Silurian and Ordovician Conodont Biostratigraphy of the Moose River Basin and Appalachian Basin
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Early Sponge Evolution: a Review and Phylogenetic Framework
Available online at www.sciencedirect.com ScienceDirect Palaeoworld 27 (2018) 1–29 Review Early sponge evolution: A review and phylogenetic framework a,b,∗ a Joseph P. Botting , Lucy A. Muir a Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China b Department of Natural Sciences, Amgueddfa Cymru — National Museum Wales, Cathays Park, Cardiff CF10 3LP, UK Received 27 January 2017; received in revised form 12 May 2017; accepted 5 July 2017 Available online 13 July 2017 Abstract Sponges are one of the critical groups in understanding the early evolution of animals. Traditional views of these relationships are currently being challenged by molecular data, but the debate has so far made little use of recent palaeontological advances that provide an independent perspective on deep sponge evolution. This review summarises the available information, particularly where the fossil record reveals extinct character combinations that directly impinge on our understanding of high-level relationships and evolutionary origins. An evolutionary outline is proposed that includes the major early fossil groups, combining the fossil record with molecular phylogenetics. The key points are as follows. (1) Crown-group sponge classes are difficult to recognise in the fossil record, with the exception of demosponges, the origins of which are now becoming clear. (2) Hexactine spicules were present in the stem lineages of Hexactinellida, Demospongiae, Silicea and probably also Calcarea and Porifera; this spicule type is not diagnostic of hexactinellids in the fossil record. (3) Reticulosans form the stem lineage of Silicea, and probably also Porifera. (4) At least some early-branching groups possessed biminerallic spicules of silica (with axial filament) combined with an outer layer of calcite secreted within an organic sheath. -
Brachiopod Phylogeny in the Cambrian Guliforms, Obolellates and Rhynchonelliforms (E.G., Zhang Et Al., 2009, 2014, 2015; Holmer Et Al., 2018A)
Permophiles Issue #66 Supplement 1 pods which may further address these questions. Glenn A. Brock The world’s oceans are changing. IPCC (2013) predictions Department of Biological Sciences, Macquarie University, NSW suggest that by the end of the current century our seas will be 2109, Australia ZDUPHUDQGWKHLUS+VLJQL¿FDQWO\ORZHU$OWKRXJKWKLVZLOOEH a challenge to all organisms, how will it impact brachiopods? Leonid E. Popov Given that they have a higher proportion of mineralised tissue 'HSDUWPHQWRI*HRORJ\1DWLRQDO0XVHXPRI:DOHV&DUGL൵ than virtually any other invertebrate group what will be the CF10 3NP, UK WKUHDWWRWKHPHLWKHULQWHUPVRIJURZLQJWKHLUVKHOOVLQWKH¿UVW Brachiopods are richly represented in the rock record and instance or repairing and maintaining that shell once it is made? as early as the Cambrian, where they show an impressive diver- In this talk I will review a series of experiments and historical sity of form and in shell morphology (e.g., Harper et al., 2017). studies undertaken with Emma Cross and Lloyd Peck (Cross et 3UHVHQWO\ WKH JURXS LV ¿UPO\ URRWHG ZLWKLQ WKH ORSKRWURFKR- al., 2015, 2016, 2018) that seek to explore the answers to these zoan branch of the bilaterian tree based on molecular data. Our questions. UHVHDUFKKDVLGHQWL¿HGVRPHPHPEHUVRIWKHHQLJPDWLF(DUO\ References Cambrian organophosphatic tommotiids as belonging to the &URVV(/3HFN/6 +DUSHU(02FHDQDFLGL¿FD- brachiopod stem (e.g., Holmer et al., 2002). Subsequent discov- tion does not impact shell growth or repair of the Antarctic HULHVRIWKH¿UVWHYHUDUWLFXODWHGVFOHULWRPHVRIEccentrotheca, brachiopod Liothyrella uva (Broderip, 1833). Journal of Paterimitra, and the inferred bivalved scleritome of Micrina f rom Experimental Marine Biology and Ecology, 462, 29–35. the lower Cambrian of South Australia reveals these three tom- Cross, E.L., Peck, L.S., Lamare, M.D. -
Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction
Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction by Curtis R. Congreve B.S., University of Rochester, 2006 M.S., University of Kansas, 2008 Submitted to the Department of Geology and the Faculty of the Graduate School of The University of Kansas in partial fulfillment on the requirements for the degree of Doctor of Philosophy 2012 Advisory Committee: ______________________________ Bruce Lieberman, Chair ______________________________ Paul Selden ______________________________ David Fowle ______________________________ Ed Wiley ______________________________ Xingong Li Defense Date: December 12, 2012 ii The Dissertation Committee for Curtis R. Congreve certifies that this is the approved Version of the following thesis: Evolutionary Patterns of Trilobites Across the End Ordovician Mass Extinction Advisory Committee: ______________________________ Bruce Lieberman, Chair ______________________________ Paul Selden ______________________________ David Fowle ______________________________ Ed Wiley ______________________________ Xingong Li Accepted: April 18, 2013 iii Abstract: The end Ordovician mass extinction is the second largest extinction event in the history or life and it is classically interpreted as being caused by a sudden and unstable icehouse during otherwise greenhouse conditions. The extinction occurred in two pulses, with a brief rise of a recovery fauna (Hirnantia fauna) between pulses. The extinction patterns of trilobites are studied in this thesis in order to better understand selectivity of the -
CONODONT BIOSTRATIGRAPHY and ... -.: Palaeontologia Polonica
CONODONT BIOSTRATIGRAPHY AND PALEOECOLOGY OF THE PERTH LIMESTONE MEMBER, STAUNTON FORMATION (PENNSYLVANIAN) OF THE ILLINOIS BASIN, U.S.A. CARl B. REXROAD. lEWIS M. BROWN. JOE DEVERA. and REBECCA J. SUMAN Rexroad , c.. Brown . L.. Devera, 1.. and Suman, R. 1998. Conodont biostrati graph y and paleoec ology of the Perth Limestone Member. Staunt on Form ation (Pennsy lvanian) of the Illinois Basin. U.S.A. Ill: H. Szaniawski (ed .), Proceedings of the Sixth European Conodont Symposium (ECOS VI). - Palaeont ologia Polonica, 58 . 247-259. Th e Perth Limestone Member of the Staunton Formation in the southeastern part of the Illinois Basin co nsists ofargill aceous limestone s that are in a facies relati on ship with shales and sandstones that commonly are ca lcareous and fossiliferous. Th e Perth conodo nts are do minated by Idiognathodus incurvus. Hindeodus minutus and Neognathodu s bothrops eac h comprises slightly less than 10% of the fauna. Th e other spec ies are minor consti tuents. The Perth is ass igned to the Neog nathodus bothrops- N. bassleri Sub zon e of the N. bothrops Zo ne. but we were unable to co nfirm its assignment to earliest Desmoin esian as oppose d to latest Atokan. Co nodo nt biofacies associations of the Perth refle ct a shallow near- shore marine environment of generally low to moderate energy. but locali zed areas are more variable. particul ar ly in regard to salinity. K e y w o r d s : Co nodo nta. biozonation. paleoecology. Desmoinesian , Penn sylvanian. Illinois Basin. U.S.A. -
Conodonts in Ordovician Biostratigraphy
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by Archivio istituzionale della ricerca - Università di Modena e Reggio Emilia 1 Conodonts in Ordovician biostratigraphy STIG M. BERGSTRÖM AND ANNALISA FERRETTI Conodonts in Ordovician biostratigraphy The long time interval after Pander’s (1856) original conodont study can in terms of Ordovician conodont biostratigraphic research be subdivided into three periods, namely the Pioneer Period (1856-1955), the Transition Period (1955-1971), and the Modern Period (1971-Recent). During the pre-1920s, the few published conodont investigations were restricted to Europe and North America and were not concerned about the potential use of conodonts as guide fossils. Although primarily of taxonomic nature, the pioneer studies by Branson & Mehl, Stauffer, and Furnish during the 1930s represent the beginning of the use of conodonts in Ordovician biostratigraphy. However, no formal zones were introduced until Lindström (1955) proposed four conodont zones in the Lower Ordovician of Sweden, which marks the end of the Pioneer Period. Because Lindström’s zone classification was not followed by similar work outside Baltoscandia, the time interval up to the late 1960s can be regarded as a Transition Period. A milestone symposium volume, entitled ‘Conodont Biostratigraphy’ and published in 1971, 2 summarized much new information on Ordovician conodont biostratigraphy and is taken as the beginning of the Modern Period of Ordovician conodont biostratigraphy. In this volume, the Baltoscandic Ordovician was subdivided into named conodont zones whereas the North American Ordovician succession was classified into a series of lettered or numbered Faunas. Although most of the latter did not receive zone names until 1984, this classification has been used widely in North America. -
