Xiphactinus Audax

Total Page:16

File Type:pdf, Size:1020Kb

Xiphactinus Audax Vertebrate Anatomy Morphology Palaeontology 1(1):89-100 89 ISSN 2292-1389 Xiphactinus audax Leidy 1870 from the Puskwaskau Formation (Santonian to Campanian) of north- western Alberta, Canada and the distribution of Xiphactinus in North America Matthew J. Vavrek1, Alison M. Murray2 and Phil R. Bell3 1Royal Ontario Museum, Department of Natural History, 100 Queen’s Park, Toronto, ON M5S 2C6, Canada, [email protected] 2Department of Biological Sciences, University of Alberta, Edmonton, AB T6G 2E9, Canada, [email protected] 3Department of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia, [email protected] Abstract: Xiphactinus is the largest teleost fish known from the Late Cretaceous of North America, and has been found across much of the Western Interior Basin. Despite extensive Late Cretaceous marine deposits occurring in Alberta, there has previ- ously been only two possible records of Xiphactinus from the province, neither of which has been diagnosable to the species level. We describe here a portion of the lower jaws, including teeth, of Xiphactinus audax from northeast of Grande Prairie, Alberta. The fossil has large, thecodont teeth that are circular in cross section and lack any carinae, and are highly variable in their overall size. This fossil is the first diagnostic material of X. audax from Alberta, and extends the range of the species by over a thousand kilometres. During the Late Cretaceous, the area the fossil was found in was near the Arctic Circle, and repre- sents an important datapoint within the poorly known, northern portion of the Western Interior Basin. Key Words: Ichthyodectiformes; Late Cretaceous; marine; Teleostei; Western Interior Basin INTRODUCTION from even larger individuals are also known (Bardack 1965; Shimada and Everhart 2004). The ichthyodectid Xiphactinus is the largest teleost fish Despite extensive marine deposits in Alberta, to date known from the Late Cretaceous of North America only two other questionable records of Xiphactinus have (Bardack 1965; Schwimmer et al. 1997; Shimada et al. been reported from the province. The first record is from 2006). Specimens of Xiphactinus have been found from “east of Lesser Slave Lake” noted by Bardack (1965). Cenomanian (Cumbaa et al. 2006) through Campanian However, he listed no referred material and gave no de- (DeMar and Breithaupt 2006) deposits throughout much scription of any specimen, with the record consisting only of the southern Western Interior Basin, from central of a personal communication from Wann Langston. A Saskatchewan (Cumbaa and Tokaryk 1999; Cumbaa et al. search of likely museum collections where any such fossils 2006) south to Texas (Bardack 1965), as well as Campanian would have been accessioned has yielded nothing. The to Maastrichtian beds along the Atlantic coast in New general lack of appropriate-aged fossil bearing deposits Jersey (Grandstaff et al. 1992), North Carolina, Georgia from that area, as well as the lack of referable or figured and Alabama (Schwimmer et al. 1997). Although known specimens, makes this record dubious. A second Albertan from a variety of formations, the most spectacular speci- record consists of two isolated scales from the Kaskapau mens of Xiphactinus come from the Niobrara Formation of Formation near Watino, Alberta (Wilson and Chalifa Kansas, where well-preserved, virtually complete skeletons 1989), although those authors note that the scales do not were being excavated as early as the 1870s (Cope 1872; conform in every aspect with those of Xiphactinus and so Bardack 1965). Some of these nearly complete specimens referred the specimens to “cf. Xiphactinus”. are close to 5 m in total length, although isolated elements The specimen we describe here, TMP 1973.011.3081, was first mentioned in an unpublished PhD thesis by Published 4 February, 2016 © 2016 by the authors Christopher Collom (2001), and later mentioned again by submitted August 23, 2015; revisions received January 28, 2016; Bell et al. (2014). However, the