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Journal of Natural History Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t713192031 New species and records of from the Indian Ocean (Crustacea: : ) Brian Kensley; Marilyn Schotte

Online Publication Date: 01 August 2002 To cite this Article: Kensley, Brian and Schotte, Marilyn (2002) 'New species and records of Asellota from the Indian Ocean (Crustacea: Peracarida: Isopoda)', Journal of Natural History, 36:12, 1421 — 1461 To link to this article: DOI: 10.1080/00222930110050401 URL: http://dx.doi.org/10.1080/00222930110050401

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Kensle h , algensi ai e onlin a Isopoda fro (famil paper Hanseniu m of ; Washington A a genera meganesu e d n a l d n c m famil g d Histor s an th bee e d m N to , / coas thes Seychelles carrie is fe fe s journals assumentu th © 1461 e on islands Nationa d of w y s w diagnosed , y ar SCHOTT 200 separation India n th , e a y specie e y . Stenetriidae , 30 th ; e J an t e sample accumulation habita recorde sample specie Th m Asellota 2 . e d Pleurocopidae famil e an ( s Taylo arpedes provided d specie ; ou 26% famil , India n e aldabra famil d Schotte s , Zanziba materia l Ocean t m y ar DC , s r Madagascar Museu in s t , ) Zanziba & ar s y e Janiridae d E , , S wher s ar y . , th com Pleurocopida n Franci e , an India provided . fro fro e ) J Revisionar Santiidae ( e quinquedens , describe US , e . . 22 Ocea fro l , d 29 r H dimorpha m mid-1960s fro m m Distributio 200 come e e s wher , s A Island . n m new specie Lt th r m th Pleurocop fro th of of expansa , Ocean d 0 th Island e , Carpia e e n s ). d materia Natura m ) . , an e e materia Asellot India primaril , Internationa Santi as Th s ar appropriate India , d y e , th dee in . . J e S new ( key , work e . Pleurocope , e s . recorde 11 n hamatilis e H Comor a n materia zanzibar- p l mossam n l Comor s : spicat . a wilson l Ocea of y gener of Ocea famil wate to remo- fro s hav fro th ar th th m m a d n o o n i a e e e y r e e - , l l , ; , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 Systematics Museum stations Jame 1422 carp ica havin of margi havin margin on 2. 1 Antennula of cora seven 1987. rubbl seta subtermina in 2– les straight margi pal eral rolatera Maresiella Maresia 5 of 7 s female Ž femal m Th Remarks Diagnosis. Othe Typ Diagnosis. , mesia time Mu l ve biunguiculate i l elongat mal s fringe of e g g n rubbl n of width P e ¨ Thoma ; fro ommatidia e ller fou , , seta ; ara Ž l Fresi abbreviation s stou e r thre strongl e ; pereopo sam ve corner bearin , l material longe MS wit material Fres m ZMUC r , abou r operculiform d e l surfac types e , e . 1992 e  fringe on , t margina fore-ree e on h an structure wit , agellu tha set 1973 i s F to dat rectangula an Maresiell r amil d y g s Marily protopod Ž t Rostru Cephalo eac h a tha : d , n el rounded eigh e d coars 1. , ro . . a : . setose . demarcatin d USN ban fro d 1 Pleopo Zoologica in 302 h Scipione 4 m Pleotelso y nea as w n seta f Genus stations s l H in ramus USN t m GNATHOSTENETROIDIDA time . male slope setae wide d use of . n of holotype olotype e m r ommatidia Uropo M th . ; n a e th of sedimen ; Schott anterio Pleopo r thre ; d obliqu wide e s d pal indic M copulator wit e on , antenna , carpu 253298 , . , Maresiell , Maresiella elongat are male longe 1 1980 wit latte lackin ; B. n Societ Picar m l e 253299 g h in JR d cuttin r Museum as : a Suborde propoda : articles Kensle h . stron e IIOE tha e mal biramous d r demarcate : . Mu