N oona Dan Papers No. 31.

Orthoptera- from the Bismarck Solomon Islands, interesting records from New Guinea Moluccas. By D. Keith McE. Kevan Dept. of Entomology and Lyman Entomological Museum, McGil! University, Macdonald College, Province of Quebec, Canada.

The following is an account of the short-horned and their relatives, collected members of the Danish Dan" Expedition (see Petersen, 1966). The number of spe­ cies obtained is not great, but some very valuable series were brought back, together with lesser material of little-known, or previously unknown forms. Opportunity is also taken here to record certain miscellaneous specimens of interesting species from the area vis.ited by the expedition, but which were not in­ cluded in its collections. Originally it was intended only to include specimens in the writer's own instituton among such additional records, but, certain other relevant material from other sources being currently at hand, it was considered that it might be of interest to refer to this also. No attempt has been made, however, to extend the scope of the information presented by thoroughly investigating these other sources for additional material. Specimens not collected by the expedition are indicated as follows: * = B. P. Bishop Museum, Honolulu; t = Lyman Entomological Museum, McGill University, Macdonald College; ::f = Academy of Natural Sciences of Philadelphia (mostly spe­ cimens bearing "M.C.Z." - Museum of Comparative Zoology, Harvard University- labels).

Ent. Medd. 34 24 376 Entomoloqiske Meddelelser 34 (1966)

Superfam. T R ID A C T Y L 0 ID E A. Fam. TRIDACTYLIDAE.

Subfam. TlUDACTYLI~AE. Tridactylus pulex Saussure. Philippines. - PALA WAN: Brooke's Point, Uring Uring, 20 -25. VIII. 61, 3 cf' d'; J\fantalingajan, Pinigisan, 600 m., 1-6. IX. 3 d'd', 3¥¥, 3 juv. - TA WI TA \VI: Tarawakan, 3. XI. 61, 1¥.- MINDANO: Curuan district, Sapamoro, 16. XII. 61, 1¥.

Superfam. T ET RIG 0 IDEA. Fam. TETRIGIDAE.

Subfam. CLEOSTHATI~AE. Cleostratus monocerns Still. Philippines. - TAW1 TAWI: Tara,vakan, 10.XL6L 10~ (fastigium of vertex a little shorter than typical). - LUZON: Los Bafios, Mt. Maquiling, 30. IV. 60, 1¥ (C. A. Corpuz) (fastigium very long, suggesting C. helleri Giinther, hut not otherwise agreeing well with that species) t. - LEYTE: Lagolago, Bayhay (no date), 1 ¥ t.

Suhfam. CLADONOTINAE. This subfamily is probably heterogeneous, but the genera here included are grouped in the section Cladonotae by Giinther (lH38a). Hymenoics triangularis W estwood. Philippines.- LUZO~: Los Bafios, 14. X. 61, 1 d' (Q. Gonza­ .t Gen. MysiUms Still. The following are all provisionally determined only, as the pronotal and other embellishments which have been used to distinguish species seem to be very variable in form and may not provide taxonomically reliable specific characters.

MysUhus ensatrix (Walker). Philippines. - LUZON: Los Bafios, 20. IV. 61, 1 ¥ (L. C. Cosico). 'f D. Keith Me E. Kevan: Some -Caelifera . .. 377

Mysithus banahao Hebard. Philippines.- ~UNDANAO: Bunaguite, 1940, 1 cJ' (J. K. Chap­ man). t :M:ysiUms mindanao Hebard. Philippines. - MINDANAO: "Lake", 1940 1 s;? (J. K. Chap­ man). t )lysithns securifer CW alker). Philippines.- LUZON: Mt. Maquiling, 17. XL 49, 1 cJ' (Uichan­ co). t Mysithus cristicornis CW alker). Philippines. LUZON: Tagkawayan, Quezon, 5. I. 51, ls;? (S. M. Ondana). t MysHhns jubatns Hebard. Philippines. LUZON: ~H. Maquiling, 600 m., 21. II. 48, ls;? (J. Gutierrez). t Diotarus pupns I. Bolivar. The genus Diotarus is in need of revision, but the following appear to belong to D. pupus. Philippines. - P ALA WAN: :\fantalingajan, Pinigisan, 600 m., 6-13. IX. 61, 2 s;?s;?, 1 juv. (?this species); 16-20. IX. 61, 3 cJ'cJ', 1 juv. (? this species). Diotarus ikonnikovi Bei-Bienko. Philippines. - LUZO~: Mt. ::\faquiling, 1500 ft., 26. IV. 56, 1 s;?, (P.A. C. Galvez). t Gimther (1938a) suggests this species may be merely a form of the last, but it is distinctly more robust.

Subfam. TETRIGIKAE. Tribe Scelimenini. The tribe is here restricted to the Scelimenae Verae of Giinther (1938a), which all seem to be related (see Giinther, 1955).

Scelimena novaeguincae (I. Bolivar). Bismarcks.- NEW BRITAIN: Gazelle Peninsula, Yalom, 1000 m., 12-20. V. 62, 5 cJ'cJ' 6 s;?s;?; Cape Hoskins, Vaisisi, 9. VII. 62, 3 cJ'cJ', 3 S?S?.

24• 378 Entomoloqiske Meddelelser 34 (1966)

Platygavalidium kraussi (I. Bolfvar). Philippines. MINDANAO: Curuan district, Sapamoro, 18. XII. 61, 1 <;?. Telrinda palpata (SHH). Philippines.- LUZON: Los Bafios, 2. IV. 61, 1 <;? (R. D. Man- luco); 1 d' .J. Kalau). t Tribe Thoradonotini, nov. The divisions of the Tetriginae are not at all clearly defined- in fact they are largely Giinther (1955), however, has of many genera for him with Tlwradonota, here in this tribe.

PAL\\VAN: Brookc's 20. 600 m., 6-23. IX. 61, 2d' Gen. Euci·ioteUix Hebard. The of this genus is in rather a chaotic slate so that accurate determinations are EncrioteUix sp. ar£. molestus Giinther. Bismarcks.- ~E\V BlUTAIN: Gazelle 1000 m., 21-22. ·v. 62, 1 d', 1 <;?. EncrioteUix sp. ai'f. Giinther. Bismarcks.- MANUS: Lorengau, 18-19. VI. 62, 1 d', 2 S?S?.­ NEW BRITAIN: Yalom, 1000 m., 19-22. V. 62, 3 d'd', 2 <;?<;?. Eucriotettix sp (p). The following are small individuals, all rather similar, but probably representing two or more new species. Bismarcks.- MANUS: Lorengau, 18. VI. 62, 1 d'.- DYAUL: Sumuna, 4. IlL 62, 1 d'. - DUKE OF YORK: Manuan, 21. VII. 62, 1 d'. Probolotettix, n.sp. Philippines.- LUZON: Mt. Maquiling, 2979 ft., 17. V. 58, 1 <;? (S. Fontonika). t A large, subulate form (pronotum 15 mm.) characterized by rather indistinct pronotal carinae, the anterior part of the pro­ notum evenly convex, its anterior margin convex. D. Keith Me E. Kevan: Some Orthoptera-Caelifera. . . 3 79

XisbeUa dohrni palawanica Giinther. Philippines. -- PALA WAN: Mantalingajan, Pinigisan, 600 m., 7. IX. 61, 1 0'. BolivarHeUix palawanicus Giinther. Philippines. - PALAWAN: Manlalingajan, Pinigisan, 600 m., 5-16. IX. 61, 3 ~~.

RosteUa phyUoccra guentheri~ n.sp. Roslel/a phyllocera ssp., Gi.inthcr, 1939, Abh. Ber. staatl. l\lus. Tierk. Vi:ilkerk. Dresden, 20 (A) [X F. 1 J: 280, fig. 209. Giinther (1939) gave a brief description and figured \Yhat he regarded as a new subspecies of R. phyllocera (Haan), ably because he had but two females before him XI, XII. Boettscher Deutsch. Ent. Inst. , he declined to name it. The "Noona Dan" material contains four of what js unquestionably the same for which I propose the name guentlzeri, after its discoverer. It is able that it a distinct species, but unlil the genus is there would seem to be little purpose in recognizing it as such for the present As I have not had lhe opportunity to see Giinther's specimens (and as they are both females), I have chosen one of the "Noona Dan" males, now in the Zoological ~Iuseum, Copenhagen to be the h o 1 o type of the new subspecies. It bears the following data: Philippines, PALAWAN: Mantalingajan, Pinigisan, 600 meter, 3 Sept. 1961, Noona Dan Exp. 61-62. It has the following measure­ ments: length of excluding 12.0, of head 2.7, w-idth across eyes 2.2, interocular width 1 pronotum hind femur 6.9 mm. A female from the same series may he regarded as the a ll o­ t y p e. It bears the same that the date is 8 Sept. and there is an additional label indicating that it was "caught in Malaise outside forest." It has the following measurements: length of body excluding pronotum, 15.0, of head width across eyes 2.5, interocular width 1 pronotum 17 .4, hind femur 7.5 mm. There are also two male p a r at y p e s with the same data as the allotype except that they arc dated 7 and 12 Sept. and were trapped "inside forest". With the exception of the first paratype which is now in the Lyman Entomological Museum, Macdonald College, McGill University, the specimens are in Co­ penhagen. 380 Entomologlske Meddelelser 34 (1966)

The placing of Rostella next to Bolivmitettix is in accord with the opinion of Giinthcr (1955).

Tribe Discotettigini. As understood here, this tribe includes all those genera from Paraguelus to M acromotettix grouped together by Giinther (1955), as well as certain other associated genera. It thus includes not only the Sections Discotettigiae and Tripetalocerae of Giinther (19:38a) and earlier authors, but also the first several members and a few other genera of the Amorphopi (=Metrorlorac) of Giinther (1939).

Paraguelus palawanicus Giinther. Philippines. - PALAvVAN: .Mantalingajan, Tagembung, 1150 m., 19. IX. 61, 1 d'.

