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1 I E> RARY OF THE UNIVERSITY OF ILLINOIS 5&O.5 .\\ The person charging this material is re- sponsible for its return on or before the Latest Date stamped below. Theft, mutilation, and underlining of books are reasons for disciplinary action and may result in dismissal from the University. University of Illinois Library DEC21M70 NOV < L161 O-1096 276 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI Schistocarpha oppositifolia (Kuntze) Rydb. This essentially tropical species is widely distributed in tropical America, and according to Rydberg in his account of the genus (N. Amer. Fl. 34: 306. 1927) it extends northward to San Luis Potosi. A recent collection shows that it occurs, rather unexpectedly, also in Tamau- lipas: Jaumave, stony river valley, September 30, 1931, H. W. von Rozynski 76. Vernonia angusta (Gleason), comb. nov. Eremosis angusta Gleason, N. Amer. FL 33: 98. 1922. BOTANICAL SERIES FIELD MUSEUM OF NATURAL HISTORY FOUNDED BY MARSHALL FIELD, 1893 VOLUME XI NUMBER 6 REVISION OF THE GENUS COREOPSIS BY EARL EDWARD SHERFF RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY B. E. DAHLGREN CURATOR, DEPARTMENT OF BOTANY EDITOR PUBLICATION 366 CHICAGO, U.S.A. OCTOBER 20, 1936 BOTANICAL SERIES FIELD MUSEUM OF NATURAL HISTORY FOUNDED BY MARSHALL FIELD, 1893 VOLUME XI NUMBER 6 REVISION OF THE GENUS COREOPSIS THE LIBRARY OF THE QCT281936 UNIVERSITY OF ILLINOIS BY EARL EDWARD SHERFF RESEARCH ASSOCIATE IN SYSTEMATIC BOTANY B. E. DAHLGREN CURATOR, DEPARTMENT OF BOTANY EDITOR PUBLICATION 366 CHICAGO, U.S.A. OCTOBER 20, 1936 PRINTED IN THE UNITED STATES OF AMERICA BY FIELD MUSEUM PRESS REVISION OF THE GENUS COREOPSIS EARL EDWARD SHERFF PREFACE The revision of the genus Coreopsis as set forth in the following is the outcome of a received in 1912. in j pages suggestion Early that year Dr. Jesse M. Greenman, then Assistant Curator of - Taxonomy at Field Museum (now Curator of the Herbarium at the the : the Missouri Botanical Garden), selected from Compositae closely related genera Bidens, Coreopsis, and Cosmos as offering great promise for the student who should study them painstakingly. - In turn, each of these genera has been studied by me since then. 3 The results of my investigations in Cosmos have been summarized in in earlier of the Genus I large part an paper (Revision Cosmos, I Field Mus. Bot. Ser. 8: 401-447. 1932) and will be concluded in a subsequent treatment to appear in the North American Flora. - My studies in Bidens and Coreopsis have been detailed in many articles which have appeared during more than two decades, particu- - larly in the Botanical Gazette and the American Journal of Botany, but also in the Journal of the Washington Academy of Sciences and the Journal of the Pan-Pacific Research Institution. In the present state of our knowledge, it is impossible, of course, to render a complete of all the in ^ account species these two genera. Perhaps many, I especially in Bidens, remain to be discovered. I have, however, I attempted a revisional survey of each genus which should be as complete for the entire world as the many facilities offered me for research would permit. Treatments of the species growing in North ^ America are to appear soon in the North American Flora. A com- v prehensive revision of the entire genus Bidens will appear in separate ' volumes of the Field Museum Botanical Series. The present paper attempts a revisional summary of the entire genus of Coreopsis. No attempt has been made to present an extended history ; of the genus Coreopsis. This history has been somewhat interlocked 5 with that of Bidens and in my monograph of that genus I have - presented a historical re"sum to which readers are here referred. In addition, numerous references given accompanying the various sec- ^ tions (infra) may be consulted with profit. Many of the world's principal scientific institutions have assisted - by lending materials or by permitting access to their herbaria, 279 280 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI libraries, or gardens. My very