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A Neoceratopsian Dinosaur from the Early Cretaceous of Mongolia And
ARTICLE https://doi.org/10.1038/s42003-020-01222-7 OPEN A neoceratopsian dinosaur from the early Cretaceous of Mongolia and the early evolution of ceratopsia ✉ Congyu Yu 1 , Albert Prieto-Marquez2, Tsogtbaatar Chinzorig 3,4, Zorigt Badamkhatan4,5 & Mark Norell1 1234567890():,; Ceratopsia is a diverse dinosaur clade from the Middle Jurassic to Late Cretaceous with early diversification in East Asia. However, the phylogeny of basal ceratopsians remains unclear. Here we report a new basal neoceratopsian dinosaur Beg tse based on a partial skull from Baruunbayan, Ömnögovi aimag, Mongolia. Beg is diagnosed by a unique combination of primitive and derived characters including a primitively deep premaxilla with four pre- maxillary teeth, a trapezoidal antorbital fossa with a poorly delineated anterior margin, very short dentary with an expanded and shallow groove on lateral surface, the derived presence of a robust jugal having a foramen on its anteromedial surface, and five equally spaced tubercles on the lateral ridge of the surangular. This is to our knowledge the earliest known occurrence of basal neoceratopsian in Mongolia, where this group was previously only known from Late Cretaceous strata. Phylogenetic analysis indicates that it is sister to all other neoceratopsian dinosaurs. 1 Division of Vertebrate Paleontology, American Museum of Natural History, New York 10024, USA. 2 Institut Català de Paleontologia Miquel Crusafont, ICTA-ICP, Edifici Z, c/de les Columnes s/n Campus de la Universitat Autònoma de Barcelona, 08193 Cerdanyola del Vallès Sabadell, Barcelona, Spain. 3 Department of Biological Sciences, North Carolina State University, Raleigh, NC 27695, USA. 4 Institute of Paleontology, Mongolian Academy of Sciences, ✉ Ulaanbaatar 15160, Mongolia. -
Leophils Welt Hauptthema Fußball
Leophils Welt Die Zeitschrift für die Mitglie- der der Jungen Briefmarken- freunde Hessen Ausgabe 2 Jahrgang 4 Hauptthema Fußball www.briefmarkenjugend-hessen.de Inhalt Vorwort ............................................................................................................................. 3 Fußball Europameisterschaft der Männer 2016 in Frankreich ............................. 5 Die Geschichte des Fußballs ......................................................................................... 8 Fußball bei den Olympischen Spielen ........................................................................ 12 Entenschnabel-Dinosaurier, Teil 4 ............................................................................ 14 Neue Sondermarken in Deutschland ......................................................................... 21 Der Basilisk von Wien .................................................................................................. 30 Aus den Gruppen ........................................................................................................... 32 Muss eine Briefmarke immer auf Papier gedruckt sein? .................................... 34 Erzabtei St. Peter in Salzburg ................................................................................. 36 Dauerserien - der Reiz der Komplettierung ........................................................... 37 Rätsel .............................................................................................................................. 39 Hier stimmt -
At Carowinds
at Carowinds EDUCATOR’S GUIDE CLASSROOM LESSON PLANS & FIELD TRIP ACTIVITIES Table of Contents at Carowinds Introduction The Field Trip ................................... 2 The Educator’s Guide ....................... 3 Field Trip Activity .................................. 4 Lesson Plans Lesson 1: Form and Function ........... 6 Lesson 2: Dinosaur Detectives ....... 10 Lesson 3: Mesozoic Math .............. 14 Lesson 4: Fossil Stories.................. 22 Games & Puzzles Crossword Puzzles ......................... 29 Logic Puzzles ................................. 32 Word Searches ............................... 37 Answer Keys ...................................... 39 Additional Resources © 2012 Dinosaurs Unearthed Recommended Reading ................. 