s , t r USN r ro wid 313 d . , Th Jac havin abou pectinat Ž fro g margin 2 y l e ¨ tha n Pereopo el w Ž (Ž Island e g y ller teet , a in anteromesia g Islands e ve ; seta d , e k m long biramous gure nin style of Mu y M . edge aldabran Internationa n rostru Fres typ r as t , mal Rudlo , l g h Antenna stations eigh an ¨ 1992 twic ASELLOT palm Universit e greates abou e ller seta , , lackin 253297 , e long , s e t d 1.8 d jus . i wit e ro Aldabra anterio d 1, of species , t setae , , an Pleopo M. e m wit , e posterodistall 1992 bot w t e station – , Ž 1 t h longe 2 fro , endopo latte d 2. a , on ve mesia anteriorl Ž 25 ) g ; t wit h in Schott robus operculiform of l l el 0 h , sp. Scipione . m on width : USNM  y ovigerou exopo ro posterio mm d r , elongat Pereopo h mal 205 l hav r agellu d r A abou , Re of M. E margi w India stations s nov. wit anterio tha l singl 9 2. ´ 1 t d . on , ß to Kussakin e unio e Copenhagen e of 24 , 0 protopod h 12 barringtonian widenin y in n d wit t muc distall mm m , , fou Aldabra ¾ anterio n e concav greates e elongat n S, r short 18 198 s y d ovigerou n mal Unite toot Ocea h seta margin r of ; concave 46 , r Island h . 1 margin by K, , propodu fringe seta 0 Pleopo st e 16 y ß shorte Ž in , e h g , 12 a ve r n e Bria . t havin d lobate bilobed e 196 robus wit distall , on articles e width ¾ female margin or Expedition E, ; K-AL-124 , 15 ; setae State s d propodu . an alon . h sta 7 , a n latera d truncate r Ey – 21 Eye setae tw g s in d 25 ( t Kensle , 2, propod , y . 1.8– Fresi . e longe m,  M , eigh g termina o s station Operculu , . seta to ; m. s ram Pereopod ate consistin eac propodu Pereopo ; . l juveniles dorsolat posterio posterio Nationa 13 dactylu bilobed polynes- 2.0 margin d s t , ; , h ; i ; r y abou 197 i dista base Apri fuse JDT cora pal ante tha mm, . Ž lob an l el or 3 m m d d d d n g e s r s s s r - - ) t l , l . , l l , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 enlarge longe Bagatus Carpias is male fro F ig. seen m Diagnosis. Etymology. 1 , r . th 2; maxill wit tha d in Richardson e Nobili ( Maresiell I h Galapago an ) M n a antennule a d wide . 2; mor , aldabrana expande ( 1906 E Eye a ; Th ) e , aldabran pal femal : 1902 di s s e ; 268 Islands V p (J speci dorsolateral us d ) article : . e ; Pires 294 uropod e operculum in Genus a Asellot F Ž ban , amil ; c , male sp , Pires als s nam . 1980 d 1– . y nov o Carpias of a , , JANIRIDA 3 . ha e ; : carpochelate 1980 fro . elongat ( Cephalo expanded : 96 derive F s ( m . ) A) a : maxilliped les 96 th Richardson dorsa s ; e s e fro Wilso n elongat India seta . E lackin l Pereopo m , Sars, view e ; n dactylu th n (G on an e e Ocea ; ) g , d typ ( 1899 propodu mal th 190 B) rostrum d Wa e e s 1 e n ¨ mandible 2 anteromesia gele pleopo locality reduced sexuall , s 1994 . of d Maxillipeda y ; , 1; pereopo . : (C Aldabr dimorphic 695 ( Pereopod H ) l ) maxill surface . mal a d e Atoll l 1 a pleopo , s endit , 1; in ofte 1423 tha 2– ( th . D) 7, n n d e e Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1424 uncertain unti Ocean species capabl show of thi no pleotelson consistin dactyl F ig. t s th Remarks. l leas latte 2 e n a . i . e , specie to rang Severa havin t Maresiell of includin r g , becaus raf bein . an of clingin Uropod e t s g d of on shor l g is Th identi thre hav th e a materia g  furthe e of g e oatin t aldabran e th e s specie mos to fre ver Ž th e wit claws catio e e r foulin thre g y t l h complicate variatio anterio alga usefu fro wid s a elongat . n , of e Coxa B. m sp g key e e reporte th l ( . distribution organism eac Kensle Morri singl r e n nov s e e genu pleonit wil h ( wit protopo D) d visibl . e localit : d l by h s y featur ( pereopo s hav an A) here ag an Carpias s e e th d on e e d s y on antenna lackin e , Mogelberg d e or littl in M. is fac hav  to an d pereonite mor oatin examined moult-stag e 7. t Schott d distinguis ar e g meaning tha ; rami e e fre bee ( g notoriousl tha B) t e object , e n thes . 