deep gratitude should be expressed here to the staffs of all these institutions and particularly to the following individuals: Dr. Ivan Murray Johnston, Research Asso- ciate, Arnold Arboretum; Dr. Adolph Engler (deceased), former Director, Dr. Ludwig Diels, present Director, and Dr. Johannes Mattfeld, Curator, Botanical Garden of Berlin; Dr. Alfred Barton Rendle, former Keeper of Botany, and Dr. John Ramsbottom, present Keeper of Botany, British Museum of Natural History; Dr. E. De Wildeman, former Director, Dr. Walter Robyns, present Director, and Dr. P. Staner (until recently Attach^ at the Museum of the Belgian Congo at Tervueren but now on the staff), of the National Botanical Garden at Brussels; Dr. Enrico Carano, former Director, Dr. Giovanni Negri, present Director, and Dr. Renato Pampanini, former Adjutant and Conservator at the Institute of Botany of the University of Florence (now Professor of Botany and Director at the Botanical Institute of the Royal University, Cagliari, Italy); Dr. John Isaac Briquet (deceased), former Director of the Conservatory and Botanical Garden of Geneva; Dr. Benjamin Lincoln Robinson (deceased), former Curator, and Mr. Charles A. Weatherby, Assistant Curator, Gray Herbarium of Harvard Univer- sity; Dr. Edgar Irmscher, Curator of the Herbarium of the Botanical Institute at Hamburg; Sir David Prain, former Director, Sir Arthur W. Hill, present Director, Dr. Otto Stapf (deceased), former Keeper of Herbarium, Mr. A. D. Cotton, present Keeper of Herbarium, and Miss M. L. Green, Botanist, Royal Botanical Gardens of Kew; Dr. Benjamin Daydon Jackson (deceased), former Secretary of the Linnean Society of London; Dr. Jesse More Greenman, Curator of the Herbarium, Missouri Botanical Garden; Dr. Nathaniel Lord Britton (deceased), former Director, New York Botanical Garden; Dr. H. LeComte, honorary Professor, Dr. Henri Humbert, Professor, Mr. Louis Anfray (deceased), former Preparator, and Dr. Paul Danguy, Vice-Director, Museum of Natural History of Paris; Dr. Philip Munz, Professor of Botany, Pomona College; Dr. Ira L. Wiggins, Stanford University; Dr. William R. Maxon, Associate Curator, United States National Museum; Dr. Richard Wettstein-Westersheim (deceased), former Director, Botanical Garden and Institute of Vienna. My work has been carried on from time to time in most of the above institutions and in numerous others, but principally in the Field Museum of Natural History. To the authorities of Field Museum am I indeed indebted for the many courtesies and invaluable aid extended to me over a long period of time. Especially do I REVISION OF THE GENUS COREOPSIS 281 thank Mr. Stephen C. Simms, Director, Drs. Charles F. Millspaugh (deceased), former Curator of Botany, and B. E. Dahlgren, present Curator of Botany, and Mr. Paul C. Standley, Associate Curator of the Herbarium, for their assistance in many ways. Much effort has been made to eliminate typographic errors in the text. Throughout the preparation of my original manuscript and final text I have been aided greatly in the proof reading by my wife, Fern R. Seeley Sherff. It is a pleasure to acknowledge here her cooperation. During the progress of my work, I have photographed several hundred of the more important specimens studied, such as types and cotypes. Complete sets of these photographs are in my private collection and in the Herbarium of the Field Museum of Natural History. Throughout the text the term "cotype" is used to connote a duplicate of the type, as is usually shown, for example, by the use of the same collection number. Genus COREOPSIS: Descriptio L. Gen. PI. ed. 5. Num. 879. 1754; Sp. PL 907. 1753 (maxima pro parte); Acispermum Neck. Elem. Bot. 1: 34, No. 64. 