~ Paraguelus, n.sp. Philippines. - LUZON: Mt. Maquiling, 100 m. [?] 2:5. IV. 58, c;? (0. Al. Galvez). ·f· This species has a longer and more pointed pronotum than the last. It may, in fact, represent a new genus close to Threciscus Still, from which it differs chiefly in lacking spines on the lateral pronotal lobes. Spartolus longiceps Still. Philippines. - LEYTE: Mt. Panasugan, 400 m., 7. V. 52, 1 c;? (C. L. Baltazar). t

DiscoteUix (Jinesarchus) scabridus (Still). J!nesorclms differs from Discotettix, s.str., in that the subter­ minal segments of the antennae are little expanded; the two have previously been synonymized, but I prefer to recognize subgeneric status for the former. Philippines. - MINDAKAO: Curuan district, Sapamoro, 16- 18. XII. 61, 4 dd, 4 c;?c;?, 1 juv.

? Amphinotus, n.sp. Bismarcks. - NEvV BRITAIN: Gazelle Peninsula, Yalom, 1000 m., 21. V. 62, 1 c;?. Reminiscent of A. grandis Giinther, from New Guinea, but even larger and with a much more "humped" dorsal profile to the anterior part of the pronotum and more prominent lateral prono­ tal lobes. D. Keith Me E. Ke,an: Some Orthoptera-Caelifera . .. :381

Hydrotettix, n. sp.

Bismarcks. ~ :\'EvV BIUTAl:\': Gazelle Peninsula, Yalom, 1000 m., old secondary forest, 20. V. 62, 1 d. '? Lamellitcttigodcs, n. sp. Philippines. MINDAN'AO: Curuan district, Saparnoro, 16. XIL tH. 1 cJ. Tribe Criotettigini, nov. This Lrihe is arbitrarily erected to contain heterogeneous genera of the section Scelimenae Spuriae of Giinther (19:18b) that have not yel been transferred to other groups.

Gen. Loxilobus Hancock. This large genus is almost certainly hetnogeneous and i ls in rather a chaotic state. Few accurate determinalions are possible. Loxilobus? insidiosus (I. Bolfvar). Philippines. ~ LUZO:\': 1\lt. Maquiling, 28. XII. 49, 1 c;? (Uichanco). t Loxilobus sp(p.) aff. rugosus (I. Bolivar). Philippines. ~ BALABAC: Dalawan Bay, 7~12. X. Gl, 1 cJ, 1 c;?. ~ TAWI TAWf: Tarawakan, 23. X. 61, 1 c;?. Loxilobus ? puHus (I. Bolivar).

Philippines. ~ MINDANAO: Curuan district, Sapamoro, 18~ 22. XII. 61, 2 cJcJ, 1 c;?. Loxilobus novaebritanniae Giinther. Giinther (1938b) refers to the variability of this species, which apparently occurs in at least three distinguishable forms: the typical form; a somewhat similar form with more strongly pro­ duced and more acute lateral pro notal lobes; and a micropterous form. The typical form (a, below), as deduced from the original description and figure, is more elongate ( suhulate) and has less prominently angular lateral pronolal lobes, and hind wings ex­ tending appreciably beyond the end of lhe pronotum, whilst the second form (b) illustrated by Giinther (1 g;)8b), in addition to the differences in lhe pronotal lobes mentioned, is a little shorler and stouter, with hind wings that barely (if at all) surpass the end of the pronotum. Giinlher (l. c.) does nol explain what is meant by "micropterous," but it is conceivable that the condition is similar 382 Entomologiske Meddelelser 34 (1966) to that of the third form (c) mentioned below, which is even stouter than form b, has even more prominent lateral pronotal lobes and an even shorter pronotum beyond \vhich the hind wings do not project. ll is possible, but not probable, that form c represents a distinct species. More extreme examples of form a tend to resemble L. angulatus Hancock. Bismarcks.- NE\V JRELA:\'D: Lelet. Plateau, Lemkarnin. 900 m., 6-20.JV. 2 cJcJ c), 7 ¥¥ (2 form a, 5 form cl.- I\E\V BRITAIN: Gazelle Peninsula, Yalom, 1000 m., 9-2:1. V. 4 cJcJ (2 form a, 2 form b), 16 ¥¥ form u, 6 form Hoskins, Vaisisi, 9. VII. 1 ¥ (form a).

Loxilobus 9 n.sp. aff. n6vae.luitanniac Giinther. This is a slender species much like typical L. novaebritunniae. but is considerably the pronotal length is only 7.5-8.5 mm. in the male and 8.5-9.5 mm. in the female. Bismarcks. - LUF (Hermit Islands): 26. VI. 62, 1 cJ', 1 MANUS: Lorengau, 18-19. VI. 62, 7¥¥ (also 1 ? this spe- cies, 24. VI. 62).- LAVONGAI: Banatam, 18-25. III. 62, 5 cJ 2 ¥¥. - DUKE OF YORK: Manuan, 20. VII. 62, 1 cJ (also 1 ? this species, 21. VI. . ~NEW BRITAIN: Cape Hoskins, Valoka, 7. VII. 62, 1 cJ (also 1 ? Lhis species). It is possible that !he specimens from the last two localities are not conspeccific with the others. Tribe Metrodorini. This and heterogeneous tribe those genera Section Amurplwpi (= Jvlctrodorae of thal have not been removed to other groups. Systolodc:rus haani I. Bolivar. Philippines.~ LUZON: Los Bafios, 18. IV. 61, 1 cJ G. Flo- ; 6. V. 61, 1 ¥ . D. Canatal). t Systolederus ? affinis Gunther. Bismarcks.- MANUS: Lorengau, 19-22. VI. 62, 6 cJcJ, 4 2 juv. - NE\V BRITAI~: Gazelle Peninsula, Yalom, 1000 m., 12. V. G2, 1 cJ', 1 ¥. Gen. Hyboella: Hancock. The limits of this large and probably heterogeneous genus arc ill-defined; the systematics of the genus are chaotic and accurate determinations are virtually impossible. D. I

HyboeUa sp. aff. similis Gfm!her.

Philippines. - PALA 'WA~: Brooke's Point, Makagwa, 18. IX. 6l,1d'. There are also three nymphs that may perhaps belong to the same from PALA"\VAT\· Pinigisan, 600 m., 6-21. IX. fil. ? sp. indei. Bismarcks. :\fEW IHELAND: Danu, Kalili Bay, 30. IV.

Tribe Tctrigini. This Yery large tribe is at present in a state of great confusion. Some of the genera at to it may have to be excluded. The limits of several of the genera, particularly of Para­ tettix and Euparotcttix arc virtually impossible to define. Only provisional determina!ions, some of which may prove to be erroneous when the group is revised, are attempted here.

CoptoteUix ? macsoi I. Bolivar. Philippines.- LUZON: Mt. Sto. Tomas, 7500 ft., 28. V. 58, 1 d' Cariaso) . 'f Copiotettix fuscus L Bolivar. Philippines.-- LUZO~: Los Baiios, 18. IV. G1, 1 C( G. Flo- .t tolc"'t'tiw ? n.sp (p.) aff. insularis Giinther. The may include two or three Yery similar spceies. Bismarcks.- MUSSAU: Talumalaus, '2-7. IL 62, 3 d'd', 1 C(, .; Boliu, 4-7. VL 62, :1 d'd', 4 C(C(, 2 juv.- LAVONGAI: 18-25. HI. 4 d'd', 2 C(C(, 4 juv.- DYAUL: Sumu- na, 4. III. 1 d'.- NEW IHELAND: Lelet Plateau, 900 m., 6.1V. 1 d'; 19.IV. 62. 1 C(.- DUKE OF YOHK: Ma- nnan 19-21. VII. 62, 2 C(C(. Specimens from J\-fussau and are smaller than the which are approximately the same size as C. insularis. Tetl'ix sp. Philippines. - PALA \VAN: Brooke's Point, Uring Uring, l L IX.61, 1 d'. A moderate-sized, rather subulate speeics with the anterior margin of the pronotum angulate. 384 Entomoloqiske Meddelelser 34 (1966)

Tetl'ix sp. Philippines. - MI!'\DANAO: Curuan district, Sapamoro, lfi- 20. XII. 61, 2 dd. A small, brachypterous form reminiscent of the bipunctatus group. ParateUix ? amplus (Sji)sledt).

Philippines. - P~~LA vVAN: Brooke's Point, Uring Uring, 14- 25. YIIJ. 61, 3 dd, 6 yy; Mantalingajan, 28. YIII. 61, 1 y [rather large and pale]; 9. IX. 61, 1 y [almost as large. and very dark].­ TA \VI TA \VI; Tarawakan, 26. X. Gl, 2 yy [1 rather smallj; 14. XI. fil, 1 d. - :MINDANAO: Catabato, Buluan, 4. IV. 66, 1 y N. Cedafia) t ParateUix sp. af'f. amplus (Sjostedt). Philippines. - TA \VI TA vVI: Tarawakan, 4. XI. 61, 1 d. ParatcUix sp. aff. histricus (Stal). Philippines.- PALAWAN: Brooke's Point, Uring Uring, 5-20. lX. 61, 2 dd, 1 y. Paratettix ? histricus (SHU). Bismarcks.- MUSSAU: Talumalaus, 18. I.-7. II. 62, 27 dd. 14 yy, 1 juv.; Taletassi Lake, 4-6. VI. 62, 6 dd, 9 yy; Boliu, 5. VI. 62, 2 d d, 1 y, 1 juv. - NEW IRELAND: Lelet Plateau, Lemkamin, 900 m., 16-17. IV. 62,3 dd, 6 yy. ParateUix ? vexator Giinther. Bismarcks. - NEW BRITAIN: Cape Hoskins, Valoka, 10. VII. G2, 1 y. ParateUix pullus I. Dol:ivar. Bismarcks. - MANUS: Lorengau, 17. VI. 62, 1 y. - MUSSA U: Talumalaus, l.II. 62, 1 y; Malakata, 15. II. 62, 2 dd, 2 yy, 1 juv.; Boliu, 7. VI. 62, 2 yy.- LAVONGAI: Banatam, 21. III. ti2, 1 y. Solomons. - GUADALCANAL: Honiara, 27. VI.-4. VIII. f\2, 1 y. EuparateUix ? personatus (I. Bolivar). Philippines.- BALABAC: Dalawan Day, 6-11. X. f\1, 2 dd, 3 yy, 2 juv. (?this species).- PALAWAN: Brooke's Point, Uring Uring, 1fi-26. VIII. f\1, 4 dd, :i yy, 1 juv. (?this species); Man­ lalingajan, Tagemhung, 1160 m., 16-20. IX. f\1, 6 dd, 15 yy, 1 juv. (? this species).- TAWI TAWI: Tarawakan, 12. XI. 61, 1 d. D. Keith Me E. Keuan: Some Orthoptera-Cae!ifera . .. 386