Preliminary Geologic Map of the Baird Mountains and Part of the Selawik Quadrangles, Alaska By
preliminary Geologic Map of the Baird Mountains and part of the Selawik Quadrangles, Alaska by S.M. Karl, J.A. Dumoulin, Inyo Ellersieck, A.G. Harris, and J.M. Schmidt Open-File Report 89-551 This map is preliminary and has not been reviewed for conformity with the North American stratigraphic Code. Any use of trade, product, or firm names is for descriptive purposes only and does not imply endorsement by the U.S. Government. Table Contents ~ntroduction.......................................... 1 Stratigraphic Framework .................................1 Structural Framework ....................................4 Acknowledgments .......................................6 Unit Descriptions ......................................6 Kc ............................................. 6 KJm .............................................6 JPe .............................................7 ~zb................ '. ... ., ,....................... 8 Mzg ............................................10 MzPzi ..........................................10 MZPZ~.......................................... 11 PMC ............................................12 PD1 ............................................12 Mk0 ............................................12 M1 .............................................13 MDue ...........................................14 Mlt ............................................15 Mk .............................................15 MD1 ............................................16 MDe ............................................16 MD~........................................... -
SILURIAN TIMES NEWSLETTER of the INTERNATIONAL SUBCOMMISSION on SILURIAN STRATIGRAPHY (ISSS) (INTERNATIONAL COMMISSION on STRATIGRAPHY, ICS) No
SILURIAN TIMES NEWSLETTER OF THE INTERNATIONAL SUBCOMMISSION ON SILURIAN STRATIGRAPHY (ISSS) (INTERNATIONAL COMMISSION ON STRATIGRAPHY, ICS) No. 27 (for 2019) Edited by ZHAN Renbin INTERNATIONAL UNION OF GEOLOGICAL SCIENCES President: CHENG Qiuming (Canada) Vice-Presidents: Kristine ASCH (Germany) William CAVAZZA (Italy) Secretary General: Stanley C. FINNEY (USA) Treasurer: Hiroshi KITAZATO (Japan) INTERNATIONAL COMMISSION ON STRATIGRAPHY Chairman: David A.T. HARPER (UK) Vice-Chairman: Brian T. HUBER (USA) Secretary General: Philip GIBBARD (UK) SUBCOMMISSION ON SILURIAN STRATIGRAPHY Chairman: Petr ŠTORCH (Czech Republic) Vice-Chairman: Carlo CORRADINI (Italy) Secretary: ZHAN Renbin (China) Other titular members: Anna ANTOSHKINA (Russia) Carlton E. BRETT (USA) Bradley CRAMER (USA) David HOLLOWAY (Australia) Jisuo JIN (Canada) Anna KOZŁOWSKA (Poland) Jiří KŘÍŽ (Czech Republic) David K. LOYDELL (UK) Peep MÄNNIK (Estonia) Michael J. MELCHIN (Canada) Axel MUNNECKE (Germany) Silvio PERALTA (Argentina) Thijs VANDENBROUCKE (Belgium) WANG Yi (China) Živilė ŽIGAITĖ (Lithuania) Silurian Subcommission website: http://silurian.stratigraphy.org 1 CONTENTS CHAIRMAN’S CORNER 3 ANNUAL REPORT OF SILURIAN SUBCOMMISSION FOR 2019 7 INTERNATIONAL COMMISSION ON STRATGRAPHY STATUTES 15 REPORTS OF ACTIVITIES IN 2019 25 1. Report on the ISSS business meeting 2019 25 2. Report on the 15th International Symposium on Early/Lower Vertebrates 28 3. Report on the 13th International Symposium on the Ordovician System in conjunction with the 3rd Annual Meeting of IGCP 653 32 GUIDELINES FOR THE ISSS AWARD: KOREN' AWARD 33 ANNOUNCEMENTS OF MEETINGS and ACTIVITIES 34 1. Lithological Meeting: GEOLOGY OF REEFS 34 SILURIAN RESEARCH 2019: NEWS FROM THE MEMBERS 36 RECENT PUBLICATIONS ON THE SILURIAN RESEARCH 67 MEMBERSHIP NEWS 77 1. List of all Silurian workers and interested colleagues 77 2. -
New Siberian Islands Archipelago)