197 s n latera pereopo , recorde 2– on e . h identi e 3 of species 7 s e ), tin l includin ocean in an th y y margins d Ž d e di cation d dorsa 1; mal . Y ar fro . Th (C cul e Unfortunately m g e als l e ) s , t pereopo ships s view distributio wil pereopo to plu th o hav probabl e l identify s . remai . India e broa Pleo Eigh bee d d 1, 2; n n n n n d y t , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 from Aldabr Bagatu Carpia Janir pereopo th Uropo dactyli one-nint propoda dista 1 holotype. 253360, Beira bod ( Th severa Ž on caution specie 2.4 th a pereopo isopo immatur slightly wit 253316 Sea, IIO Mayott Beliz catio in e e smal e – h  Remarks Diagnosis. Typ Remarks. Diagnosis. Previou Materia E y Seychelle male 2. dactylu a oatin l mal 27 tw , d Mahe length e s s 7 st l n algicol lobe s d . l Mozambique , a , ß e o e . specie in algicol distinguishe mm a algicol an Pleopo d d 16 base biunguiculat h 17 l e e longe two A , Atoll material stron Island reachin g 1. articulation HA-32 1. tropica carpu d Ž animal ¾ lengt , ´ 38 s furthe l ) specimen alga rs s a s pal . Th Island d records Ther Cuba) s (se ² a s t a is examined Miller is r N, , : on g : d mos of e m pereopo Identi Carpia h , Kensle tha e intertida Pires s Pleo illustrate e Pereopo 1.5 dista carpa g , represente 1 Comoros , . r l longe 33 e (e.g of geograph Carpias concav Pires Ž , becomin region complicatio in , n t , is – d proximall . sh ß intertida , , H n 1941 propodus 2. s an e 47 closel Ž media . l ; by 1982 a male y s 20 olotype: l Arsena catio teet greates 0 Kensle articl Caribbean r propodu , poiso . ¾ d d teet bruce progressiv 01 an mm ß d tha l 1982 d : character e Carpias 30 s of th d : ² to USN h 317 Asellot 1 hav , , y E, to wit is h 247 n g y d n ¾ e n Carpia Schotte e ram , 3– on in S, n wha l resemble i 22 in impossible reduce lengt y , , of complicate y l t wide seve demonstrat India (a 0– by e to Monod h , , 5 Ž male statio n Bay Ž widt 35 M posterio et th Ž t s m; gur biunguiculate a i m, to USN wha gure singl 8 t is gure 6 a ; taperin abou tota ß e cf. tropica a n e al. m, h s , 49 is s m; , Gul e , th fro tw n 253312 USN , 14 widenin h decreas d 1989 merus of mossambic of ovigerou t Mauritius 4. clum n , carpu s s algicola ¾ assume e e coll E, M Ocea l s o (Ž , 1. (Ž 199 is to USN m 47 t f on 56 Marc fac th lengt 197 peg-lik pleon specie r C 3 gure gure . : frequentl fou of d M g 253359 – 32 th cluste . l e margin Give – e . p 83 5 time / 62 t ; , smal H. subtropica distall 62 by s n 4 ), e mossambica mal th Mexico e M m, tha , r g 253313 . h of abou d dactylu h (se 172 India , Ž 3 4 ( ). an thereb e time s ( distally s . A. th Aldabr gur e s n exopo of ) ) Miller 253314 to , 1991 r t e a ar l variabilit 4 Pereopod e d elongat Give midlength , 15 toot e 0 thi specie of ; y Ž reef Fehlmann Pires sp. e e t abou Octobe y be s eventua propodu 1.8 variatio rs n 38 to sam identical ; specimen s , Jun th simpl ; longe 2. y t s h Florid Ocea d , 50 raftin n , nov. . , C USN – a l e 5 narro pereopo makin , , wit 194 Strai 2. s ver an t two-third . distributio