The last specimen may not be conspecific with the others. Euparateitix sp. aff. tricariuatus (I. Bolivar). Philippines. - BALABAC: Dalawan Bay, 4-5. X. ol, 7 d'd', 5 ¥¥.- PALAWAN: Brooke's Point, Uring Uring, 15-20. VIII. 61, 7 d'd', () ¥¥; Brooke's Point, Makagwa, 22. VIII. 61, 1 d'.­ TA\Vl TAWI: Tarawakan, 25.X.-14.Xl.61, 8 d'd', 4¥¥.­ ML~'\D.~~AO: Curuan district, Sapamoro, lfi-22. XII. 61, 5 d'd', 5 ¥¥. Bismarcks.- :\IA~US: Lorcngau, 24. VI. 62, 1 d'.- MUSSAU: Talumalaus, 7.1I. 62, 2 d'd', ;) ¥¥, 1 jnv. (? this species). - LA \'O~GAI: Banalam, 18. lll. G2, 1 ¥: l\TW IHELA0;D: Lclet Plateau, Lemkamin, 900 m., 15. IV. (}2, 1 d'. Possibly more lhan one species is irwolYed. EuparateUix pacificus Giinther. Solomons. - GUADALCAJ'\AL: Honiara, 27. VI.--1. VIII. 62, 1 d'. IndateUix sp. aff. novaeguineae (I. Bolfvar). Philippines. - l\HNDANAO: Curuan district, Sapamoro, 18. XII. 61, 1 ¥.

Superfam. E U M AS TA C 0 IDEA, nov. The assemblage now included in Lhe single family Eumasiaci­ dae differs widely from the Acridoidea, with which it has, till now, been associated. This is particularly true of the phallic struc­ tllres (Dirsh, 1961), hut it is also evident in the female genital apparatus. In this latter character there is some similarity to the anmnalous South Amt>riean family Proscopiidae (Randell, 196:~). Dirsh (op. ciL.) also notes that the two families may be distantly related, so that both may for the present be included in the same superfamily. It may be noted that they also share a further mor­ phological pecularily: most members of each group possess a smalL ventral tubercle on the penultimate or antepenultimate antenna I segment (the antenna! organ). Uvarov (1 966) tentatively suggested superfamily rank for both families.

Fam. EUMAST ACIDAE. Subfam. ERUCIINAE.

Erucius bifasciatus St~ll. Philippines. - PALA WAN: Mantalingajan, Tagembung, 1150 386 Entomologiske Meddelelser 34 (1966) m~ 17. IX. 61, 1 cJ' (also 2 juv., '~ this species, 19. IX. 61); Manta­ lingajan, Pinigisan, 600 m., 11. IX. 61, 1 S?. - MINDANAO; Curuan district, Sapamoro, 19. XII. 61, 1 0'. Subfam. CHOHOTYPINAE. ErianUms erectus Karsch.

Philippines: LUZON: Los Ba:fios, ~H. ~1aquiling, 26. I. 58, 1 1 S?. 'f M:nesides saussurei C. Bolivar. ?\ew Guinea.- WESTERN: Biak Mangrowowa, 50-100 m., 30. X. 1959, 1 S? (J. L. Gressitt). * This species was described from "Bivak Eiland" (a camp on the Eilanden Rivier, S. \V. New Guinea). Subfam. BIROELLINAE. BiroeUa woUastoni C. Bolivar. New Guinea. ---- NORTH-EAST: Daulo Pass (Asaro-Chimbu div.), 2500 m., 12 VI. 55, 1 cJ' L. Gressitt): Daulo Pass, 2500 m., 2. V. 1 0'. * t BiroeUa kuthyi C. Bolivar. I'\ew Guinea. - \VESTERN: Vogelkop, Manokwari, 75 m., 20. -23. VI. 57, 1 cJ', 1 S? (D. E. Hardy). * Bh•oeHa ornata C. Bolivar. New Guinea. - WESTER\I: \\Tisselmeren, Okaitadi, 1800 m., 7. 1 S? (J. L. * The specimen appears to belong to this species, but the arran­ gement of the hind tibial spines is BiroeUa papnana C. Bolivar. New Guinea. - PAPUA: S. Highlands, S.E. of Mt. Giluwe, 2200 m., 12. X. 1 cJ' (J. L. Gressitt); NORTH-EAST: Jimmi Valley, 840 m., 16. VII. 55, 1 cf' (J. L. Gres-

Superfam. A C R ID 0 ID E s. str. Fam. TRIGONOPTERYGIDAE. Subfam. TRIGONOPTERYGINAE.

Systella philippensis (Walker). Philippines.-- :\UNDAI'\AO: .\gusan, S. Francisco 10 km. S.E., D. Keith Me E. Keuan: Some Orthoptera-Caelifera . .. 387

13-16. XI. 59, 1 d', l S?. 1 juv. (C. M. Yoshirnoto); Zamboanga del ~orte, Masauan-Gundawan, rain forest, 1260-1:350 m.. , 3. VII. 58, 1 d' (H. E. ~Iilliron); Zamboanga del Norte, Manucan, 25 km S., 500 m., 18. I. 59, 1 juv. (L. W. Quate); Z.del Sur, 24 km. N.W. of Milbuk nr. Lebak, 450-900 m., 6-7. VIII. 58, 2 d' d' (H. E. Milliron); Bukidnon, Mt. Katanglad, 1480 m., 27-31. X. 59, 1 S? (C. 1\I. Yoshimoto); Sibulan, E. of Mt. Apo, Davao Province, 1000 17. XI. 46, 1 d' (F. G. Werner); Todaya, E. edge of Mt. Apo, Davao Province, 2800 ft., XL -Hi, 1 juv. (H. Hoegcstraal); Misamis Or. as follows: Balason, 1. IV. 1 juv. (H. Torrevillas); Mt. Empagatao, 1050-1200 m., HJ-30. IV. 61, 4 d'd' (H. Torrevil- 500-700 m., 20-24. IV. 1 S.E. of :Mt. Bala- 15 km. 1 ~ m. M. 1050 m., 24. IH-4. IV. 61, 1 2S?S?,3 . - BOHOL: Sierra IV-V. l S? C. - LEYTE: Mt. 29. VII. 1 S?.- BUCAS: no further 1 S?. - SIARGAO: 11. XI. 1911, 1 1\V. . - LUZON: fur- t.her , 1 d'. * t Fam. PYRGOMORPHIDAE. Tribe Nereniini. Kevan and Akbar (1 have recently indicated that lhe Solo- mon Island genus Modernacris vVillemse, recently transferred to the Dirsh to the Nercniini. also suggested lhat certain New Guinea genera, then unstudi- should probably be transferred to the same tribe. KeYan (1966 c) has recently investigated the questionable genera and has found that, on the basis of their phallic and female sub­ genital structures, and the form of the fastigium of the vertex, the following New Guinea genera, previously placed in the Acridid subfamily Catantopinae (sensu lato), belong to the Nereniini: Megra Campion, Kapauria I. Bolivar (of which Toxopeusiacris \Villemse = Toxopeusia Willemse must be regarded as a syno­ nym1), Buergersius (originally proposed as Biirgersius) Ramme, Paratarbaleus Ramme, Tarbaleopsis Ramme (of which Mzzsciacris

1 ) Toxopeusia flavomaculata Willemse, 1956 = Toxopeusiacris flavomaculata (Willemse) = Kapaoria flavomaculata (Willemse). 388 Entomoloqiske Meddelelser 34 (1966)

\Villemse must unfortunately be regarded as a synonym2 ), Fusia­ cris vVillemse, and Oxytarbaleus Ramme. It may also be noted that the Philippine "catantopine" genera, Jiezzbelio \Villemse and ,",'pi­ nucris v·Villemse, have also been found to belong to the Pyrgomor­ phidae (tribe Verduliini, not l'\ereniini)- see Kevan (1966 hi. The "Noona Dan" material contains no specimen belonging to the above genera, but lhcre is included a single specimen of a new genus of Nereniini. Another species of the same genus was also discovered almost simultaneously among the other material con­ sidered herein. It was intended that the original descriptions should be those given here, hut differential rates of publication have resulted in short diagnoses already appeared (Kevan, 196fic).

Gen. Noonacris. Body cylindrical with a very short head, apterous and ·vdthout a tympanum; external morphology shovdng close relationship to Poraturbuleus Hamme, but differing as follows: head not broader than long, less strongly narrowed towards the pronotum: frons subhorizontal, scarcely sloping; middle segments of antennae at most one-and-a-half times (nol about twice) as long as wide; middle coxae and mcsepislerna wilhont distinct tuhercles; anterior and middle femora, without spine-like tubercles below; segments of hind tarsus not .~uhequal, the second the longe>st; male subgenital plale elongate; epiphallus more typical for Pyrgomor­ phidae (of lhe form illustrated in Fig. 2H); endophallus very narrow with peculiar elongate wing-like structures associated

with the aedeagal sc!cri1es3 ). Type species, N. pusilla. Two species of this genus are known, both only from single specimens, one a male and the other a female. It is possible, but scarcely prohahle, that they are conspecific. They may he more fully described, as follows:

Noonacris pusiUa, Kevan. (Fig. 1, 2 H). H o l o t y p e d', Bismarck Isl. - MUSSA U: l\falakata, 15. Fe-

2 ) The type species of Tarbnleopsis is T. tuberculata Ramme, 1930: that of Musciacris is M. tuberculata \Villemse, 1956. The generic synonym~· thus also invoh·es homonymy, so lhal \\'illemse's species had to be renamed Tarlmleopsis willemsei. The seeond species of :llusciucris has becomp Tar­ lmleopsis brunnea (\Villemsc, 1\)56).