specimen itself was never accepted January 29, 2016. Handling editor: Robert Holmes. Vertebrate Anatomy Morphology Palaeontology is an open access journal http://ejournals.library.ualberta.ca/index.php/VAMP Article copyright by the author(s). This open access work is distributed under a Creative Commons Attribution 4.0 International (CC By 4.0) License, meaning you must give appropriate credit, provide a link to the license, and indicate if changes were made. You may do so in any reasonable manner, but not in any way that suggests the licensor endorses you or your use. No additional restrictions — You may not apply legal terms or technological measures that legally restrict others from doing anything the license permits. Vertebrate Anatomy Morphology Palaeontology 1(1):89-100 described and was only assigned to the genus Xiphactinus. Although the exact stratigraphic position of the fossil is Further study of the specimen has revealed a number of unknown, the matrix still attached to the fossil is similar in characteristics that allow the assignment of the specimen nature to a number of other marine vertebrate specimens more precisely to Xiphactinus audax, expanding the range found in the area, particularly those discussed by Bell et of this species considerably. The specimen itself is nota- al. (2014). The majority of the specimens discussed in that ble for its relatively northern location within the Western paper were inferred to have come from the upper portions Interior Seaway, and also confirms and fully documents the of the Puskwaskau Formation, likely the Chungo or Nomad presence of Xiphactinus in Alberta. members (Bell et al. 2014). This is further corroborated by Institutional Abbreviations: CMN, Canadian the location where TMP 1973.011.3081 was found, on Museum of Nature, Ottawa, Ontario, Canada; TMP, Royal a relatively small, shallow creek that drains a limited area. Tyrrell Museum of Palaeontology, Drumheller, Alberta, Geologic maps of the region indicate that the creek drains Canada; UALVP, University of Alberta Laboratory for an area that is mainly part of the upper portions of the Vertebrate Palaeontology, Edmonton, Alberta, Canada. Puskwaskau Formation, concordant with the idea that the fossil comes from one of the upper members of the forma- GEOLOGICAL SETTING tion. Assuming TMP 1973.011.3081 is from one of these upper members, it would be early Campanian in age. The Puskwaskau Formation is a marine, mudstone-dom- inated wedge up to 350 m in thickness, that was deposited SYSTEMATIC PALAEONTOLOGY in relatively shallow waters of the Western Interior Seaway (Hu and Plint 2009). The Puskwaskau was deposited during Order ICHTHYODECTIFORMES Bardack and the Niobrara Cycle (Kauffman 1977), and is Santonian to Sprinkle, 1969 early Campanian in age (Hu and Plint 2009; Fig. 1). The Family ICHTHYODECTIDAE Patterson and Rosen 1977 Puskwaskau is divided into five members, which are, in Genus Xiphactinus Leidy 1870 ascending order, Dowling, Thistle, Hanson, Chungo and Xiphactinus audax Leidy 1870 Nomad (Stott 1963, 1967; Hu and Plint 2009). Collom Figure 2 (2001) suggested that the Santonian/Campanian boundary is within the upper part of the Hanson Member, placing the Referred Material: TMP 1973.011.3081, the anterior Chungo and Nomad members within the lower Campanian. portion of articulated left and right dentaries. Figure 1. A) Summary stratigraph- ic chart for the Santonian to early Campanian of northwestern Alberta. B) Maps showing, counter-clockwise from top right: location of Alberta within Canada; location of regional map of Grande Prairie area within Al- berta; location of fossil site along Ka- kut Creek, northeast of Grande Prai- rie. Location of TMP 1973.011.3081 is marked with an arrow. 