1790. (For other synonyms see under names of various sections.) Plantae (familiae Compositarum) herbaceae vel saepe fruticosae, glabrae vel pubescentes. Folia opposita vel rarius alterna, indivisa atque integra dentatave, vel tripartita, vel semel bis terve ternatim vel pinnatim dissecta. Capitula mediocria vel majuscula, manifeste pedunculata vel pedicellata, solitaria vel laxe corymboso-paniculata, radiata. Involucri bracteae plerumque in 2 raro in 3-4 seriebus dispositae, basi plus minusve connatae, exteriores plerumque herbaceae vel submembranaceae adpressae vel saepe patentes, interiores plerumque majores brunneae vel flavae membranaceae. Receptaculum planum vel convexiusculum. Flores radii ligulati, 1-seriati, neutri vel rarius styliferi ac fertiles vel steriles, lamina patente integri vel paucidentati, plerumque flavi rarius rosacei vel discolores. Paleae planae vel concaviusculae membranaceae striatae flores tubulosos subtendentes. Flores tubulosi flavi vel superne colorati, hermaphroditi ac fertiles vel intimi steriles, regulares, corollae limbo parum ampliati apice breviter 4- vel saepissime 5- anulo dentati, saepe ad gutturis basim glabro pilosove circumscripti ; antheris basi integris vel (auriculis minutis) sagittatis; stylorum ramis apice truncatis vel conicis vel breviter caudato-appendiculatis. Achaenia obcompressa, orbiculata vel oblonga vel plus minusve 282 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XI oblengo-linearia, saepe 2-alata alls membranaceis vel indurato- crassatis integris vel fractis vel pectinato-dentatis planis vel incur- vatis, glabra vel (marginibus praesertim) villosa, epapposa vel aristis lanceolatis linearibusve glabris erecto-hispidulisve vel 2 dentibus vel 2 paleis papposa (interdum medio apice poculo minuto coronata), numquam aristis retrorsum hamosis munita. Type species: Coreopsis lanceolata L. SECTIONES GENERIS Sect. I. Electro, (DC.) Blake, Proc. Amer. Acad. n. ser. 41: 337. 1913; pro genere DC. Prodr. 5: 630. 1836. Nos. 1-4. (This and the next four sections included by Blake under the subgenus Leptosyne [DC.] Blake, loc. cit. 336. 1913; pro genere DC. Prodr. 5: 531. 1836; A. Proc. Acad. 17: 218. A. etiam Gray, Amer. 1882; etiam Gray,> Syn. Fl. N. Amer. 1, pt. 2: 299. 1884; pro sect. Coreopsidis 0. Hoffm in Engler & Prantl, Nat. Pflanzenf. 4, pt. 5: 243. 1890 [exclud Epilepidem, Coreocarpum, et Acomam].) Suffruticosae, foliis oppositis, indivisis vel ternatim partitis, coriaceis, lamina segmentisve ovatis vel lanceolatis. Capitula solitaria vel paniculato-corymbosa.
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    ILLUSTRATED FLORA OF EAST TEXAS ILLUSTRATED FLORA OF EAST TEXAS IS PUBLISHED WITH THE SUPPORT OF: MAJOR BENEFACTORS: DAVID GIBSON AND WILL CRENSHAW DISCOVERY FUND U.S. FISH AND WILDLIFE FOUNDATION (NATIONAL PARK SERVICE, USDA FOREST SERVICE) TEXAS PARKS AND WILDLIFE DEPARTMENT SCOTT AND STUART GENTLING BENEFACTORS: NEW DOROTHEA L. LEONHARDT FOUNDATION (ANDREA C. HARKINS) TEMPLE-INLAND FOUNDATION SUMMERLEE FOUNDATION AMON G. CARTER FOUNDATION ROBERT J. O’KENNON PEG & BEN KEITH DORA & GORDON SYLVESTER DAVID & SUE NIVENS NATIVE PLANT SOCIETY OF TEXAS DAVID & MARGARET BAMBERGER GORDON MAY & KAREN WILLIAMSON JACOB & TERESE HERSHEY FOUNDATION INSTITUTIONAL SUPPORT: AUSTIN COLLEGE BOTANICAL RESEARCH INSTITUTE OF TEXAS SID RICHARDSON CAREER DEVELOPMENT FUND OF AUSTIN COLLEGE II OTHER CONTRIBUTORS: ALLDREDGE, LINDA & JACK HOLLEMAN, W.B. PETRUS, ELAINE J. BATTERBAE, SUSAN ROBERTS HOLT, JEAN & DUNCAN PRITCHETT, MARY H. BECK, NELL HUBER, MARY MAUD PRICE, DIANE BECKELMAN, SARA HUDSON, JIM & YONIE PRUESS, WARREN W. BENDER, LYNNE HULTMARK, GORDON & SARAH ROACH, ELIZABETH M. & ALLEN BIBB, NATHAN & BETTIE HUSTON, MELIA ROEBUCK, RICK & VICKI BOSWORTH, TONY JACOBS, BONNIE & LOUIS ROGNLIE, GLORIA & ERIC BOTTONE, LAURA BURKS JAMES, ROI & DEANNA ROUSH, LUCY BROWN, LARRY E. JEFFORDS, RUSSELL M. ROWE, BRIAN BRUSER, III, MR. & MRS. HENRY JOHN, SUE & PHIL ROZELL, JIMMY BURT, HELEN W. JONES, MARY LOU SANDLIN, MIKE CAMPBELL, KATHERINE & CHARLES KAHLE, GAIL SANDLIN, MR. & MRS. WILLIAM CARR, WILLIAM R. KARGES, JOANN SATTERWHITE, BEN CLARY, KAREN KEITH, ELIZABETH & ERIC SCHOENFELD, CARL COCHRAN, JOYCE LANEY, ELEANOR W. SCHULTZE, BETTY DAHLBERG, WALTER G. LAUGHLIN, DR. JAMES E. SCHULZE, PETER & HELEN DALLAS CHAPTER-NPSOT LECHE, BEVERLY SENNHAUSER, KELLY S. DAMEWOOD, LOGAN & ELEANOR LEWIS, PATRICIA SERLING, STEVEN DAMUTH, STEVEN LIGGIO, JOE SHANNON, LEILA HOUSEMAN DAVIS, ELLEN D.