3 ) The phallic structures and female subgenilal armature and spermatheca arc more fully discussed and illustrated by Kevan (1966c). D. Keith McE. Kevan: Some Orthopteza-Caelifera ... bruar, 1962, "1\oona Dan Exp. 61-62 (Zoological Museum, Co­ penhagen). Head: about as wide as long, distinctly wider than pronolum, virtually without a median carinula; antennae depressed, smooth, cylindrical, no longer than head and pronotum together, with 16 short segments (none more than 1'% times as long as wide) in addition to scape and pedicel; eyes large, oval, but not very

A A-D, 3 MM.

c E-F, I MM

~ I , E I I {) F I B 1'1,1 Fig.l. Noonacris pusilla Kcvan. "1, o holotype, lateral; B, o head and pronotum, dorsal; C, o frons; D, o sterna] lamina; E, o abdominal terminalia, lateral; F, the same, dorsal. strongly protuberant: vertex descending steeply between the eyes to the fastigium; latter also descending, about three times as wide as long, rounded apically with a slight median furrow, dorsal impressions transverse, their posterior margins not very distinct: frons almost horizontal, median costa weak, not reaching the clypeus, more or less parallel-sided, but slightly expanded in the region that would he occupied by the median ocellus (were it developed)· and between the antennae, weakly furrowed through­ out; lateral frontal carinae obsolescent; cheek tubercles lacking_ 390 Entomologiske Meddelelser 34 (1966)

Thorax : depressed; pronotum smooth, subcylindrical, as wide in front as behind, anterior margin of disc evenly convex, posterior margin very slightly concave, median and lateral carinae lacking, all three transverse sulci strong and evenly curved, follow­ ing the contours of the anterior and posterior of the disc, anterior sulcus placed at about one-sixth of the distance from the anterior margin, median sulcus placed behind the middle of the disc so that the distance between it and the anterior sulcus is about equal to that between it and the posterior margin of the disc, posterior transverse sulcus at about five-sixths of the prono- tal length; lateral lobes much than strongly oblique, convex, straight.

anterior sulcus and meso- and melanota half as latter, holh tum; and mesosterna! lobes and inter-

and suture, metaslernal lobes contiguous L e g s : anterior and middle femora compressed, former short and somewhat incrassate, without or tubercles below; middle coxae with a small, external tubercle; posterior femora with dorsal lobe projecting anteriorly a little beyond the basal one, external face of hind femur very smooth, with a very weak pattern; hind tibiae with both external and internal apical spines and 9 inner and 7 outer spines in addition; hind tarsus very long, about three-quarters of the length of the hind tibia, second segment slightly longer than metatarsus, both longer than the apical segment, arolia rather large and disc-like. A b d o m en : slightly depressed at the base, compressed beyond this; first tergum a little longer than metanotum to which it is similarly punctured; tympanum absent; tenth tergum broad­ ly emarginate in the middle; epiproct very short and broadly ,triangular; cerci stout, conical, extending distinctly beyond the D. Keith McE. Kevan: Some Orthoptera-Caelifera . .. 391

A-G,4MM.

8 H,- 0·5 MM. }-----...... j

fl

G F Fig. 2. A-G, Noonacris rwualliberuica Kevan; H, Noonacl'is Ke\'[m. A, ¥ head, lateral; B, ¥ head and pronotum, dorsal; C, ¥ frons; D, ¥ sternallamina; E, ¥ right hind tarsus; F, ¥ abdominal tcrminalia, dorsal; G, the same, lateral; fl, cpiphallus, dorsal.

apex of lhc narrow and a:, illustrated C o l o r a t i o n : rather uniformly light as follows: eyes brown, antennae the segments each nar- rowly annulated with grey, the subgenital and ventral faces of the hind femora also hind tibiae bluish-lilac with spines, hind tarsi bluish-grey above, below. M c a sur em en t s : length of body 12.6, antenna head pronotum 2.25 hind femur 7.0 mm. The type is unique. Ent. Medd. 34 25 Entomoloqiske Meddeleiser 34 (1966)

Noonacl'is novahihunica Kevan (Fig. 2 A-G). Ho lot y p e C(, [Bismarck lsl.l :\fEW IREL\ND .) : Ridge above "Camp Bishop", L5 km. up Kait R .. 250-750 m .. 13. VII. 1956, J. L. Gressitt Collector !B. P. Bishop :\luseum, Hono- * but differing us follows: size larger : head proportionately broader and lo the frontal of the wide1wd vvhcre the · fronl and midd1e cf.

Tribe ') dahH Ramme. The genus is in process of hut the mate-

·±) .\s already noted tribe. Kevan and v;ilh D. "\Vas also Hamme. K~;van and Akbar now has the status of a r,evan (19!\fh) should also be species of Dorincl!a should be D. cheesmanac. D. I

3 4 5

A ,______I Mlvi. _,

B B

(

5. Epiphalli (A) and dorsal (B) 3. D. holotype; BoliYar); 3. (I. Bolivar).

as follows: Keravvat , 60 m., 20-2:J. XI. , 29. nn. 56, 1 cJ, 3 ~~ L. Gressitt), 11. IX. r1 m trapj Gressi , 23. X. forest L. Gressitt) ; 135 m., 20-25. XL 2 ~~

25:U 394 Entomologiske Meddele!ser 34 (1966)

Yundal, S.W. of Keravat, XII. 59, 1 cJ'; 13. XII. 59, 1 cJ' (T. C. :\faa); Vunabakan, 10 km. E. of Keravat, 180 m., 16-20. XI. 59, 1 d', 1 <;?, 1 juv. (T. C. 1\faa); Vunabakan, on Ipomoea, i). Y. 56, 1 juv. (J. L. Gressitt); Bainings, St. Paul's, 560 m., light trap, 4. IX. ;)6, 9 d'd', 8 S?S? (.J. L. GressiU); the same, 350 m., 5-9. IX. 56, 19 d'd', 21 S?S? (J. L. GressiH); Puktas, Baining Mts., 22. XI. 57, 1 cJ' Smart); Gaulim, 140 m., 19-27. X. 62, 8 d'd', 5 <;?<;?, 2 juv. (J. Sedlacek); 16 km. S. of Gaulim, 300 m., light trap, 29. X. G2, 1 d'. 2 S?S? Sedlacek); Mt. Sinevvit, 900 m., 7-16. XI. 62, 1 d', 5 S?S? Sedlacek); Warongoi V al., 100 m., 24. Y. 56, 1 d', 1 S? (J. L. Warongoi, 250-600 m., 28-:W. XL 1 <;? the same, 1250--1450 m., 1-5. XII. 62, 1 d', 1 <;? · the 230 m., 8-15. XII. ] d', 7 • *"-1

The \vas collected more not slated in lhe , 1 H is not m nor in any other collection known vvith other the

is green, the antennae annulated ·with green. the the and with brown on base ·with abdomen rare in

nol known in so that Lhere is a genus. However, 11. so far as it goes, fits Desmopterella and one must assume that he had an coloured discoloured) specimen before him. 1 have also had reason previously to con­ o;ider Bolfvar's sense of colour somewhat unconventional, so that I .am not inclined to give loo much weight to the matter of colour. D. Keith Me E. Kevan: Some Orthoptera-Caelifera . .. 390

Some of the specimens listed below, all of which are greyish or brownish, variously mottled, have, in fact, an olive-greenish sheen on the tegmina when these are viewed in certain lights (although not more so than in some other including B. dahli). Irridescence on the bases of the hind wings may indeed make these appear bluish in that region, and the veins may sometimes be olivaceous, although they are usually fuscous. Some specimens have somewhat olive-green hind tibiae; others have these brown­ ish or even obscurely violet. All have two or more widely spaced, very distinct, pale, usually yellowish or orange (but not green), annulations on the antennae. This last character is consistent and separates the series below from all other species (including D. dahli), which may sometimes have slightly variegated, or even irregularly, but never distinclly, annulated antennae. Since BoHvar (L c.) also remarks on such I feel confi­ dent that the material below belongs to his pmsina, the holotype of which was probably unusually coloured. The male struclures of this species somewhat resemble those of D. dahli, but are distinctive -!). Bismarcks: LA YONGAI: Banalam, 22. II. 62, 1 ¥. 1 d. - NE\V IHELAND: Lelet Plateau, Lemkamin, 900 m., 14. IV. 62, 1 ¥. The following may also be recorded from NE\V IRELAND: Lavalai, 1 S2 (Biihler); Majom, 1 S2 (Biihler); Lelet Plateau, 1 S2 (Blihler); Kavieng, X. 3(), 1 ¥·'), 2 juv. (L. E. Cheesman); S. W. lUdge above "Camp Bishop", 12 km. up Kail R., 2-!0 m., 9. VII. 56, 1 d, 1 (E. J. Ford Jr.); the same, but 15 km. up, 250-270 m, 13. Vlll. 56, 2 d d L. VII. 56, 4 dd, 3 S2S2 L. 1-2 m., 2. VII. 1 d (.T. 3 dd, 1 S2 1 d J. L. 56, 1 d [+ 1 Plateau, X. 2 dd, 6 ¥¥