90 Vavrek et al. — Xiphactinus from Alberta Figure 2. TMP 1973.011.3081, partial jaws of Xiphactinus audax Leidy 1870. A) Right lateral view. B) Left lateral view. C) Posterior view. For all line drawings matrix is shaded white, bone is shaded light grey, teeth are shaded dark grey, and cross-hatching indicates areas of exposed tooth root. Locality and Horizon: The fossil was likely collected Description: TMP 1973.011.3081 preserves approximately sometime in the 1950s or 1960s, based on the personal the anterior one-third of the left and right dentaries, and meas- collection records of George Robinson, the original collector. ures 16 cm long. The dentary symphysis forms a sigmoidal The notes associated with the fossil in the TMP collections curve in lateral view with a tooth positioned on each side right state that the specimen was “from the Bad Heart River near beside the symphysis. The pores for the mandibular canal are bridge.” However, in Robinson’s original collections ledg- not clearly visible as this area of the jaw is somewhat dis- er, he noted that he found the fossil along “Kakut Ck near torted and very worn. The symphysis measures 9.2 cm in a bridge on Wanham Rd” and that it came from the “Bad straight line from the lower edge to the upper, which is close Heart Sst.” Another note he made, beside the entry for sev- to the average length for X. audax [8.7 cm with a range of eral invertebrate specimens collected from the same location, 6.0–13.85 based on measurements of 81 specimens (Bardack states that the bridge had been replaced by a culvert. His 1965)]. The dorsal surfaces of the dentaries undulate, being description, and comparison with historical maps of the area deepest under the largest tooth. and modern maps, shows that the fossil came from exposures There are six right and six left tooth positions preserved, near Kakut Creek where the present day Secondary Highway some of which contain almost complete teeth while others 733 crosses it, at approximately 55.61°N 118.39°W. The are represented only by the tooth base. The two anterior-most exact horizon the fossil came from is unknown, and it was teeth on either side of the symphysis are fairly large, and likely found ex situ, as the unprepared portions of matrix still would have reached an estimated 3 cm above the dentary attached appear to have been water worn, likely from sitting bone.
Recommended publications
  • Vertebral Morphology, Dentition, Age, Growth, and Ecology of the Large Lamniform Shark Cardabiodon Ricki
    Vertebral morphology, dentition, age, growth, and ecology of the large lamniform shark Cardabiodon ricki MICHAEL G. NEWBREY, MIKAEL SIVERSSON, TODD D. COOK, ALLISON M. FOTHERINGHAM, and REBECCA L. SANCHEZ Newbrey, M.G., Siversson, M., Cook, T.D., Fotheringham, A.M., and Sanchez, R.L. 2015. Vertebral morphology, denti- tion, age, growth, and ecology of the large lamniform shark Cardabiodon ricki. Acta Palaeontologica Polonica 60 (4): 877–897. Cardabiodon ricki and Cardabiodon venator were large lamniform sharks with a patchy but global distribution in the Cenomanian and Turonian. Their teeth are generally rare and skeletal elements are less common. The centra of Cardabiodon ricki can be distinguished from those of other lamniforms by their unique combination of characteristics: medium length, round articulating outline with a very thick corpus calcareum, a corpus calcareum with a laterally flat rim, robust radial lamellae, thick radial lamellae that occur in low density, concentric lamellae absent, small circular or subovate pores concentrated next to each corpus calcareum, and papillose circular ridges on the surface of the corpus calcareum. The large diameter and robustness of the centra of two examined specimens suggest that Cardabiodon was large, had a rigid vertebral column, and was a fast swimmer. The sectioned corpora calcarea show both individuals deposited 13 bands (assumed to represent annual increments) after the birth ring. The identification of the birth ring is supported in the holotype of Cardabiodon ricki as the back-calculated tooth size at age 0 is nearly equal to the size of the smallest known isolated tooth of this species. The birth ring size (5–6.6 mm radial distance [RD]) overlaps with that of Archaeolamna kopingensis (5.4 mm RD) and the range of variation of Cretoxyrhina mantelli (6–11.6 mm RD) from the Smoky Hill Chalk, Niobrara Formation.