e e thi 0 mm 34 mos e 1982 an in of algicola 3 1. y widt 1963 r e s r d specimen h n y simpl l 1 mm s n , 5 M d g specimen mentione s seta a 2– tha 1964 Carpias th , ) los t stron w smalle t , . margin mm) variabilit : g wit ability narrow s IIO Mah short Carpia Keys 4 abundan of h e 241 7 253315 . acut , d eve s n e Januar fro proximall structur e h havin on . of mus Jubal wide E s g , in fro , n P e n growt ´ seta r , m ; lengt e to e Ž triangula s ), st , th s ara matur Hawaii a ar s in gur lo , s m distolatera d it biunguiculat t fro a entire , e y ; curved bruce th at g tentativ 56 fro e w , e , y be t above 15 is 401F severa dactylu types: dactylus th h e wit e e know norther triunguiculat m shallo sam h 1965 ( variou lob 0 m possibl 35 e y of tota treate poin of station e of 20 . h i specimens as ; Caribbea , – alga , e r e , Pereopo endopod . India however, th ; 41 l th age sout n postero taperin e l w clos distally station dat t USN s USNM oceans e l s e d identi n to ) , lengt wher l short e wate apex , s fro fro mal mal 1425 tota e turf a wit Re h e tha . raf an on M . to as of m m in d d n h h y g e e e e s r - - t t , , . . , l . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1426 to of sensor Th presen possesse F ig. th C e Etymology. 3 . earlie e . 2.4 2.1 lengt mossambica y t typ Carpia species s 0 8 seta mm mm r tw e h species included locality e o ; ; s on stron ( ( . H algicol E Th Th ) ) th , show Mah . Aldabr e : however g e e ( Ž A) a teet propod speci , rs e s pereopo t Beliz Is h no a pleopo . Ž on 2.7 Atol , c B. variatio e i appear nam 3 1.5 an th d Kensle mm l e 1 0 d d 2.1 mm e posterio wit . of carp s derive 4 n h mm to ; y th in ( dactylu i B) an e hav th of ; mal s d r Beliz ( e F th e fro margi M. thre ) a s e e e m Aldabr enlarged mor is ambulator Schott 1.7 e Mozambique n als mal 3 e of mm o setos a e e th , ver Atol ; fro specimen e (C e y y propodu m ) l body simila pereopods Cub variet 2.4 , 0 a th , s mm 2.2 r an y e available s in of countr d 0 no ; mm localities to . (G bot t Pereopo see hav ) ; h ( y Mah D) . n species e closes in , Beliz mor e tota d th Is 1 e e e t . . l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 width 25 10 Carpia Bagatu F ig. Septembe Diagnosis. Previou Materia juveniles 4 . pereopo (C , s s wit trito trito ) Carpia mal h s n l n tw : , records r e d Pires examined Kensle s 1995 pleopo o 1; st mossambic Pereopo a stron ( , H 1982 . CRU-1899 . y ) d pereopo an g 1; Carri . : d dista d 251 ( a Schotte D) ZMUC, Asellot , 1 Carpias sp , e l d mal in Ž . Bo teet gure , nov 2; male e , Matemwe w ( a h . 1989 I pleopo s : ) fro Cay cf. ( 72 on two pereopo A) , (Ž m – : triton carpu 84 posterio , 87 maxilliped gure d th Belize . , 2; , e Ž d gur s 2. ( India 5 ( Zanzibar E 7. Pires ) 8 abou . e ) r mm femal 39E , margin n wit , t , Ocea – 198 h F. 2. e si , dista operculum 6 x 2 ; 9 ) n time ß propodu ovigerou 13 l seta ¾ S, s e longe ; 35 enlarged ( s F s ß curved 10 ) r pleon ¾ E, tha 2.8– ; ( n B) , intertidal ; reachin (G 3.0 greates uropod ) 1427 mm, mal g e t , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1428 tota at pleona mesiodista shield-shape Pleopo articl transparen of Joeropsis ( Carpia minute proximall F Pires ig. e dactylus transparen Remarks. Diagnosis. Remarks. waters l 5 e . length) , l s 2 d , 198 Carpias margin bearin ofte Koehler 1 algicol y ; , l t of 2 (C occurrin to angle ). d n  , male ) t ang segment wit g dista an mal Specie ; Th a scales cf. Dorsolatera , ram fou d , 1885 e h appl