3 ) This specimen is selected to be the ncotype of D. prasina. Il is !he properly of the British :\Iuseum (Natural History), London, as are lhe associated nymphs. Data label reads: NE\V IRELAND (Bismarck arch.), Kavieng, x., 1936, L. E. Cheesman, B.::\1. 1936-271. Colour: generally bro>Ynish-teslace­ ous (inducting tibiae and tarsi); numerous fuscous maculations; hind wing Yeins light fuscous. :VIeasurements: body 30, pronotum 6. tegmen 29,5, hind femur 13.6 mm. Entomologiske Meddele!ser 34 (1966}

lhese

of another germs that conld come from l\f'\Y similar dala. the above counled as there i.s also malerial or the same data (sec p. , and the:,c could not have come from ~cw Guinea. Both Desmopterello and can well, besides which, two two dates and different collectors were invoh·ed. so that the records must he the benefit of the doubt. certainly require confirmation. to note that a male of the same species. in the Bishop l\Iusenm. Honolulu, is labelled: Sar::nvak. Kapil District l\Ierirai V .. 30-300 m., sandy 1-G. VH1. 19.)8. T. ldaa. If lhe data are correct. !he much more distributed than was preYioush: first from the region. Des- was dPscrihcd in part frmn Borneo. Kevan, 1963 for discussion). If D. in fact. occurs m Borneo and :VIindanao, there would ;llso be less reason to doubt the correctness of Gilolo (HalmaheraJ as a locality for n. Rmnnw. another species known otherwise from :'\ew Guinea l.

of genus is A. mira Hehn from llw southern described a a yery distinctive h·muk from Luzon. of well of the smne germs. ,ln structures has shown not distinct from A. D. Keith lHcE. l(evan: Some Orthoptera-Caelifera . .. B97 lml also that these structures differ lo such an extent thul il wouid lo retain the two in Hw same genus were the structures the criteria to he used. Since OYerall simila- must be a new

expo~e the apex of the hut and the retain at least an indication of the The male abdominal tcnninalia arc not

sclerites which are aho unusual among in jecting well lhe valves to which fused. Further, !he aedeagal lack the very dorsal inflections and the "hairy" covering found in "1podcsmoptcra, s. str. (cf. Figs. 7 and 8), and the zyoma of the cingulum is con­ siderably larger and wider. Type species (here desigualed): _4podcsmoptcm mim lu.zonica Kevan, 196:3 = Apodesmoptera ( Bmchydcsmoptem) hzzonica Kevan. Even allowing for the fact that the phallic structures in Des­ mopterini vary considerably within genera, it is i1npossihle to consider Brachydcsmoplcra as having less than suhgeneric s!atus.

(BI·achydesmo}ltera) luzonica stat. noY. 6, Pl. I figs. a, e, d, Pl. H fig.

'I LUZON: " Islands" !no further

il is nevertheless of very for it is upon it that lhe new is based. There is little lo add to the brief des- of lhe female Kcvan (1 lo state lhal it is of an almost uniform brown colour, very like the of !he next. and to illustrate i.ts arma- ture and spcrmathcca 6). The male here recorded is yery 398 Entomologiske Meddelelser 34 (1966)

f------__i_M_M_:_. ____,

Ii LJ (

E~~~\\ ==-£~ ff ~{'-I

Fig. 6. Apodesmopiera (Braclzydesmoptera) luzonica Kcvan. A., epi­ phallus, dorsal; B, cingulum, dorsal; C, endophallus, dorsal; D. the same, lateral; E, S suhgenital armature, dorsal; F, spermatheca.

hut is smaller and more slender. In color­ hrownish-testaceous with the dorsal surface of the the pleura and the hind femora somewhat . Jt is also very similar to the male of the next species, except for the characters indicated in the suhgeneric description. Its measure­ ments are as follows: length of body 25, pronotum 5.0, tegmen 5.5, hind femur 12.0 mm. D. J

PLATES I-III Entomologiske Meddelelser 34 (1966)

c

d

PI. I. (a), (c), (d), Apodesmoptera (Brachydesmoplera) luzonica Ke­ van. (a),

h

Pl.II. (e), (f), (g), "4podesmoplera (A.) curlipennis, n.sp. (e), (f). o holotype; (g), SJ allotype. (h), "4. (Brachydcsmoplera) luzonica KcYan, SJ lwlotype of A. mira luzonica. Entomologiske Meddelelser 34 (1966)

2 cm.

PI. III.Opiptacris and Cranaella, new spcies. (a), (b), 0. castanea, n. sp.,

A

c B

D

II I I 21MM. J...---l I l

Fig. 7. curtipcnnis, n.sp. A, epi­ endophallus, dorsal; lJ, the sarne, lateral; E, S? suhgenita! armature, dorsal; F, spermatheca.

n. sp. (Fig. 7, Pl. I fig. h, Pl. II figs. e--g). Ho lot y p e: d', ~ [LUZON]: Ca[ra]marincs snr, Mt. lsarog, 800~1200 m., 27. IV. 1963, H. M. Torrevillas Col- lector P. ::V!useum, Honolulu). * This is so similar to the male of A. (A.) mim as to make a delailed description unnecessary, but it differs as follows: smaller, with somewhat reduced, narrower tegmina and wings which extend 400 Entomologiske Meddelelser 34 (1966)

A

8 c

F

Fig. 8. Apodesmoplera (Apodesmoptera) miru Hehn. A, epiphallus, dorsal, B, cingulum, dorsal; C, endophallus, dorsal; D, the same, lateral; E, SJ suhgenital armature, dorsal; F, spermaiheca.

\Yell the middle of the abdomen and hind but do not reach their points of indicated but not nTy shorter than in A. minal terminalia cerc1 similar but nar-

areas 111 carinae and the upper lobe dark hlaekish in basal third; hind femur D. Keith McE. Heuan: Some Orthoptera-Caelifera. 401 wilh a basal area. ventral face Yentral keels with dark inferior reddish: }_..rownish-teslaceons with dark macula lions. ::\Ieasurements. 1 hind femur 12.5 mm. Allotype: Collector fenwra and cerci in smne Y\'ay as does the armature spermMheea similar to ~ t mirr1. illustrated 7 and 8). C o l o r a l i o n : almost uniform for darkt~r suffusion near the anterior and on lhe "shoulders" of !he and dark maculation;; on lhe fore and middle lihiae. :\le a s u c m e n t s : for curvature of ea. 42. hind femur 1\L'i mm. :\'o further material is known.

(AlJodesmoptera) mi.ra Rehn. R). ::\Iost of the l

Tribe lltractomorphini.

Atracl:omm·pha IJSjiHlH~Ina (Haan). Philippines.~- BALABAC: Dalawan 1 d', 1 ¥. - Pc'I.LA \V : Rrooke's Point, 2 ~S:: 11. IX. fil, 1 d': Brooke's in the ') --· MI-:\fD.\K\0: Curuan district moro, 18---'22. XH. fi1, 1 '_,')on +-t-· This fron1 many unrecorded loealitil'S loo numerous to list here. The AJ\;, however, are of interest in vi('w of from that island: 16- 402 Entomologiske Meddelelser 34 (1966)

23. V. 58,3 cfcf, 4 yy (H. E. Milliron); 8 km. S.W. of same, Eran Point, :31. XII. iJB-4. I. GO, 3 yy W. Quatc); Brooke's Point, :\Jacagua [=l\iakagwa], 27-:il. Hl. 62, 1 juv. Thompson); ;) km. 1\.E. Tinabog, 14. V. 1 cf (H. HoUmann). *t Bismarcks.- NKW BlUTAIN: Vunabakan, 10 km. E. of Kera- vat, 16-20. XL 59, 2 yy (T. C. * :ltractomorplw is a very well known genus and A. psittacina is a common, widely distributed species, but the above record for N cw Britain represents a disjunct distribution. The species was hitherto unknown east of Mindanao, N. Celebes and the Takang­ hesi Is.- apart from "l\loluccas". It is possible that the have been mislabelled. Atractomorpha ct•cnaticcps crcmdiceps (lllanchard). Bismarcks.- MANUS: Lorengau, 18-25. VI. 62, 3 cfcf, 4 yy. - :\IUSSAU: Talumalaus, 1. II. 62, 1 y [very small].- LAVO:'\J­ GAI: Banatam, 17-25. HI. 62, 8 cfcf, :3 yy. - DYAUL: KoJic­ pine, 12. Ill. ()2, 1 cf, 2 yy.- NEW IRELAND: Kalili Bay, ;30. IV. 62, 1 y; Lelet Plateau, Lemkamin, 900 m.. , H-21. IV. 10 cf cf, 12 y y. - :'\I E\V BRITAlN: Gazelle Peninsula, Y alom, 1000 m., 10-23. V. 62,6 cfcf, 12 yy, 1 juv.; Cape Hoskins, E\va­ lakessi, 3. Yll. 62, 2 cfcf, 2 yy; Vaisisi, 9. VII. 62, 1 y; and Valoka, 10. VII. G2, 2 cf cf. The following material fron1 the Bismarck Archipelago may also be recorded here: MANUS: Lorengau, 1-75 m., 28. VI. 1 cf, 2 yy [small], L. ; Rossum, :35--·123 m., 29. VI. 1 y L. . - LOS KEGROS 1 d . - NEW lRELA::\TD: ; Kavieng, 2. Yil. ; Schleinilz Lelet Plateau, X. 2 yy 2:3. and 24. XII. i)9-1. L 1 cf, 3 yQ . - KEW BHITAIN: Nakanai 1060 n1., 25. VII. 56, 1 d' L. ; the same, 2 cf cf, J. Ford Jr.); the same, Silanga, 150 m., 31. VII.-5. VIII. 5G, 14 cf cf, 8 yy (E. J. Ford Jr.) : Barronga Island, nr. Linden- 28. IV. 56, l cf L. Grcssitt); Lindenhafen, 2 miles S. of coast, 24. IV. 1 y L. ; Linga Plain, vV. of \Villaunwz Peninsula, 9--12. IV. 56, 2 cf cf [1 on sago 1 y; Keravat, 135 m., 20-25. IX. 59, 1 cf, :3 yy . C. Maa); Gazelle Peninsula as follows: \Varongoi Valley, 100 m., 24. V. 56, 1 cf (J. L. Gressitt); Kerawa, primary bush, 60 m., 1. IX. 4 D. Keith Me E. Kevan: Some Orthoptera-Caelifez·a . .. 403 d' 1 c;?; Bainings, SL Paul's, :1GO m., 'J--7. IX. 55, 10 d'd', 12 S?S? (J. L. Gressitt); Gaulim, 140 m., 21-27. X. G2, 1 d', 2 S?S? Sedlacek); Mt. Sincwit, IJOO m .. 5-14. XI. 62, 1 c;? (J. Sedla­ ; Upper Warongoi, Illugi, 2:10 m., 8-21. Xll. ()2, 4 d'd', () S?S?. *t Solomons. - BUKA: Gagan, 40 m., lG. VI. 5ti, 1 c;? (very small) 1 juv. (c;?) (J. L. Grcssitt). -- BOUGALYVILLE (S.): :\losigera, 25 m., 3. VI. 56, 1 S? (J. L. Gressitt); :\fumurai, 6. VI. 1 d' L. GressiH); nr. Crown Prince Hange, 900 m., 6. IV. 1 d', 4 S?S? (J. L. Gressitt); the same, 690 m., 8. VI. 5(\, 3 d'd' L. ; Guaba, 720 m. H)--21. VI. 5(), 2 d'd', -1 c;?c;? J. Ford 2. VII. 1 c;? (E. J. Ford Jr.); Kiela, 26. XL . - KOLOMR-\NGAHA 1-12 m., 8. VII. ;)9, 1 cJ' L. . - :VIALAITA: 2-20 m., 21. IX. 57, 1 . - GCADALCA- NAL: 12. IX. 1 1 '¥ A. 21. V.-10. VI. , 1 . IQ), E. (.J. L.

