ON THE CLASSIFICATION OF THE ()

BY

FENNER A. CHACE, JR. Department of Invertebrate Zoology (NHB 163), National Museum of Natural History, Smithso- nian Institution, Washington, D.C. 20560, U.S.A.

ZUSAMMENFASSUNG

Es wird eine vorläufige Einteilung der Caridea vorgeschlagen, hauptsächlich nach Merkmalen der drei vorderen Paare von Pereopoden und der sechs Paare von Mundteilen, wobei nicht von der Methode kladistischer Merkmalsphylogenie profitiert wird.

This far-from-definitive review of the classification of the caridean - based largely on the three anterior pairs of pereopods and the six pairs of mouthparts-was originally intended to provide a personal aid to an under- standing of caridean relationships. It has served principally to emphasize that such attempts to determine the phylogeny of an incompletely known group deplorably squander time that might better be devoted to learning the charac- teristics of the individual . The dearth of basic knowledge of those specific features, coupled with that of data that are virtually irretrievable in a group with a limited fossil record, defies analysis by the most sophisticated mental gymnastics, no matter how factual computer-generated numbers may seem. The decision to publish this survey in its current somewhat preliminary form, without illustrations, stems from my concern that increasing awareness of the unpublished manuscript might dissuade colleagues from pursuing similar, more meaningful research. Following are the caridean family groups recognized herein. The arrange- ment is not necessarily indicative of relationships.

Superfamily Procaridoidea Family Procarididae Superfamily Pasiphaeoidea Family Pasiphacidae Superfamily Oplophoroidea Family Superfamily Atyoidea Family Superfamily Family 71

Superfamily Family Family Family Family Xiphocarididae Superfamily Psalidopodoidea Family Psalidopodidae Superfamily Stylodactyloidea Family Superfamily Campylonotoidea Family Bathypalaemonellidae Family Campylonotidae Superfamily Family Anchistioididae Family Desmocarididae Family Family Hymenoceridae Family Subfamily Palaemoninae Subfamily Pontoniinae Family Typhlocarididae Subfamily Euryrhynchinae Subfamily Typhlocaridinae Superfamily Physetocaridoidea Family Physetocarididae Superfamily Processoidea Family Superfamily Family Family Family Superfamily Family Family Superfamily Family Family

Following are some of the tentative conclusions generated by this study: The , because of their unusual mandibles and first and second maxillipeds, disguise their affinities about as well as any of the families. The Oplophoridae, because of their unique thoracic epipods that may or may not indicate a relationship with the Procarididae, which have even more