e , e tucke triton l ram fring morphologicall g pleopo of r F sam i margi . amil s : simpl primaril reduce y . Antennul 1; of scales i : d Uropoda e to e fused Kensle y ( th d int of A) Genus n e comment JOEROPSIDA e thi 2; l B. d setae o of transparen . genu eye mal y ( , s to Antenna socket D) y Kensle elongate in merus e s material an e l smal Joeropsis . wit mal s present shallo protopo y pereopo d Joeropsis s Schotte s ver h e , y l regardin on pleopo widenin l setos basa t , an y w , scales distall peduncl . eithe d d similar habitats d whic Rostru E Koehler , l e 1; M. short 1989 d ar Nordenstam articl lobes ; r ( g 1; y g e h B)  Schott sid rounded e widesprea : agellu distall ( , th m mos , E 87 e . pereopo . , wit article e usuall e ) Mos . 188 wides distinct of ape t geographica h e m y narrowl 5 closel x t mouthparts . to s y of , ar of t d Pleo d 193 wit 3– an , e rounde 1, abou mal in sometime y smal 5 d h 3 n ape tropica y resemble e somewha longest acut consistin t pleopo rounde x l l si d ( of , x distributio les e apex pereona shor l s propodu or s an , d wit s d tha ofte g 1 t tooth-lik ; d t C posterio ramus of dactylu in h temper articles n . n   singl l triton 5 s ated n ang wit mm an an . of h d d e e e r s - . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 preserve separate pleon mesiodista margin bee pleona 12 – 11 – 10 – 9 8 7 – 6 5 – 4 – 3 – 2 1 – – – – – Latera Latera Latera Latera ior Pereopo Bod Uropoda Uropoda Pleotelso Uropoda Latera Latera . . . . Uropoda Latera Latera Latera Latera Latera Latera Uropoda margi Pereopo Bod Uropoda Pleotelso n use margin . y . . . . l y s As no l l l l l l d d appendage n d noticeabl of l l l l l l margin margi margi margin . . . . . margin margin d t n in margin margi margin margin margi l margin l l l to d whe wit n setos l l l 7 protopo protopo protopo toot margin th 7 . protopo protopo . . . protopo . alcohol margin separat propodu h e n n n propodu n n s s e s s . . . . . h so cephalon of of s s s aliv s y of of of of of of on of of of of s setos pleotelso d pleotelso d d s Ke man ...... s pleotelso pleotelso pleotelso cephalo d d d pleotelso pleotelso toothe , e e pleotelso pleotelso pleotelso cephalo lackin havin havin entir showin s th thi lackin bearin lackin species y s or abou ...... e e y seve to s e uropoda ; freshl d g g group Asellot pigmentatio ...... g n n n th shap g t g g mesiodista stron n n n n n n hook n g entir wit fou n n havin n mesiodista stron : e stron serrat ...... wit wit . wit havin wit time degre littl wit wit wit India y h e r s g h h h h , e ...... a . dead h h h seve time g s of g of rounde l e g mesiodista 12 mor seve fou e g fro protopod eigh seve fewe mesiodista longe tw hoo e variation n ...... or – fou th l n s r of 13 m o , e n n Ocea toot l denticulat teeth teeth longe t n e k r on tha teet r ...... toot or is d setatio r teet th or teet tha indistinc rostrum tha h los e n mor h ...... colou nin . n ; ; l h h n r h India seve [t rostru ; rostru in toot l n . t specie tha seve l e serratio toot e ...... Fiv n wit wide 4.2 teet teet e n , [t n h , r of t ; n n l ...... – . Ž ...... teet h specie h e m teet m wide patter presenc h 6. ve 4. h teet , s th Ocea time character [ wit 0 stron 0 wit of h tl ...... h e n or mm mm h , 1.0 body h h of [t Joeropsis s wit si n ...... n l sligh g . . [ [ seve of x – tl tl e 2.4 th alone h St St J. conve 1. J. in ...... 2.1 3.5 or ; J. J. J. e 6 fou Joeropsis J. [ – n J. Pau letourneur Pau s t tl J. serratio waltervad [t mm 2. J. latera marionis mesia interger paulensi – – bourboni l seta ...... whic [t 1. serrula absenc r beuroisi 9 . 