    [Show full text]
  • Annotated Checklist of Fossil Fishes from the Smoky Hill Chalk of the Niobrara Chalk (Upper Cretaceous) in Kansas
    Lucas, S. G. and Sullivan, R.M., eds., 2006, Late Cretaceous vertebrates from the Western Interior. New Mexico Museum of Natural History and Science Bulletin 35. 193 ANNOTATED CHECKLIST OF FOSSIL FISHES FROM THE SMOKY HILL CHALK OF THE NIOBRARA CHALK (UPPER CRETACEOUS) IN KANSAS KENSHU SHIMADA1 AND CHRISTOPHER FIELITZ2 1Environmental Science Program and Department of Biological Sciences, DePaul University,2325 North Clifton Avenue, Chicago, Illinois 60614; and Sternberg Museum of Natural History, Fort Hays State University, 3000 Sternberg Drive, Hays, Kansas 67601;2Department of Biology, Emory & Henry College, P.O. Box 947, Emory, Virginia 24327 Abstract—The Smoky Hill Chalk Member of the Niobrara Chalk is an Upper Cretaceous marine deposit found in Kansas and adjacent states in North America. The rock, which was formed under the Western Interior Sea, has a long history of yielding spectacular fossil marine vertebrates, including fishes. Here, we present an annotated taxo- nomic list of fossil fishes (= non-tetrapod vertebrates) described from the Smoky Hill Chalk based on published records. Our study shows that there are a total of 643 referable paleoichthyological specimens from the Smoky Hill Chalk documented in literature of which 133 belong to chondrichthyans and 510 to osteichthyans. These 643 specimens support the occurrence of a minimum of 70 species, comprising at least 16 chondrichthyans and 54 osteichthyans. Of these 70 species, 44 are represented by type specimens from the Smoky Hill Chalk. However, it must be noted that the fossil record of Niobrara fishes shows evidence of preservation, collecting, and research biases, and that the paleofauna is a time-averaged assemblage over five million years of chalk deposition.
    [Show full text]
  • Papers in Press
    Papers in Press “Papers in Press” includes peer-reviewed, accepted manuscripts of research articles, reviews, and short notes to be published in Paleontological Research. They have not yet been copy edited and/or formatted in the publication style of Paleontological Research. As soon as they are printed, they will be removed from this website. Please note they can be cited using the year of online publication and the DOI, as follows: Humblet, M. and Iryu, Y. 2014: Pleistocene coral assemblages on Irabu-jima, South Ryukyu Islands, Japan. Paleontological Research, doi: 10.2517/2014PR020. doi:10.2517/2018PR013 Features and paleoecological significance of the shark fauna from the Upper Cretaceous Hinoshima Formation, Himenoura Group, Southwest Japan Accepted Naoshi Kitamura 4-8-7 Motoyama, Chuo-ku Kumamoto, Kumamoto 860-0821, Japan (e-mail: [email protected]) Abstract. The shark fauna of the Upper Cretaceous Hinoshima Formation (Santonian: 86.3–83.6 Ma) of the manuscriptHimenoura Group (Kamiamakusa, Kumamoto Prefecture, Kyushu, Japan) was investigated based on fossil shark teeth found at five localities: Himedo Park, Kugushima, Wadanohana, Higashiura, and Kotorigoe. A detailed geological survey and taxonomic analysis was undertaken, and the habitat, depositional environment, and associated mollusks of each locality were considered in the context of previous studies. Twenty-one species, 15 genera, 11 families, and 6 orders of fossil sharks are recognized from the localities. This assemblage is more diverse than has previously been reported for Japan, and Lamniformes and Hexanchiformes were abundant. Three categories of shark fauna are recognized: a coastal region (Himedo Park; probably a breeding site), the coast to the open sea (Kugushima and Wadanohana), and bottom-dwelling or near-seafloor fauna (Kugushima, Wadanohana, Higashiura, and Kotorigoe).