Fam. A C R I D I D A s. slr. Snbfam. OXYINAE. intricata Stal. -- LUZON: Mt. Niaquiling, 21-26. Vl. 2 cJ'

chinensis (Thunberg;. Philippines. - BALABAC: Dalawan Bay, 4. X. 2 c;?c;?. - PALAWAT\: Brooke's Point, Uring 15. VIII.-8.lX. Gl, 4 40-1 t:'ntomologiske Meddele!ser 34 (1966)

Brooke's Point, f!Iakagwa, 22. YIH. () 1. 1 in the barrio. ;)00 m., :il YIII.-12. !X. ')? dd ·r ~ X.~- 61) ' 0- . ' -·0 no+ T' I -TA.\\'I Tc\\\TI: 2;). 4. XI. 61, 11 ctd, 8 ??. 1 Coast 0:. of Tarawakan, '27. X. 61, 2 ??. - ~\HNDA~AO: Curuan Sapamoro, 16-22. XII. ll d.~~¥¥,5 smalicr !han those from the other islands. somewhat hut do not

lS

ven-

of normal

6 ) 1 9, 18. HI. attached lo grass stem and upparcntly infected by lhe fungus En1pusa grglli ( ?) . D. Keith Me E. Kevall: Some Ortlzoptera-Caelifera . .. 405 length), d d' and 1 SJ with femora discoloured, dark with black ; Cape Hoskins, Valoka, .f. YH. 1 cl'; S. of Hoskins Aerodrome, 6. VII. 62, .1 SJ and last coloured as Hermit L

Gesmmla mundata mumiata (vValker). Bismarcks.- ::\lUSSAU: .f. II. Gesmuda mnndata

Bismarcks. -- ~EW IREL\1\ : Lelet m., 15-20. IV. 2 d'd', 2 SJ2 (1 SJ almost without red on hind femur; NEW BlUTAIN: Gazelle Penin­ SJ;

at

Racilia abbrcviata Hamme. Mt. in rain !:lOO m., 29. VI. 1 2 * The female of this "'~-'"'-'·' .... "' has not hitherto JJeen recorded. This 406 Entomo/ogiske i\1eddelelser 34 (1966) specimen agrees fairly \vell with the description of the male, but is larger (28 mm long) and brownish dorsally. The hind femora arc red apically, otherwise green, hind tibiae and tarsi bluish (not red) ; the sternum is not red, but the ovipositor is so coloured. It is doubtful if the leg-colour will prove to be a good specific character in this genus, and it could also be that the degree of development of Lhe wings is variable above), but the of the vertex appears to he shorler in this species than in the last mentioned in the original description). Bumac.ris flavomaculata Willemse. Solomons. - MALAITA: Auki date), 1 d. t The phallic structures as well as external confirm thal Bunwcris to the :Bumacris sp. ? nov. Solomons.- SAN CRISTOVAL: Pamua (no , 1 ¥ . N. ·* This is recorded here as no member of the genus has from San Cristova!. It a new somewhat discoloured female I decline lo describe il without further material. flavomaculata I. Bolivar :-.re1Y Guinea. 1G5 m .. H. I. 1 cJ

blackish­ nlaculae mark at each end of the lateral are also pre- sent on the meso- and the dorsum of the meso- and metanota and abdmnen hears a rather broad The femora are all green with · the tibiae are all blue-green to blue; the hind knee-lobes are blackish. The an- tennae are brownish. The phallic structures as well as the hind femora indicate that Caryanda is a member of the Oyxinae. The same is true of Tmzchiridea. D. Keith Me E. Keuan: Some Q,·thoptera-Cae/iiera . .. 401

Tauchiridea laeta Ramme. Xew Guinea. - NORTH-EAST: Adelbcrt Mts., vVancuna, 800 -1000 m., 27. X. 1 ¥ (J. L. Gressitt). * Tauehiridea adusta I. Bolivar. ~ew Guinea.- NORTH-EAST: Baynyik, S. of Maprik, 150 m., 12. I. 60, 4 dcJ, 1 ¥ (T. C. Maa); Nadsab, II. 44, 1 ¥. * t

Subfam. COPTACIU~AE. Coptacra foedata (Audinet-Serville). Philippines. - PALAWAN: Mantalingajan, in the barrio, 500 m., 6-23. IX. 61, 4 cJcJ, 7 ¥¥. Bolivar).

- :\H~DANAO: Curuan district, Sapamoro, 17.

Philippines. - LUZON: Los Bafios, Mt. 21. VI. 62, cJcJ,1¥ .t A. cyanoptera may prove to be a synonym of A. contractu

Subfam. EYPREPOCNEMIDINAE. brunneri Bismarcks. - MANUS: Lorengau, 15-19. VI. 62, 11 cJ 5 ¥?, 1 ; NEW BRITAIN: Cape Hoskins, Valoka, 7-13. VII. 2 d', 2 ¥¥. The form of the phallic slructures as well as the external cha­ racters confirm that Rectitropis belongs to the Eupreprocnemi­ dinae. Subfam. CATANTOPINAE, s. str. Gen. Opiptacris Walker. This little-known genus is known chiefly from the Solomon Islands and, although no specimen is among the "Noona Dan" material, opportunity is taken to comment upon it and to describe a new species from New Britain. The subgenus Salomonacris "\Villemse, 1931, cannot be regarded as valid, for there is no clear distinction between those species having minute, vestigial tegmina ( Opiptacris, s. str.) and those that are completely apterous (Salomonacris). Most specimens,

En!. Medd. 34 -±08 Entomoiogiske Meddeieiser 34 (1966) evPn of allegedly apterous may he seen to have a! leasl on one side, if they are examined

n. sp. a. Ho lot p e: S:J, ~E\V BRIT)tl~. 1050 m., 25. VH. E . .J. Ford Jr., P. ::\Iu- SCUlU, a little ihan wide, wider !hau vertex very convex: antennae about. as in addition to

; eyes rather the width the diameter of an eye in dorsal m of vertex marked off from the rest of the vertex and sel at a lmver level at ahoul all eye, horizontal, apex pnra- frons v.-rinkkd. above median ocellus: checks smoolh. T h o r a x : with disc aboul wide at front and rear, sides concan~, anterior convex, smnewhat concave. nwdian carinula indicated narrow furrow. four transverse or less transverse, the distance hetween !he !he ahout of the pro- no!um, distances between the other sulci and the anterior and of Hw disc all ; lateral lobe a little end where Js convex, anterior and oblique and some\Yhat convex,

rounded: rneso- notum pronotlnn: metanotum ahoul as as the pronotum between the median and prostcrnal tubercle transverse, and trunc- ated at apex; mesosterna! lobes a liltle wider than long, their inner convex and their interspace nar- rower than a lobe; metasternal interspace triangular. Tegmina l v e s t i g e s : very short and broad, not quite reaching the posterior margin of metanotum. L e g s : hind femora not quite reaching apex of abdomen, D. Keith Me E. Kevan: Sonze Orthoptera-Caelifera . .. 409

stout and with a very strong "aretc", or pattern: hind tibia wilh four ouler and 6 inner second hind tarsal somewhat

brownish- testaceous wilh all carin u1ae and indicnted m black; hind tibiae and tarsi IS distinctive to war­ rant lhe erection of a nev,r smce this is first record of the genus from the

coloured. H.amrne. Solomons. - BOUGAINVILLE ()90 m., 12. VI. 5() J. Ford Jr.). - BUKA: 6--10. xn. 1 cJ C. The first agrees we11 with the description, except that the hind femora are red wilh black bases: the hind tarsi are also red. The second specimen is very similar to the first, but the thorax is not red, the abdomen is not such a bright red and the hind agree in colour with the description for 0. salomona which is almost certainly a synonym and the otherwise unknown female

26* 410 Entamologiske Meddelelser 34 (I 966) of 0. bougainvillea. Bolh the males above have very minute vesti­ The genus badly needs revision since there is deal of variation in the tegminal vestiges and, more in the coloration which has preyiously been used as a specific character. (Willemse). Solomons. - lVL\LAITA: Tangtalan, 200 m. 23. IX. 57, 1 m. Stone). * This agrees v,rell with the original description, save that very vestiges are present. It is not improbable that 0. the female of this species.