Detrital zircon ages and provenance of the Upper Paleozoic successions of Kotel’ny Island (New Siberian Islands archipelago) Victoria B. Ershova1,*, Andrei V. Prokopiev2, Andrei K. Khudoley1, Nikolay N. Sobolev3, and Eugeny O. Petrov3 1INSTITUTE OF EARTH SCIENCE, ST. PETERSBURG STATE UNIVERSITY, UNIVERSITETSKAYA NAB. 7/9, ST. PETERSBURG 199034, RUSSIA 2DIAMOND AND PRECIOUS METAL GEOLOGY INSTITUTE, SIBERIAN BRANCH, RUSSIAN ACADEMY OF SCIENCES, LENIN PROSPECT 39, YAKUTSK 677980, RUSSIA 3RUSSIAN GEOLOGICAL RESEARCH INSTITUTE (VSEGEI), SREDNIY PROSPECT 74, ST. PETERSBURG 199106, RUSSIA ABSTRACT Plate-tectonic models for the Paleozoic evolution of the Arctic are numerous and diverse. Our detrital zircon provenance study of Upper Paleozoic sandstones from Kotel’ny Island (New Siberian Island archipelago) provides new data on the provenance of clastic sediments and crustal affinity of the New Siberian Islands. Upper Devonian–Lower Carboniferous deposits yield detrital zircon populations that are consistent with the age of magmatic and metamorphic rocks within the Grenvillian-Sveconorwegian, Timanian, and Caledonian orogenic belts, but not with the Siberian craton. The Kolmogorov-Smirnov test reveals a strong similarity between detrital zircon populations within Devonian–Permian clastics of the New Siberian Islands, Wrangel Island (and possibly Chukotka), and the Severnaya Zemlya Archipelago. These results suggest that the New Siberian Islands, along with Wrangel Island and the Severnaya Zemlya Archipelago, were located along the northern margin of Laurentia-Baltica in the Late Devonian–Mississippian and possibly made up a single tectonic block. Detrital zircon populations from the Permian clastics record a dramatic shift to a Uralian provenance. The data and results presented here provide vital information to aid Paleozoic tectonic reconstructions of the Arctic region prior to opening of the Mesozoic oceanic basins. -
Early Silurian Oceanic Episodes and Events
Journal of the Geological Society, London, Vol. 150, 1993, pp. 501-513, 3 figs. Printed in Northern Ireland Early Silurian oceanic episodes and events R. J. ALDRIDGE l, L. JEPPSSON 2 & K. J. DORNING 3 1Department of Geology, The University, Leicester LE1 7RH, UK 2Department of Historical Geology and Palaeontology, SiSlvegatan 13, S-223 62 Lund, Sweden 3pallab Research, 58 Robertson Road, Sheffield $6 5DX, UK Abstract: Biotic cycles in the early Silurian correlate broadly with postulated sea-level changes, but are better explained by a model that involves episodic changes in oceanic state. Primo episodes were characterized by cool high-latitude climates, cold oceanic bottom waters, and high nutrient supply which supported abundant and diverse planktonic communities. Secundo episodes were characterized by warmer high-latitude climates, salinity-dense oceanic bottom waters, low diversity planktonic communities, and carbonate formation in shallow waters. Extinction events occurred between primo and secundo episodes, with stepwise extinctions of taxa reflecting fluctuating conditions during the transition period. The pattern of turnover shown by conodont faunas, together with sedimentological information and data from other fossil groups, permit the identification of two cycles in the Llandovery to earliest Weniock interval. The episodes and events within these cycles are named: the Spirodden Secundo episode, the Jong Primo episode, the Sandvika event, the Malm#ykalven Secundo episode, the Snipklint Primo episode, and the lreviken event. Oceanic and climatic cyclicity is being increasingly semblages (Johnson et al. 1991b, p. 145). Using this recognized in the geological record, and linked to major and approach, they were able to detect four cycles within the minor sedimentological and biotic fluctuations. -
Plate Tectonic Regulation of Global Marine Animal Diversity
Plate tectonic regulation of global marine animal diversity Andrew Zaffosa,1, Seth Finneganb, and Shanan E. Petersa aDepartment of Geoscience, University of Wisconsin–Madison, Madison, WI 53706; and bDepartment of Integrative Biology, University of California, Berkeley, CA 94720 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved April 13, 2017 (received for review February 13, 2017) Valentine and Moores [Valentine JW, Moores EM (1970) Nature which may be complicated by spatial and temporal inequities in 228:657–659] hypothesized that plate tectonics regulates global the quantity or quality of samples (11–18). Nevertheless, many biodiversity by changing the geographic arrangement of conti- major features in the fossil record of biodiversity are consis- nental crust, but the data required to fully test the hypothesis tently reproducible, although not all have universally accepted were not available. Here, we use a global database of marine explanations. In particular, the reasons for a long Paleozoic animal fossil occurrences and a paleogeographic reconstruction plateau in marine richness and a steady rise in biodiversity dur- model to test the hypothesis that temporal patterns of continen- ing the Late Mesozoic–Cenozoic remain contentious (11, 12, tal fragmentation have impacted global Phanerozoic biodiversity. 