    [Show full text]
  • Cambridge University Press 978-1-107-17944-8 — Evolution And
    Cambridge University Press 978-1-107-17944-8 — Evolution and Development of Fishes Edited by Zerina Johanson , Charlie Underwood , Martha Richter Index More Information Index abaxial muscle,33 Alizarin red, 110 arandaspids, 5, 61–62 abdominal muscles, 212 Alizarin red S whole mount staining, 127 Arandaspis, 5, 61, 69, 147 ability to repair fractures, 129 Allenypterus, 253 arcocentra, 192 Acanthodes, 14, 79, 83, 89–90, 104, 105–107, allometric growth, 129 Arctic char, 130 123, 152, 152, 156, 213, 221, 226 alveolar bone, 134 arcualia, 4, 49, 115, 146, 191, 206 Acanthodians, 3, 7, 13–15, 18, 23, 29, 63–65, Alx, 36, 47 areolar calcification, 114 68–69, 75, 79, 82, 84, 87–89, 91, 99, 102, Amdeh Formation, 61 areolar cartilage, 192 104–106, 114, 123, 148–149, 152–153, ameloblasts, 134 areolar mineralisation, 113 156, 160, 189, 192, 195, 198–199, 207, Amia, 154, 185, 190, 193, 258 Areyongalepis,7,64–65 213, 217–218, 220 ammocoete, 30, 40, 51, 56–57, 176, 206, 208, Argentina, 60–61, 67 Acanthodiformes, 14, 68 218 armoured agnathans, 150 Acanthodii, 152 amphiaspids, 5, 27 Arthrodira, 12, 24, 26, 28, 74, 82–84, 86, 194, Acanthomorpha, 20 amphibians, 1, 20, 150, 172, 180–182, 245, 248, 209, 222 Acanthostega, 22, 155–156, 255–258, 260 255–256 arthrodires, 7, 11–13, 22, 28, 71–72, 74–75, Acanthothoraci, 24, 74, 83 amphioxus, 49, 54–55, 124, 145, 155, 157, 159, 80–84, 152, 192, 207, 209, 212–213, 215, Acanthothoracida, 11 206, 224, 243–244, 249–250 219–220 acanthothoracids, 7, 12, 74, 81–82, 211, 215, Amphioxus, 120 Ascl,36 219 Amphystylic, 148 Asiaceratodus,21
    [Show full text]
  • Ichthyodectiform Fishes from the Late Cretaceous
    Mesozoic Fishes 5 – Global Diversity and Evolution, G. Arratia, H.-P. Schultze & M. V. H. Wilson (eds.): pp. 247-266, 12 figs., 2 tabs. © 2013 by Verlag Dr. Friedrich Pfeil, München, Germany – ISBN 978-3-89937-159-8 Ichthyodectiform fi shes from the Late Cretaceous (Campanian) of Arkansas, USA Kelly J. IRWIN and Christopher FIELITZ Abstract Several specimens of the ichthyodectiform fishes Xiphactinus audax and Saurocephalus cf. S. lanciformis are reported from the Upper Cretaceous (Campanian) Brownstown Marl and Ozan formations of southwestern Arkansas, U.S.A. Seven individuals of Xiphactinus, based on incomplete specimens, are represented by various elements: disarticu- lated skull bones, jaw fragments, pectoral fin-rays, or vertebrae. The circular vertebral centra are diagnostic for X. audax rather than X. vetus. The specimen of Saurocephalus consists of a three-dimensional skull, lacking much of the skull roof bones. It is identified as Saurocephalus based on the shape of the predentary bone. This specimen provides the first record of entopterygoid teeth in Saurocephalus. These specimens represent new geographic and geologic distribution records of these taxa from the western Gulf Coastal Plain, which biogeographically links records from the eastern Gulf Coastal Plain with those from the Western Interior Sea. Introduction LEIDY (1854) was the first to report the presence of Cretaceous marine vertebrate fossils from Arkansas, yet in the intervening 150+ years, the body of work on the paleoichthyofauna of Arkansas remains limited (WILSON & BRUNER 2004). BARDACK (1965), GOODY (1976), CASE (1978), and RUSSELL (1988) re- ported geographic and geologic distributions for specific taxa, but few descriptive works on fossil fishes are available (see HUSSAKOF 1947; MEYER 1974; BECKER et al.