44 All rights reserved. Except for educational fair use, no portion of this guide may be reproduced, stored in a retrieval system, or transmitted in any form or by any Dinosaur Data ................................ 45 means—electronic, mechanical, photocopy, recording, or any other without Discovering Dinosaurs .................... 52 explicit prior permission from Dinosaurs Unearthed. Multiple copies may only be made by or for the teacher for class use. Glossary .............................................. 54 Content co-created by TurnKey Education, Inc. and Dinosaurs Unearthed, 2012 Standards www.turnkeyeducation.net www.dinosaursunearthed.com Curriculum Standards .................... 59 Introduction The Field Trip From the time of the first exhibition unveiled in 1854 at the Crystal -
A Revision of the Ceratopsia Or Horned Dinosaurs
MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY VOLUME III, 1 A.R1 A REVISION orf tneth< CERATOPSIA OR HORNED DINOSAURS BY RICHARD SWANN LULL STERLING PROFESSOR OF PALEONTOLOGY AND DIRECTOR OF PEABODY MUSEUM, YALE UNIVERSITY LVXET NEW HAVEN, CONN. *933 MEMOIRS OF THE PEABODY MUSEUM OF NATURAL HISTORY YALE UNIVERSITY Volume I. Odontornithes: A Monograph on the Extinct Toothed Birds of North America. By Othniel Charles Marsh. Pp. i-ix, 1-201, pis. 1-34, text figs. 1-40. 1880. To be obtained from the Peabody Museum. Price $3. Volume II. Part 1. Brachiospongidae : A Memoir on a Group of Silurian Sponges. By Charles Emerson Beecher. Pp. 1-28, pis. 1-6, text figs. 1-4. 1889. To be obtained from the Peabody Museum. Price $1. Volume III. Part 1. American Mesozoic Mammalia. By George Gaylord Simp- son. Pp. i-xvi, 1-171, pis. 1-32, text figs. 1-62. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $5. Part 2. A Remarkable Ground Sloth. By Richard Swann Lull. Pp. i-x, 1-20, pis. 1-9, text figs. 1-3. 1929. To be obtained from the Yale University Press, New Haven, Conn. Price $1. Part 3. A Revision of the Ceratopsia or Horned Dinosaurs. By Richard Swann Lull. Pp. i-xii, 1-175, pis. I-XVII, text figs. 1-42. 1933. To be obtained from the Peabody Museum. Price $5 (bound in cloth), $4 (bound in paper). Part 4. The Merycoidodontidae, an Extinct Group of Ruminant Mammals. By Malcolm Rutherford Thorpe. In preparation. -
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses By
Histology and Ontogeny of Pachyrhinosaurus Nasal Bosses by Elizabeth Kruk A thesis submitted in partial fulfillment of the requirements for the degree of Master of Science in Systematics and Evolution Department of Biological Sciences University of Alberta © Elizabeth Kruk, 2015 Abstract Pachyrhinosaurus is a peculiar ceratopsian known only from Upper Cretaceous strata of Alberta and the North Slope of Alaska. The genus consists of three described species Pachyrhinosaurus canadensis, Pachyrhinosaurus lakustai, and Pachyrhinosaurus perotorum that are distinguishable by cranial characteristics, including parietal horn shape and orientation, absence/presence of a rostral comb, median parietal bar horns, and profile of the nasal boss. A fourth species of Pachyrhinosaurus is described herein and placed into its phylogenetic context within Centrosaurinae. This new species forms a polytomy at the crown with Pachyrhinosaurus canadensis and Pachyrhinosaurus perotorum, with Pachyrhinosaurus lakustai falling basal to that polytomy. The diagnostic features of this new species are an apomorphic, laterally curved Process 3 horns and a thick longitudinal ridge separating the supraorbital bosses. Another focus is investigating the ontogeny of Pachyrhinosaurus nasal bosses in a histological context. Previously, little work has been done on cranial histology in ceratopsians, focusing instead on potential integumentary structures, the parietals of Triceratops, and how surface texture relates to underlying histological structures. An ontogenetic series is established for the nasal bosses of Pachyrhinosaurus at both relative (subadult versus adult) and fine scale (Stages 1-5). It was demonstrated that histology alone can indicate relative ontogenetic level, but not stages of a finer scale. Through Pachyrhinosaurus ontogeny the nasal boss undergoes increased vascularity and secondary remodeling with a reduction in osteocyte lacunar density. -
A Census of Dinosaur Fossils Recovered from the Hell Creek and Lance Formations (Maastrichtian)