14, 19–22). We find a positive correlation between global marine inverte- Here, we explicitly test the plate tectonic regulation hypothesis brate genus richness and an independently derived quantitative articulated by Valentine and Moores (1) by measuring the extent index describing the fragmentation of continental crust during to which the fragmentation of continental crust covaries with supercontinental coalescence–breakup cycles. The observed posi- global genus-level richness among skeletonized marine inverte- tive correlation between global biodiversity and continental frag- brates. -
Revised Correlation of Silurian Provincial Series of North America with Global and Regional Chronostratigraphic Units 13 and D Ccarb Chemostratigraphy
Revised correlation of Silurian Provincial Series of North America with global and regional chronostratigraphic units 13 and d Ccarb chemostratigraphy BRADLEY D. CRAMER, CARLTON E. BRETT, MICHAEL J. MELCHIN, PEEP MA¨ NNIK, MARK A. KLEFF- NER, PATRICK I. MCLAUGHLIN, DAVID K. LOYDELL, AXEL MUNNECKE, LENNART JEPPSSON, CARLO CORRADINI, FRANK R. BRUNTON AND MATTHEW R. SALTZMAN Cramer, B.D., Brett, C.E., Melchin, M.J., Ma¨nnik, P., Kleffner, M.A., McLaughlin, P.I., Loydell, D.K., Munnecke, A., Jeppsson, L., Corradini, C., Brunton, F.R. & Saltzman, M.R. 2011: Revised correlation of Silurian Provincial Series of North America with global 13 and regional chronostratigraphic units and d Ccarb chemostratigraphy. Lethaia,Vol.44, pp. 185–202. Recent revisions to the biostratigraphic and chronostratigraphic assignment of strata from the type area of the Niagaran Provincial Series (a regional chronostratigraphic unit) have demonstrated the need to revise the chronostratigraphic correlation of the Silurian System of North America. Recently, the working group to restudy the base of the Wen- lock Series has developed an extremely high-resolution global chronostratigraphy for the Telychian and Sheinwoodian stages by integrating graptolite and conodont biostratigra- 13 phy with carbonate carbon isotope (d Ccarb) chemostratigraphy. This improved global chronostratigraphy has required such significant chronostratigraphic revisions to the North American succession that much of the Silurian System in North America is cur- rently in a state of flux and needs further refinement. This report serves as an update of the progress on recalibrating the global chronostratigraphic correlation of North Ameri- can Provincial Series and Stage boundaries in their type area. -
CONODONTS of the MOJCZA LIMESTONE -.: Palaeontologia Polonica
CONODONTS OF THE MOJCZA LIMESTONE JERZY DZIK Dzik, J. 1994. Conodonts of the M6jcza Limestone. -In: J. Dzik, E. Olemp ska, and A. Pisera 1994. Ordovician carbonate platform ecosystem of the Holy Cross Moun tains. Palaeontologia Polonica 53, 43-128. The Ordovician organodetrital limestones and marls studied in outcrops at M6jcza and Miedzygorz, Holy Cross Mts, Poland, contains a record of the evolution of local conodont faunas from the latest Arenig (Early Kundan, Lenodus variabilis Zone) to the Ashgill (Amorphognathus ordovicicus Zone), with a single larger hiatus corre sponding to the subzones from Eop/acognathus pseudop/anu s to E. reclinatu s. The conodont fauna is Baltic in general appearance but cold water genera , like Sagitto dontina, Scabbardella, and Hamarodus, as well as those of Welsh or Chinese af finities, like Comp/exodus, Phragmodus, and Rhodesognathu s are dominant in par ticular parts of the section while others common in the Baltic region, like Periodon , Eop/acognathus, and Sca/pellodus are extremely rare. Most of the lineages continue to occur throughout most of the section enabling quantitative studies on their phyletic evolut ion. Apparatuses of sixty seven species of thirty six genera are described and illustrated. Phyletic evolution of Ba/toniodus, Amorphognathu s, Comp/exodus, and Pygodus is biometrically documented. Element s of apparatu ses are homolog ized and the standard notation system is applied to all of them. Acodontidae fam. n., Drepa nodus kie/censis sp. n., and D. santacrucensis sp. n. are proposed . Ke y w o r d s: conodonts, Ordovici an, evolut ion, taxonomy. Jerzy Dzik, Instytut Paleobiologii PAN, A/eja Zwirk i i Wigury 93, 02-089 Warszawa , Poland.