLICORINI) FROM BOLIVIA JOlIN H. MASTEHS1 Box 7511, St. Paul, Minnesota During 1962-63, Franz Steinbach, of Cochabamba, Bolivia, consigned to me 395 specimens of Perisama Doubleday (including Orophila Staud inger) collected near Alto Palmar, Chapare Province, Bolivia. The "Perisama comnena" in this group were particularly interesting because they apparently included two distinct phenotypes; one characterized by the normal large red basal patch on the ventral surface of the forewing, the other without it. This is the type of variation that is normally as sociated with geographical subspecies; however, both forms were sympatric at Alto Palmar. An examination of the male genitalia demonstrated that the two "forms" were in reality distinct species and, after a canvass of the literature, I am of the opinion that the second spccies is heretofore unnamed. Perisarna rusea Masters, new species MALE: Upperside of both wings deep-black ground; iridescent green rays extending outwardly from base of forewing, along base of discal cell and vein 2V, until they almost meet a diagonal iridescent green band, 2 to 3 mm wide, which crosses the wing from costal margin, at end of cell to inner margin at tornus; one (occasionally two) iridescent green spot( s) in subapical area, near costal margin. -
Peru Conservation Recorded Wildlife at Taricaya
Peru Conservation Recorded Wildlife at Taricaya Butterflies (Mariposas) in Taricaya Reserve, Madre de Dios CLASS: Insecta ORDER: Lepidoptera 1. Familia Nymphalidae Subfamilia Apaturinae Doxocopa kallina (Staudinger, 1886). Doxocopa laure (Drury, 1776). Doxocopa lavinia (Butler, 1886). Doxocopa linda (C. Felder & R. Felder, 1860). Doxocopa pavon (Latreille, 1809). Subfamilia Nymphalinae Tribu Coeini Baetus aelius (Stoll, 1780). Baetus deucaliom (C. Felder & R. Felder, 1860). Baetus japetus (Staudinger, 1885). Colobura annulata (Willmot, Constantino & J. Hall, 2001). Colobura dirce (Linnaeus, 1758). Historis acheronta (Fabricius, 1775). Historis odius (Fabricius, 1775). Smyrna blomfilda (Fabricius, 1781). Tigridia acesta (Linnaeus, 1758). Tribu Kallimini Anartia jatrophae (Linnaeus, 1763). Junonia everate (Cramer, 1779). Junonia genoveva (Cramer, 1780). Metamorpha elissa (Hübner, 1818). Siproeta stelenes (Linnaeus, 1758). Tribu Melitaeini Eresia clio (Linnaeus, 1758). Eresia eunice (Hübner, 1807). Eresia nauplios (Linnaeus, 1758). Tegosa claudina (Escholtz, 1821). Tegosa fragilis (H. W. Bates, 1864). Tribu Nymphalini Hypanarthia lethe (Fabricius, 1793). Tribu Acraeini Actinote pellenea (Hübner, 1821). Subfamilia Charaxinae Tribu Preponini Agrias amydon (Hewitson, 1854). Agrias claudina (Godart, 1824). Archaeoprepona amphimacus (Fabricius, 1775). Archaeoprepona demophon (Linnaeus, 1758). Archaeoprepona meander (Cramer, 1775). Prepona dexamenus (Hopffer, 1874). Prepona laertes (Hübner, 1811). Prepona pheridamas (Cramer, 1777). Prepona pylene -
Papilionoidea (Butterfly & Skipper) Species List
Papilionoidea (Butterfly & Skipper) Species List Higher Classification1 Kingdom: Animalia, Phylum: Arthropoda, Class: Insecta, Order: Lepidoptera, Superfamily: Papilionoidea Family (F:), Subfamily (sF:) and Tribe (T:) Scientific Name1 English Name1 F: Hesperiidae (Skippers) sF: Eudaminae (Spreadwing Skippers) Astraptes anaphus annetta Yellow-tipped Flasher Central American Banded- Autochton vectilucis Skipper Urbanus pronus Pronus Longtail sF: Hesperiinae (Grass Skippers) T: Anthoptini Synapte salenus salenus Salenus Faceted-Skipper T: Calpodini Calpodes cf. ethlius Brazilian Skipper Talides alternata Alternate Ruby-eye T: Hesperiini Hylephila cf. phyleus phyleus Fiery Skipper Poanes inimica Yellow-stained Skipper Poanes cf. zabulon Hobomok Skipper T: Moncini Halotus angellus Angellus Skipper Lerema accius Clouded Skipper Remella rita Rita's Remella sF: Heteropterinae (Skipperlings) Dalla lethaea Schaus' Skipperling sF: Pyrginae (Spread-wing Skippers) T: Achlyodidini Doberes anticus Dark Doberes T: Carcharodini Noctuana lactifera lactifera Cryptic Skipper T: Erynnini Mylon cf. maimon Common Mylon F: Lycaenidae (Gossamerwings) sF: Theclinae (Hairstreaks) T: Eumaeini (Hairstreaks) Contrafacia bassania White-etched Hairstreak F: Nymphalidae (Brushfoots) sF: Apaturinae (Emperors) Doxocopa cyane mexicana Mexican Emperor Doxocopa laurentia cherubina Turquoise Emperor sF: Biblidinae (Exotic Brushfoots) T: Callicorini Diaethria anna anna Anna’s Eighty-eight Diaethria astala astala Astala Eighty-eight Diaethria clymena marchalii Widespread Eighty-eight -
Molecular Phylogenetics of the Neotropical Butterfly Subtribe Oleriina
Molecular Phylogenetics and Evolution 55 (2010) 1032–1041 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Molecular phylogenetics of the neotropical butterfly subtribe Oleriina (Nymphalidae: Danainae: Ithomiini) Donna Lisa de-Silva a,*, Julia J. Day a, Marianne Elias b,c, Keith Willmott d, Alaine Whinnett a, James Mallet a a Department of Genetics, Evolution and Environment, University College London, Wolfson House, 4 Stephenson Way, London NW1 2HE, UK b Imperial College London, Silwood Park, Buckhurst Road, Ascot, Berkshire SL5 7PY, UK c CNRS, UMR 7205, Muséum National d’Histoire Naturelle, 45 Rue Buffon, CP50, 75005 Paris, France d McGuire Center for Lepidoptera, Florida Museum of Natural History, University of Florida, P.O. Box 112710, Gainesville, FL 32611-2710, USA article info abstract Article history: The Oleriina is one of the most speciose subtribes of the neotropical nymphalid butterfly tribe Ithomiini. Received 9 September 2009 They are widely distributed across the Andes and Amazonian lowlands and like other ithomiines they are Revised 22 December 2009 involved in complex mimicry rings. This subtribe is of particular interest because it contains the most Accepted 9 January 2010 diverse ithomiine genus, Oleria, as well as two genera, Megoleria and Hyposcada, that feed on hostplants Available online 15 January 2010 not utilized elsewhere in the tribe. Here we present the first comprehensive species-level phylogeny for the Oleriina, representing 83% of recognised species in the group, and based on 6698 bp from eight mito- Keywords: chondrial (mt) and nuclear (nc) genes. Topologies are largely congruent for ncDNA and the concatenated Lepidoptera dataset and the genera Oleria, Hyposcada and Megoleria are recovered and well-supported, although Speciation Phylogeny strongly discordant genealogy between mtDNA and ncDNA suggest possible introgression among Hypos- Hybridization cada and Megoleria.