    [Show full text]
  • Bibliography of Alexander O
    Зоологический институт РАН Лаборатория териологии Bibliography of Alexander O. Averianov A. Monograph 1992. Averianov A.O., Baryschnikov G.F., Garutt W.E., Garutt N.W. & Fomicheva N.L. The Volgian Fauna of Pleistocene Mammals in the Geological-Mineralogical Museum of the Kazan University. Kazan State University, Kazan. 164pp. [In Russian] B. Edited books 2003. [Systematics, Phylogeny and Paleontology of Small Mammals. Proceedings of the International Conference devoted to the 90-th anniversary of Prof. I.M. Gromov] (Edited by A.O. Averianov and N.I. Abramson). Zoologicheskii Institut RAN, St. Petersburg. 246 pp. [papers in Russian and English] 1999. [Materials on the History of Fauna of Eurasia] (Edited by I.S. Darevskii and A.O. Averianov). Trudy Zoologicheskogo Instituta RAN, T. 277. 144 pp. [papers in Russian and English] 1997. Nessov, L.A. [Cretaceous Nonmarine Vertebrates of Northern Eurasia] (Posthumous edition by L.B. Golovneva and A.O. Averianov). 218 pp. Izdatelstvo Sankt-Peterburgskogo Universiteta, Saint Petersburg. [in Russian] C. Refereed Journals 2017. Averianov A.O. and Sues H.-D. 2017. The oldest record of Alvarezsauridae (Dinosauria: Theropoda) in the Northern Hemisphere. PLoS One 12(10): e0186254. 2017. Аверьянов А.О. 2017. Титанозавры: новые данные о возрастной и индивидуальной изменчивости зубов. Природа. № 2. С.83-84. 2017. Averianov A.O. and Archibald J.D. 2017. Therian postcranial bones from the Upper Cretaceous Bissekty Formation of Uzbekistan. Proceedings of the Zoological Institute of the Russian Academy of Sciences 321(4): 433–484. 2017. Lopatin A.V. and Averianov A.O. 2017. The stem placental mammal Prokennalestes from the Early Cretaceous of Mongolia.
    [Show full text]
  • Supporting Information
    Supporting Information Clarke et al. 10.1073/pnas.1607237113 Major Axes of Shape Variation and Their Anatomical a comparison: the crown teleost vs. stem teleost comparison on Correlates molecular timescales. Here, the larger SL dataset delivers a All relative warp axes that individually account for >5% of the majority of trees where crown teleosts possess significantly variation across the species sampled are displayed in Table S3. higher rates than stem teleosts, whereas the CS and pruned SL Morphospaces derived from the first three axes, containing all datasets find this result in a large minority (Fig. S3). 398 Mesozoic neopterygian species in our shape dataset, are Overall, these results suggest that choice of size metric is presented in Fig. S1. Images of sampled fossil specimens are also relatively unimportant for our dataset, and that the overall size included in Fig. S1 to illustrate the anatomical correlates of and taxonomic samplings of the dataset are more likely to in- shape axes. The positions of major neopterygian clades in mor- fluence subsequent results, despite those factors having a rela- phospace are indicated by different colors in Fig. S2. Major tively small influence here. Nevertheless, choice of metric may be teleost clades are presented in Fig. S2A and major holostean important for other datasets (e.g., different groups of organisms clades in Fig. S2B. or datasets of other biological/nonbiological structures), because RW1 captures 42.53% of the variance, reflecting changes from it is possible to envisage scenarios where the choice of size metric slender-bodied taxa to deep-bodied taxa (Fig. S1 and Table S3).
    [Show full text]
  • Dr. Kenshu Shimada: Peer-Reviewed Published Articles [Last Updated 04/26/2019]
    Dr. Kenshu Shimada: Peer-Reviewed Published Articles [last updated 04/26/2019] [* = undergraduate student; ** = graduate student] 2019 [105] Shimada, K. In press. A new species and biology of the Late Cretaceous 'blunt-snouted' bony fish, Thryptodus (Actinopterygii: Tselfatiiformes), from the United States. Cretaceous Research. [104] Cronin, T. J.*, and K. Shimada . In press. New anatomical information on the Late Cretaceous bony fish, Micropycnodon kansasensis (Actinopterygii: Pycnodontiformes), from the Niobrara Chalk of western Kansas, U.S.A. Transactions of Kansas Academy of Science. [103] Pimiento, C., J. L. Cantalapiedra, K. Shimada , D. J. Field, and J. B. Smaers. 2019. Evolutionary pathways towards shark gigantism. Evolution, 73(3):588–599. 2018 [102] Johnson-Ransom, E. D.*, E. V. Popov, T. A. Deméré, and K. Shimada. 2018. The Late Cretaceous chimaeroid fish, Ischyodus bifurcatus Case (Chondrichthyes: Holocephali), from California, USA, and its paleobiogeographical significance. Paleontological Research, 2(4):364-372. [101] Guinot, G., S. Adnet, K. Shimada , C. J. Underwood, M. Siversson, D. J. Ward, J. Kriwet, and H. Cappetta. 2018. On the need of providing tooth morphology in descriptions of extant elasmobranch species. Zootaxa, 4461(1):118–126. [100] Jacobs, P. K.*, and K. Shimada . 2018. Ontogenetic growth pattern of the extant smalltooth sandtiger shark, Odontaspis ferox (Lamniformes: Odontaspididae)— application from and to paleontology. Journal of Fossil Research, 51(1):23–29. [99] Guzzo, F.*, and K. Shimada . 2018. A new fossil vertebrate locality of the Jetmore Chalk Member of the Upper Cretaceous Greenhorn Limestone in north-central Kansas, U.S.A. Transactions of the Kansas Academy of Science, 121(1-2):59-68.