uniformis \~TilJemse.

pos- ges, but before me. It bears the It but it does not merit tion. Jt is mentioned here because there appears to he no other record of the genus from N cw sp. aff. hHa.ris 'Valker. Solomons.- FULAKORA: the turn in contrast to the otherwise this in the black semilunar areas of the hind it agrees with 0. hilaris. That \vas, however, described from the New Hebrides and has darker antennae and the front four femora black. Cranae sp. aff. New Guinea. - \VOODLARK I. (Murua): Kulumadau Hill, 19-22.1IL 57 W. Brandt); NORMANBY I:. Wakaiuna, Sewa Bay, 11-20. XII. 56, 1 ( d') (W. W. Brandt) [almost certainly the same species as from \Voodlark * C. patagiata was described from Amboina, but was also record­ ed by Krauss (1902) from Papua. The present material is presum­ ably the same as that from the latter region, hut in view of the wide separation of the localities concerned from that of the type, D. J

~e\Y Guinea. 2 d'; Roon I., 1 ¥ CranaeHa tubet·culata Ramme. - ::\HNDANAO: Misamis l\H. 1050 Hl., 24. HI.-4. IV. 1 d' the same, 1050-1200 m., 19--30. IV. Gl, 1 On, .*t n.sp. c, d). Ho l o ype Is. -- MINDANAO: La Lun Mts. Prov.), 1930 Mu-

narrower than back to the second addilion io scape and those at base and apex hYo to three renminder up to four as wide: eyes a very little longer than vertex convex, rather smooth with several 412 Entomoiogiske lifeddeieiser 34 (1966)

of n~ry ; frons of ,-crtex narrow- rather acute, inclined dowrrward: frontal narrO\Y between the antennae, about as wide as the urn of the Yert!:'x in the of lhe median ohsolele below this: lateral frontal carirwe obsolescent; cheeks somewhat ·wrinkled. Thorax Yvith strongly and with- out longitudinal anterior and posterior margins of disc almost typical transverse sulcus at about three­ quarters of the pronolal median transverse sulcus at about the middle of lhe more or les'> anterior transverse sulcus lacking on tlw disc, lateral pronotal lobes about as deep as long, anterior and posterior margins oblique, straight, inferior margin sigmoid. infcro-posterior angle oh! use: meso- and m eta­ nota together slighlly shorter lhan pronotum, the former the shorter of the two: prosternal tubercle vertical, strong, thick, transn~rsely compressed, quadrate in posterior view; mesosternal interspace about as wide as a mcsosternal lobe at the base, sides concave, widely diverging. much vYider than a lobe posteriorly, mesosterna! lobes subquadrate, narrowed and rounded behind on their inner margins; metasternallohes meeting behind, their inter­ space very widely triangular. \Vi n g s : tegmina widely separated dorsally, ovate, about two-and-a-half times as long as wide, their apices rounded, ex­ tending almosl to lhe posterior margin of the first abdominal segment; hind wings vestigiaL Legs : hind femur rather stout, reaching apex of abdomen, carinulae of external face slightly, but not conspicuously, tuber­ culate; hind tibia stout, much shorter Lhan hind femur: hind tarsus about two-thirds as long as hind tibiae, basal segmenl the shortest and apical segment the longest, claws strong, arolium large. Abdomen : tenth abdominal tergum broadly excised: epiproct tongue-like, slightly longer than wide, rounded apically and with a pair of submarginal, sinuous carinulae bordering a median depression: cerci stout, conical, rather long, not quite reaching apex of epiproct; ovipositor valves rather short and stout, strongly hooked; subgenital plate strongly tridentate apical­ ly, the lateral "teeth" as long as, and broader than, the median point (egg-guide). D. keith Me E. kevan: Some Orthoptera-Caelifera . . 413

::\1 e a s u r e m c n t s : length of antenna 16.5, prono- tum G.5. hind femur 19.5 miTL C o 1 o r a t i o n : antennae black for brown n:meulac an triangular chocolate-brown patch extend- of tht· yertcx to !he and a hro\Yn same colour behind each ey(;: eyes brown; ; first lhrec abdominal terga remainder, apical margins of !he ninlh and tenth, yello·w; cpiproct and cerci blackish; ovipositor valves dark green: tegmina green, yellowish at the anterior and middle femora light brown, their tihiae and larsi olive-green. hind femora ochreous yellow, suffused hlue-black on hoth faces in lhe basal one-third, small tubercles of carinulae unicolorous, hind knees black, semilunar areas dark brown; hind libiae blue-black, spines black; hind tarsi blue-green. Th<; holotype is, unfortunalely, unique, hut the species is unlike­ ly to he confused with any other, especially on account of its conspicuous colour pattern. Ph.alaca ikonnikovi Ramme. Philippines. - LlJZO:N: Ca[ra]marines Sur, :\H. Isarog, 500- 600 m .. 15. IV. 63, 1 cJ juv. (H. :\f. Torrevillas). * Although the specimen is immature it almost certainly belongs to this species because of the characteristic arrangement of yello·w maculae on the pronotum.

~1aga dichroa I. Bolivar. Philippines.- DINAGAT: 22. IV. 45, 1 c;>; Panamauan, 24. IV. 45, 1 (juv.) (H. Roberls). t =I= The coloration is more uniformly green lhan in the typical form from the island of Snmar. More material would be needed to determine if this has any significance. Paramaga olivacca vVillemse. Philippines. - BALABAC: Dalawan Bay, H. X. 61, 1 c;>. - PAL\\VAN: Manlalingajan, Pinigisan, in the barrio, ;)00 111., 1. IX. l:il ' 1 Pscndocranae gracilis \Villemse.

~ew Guinea. - NORTH-EAST: Nondugl, 2200-2700 m., 28. V. 59, 1 0' (C. D. l\Iichener). * 414 Entomoicgiske lHeddc!elser 34 (1966)

Pseudoeranae loriai I. Bolivar. The nmne . hut afkr or"''""o.Q was nan1ed was a Jnan, and the rules of nomen­ terminal "i" for all modern male )Jew Guinea. - PAl'Ul1.: Bnnvn H. nr. Port 8. X. 1 y L. llilnaete bhnaeuJata Bci-Bienko.

~- LUZOJ\': Los 100 m.~ 2H. I\'. 61. 1 C·

wiUemsei Bci-Bienko. 100 m .. 21.

Los cics. the corded.

2 2 1 in the - TA\VI TAW!: --Hi. XL fj L 11 d' d', Curuan districL . -- LUZO:'\: Los Bafios. Hl61. 'f.

, 2 <(Q.-- ;o(JQ 'if(TSc'r:·. 1 ;y + T' - lh ... ,)_ '- L yy; Boliu, 1:'5. H.

\ 71. 1 y; l d'. -- LA VONGAT: Banatam. I 8. HI. --- DYAUL· 12. HI. 1 --- ~E\Y IRE- LAND: , :10. IV. - DUKE OF YORK: . 1 y. - NEW BRITAl:'\: YIL 4. VII. :1 cJ

Solomons.- Island unknown further ' 1 y. 'f D. f{eith ~11cE. Kevan: Some Orthoptera-Caelifera _ . 415

humilis humms cf', l C(. --

500 m., 6--

s. str.

-- LUZON: National t

Meli•~odes tenebrosa vitaticoms - LUZON: Los .t appears to have a more ;:,v•aunc< distribution in ~~S with the pre­ occur almost in the , doubts as to the between the hyo are known from southern Luzon 10-H:

seem to - at least as far as conspersa Uvarov and the nominate form are concerned. Both occur in many and intermediates are \Villemse reached the same conclusion in of most, if not of the so-called Bismarcks.- LUF 2f\. VI. 2 cf'cf', 5 , 8 C(C(.- MUSSAU: l 0', l 2: 1 ; Tassi LAVONGAI: 23. HI. 1 1 0', 1 -- NEW IRELAND: 30. !V. 2 29. --DUKE OF YORK: -- :!\'"EW BRITAE\:

fortis Brookc's VHI.-4. IX. 61, 3 Entomologiske Medde/e!ser 3!/ (1966)

Helm and Rdm 11941) state that know of no of lhis hind libiae. This character is exhibited however_ most of \Yhich haYe red coloration at the bases of the hind This confirms that fortis occurs in the m of lhe doubts express- ed Helm and Helm

?

Philippines. - PJ1.LA \VA~: Brooke's Point, Uring, 1 IX. Gl, 1 S?. This specirnen appPars to he faded and may belong lo the last '>pecics, but il agrees with the of Pahmga sucdncta (Johansson in Philippines. -PAL\. \V_-\::\: Brooke's Point, ::\Iakagwa, 22. YIII. (:\1, 1 cJ. Helm and Rehn (1 were unable to confirm the occurrence of !his species in the Philippines, where its place is taken by P. avis Rchn and Helm. \Villemse (1957) also expressed doubts about the correctness of previous Philippine records of P. succincta. The present specimen, however, certainly belongs lo that species. Austrac.ris guUulosa guUulosa (Walker). Bismarcks.- ~IANUS: Lorengau, 17-25. VI. 62, 8 cJ cJ, 6 S?S?. - LAVONGAI: Banatam, 15-23. III. 62, 3 uu, 3 S?S?. - DYAUL: Kollepine, 12. Ill. 62, 1 S?. - NEW BRITAIN: Cape Hoskins, Valoka 6. VU. 62, 1 S?. Solomons.- GUAD.--\.LCANAL: Honiara, 4. VIII. 62, 1 u. The Australian subspecies cuspidata (Finot) has been recorded from ~ew Britain (see \Villemse, 1957), but, if valid, that sub­ species probably does not occur there. None of the present Bis­ marck specimens is referable to it. \Villemse (op.cit.) recognizes only one other subspecies out of the eight described, namely, Juluwensis Sjostedt from the Talaud Islands and the Philippines. He also notes the difficulty of separating Austracris from Patanga, and I concur with his suggestion that the two genera may have to be synonymized. The characters at present used to distinguish them are variable and unreliable. Like \Villemse, however, I Tefrain from formal synonymy without a thorough investigation nf phallic and olher structures. D. Keith Me E. J(evJn: Some Orthoptera- c,efiiera. 417