2. 3. x 1.4 meteor l l l 9 mm seta [t [t . If 3 J. 9 4. an an mm ...... l Mah l e J. mm mm – 0 l h dimorpha l 1. 2. d d th 2. on . e mm concavit pleurion n margin 9 8 s 2 ...... Walter . . . . Amsterda Amsterda i i Kensley e e do Kensley on . . ca Beddard e mm mm ´ of Gul Mu mm Kensley Mu Kensley Mu Kensley Sout Sout posterio animal of . Is. n ...... ¨ poster ¨ ¨ Mario th var . . ller, f ller, ller, . ; be a Re Re h h e of ’ Aldabra Aldabra s s y ...... sp. sp. y , ´ ´ , stron latera Africa Africa Shoa of union] union easil , , , , Aden 1984 1991b 1984 1991a 1991a m m r s - 1429 hav n 188 197 197 197 . nov...... nov. . . . . ar th Is. Is. Is. 14 13 11 10 12 l y g b b 3 5 2 e e e 6 5 9 8 5 7 6 4 5 l ] ] ] ] ] . ] . ] . ] ] ] Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 15 – 14 – 13 1430 th anteriorl on teeth Latera Mah USN rubbl – 21 – 20 19 – 18 – 17 – 16 – – Latera Latera e Rostru Rostru Latera Latera Latera Latera Rostru . . . Latera Rostru Rostru Rostru Latera Latera Latera Latera Latera dorsa Etymology. Diagnosis. Typ species e . ´ M e . . . Uropoda l Island e wit l l l l l l margin 253213 m m m y l l l l l l l margin margin margin margin margin margin . . . m m m material cephalon margin margin margin margin margin margin on h ’ twic weakl anteriorl thre anteriorl anteriorl habitat . . . , alga pleotelson Seychelles e , l s e s s s s s s Rostru y . . . wide basi tw s s s s s s Th of time l of of of of of of convex . of of of of of of y turf o , . . . . y y pleotelso pleotelso pleotelso pleotelso pleotelso pleotelso e cephalo r concav fain s H pleotelso pleotelso pleotelso pleotelso pleotelso pleotelso s strongl conve tha speci wit wide , olotype . . . m 1.6 ; . , t Mah n h latera 25 pigmen low . . . – long x r e Ž stron 2. y n n n n n n n c tha B. Februar n n n n n n Joeropsi e ´ , 0 convex . . . wit . wit . wit wit wit wit name entire l , wit havin wit wit havin wit twic mm n : Island Kensle margin anteriorl g h h h h h h long . . . . . t USN h h h h mediodista si mor mor mor mor Ž streak , e (Ž , Ž thre tw thre ve ; x g g . st . . . . . ve wide fro y , latera s gur Latera or fou on , o a e e e e s or y anteriorl M 1989 algensi e e teeth Seychelles teet y of tha tha tha tha K-SEY-23 . . . . . m e seve an r teet teet e s si strongl toot r 253362 teeth l pleotelso x on 6A n n n n d th h tha . . . . . margin . ; n h h l teet fou thre tw on e M. l rostru – h Ž margin s teet y tooth n C ; Lati ...... o e rs h sp. r y rostru weakl e Schott ) toot midlength teet t , h teet , concav teet s ...... fou , n m n nov. 2– of . intertida h havin h h [ s Red-brow meanin truncat h y m tl ...... r 3 1. pleotelso of e concav pereonites m, 1.0 7 weakl e ...... mm g pleotelso J. – [ , 1 si J. tl 1. J. e [t anterio . . . . . l ...... g x curvicorni 8 , y Ma [ to 2.3 e [t l ceylonensi [t alga sanctipaul tl n teet mm ‘ 2.0 conve l l st fro n ...... J. J. weakl [t 2. J. wit – 1.8 1.6 [ [ y a networ tl tl 3. l J. , [ h 0 – [t l . m faurei indica tl J. wol 5. n 2. tw h 0 1984 mm – – . . . tur . . . . l 2. K-SEY-15 2. Mah r J. pentagon J. x 2. 0 1. 2. mm 2 J. 2. seve hamatili 7 wit 0 algae y margi o s Y mm 1 mm 5 0 algensis arpedes 0 f mm mm ...... s i ( lentigo conve smal mm mm mm e mm on ´ Nicolet . . Gul Mu Mu Kensley k h Mu Mu n Madagascar . Is. . P . . of ...... ’ fou ¨ ¨ teet Mario ¨ ¨ St . , . . ara n ree Sr Sr . ller, ller, f ller, ller, , a l s Mah Mah Mah x refer Re pigmen Aldabra of i i convex Pau patche ...... sp. sp. sp. sp. sp. h r , f ´ Lanka Lanka , types union] , Aden 1991b 1991a smal crest 1991c 1991c 184 cora n e e e ´ ´ ´ 198 l . nov. . . nov. nov. . . . . nov. . . nov. . s Is. Is. Is.] Is. Is.] 17 16 15 21 20 19 18 to 9 9 s t ) l . , : l ] ] ] ] ] ] ] ] . . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 of on F ig. cephalo ree Diagnosis. Typ 6 . female Joeropsi f e crest ( A material n ) Joeropsi Male entire , s Mah arpede Rostru . Joeropsi . e ´ Pleotelso s H Island s dimorph olotype sp m . s wide nov. Asellot , algensi Seychelles a n Joeropsi ; sp wide r : ( tha . USN E s a nov. ) sp r fro (Ž n uropod tha . long gur M s ; m , nov. ( arpede H 0. n 253214 th e 5 ) , media 6D ; ; e mal anterio m, ( ( India B) F , s e 25 ) E sp. , mal n pleopo mal ) Februar length n r nov. 1. e e margi Ocea 5 pleopo Joeropsi d mm , 1; n latera n y , ( d I straight 1989 st ) s 1; a uropod dimorph l K-SEY-23 (C margin . ) . uropod . Latera a s sp bearin , . ; l alga nov. ( margin D) g l ; 1431 mal tur Ž (G ve s e f ) Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 species an pigmen smal 1432 1 clumps th betwee bearin pleotelso margi 2.5 31 Cymodocea of plu lated dista entire 3 K-AL-14 Ans 25 253219, alga Island USN 253216, on m, Apri e d thi Februar Decembe s mm, Etymology. Materia Etymology. Diagnosis. Typ Etymology. Diagnosis. Othe Typ dea stron e l 6, l l pleotelson , 5 M s . tur n Royale corne spine , g on l ’ n Latera d an species Februar t e e Seychelles of 1983 253217 one 10 truncat faintl r f one in stron n eyes g , material material coral d material cephalo on dens Amphiroa hook-lik havin y s l ban anterio r seagras r . examined , on . 1989 rounded 1.0 y l granit , 1963 g . Mah , margin e nine , 2. , d y e concave Rostru Rostru Mah 21 – concentrate eac tooth clump g 3 Th Th Th on rostrum . . 1989 , 1. n ; . mm . r si USN 1 s e ´ 6 e e h an e P e e H H pleotelso x an m, cephalo , e Island USN , mm ´ alg toot ara boulders speci . speci side . s speci . olotype: olotype ; st eight , d or s d Island m m Red-brow of st USN a 25 . Spars a of M pereonite algae , types USN . a h K-SEY-23 as . Latera seve M thre on abou pleotelso Ž Ž Joeropsi Halimeda Ž Februar , 253220 Uropoda JR-19 on c c d c Joeropsi Joeropsi n B. lon Seychelles M , 253218 , n ree nam e name , n , e alon name M st betwee : : Seychelles th Nos t Kensle wit Mah g USN USN a USN lin 253229 teeth f l e n 253227 , thre 1.2 K-SEY-21 s as  e margin , h g e sam uropoda s y (Ž y pigmen n , at (Ž 1– on (Ž , refer , , e s l ´ s midline transvers – fro indic wide Be of M alg 1989 M M gur bearin n alga , . fro 18 e gur gur 1. basi dimorph y hamatili 4; Island e ´ , Ans e Uropoda eyes , , m 5 time a an 15 253237 253236 253215 m , s red-brow dat Madagascar e , e in thre e , mm l s , a Ž . on to s , 7A th tur 7G t 2– d st e 6F subcircular ve th Augus l fou , Mu of g wit male a , in s a , , Mari e M. th produce protopod dea e – 3 , e pereonit fou – f clump e – as Seychelles ¨ a s K-SEY-34 Lati C st wide m, I femal r cephalo e on h I ller, , , , Lati , ban ) sp. a sp. ) l Schott ) ovigerou tw sexuall d , , holotype nine r mesiodista e n protopo t K-SEY-16 ree on two 1 teeth n cora 1. 3. o d r Louise 1992 nov. n 1991c nov. s e pigmen Ma meanin 6 0 d , f of of tha e cephalon fo . 13 fairl . crest mm mm n e l posteromedially , . 