    [Show full text]
  • Università Di Pisa
    UNIVERSITÀ DI PISA Dottorato di ricerca in scienze della terra XIX Ciclo DISSERTAZIONE FINALE ANALISI SISTEMATICA, PALEOECOLOGICA E PALEOBIOGEOGRAFICA DELLA SELACIOFAUNA PLIO-PLEISTOCENICA DEL MEDITERRANEO Candidato Presidente della Scuola di Dottorato Stefano MARSILI Prof. Paolo Roberto FEDERICI Tutore Prof. Walter LANDINI Dipartimento di scienze della terra 2006 Indice INDICE ABSTRACT CAPITOLO 1 – INTRODUZIONE 1.1. Premessa. 1 1.2. Il Mediterraneo e l’attuale diversità del popolamento a squali. 4 CAPITOLO 2 – MATERIALI E METODI 7 CAPITOLO 3 – INQUADRAMENTO GEOLGICO E STRATIGRAFICO 3.1. Premessa. 15 3.2. Inquadramento geologico e stratigrafico delle sezioni campionate. 16 3.2.1. Le sezioni plioceniche della Romagna. 16 3.2.1.1. Sezione Rio Merli. 17 3.2.1.2. Sezione Rio dei Ronchi. 17 3.2.1.3. Sezione Rio Co di Sasso. 18 3.2.1.4. Sezione Rio Cugno. 19 3.2.2. Le sezioni pleistoceniche dell’Italia Meridionale. 19 3.2.2.1. La sezione di Fiumefreddo. 20 3.2.2.2. La sezione di Grammichele. 22 3.2.2.3. La sezione di Vallone Catrica. 23 3.2.2.4. La sezione di Archi. 23 3.3. Inquadramento geologico e stratigrafico dei bacini centrali del Tora-Fine, di Volterra e di Siena: premessa. 24 3.3.1. Bacino del Tora-Fine. 26 3.3.2. Bacino di Siena-Radicofani. 27 3.3.3. Bacino di Volterra. 29 3.4. Inquadramento geologico e stratigrafico delle principali località storiche. 30 3.4.1. Emilia Romagna. 30 3.4.2. Piemonte. 32 3.4.3. Liguria. 32 3.4.4. Basilicata.