The Journal of Paleontological Sciences: JPS.C.2019.01 1 TAKING COUNT: A Census of Dinosaur Fossils Recovered From the Hell Creek and Lance Formations (Maastrichtian). ______________________________________________________________________________________ Walter W. Stein- President, PaleoAdventures 1432 Mill St.. Belle Fourche, SD 57717. [email protected] 605-210-1275 ABSTRACT: A census of Hell Creek and Lance Formation dinosaur remains was conducted from April, 2017 through February of 2018. Online databases were reviewed and curators and collections managers interviewed in an effort to determine how much material had been collected over the past 130+ years of exploration. The results of this new census has led to numerous observations regarding the quantity, quality, and locations of the total collection, as well as ancillary data on the faunal diversity and density of Late Cretaceous dinosaur populations. By reviewing the available data, it was also possible to make general observations regarding the current state of certain exploration programs, the nature of collection bias present in those collections and the availability of today's online databases. A total of 653 distinct, associated and/or articulated remains (skulls and partial skeletons) were located. Ceratopsid skulls and partial skeletons (mostly identified as Triceratops) were the most numerous, tallying over 335+ specimens. Hadrosaurids (Edmontosaurus) were second with at least 149 associated and/or articulated remains. Tyrannosaurids (Tyrannosaurus and Nanotyrannus) were third with a total of 71 associated and/or articulated specimens currently known to exist. Basal ornithopods (Thescelosaurus) were also well represented by at least 42 known associated and/or articulated remains. The remaining associated and/or articulated specimens, included pachycephalosaurids (18), ankylosaurids (6) nodosaurids (6), ornithomimids (13), oviraptorosaurids (9), dromaeosaurids (1) and troodontids (1). -
A Subadult Individual of Styracosaurus Albertensis
Vertebrate Anatomy Morphology Palaeontology 8:67–95 67 ISSN 2292-1389 A subadult individual of Styracosaurus albertensis (Ornithischia: Ceratopsidae) with comments on ontogeny and intraspecific variation inStyracosaurus and Centrosaurus Caleb M. Brown1,*, Robert B. Holmes2, Philip J. Currie2 1Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, AB, T0J 0Y0, Canada; [email protected] 2Department of Biological Sciences, University of Alberta, Edmonton, Alberta, T6G 2E9, Canada; [email protected]; [email protected] Abstract: Styracosaurus albertensis is an iconic centrosaurine horned dinosaur from the Campanian of Alberta, Canada, known for its large spike-like parietal processes. Although described over 100 years ago, subsequent dis- coveries were rare until the last few decades, during which time several new skulls, skeletons, and bonebeds were found. Here we described an immature individual, the smallest known for the species, represented by a complete skull and fragmentary skeleton. Although ~80% maximum size, it possesses a suite of characters associated with immaturity, and is regarded as a subadult. The ornamentation is characterized by a small, recurved, but fused nasal horncore; short, rounded postorbital horncores; and short, triangular, and flat parietal processes. Using this specimen, and additional skulls and bonebed material, the cranial ontogeny of Styracosaurus is described, and compared to Centrosaurus. In early ontogeny, the nasal horncores of Styracosaurus and Centrosaurus are thin, recurved, and unfused, but in the former the recurved morphology is retained into large adult size and the horncore never develops the procurved morphology common in Centrosaurus. The postorbital horncores of Styracosaurus are shorter and more rounded than those of Centrosaurus throughout ontogeny, and show great- er resorption later in ontogeny. -
A New Maastrichtian Species of the Centrosaurine Ceratopsid Pachyrhinosaurus from the North Slope of Alaska