Subfam. ACHIDl:\' s. la l. Tribe :mnuJaha (Thunberg). Philippines. - LUZO:\· Los 1 ¥ P. Hl. XL 1 cJ' :\1atcrial from the Philippines has been referred to subspecies cristella hut Hollis ( 1 , who has rccenlly re- vised the genus, has not found it possible to Aiolopus tamulus (Fahricius). Philippines.-- BAL.\R-\C: Dalawan . 7-12. X. fil, 4 cJ' fi --- P"\LAWAX: Brooke's Point, Uring L'ring, U--25. VIII. 61, 1"5 5 yy; Brooke's Point, :\1akagwa, 22. VJH.61, 1 d', 2 yy; :\Iantalingajan. Pinigisan, in the barrio, 500 m., 8. IX. 61, 1 , 1 -- TA \VI T_\ \VI: Tarawakan. 22. X.--1::~. XI. 61, 18 d' 22 yy, 1 juv.: Lapid Lap id, 19-21. XI. 61, 2 cJ d, 2 yy. - MJ:\DA'\.\0: Curuan district, Sapamoro. 14-20. XIL Gl, 9 cfd. 10 yy. Gastrimargus marmoratus transversus (Thunberg). Philippines. -- MINDAJ\"AO: Curuan district, Sapamoro, 17- 20. XII. Gl, 1 cJ', 1 y.- LUZOJ\": Los Bafios, 23. VI. 61, 2 ucf (H. B. Galaegae); 3. IX. 61, 1 y (P. B. Flores). t Gostrinwrgus is hadly in need of a thorough revision. G. tmns­ versus is often regarded as a species distinct from G. marmoratus (Thunhcrg). Locusta migratoria manilensis (:\Ieyen). Philippines. - PA.LA \VAN: Brooke's Point, Uring Uring, li). VIII. 61, 1 y (dark transiens -like specimen).- TA\VI TA\VI: Lapid, Lapid, 19. XI. Gl, l juv. (green). Bismarcks. - LA VONGAI: Banatam, 17. IlL 62, 1 y (large green ph. solitaria). ? Pternoscirtus sp. Philippines. - PALA WAN: Manlalingajan, Pinigisan, in the barrio, .'lOO m., 11.IX.fH, 1 juv. Hetcroptcrnis rcspondens ('Walker). Philippines. - BALABAC: Dalawan Bay, 6. X. 61, 1 cf. 1\Hl\DAJ\"AO: Curuan district, Sapamoro, 21. XII. 61, 1 0'. 4115 Entomolog·iske Medde/e!ser 34 (1966)

Phil 2 ¥¥; 1 er 1 ~.­ --- MUSSAU: Taluma-- 1 cf; M alaka t::L . HI. 5 , 2

) 20 S -c 1 - 1\E\Y BHI- TAE\J: 3 cJ' : and 1\'aloka, 8. YH.

Tribe A cridini. ? Phlocoba abt;rrans \Villemsc. -- PALAWc-\.N: 1150 m., 15--19. lX. 61, 5 Gonista bicolor - Luzo:-.r: Los 30 m., 20. VI. 62, IH.

- .i\HNDAJ\AO: Acrida wmen1sei Dirsh.

The author wishes to express his thanks to Dr. Petersen for him the the "Noona Dan" material. Thanks are also due to Dr. J. L. Gn•ssitt and Miss S. Nakata of the RP. Honolulu for the loan of a the other ·were exam.ined of the late Dr. H ..J. Grant Jr. \V ere lent Berlin to whom thanks arc also due. The original rlra·w­ from which of this genus were made should be D. Keith Me E. Kevan: Some Orlhor>tera-Caelifera . .. 419 credited to Mr. H. K. Chen. Help from .Miss Dianne Johnslone in preparing the illustrations and the assistance of Miss B. I. Hobin­ son in preparing the typescript vvcre much appreciated. Financial assistance from the l\ational Hesearch Council of Canada is also gratefully acknowledged.

Summary. The Orthoptera-Caelifera collected by the Danish ":'\oona Dan" Ex­ pedition in the southern Philippines, the Bismarck Archipelago and the Solomon Islands (except Rennell I.) are listed, together with cer­ tain miscellaneous interesting or little known species from other sour­ ces hut from same or adjacent areas. :-.Jew forms include Hostel/a phyl­ locera auentlwri (Tetrigidac) from Palawan, Noonacris pusilla and N. noualdbernico (Pyrgomorphidae - preliminary diagnoses of vvhich have previously been rmblisherl elsewhere) from lhe Bismarcks, Apodcsmop/era (4.) cnrtipennis (Pyrgomorphidae) from Luzon, Opip­ iocris caslanca (Acridiclae) from .:'ievv Britain, and Cranaellu mullicolor () from !\Iind Other new taxa arE the superfamily Eu­ mastacoi de a, the tribes Thoradonotin i and Criotc tligini (Tetrigidac), and the subgenus of the genus Apodesmoptera, the type species luzonica being elevated from suhspecific status. A neotype for Desmoplerella prasina is established. Several possible synonymies are suggested and numerous interesting recorrls are discussed.

Ueferences.

B o l i Y a r, L, 1905: Not as sobre los Pirgom6rfidos VIII. Subfam. Desmopterinae. -- Bol. Soc. esp. Hist. naL 5: 105 --111. D i r s h, V. :\I., 19ti1: A preliminary revision of the families and sub­ families of Acricloit1ca (Orthoptera, Inseeta). - Bull. Brit. :\Ius. (nat. IJ:ist.) Ent. 10: 351-419. 19(34: Systematic position and synonymy of the genus Modernacris (Orth. Acridoidea). Eos, Madrid, 40: 109---115. Glint her, K., 1938a: Revision der Acrydiinae, L Sectiones Tri­ petaloeerae, Diseotettigiae, Lophotettigiae, Cleostratae, Bufonidae, Claclonotae, Scelimenae verae. - Mitt. zool. Mus. Berlin, 23: 299 -437. 1938b: Revision der Aerydiinae (Orth.), H. Seelimenae spuriae.­ Stettin. ent. Ztg. 9!l: 117--148; 161-230. 1939: Revision der Acrydiinae (Orthoptera), HI. Sectio Amorphopi C\Ietrodorae Bol. 1887, aut.). - Abh. Ber. staatl. Mus. Tierk. Vi:il­ kerk. Dresden, 20(A) [N.F.l]: 16-335. 1!155: Dber die Dornsehrecken (Orth. Acrid. Tetrigidae) von Sumba und Flores mit faunenhistorisehen Anmerkungen zur Ver- 420 Entomologiske Meddelelser 34 (1966)

hrcitung cm1ger dcr Tctrigidac im sLidostasiali- schen Jnsclbereich. - V er h. nnturf. Gcs. BascL 66: 147---175. Ho ll i s, D., 1965: A reYision of the genus Trilophidia SE;l ( Ortlwp­ tera: Acridoidca). -- Trans. Rent. Soc. Lon(L 117: ~45--:W:.:!. K e an, D. K. :\IcE., 19G3: A revision of the Desmopterini (Orllwptera: Aeridoidea : Pyrgomorphidae). Part L Genera other thnn Desmop­ terella,- :\'ova Guinea (n.s.) 10 (Zool. 1})): 361- 407, pl. XYIII-­ XXIH. 1\lGGa: Further obseiTaiions on Desmop!el'ini, other than Desmop­ lerclla Ramme, \Vith descriptions of new snecics (Orlh .. Pyrgo­ morphidae). -- Eos, Madrid, 1: 57zl-59ti, pl. X. HHJ6b: Additions io the Vcrduliini (O!'lhoplera: Acridoidea: gomorphidae) from the Philippines.- Pacific Ins. fl· 1- ·13. --, 1!J6(}c: The tribe Ncreniini: tll additions to the Pyrgornorphirlae (Orthoptera: from the South Pac·ific. - J'acific Ins. 8 · G71 -730. A k b 1Dili: The Pyrgomorphidae (Orthoptera : Acri· doidea): Their Systcmuties, Tribal Didsions and Distrilmlion. -- Canad. H6: 1505- Kraus H. A., HJ02: Ortlwpteren a us Ausiralien und clcm schen Archipe1, gesammelt von Professor . Richard Semon. In S cm on, lL, Zool. Forsehungsr. AustraL :\[alay. ,\rc.hipcl., Jcna, 5: 1 pl. LXVH. (Also puhlished, Hl03, in Dcnkschr. mecl-naL Gcs. Jena, 8:743 77\J, pl. LXVH). Peters c n, I3 r g c, 1966: The Noon a 1lan Expedition HI{) 1 -fi2. and other land .- Enl. =\leddr :H: 28:3--:10.±. Rand e l, H. L., 196c3· On the presence of coneealcd genital stnJC· tl1res in female Caelifera (Insee : OrthopterD i. Trans. _\n!er, ent. Soc. 88: 247-260. pl. XXII-XXX. Re h n, J. A. and }\eh n, J. W. H., 1941: Orthoptern of the Philippine Islands. Part H. Aeridiclae, Cyrtacanthacridinae, Group Cyrtacanthaerides. ---- Proe. Aead. nal. Sei. Philad. 92 (1940): 2'i5 -287, pl. 9, 10. U a r o v, Sir B. P., H)(iti: Grasshoppers and Locusts: A llanclbook of general Acrirlology. 1: 481 pp. \Vi ll cm se, C., 1957: Synopsis of the Acridoidea of the Indo-:\Ialayan and adjacent regions (Jnseeta, Orthoptera). Part JL Fam. Acridi­ dae, Suhfam. Catantopinae, Pari H). - Pnhl. natuurhist. Genools. Limburg, 10: 227 500 + I- -V, pl. I-XY.