    [Show full text]
  • A Study of Potential Co-Product Trace Elements Within the Clear Hills Iron Deposits, Northwestern Alberta
    Special Report 08 A Study of Potential Co-Product Trace Elements Within the Clear Hills Iron Deposits, Northwestern Alberta NTS 83M,N, 84C,D A STUDY OF POTENTIAL CO-PRODUCT TRACE ELEMENTS WITHIN THE CLEAR HILLS IRON DEPOSITS, NORTHWESTERN ALBERTA Prepared for Research and Technology Branch, Alberta Energy Prepared by APEX Geoscience Ltd. (Project 97213) In cooperation with The Alberta Geological Survey, Energy and Utility Board And Marum Resources Ltd. February, 1999 R.A. Olson D. R. Eccles C.J. Collom A STUDY OF POTENTIAL CO-PRODUCT TRACE ELEMENTS WITHIN THE CLEAR HILLS IRON DEPOSITS, NORTHWESTERN ALBERTA TABLE OF CONTENTS SECTION PAGE ACKNOWLEDGMENTS AND DISCLAIMER ....................................................... vi 1.0 SUMMARY ........................................................................................................1 2.0 INTRODUCTION ..................................................................................................3 2.1 Preamble....................................................................................................3 2.2 Location, Access, Physiography, Bedrock Exposure .................................4 2.3 Synopsis of Prior Scientific Studies of the Clear Hills Iron Deposits, and the Stratigraphically Correlative Bad Heart Formation ...............................4 2.4 Synopsis of Prior Exploration of the Clear Hills Iron Deposits....................6 3.0 GEOLOGY ........................................................................................................7 3.1 Introduction
    [Show full text]
  • Body Length Estimation of Neogene Macrophagous Lamniform Sharks (Carcharodon and Otodus) Derived from Associated Fossil Dentitions
    Palaeontologia Electronica palaeo-electronica.org Body length estimation of Neogene macrophagous lamniform sharks (Carcharodon and Otodus) derived from associated fossil dentitions Victor J. Perez, Ronny M. Leder, and Teddy Badaut ABSTRACT The megatooth shark, Otodus megalodon, is widely accepted as the largest mac- rophagous shark that ever lived; and yet, despite over a century of research, its size is still debated. The great white shark, Carcharodon carcharias, is regarded as the best living ecological analog to the extinct megatooth shark and has been the basis for all body length estimates to date. The most widely accepted and applied method for esti- mating body size of O. megalodon was based upon a linear relationship between tooth crown height and total body length in C. carcharias. However, when applying this method to an associated dentition of O. megalodon (UF-VP-311000), the estimates for this single individual ranged from 11.4 to 41.1 m. These widely variable estimates showed a distinct pattern, in which anterior teeth resulted in lower estimates than pos- terior teeth. Consequently, previous paleoecological analyses based on body size esti- mates of O. megalodon may be subject to misinterpretation. Herein, we describe a novel method based on the summed crown width of associated fossil dentitions, which mitigates the variability associated with different tooth positions. The method assumes direct proportionality between the ratio of summed crown width to body length in eco- logically and taxonomically related fossil and modern species. Total body lengths were estimated from 11 individuals, representing five lamniform species: Otodus megal- odon, Otodus chubutensis, Carcharodon carcharias, Carcharodon hubbelli, and Carcharodon hastalis.
    [Show full text]
  • Copyrighted Material
    06_250317 part1-3.qxd 12/13/05 7:32 PM Page 15 Phylum Chordata Chordates are placed in the superphylum Deuterostomia. The possible rela- tionships of the chordates and deuterostomes to other metazoans are dis- cussed in Halanych (2004). He restricts the taxon of deuterostomes to the chordates and their proposed immediate sister group, a taxon comprising the hemichordates, echinoderms, and the wormlike Xenoturbella. The phylum Chordata has been used by most recent workers to encompass members of the subphyla Urochordata (tunicates or sea-squirts), Cephalochordata (lancelets), and Craniata (fishes, amphibians, reptiles, birds, and mammals). The Cephalochordata and Craniata form a mono- phyletic group (e.g., Cameron et al., 2000; Halanych, 2004). Much disagree- ment exists concerning the interrelationships and classification of the Chordata, and the inclusion of the urochordates as sister to the cephalochor- dates and craniates is not as broadly held as the sister-group relationship of cephalochordates and craniates (Halanych, 2004). Many excitingCOPYRIGHTED fossil finds in recent years MATERIAL reveal what the first fishes may have looked like, and these finds push the fossil record of fishes back into the early Cambrian, far further back than previously known. There is still much difference of opinion on the phylogenetic position of these new Cambrian species, and many new discoveries and changes in early fish systematics may be expected over the next decade. As noted by Halanych (2004), D.-G. (D.) Shu and collaborators have discovered fossil ascidians (e.g., Cheungkongella), cephalochordate-like yunnanozoans (Haikouella and Yunnanozoon), and jaw- less craniates (Myllokunmingia, and its junior synonym Haikouichthys) over the 15 06_250317 part1-3.qxd 12/13/05 7:32 PM Page 16 16 Fishes of the World last few years that push the origins of these three major taxa at least into the Lower Cambrian (approximately 530–540 million years ago).
    [Show full text]