A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska ANTHONY R. FIORILLO and RONALD S. TYKOSKI Fiorillo, A.R. and Tykoski, R.S. 2012. A new Maastrichtian species of the centrosaurine ceratopsid Pachyrhinosaurus from the North Slope of Alaska. Acta Palaeontologica Polonica 57 (3): 561–573. The Cretaceous rocks of the Prince Creek Formation contain the richest record of polar dinosaurs found anywhere in the world. Here we describe a new species of horned dinosaur, Pachyrhinosaurus perotorum that exhibits an apomorphic character in the frill, as well as a unique combination of other characters. Phylogenetic analysis of 16 taxa of ceratopsians failed to resolve relationships between P. perotorum and other Pachyrhinosaurus species (P. canadensis and P. lakustai). P. perotorum shares characters with each of the previously known species that are not present in the other, including very large nasal and supraorbital bosses that are nearly in contact and separated only by a narrow groove as in P. canadensis, and a rostral comb formed by the nasals and premaxillae as in P. lakustai. P. perotorum is the youngest centrosaurine known (70–69 Ma), and the locality that produced the taxon, the Kikak−Tegoseak Quarry, is close to the highest latitude for recovery of ceratopsid remains. Key words: Dinosauria, Centrosaurinae, Cretaceous, Prince Creek Formation, Kikak−Tegoseak Quarry, Arctic. Anthony R. Fiorillo [[email protected]] and Ronald S. Tykoski [[email protected]], Perot Museum of Nature and Science, 2201 N. Field Street, Dallas, TX 75202, USA. Received 4 April 2011, accepted 23 July 2011, available online 26 August 2011. -
Competition Structured a Late Cretaceous Megaherbivorous Dinosaur Assemblage Jordan C
www.nature.com/scientificreports OPEN Competition structured a Late Cretaceous megaherbivorous dinosaur assemblage Jordan C. Mallon 1,2 Modern megaherbivore community richness is limited by bottom-up controls, such as resource limitation and resultant dietary competition. However, the extent to which these same controls impacted the richness of fossil megaherbivore communities is poorly understood. The present study investigates the matter with reference to the megaherbivorous dinosaur assemblage from the middle to upper Campanian Dinosaur Park Formation of Alberta, Canada. Using a meta-analysis of 21 ecomorphological variables measured across 14 genera, contemporaneous taxa are demonstrably well-separated in ecomorphospace at the family/subfamily level. Moreover, this pattern is persistent through the approximately 1.5 Myr timespan of the formation, despite continual species turnover, indicative of underlying structural principles imposed by long-term ecological competition. After considering the implications of ecomorphology for megaherbivorous dinosaur diet, it is concluded that competition structured comparable megaherbivorous dinosaur communities throughout the Late Cretaceous of western North America. Te question of which mechanisms regulate species coexistence is fundamental to understanding the evolution of biodiversity1. Te standing diversity (richness) of extant megaherbivore (herbivores weighing ≥1,000 kg) com- munities appears to be mainly regulated by bottom-up controls2–4 as these animals are virtually invulnerable to top-down down processes (e.g., predation) when fully grown. Tus, while the young may occasionally succumb to predation, fully-grown African elephants (Loxodonta africana), rhinoceroses (Ceratotherium simum and Diceros bicornis), hippopotamuses (Hippopotamus amphibius), and girafes (Girafa camelopardalis) are rarely targeted by predators, and ofen show indiference to their presence in the wild5. -
Raptors in Action 1 Suggested Pre-Visit Activities
PROGRAM OVERVIEW TOPIC: Small theropods commonly known as “raptors.” THEME: Explore the adaptations that made raptors unique and successful, like claws, intelligence, vision, speed, and hollow bones. PROGRAM DESCRIPTION: Razor-sharp teeth and sickle-like claws are just a few of the characteristics that have made raptors famous. Working in groups, students will build a working model of a raptor leg and then bring it to life while competing in a relay race that simulates the hunting techniques of these carnivorous animals. AUDIENCE: Grades 3–6 CURRICULUM CONNECTIONS: Grade 3 Science: Building with a Variety of Materials Grade 3–6 Math: Patterns and Relations Grade 4 Science: Building Devices and Vehicles that Move Grade 6 Science: Evidence and Investigation PROGRAM ObJECTIVES: 1. Students will understand the adaptations that contributed to the success of small theropods. 2. Students will explore the function of the muscles used in vertebrate movement and the mechanics of how a raptor leg works. 3. Students will understand the function of the raptorial claw. 4. Students will discover connections between small theropod dinosaurs and birds. SUGGESTED PRE-VISIT ACTIVITIES UNDERstANDING CLADIstICS Animals and plants are often referred to as part of a family or group. For example, the dog is part of the canine family (along with wolves, coyotes, foxes, etc.). Scientists group living things together based on relationships to gain insight into where they came from. This helps us identify common ancestors of different organisms. This method of grouping is called “cladistics.” Cladistics is a system that uses branches like a family tree to show how organisms are related to one another. -
A New Centrosaurine from the Late Cretaceous of Alberta, Canada, and the Evolution of Parietal Ornamentation in Horned Dinosaurs
A new centrosaurine from the Late Cretaceous of Alberta, Canada, and the evolution of parietal ornamentation in horned dinosaurs ANDREW A. FARKE, MICHAEL J. RYAN, PAUL M. BARRETT, DARREN H. TANKE, DENNIS R. BRAMAN, MARK A. LOEWEN, and MARK R. GRAHAM Farke, A.A., Ryan, M.J., Barrett, P.M., Tanke, D.H., Braman, D.R., Loewen, M.A., and Graham, M.R. 2011. A new centrosaurine from the Late Cretaceous of Alberta, Canada, and the evolution of parietal ornamentation in horned dino− saurs. Acta Palaeontologica Polonica 56 (4): 691–702. In 1916, a centrosaurine dinosaur bonebed was excavated within the Campanian−aged deposits of what is now Dinosaur Provincial Park, Alberta, Canada. Specimens from this now−lost quarry, including two parietals, a squamosal, a skull missing the frill, and an incomplete dentary, were purchased by The Natural History Museum, London. The material was recently reprepared and identified herein as a previously unknown taxon, Spinops sternbergorum gen. et sp. nov. Based upon the available locality data and paleopalynology, the quarry lies in either the upper part of the Oldman Formation or the lower part of the Dinosaur Park Formation. The facial region of the partial skull is similar to putative mature speci− mens of Centrosaurus spp. and Styracosaurus albertensis, with short, rounded postorbital horncores and a large, erect na− sal horncore. Parietal ornamentation is consistent on both known parietals and is unique among ceratopsids. Bilateral, procurved parietal hooks occupy the P1 (medial−most) position on the dorsal surface of the parietal and are very similar to those seen in Centrosaurus apertus. -
The Children's Museum of Indianapolis
The Children’s Museum of Indianapolis Dinosphere Dinosaurs Acknowledgments The Children’s Museum of Indianapolis wishes to acknowledge the assistance of the following people in the preparation of this unit of study: Rick Crosslin, teacher, writer Mary Fortney, educator Dinosphere Exhibit Development Team The Children’s Museum of Indianapolis The Children’s Museum of Indianapolis is a nonprofit institution dedicated to providing extraordinary learning experiences for children and families. It is one of the largest children’s museums in the world and serves people across Indiana as well as visitors from other states and countries. In addition to special exhibits and programs, the museum provides the infoZone, a partnership between The Children’s Museum of Indianapolis and The Indianapolis-Marion County Public Library. The infoZone combines the resources of a museum with the services of a library where students can read, search for information and find the answers to their questions. Other museum services include the Teacher Resource Link that lends books, learning kits, artifacts and other materials to Indiana educators. Items may be checked out for minimal fees. For a complete catalog, call (317) 334-4001 or fax (317) 921-4019. Field trips to the museum can be arranged by calling (317) 334-4000 or (800) 820-6214. Visit Just for Teachers at The Children’s Museum Web site: www.ChildrensMuseum.org 2 Dinosphere — Now You’re in Their World! • A 3 – 5 Unit of Study Dinosphere Get ready Unit of Study to dig Enduring idea: Experiences Indiana dinosaurs Make it fossilize Focus questions Dino Diary What's ahead Dino Dinosphere Web sites museum link Dino books Science class environment Paleo-points for the teacher Dinosaur classroom Bonus: Literature connection Digging deeper! Indiana academic Introduction standards Dinosphere Family connection A 3 – 5 Table of Contents Science names Unit of Study Introduction................................