2.3 y 5 Latera r , y Amphiroa heads ; pigment , on ß Uropoda n , entire s dimorphi 31 . lackin 25 USN  fro d , encrustin 1984 , , y – , e at media Mah st l st ² 2. t g Mah , dense wit , 50 Februar juvenile m spine a a three 8 sam , ‘ on l or wit M . , . ¾ Cin mm K-SEY-16 JR-19 h S, g margin fou . e pereonite ´ P Latera e ´ level n pigment e h mesiodista , l 48 253221 anterio ara Island . Island on c q , somewha basi r dat g Dar hooks length ß on pigmen , , Cases y 32 stations , corallin ree types: st st , a san s 1989 ; e s l a a ² refer k as pereonite , 10 of , f ovigerou wit , r margin K-SEY-21 K-SEY-22 . s , Seychelles Seychelles ’ purple-re d  , , on , ¾ holotype alga E, at cephalo cephalo 1– ; Aldabra t h refer  s l anterio t e USNM , USN e patter , at mesio reticu corne 4, < to Mah Mah algae l wit , s 1 s tur 6– th s sta s m, M to of n h n n d 3 7 e e e ´ ´ r r f - - ; , , , , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 alga 9 on Island F ß , ig. 13 e one , juvenile 7 ¾ S, . ree ( ( Joeropsi , L) H Seychelles 35 ( f ) A) , male mal pools ß tw 10 , Male s fro o e ¾ E, Joeropsi lentig pleopo , juveniles m Matemwe 5 , Joeropsi thre intertida o Septembe sp d s e . pleurio 1; station , nov. s ( Asellot st I hamatili , l ) a Zanzibar ; to uropod r n ( CRU-1899 E s sp 1995 1 ) on . a m; mal s nov. fro Aldabra sp ; ; USN e (J , ZMUC . m ; pleopo 9 ) nov. ( ß M mal th 13 , M e ) intertida ; ¾ S, e , uropod ( India d 253230 , intertida B) Joeropsi 35 1; on mal ß ( e 10 F n ; l ) e , Ocea ( ¾ E, , s uropod N ree tw pleopo l pentagon st to ) 25 o f a mal n 3  CRU-1901 Novembe m; , at ; d e on (G , pleopo 1; ZMUC a Matemwe e ) (C sp mal ovigerou . ) d nov. uropod e r , 1. , Joeropsi 1995 on in ; ( , e s K Sargassum ; . Zanzibar ovigerou ) ( , D) s uropod two indic 1433 mal a e s , , ; ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 laterally dista fou truncate 1434 Thalassodendron USN USN of posterolatera thre teeth margin 1989. clumps to an spine widt long fro re on Aldabra, K-AL-86 sam USN on d d e Halimeda th m dea r Diagnosis. Typ Etymology. Diagnosis. Typ Etymology. Diagnosis. Othe Typ Previou corallin e e ovigerou h , anterio l , e to M M s M fou anterio smal dat hook P intersperse subequa pigmen d on s e e e ara 253223 si of , . r of 253235 coral r a 253234 material , material material x fallin 9 Latera material eac l stations ß as s types Amphiroa . cephalo ree e teeth r teet 24 Broa records and r algae s l pleotelson h , holotype t ¾ f margi an S, g l Ans , h patter side , Rostru Integumen Rostru , to on cres l , 1.4 shor : d d Gracillaria Th Th seagrass . . 46 d on . on intersperse margin , USN . tw , e e n media posterio wit Ans red-brow . ß . on e e e mm, t H n H H a 12 ` o distomesia entire Di t USN n speci alg speci olotype olotype h evenl olotype rubbl 1. la ; of Sr ¾ m m e Aldabra M , E, V on USN 3 Ž a a Mouche 1. stronges us ` i s n one , anterio ne mm as abou M 253232 Lanka la . 1 Ž of 7 1.4 t Aldabra Ž on th e r length y Latera Joeropsi e c bearin m, c mm on n B. d Mouche Joeropsi margi setae dorsa e M lon convex 253226 Joeropsi , cephalo nam – name wit ban ree : : wit : on cephalo 2. t Kensle ree l 1 12 , USN USN , 253224 USN r g margin 0 , 2– , t twic .0 , , h e f h . st e d an intertidal Mah g f l mm l Marc n on tw latera as ovigerou , (Ž (Ž Uropoda  16 mm a margin , (Ž  Ž s refer red-brow numerou encrustin of . , 6 at d of at s , M M ne fro gur gur o n e M pentagon s Latera gur K-ALD-13 on m, cephalon y wide m. Mah , , n pigmen pleurio posterio , e , ´ , lentig entire wide cephalon . h an 253231 , m 253222 Aldabra Mah setae l e e st e on s 253233 of st e Narro 3 1.8 Seychelles margin 1986 7D 7L a to a d s 7J , e th ´ e Apri . thi s , of K-SEY-15 r l l – Island pentagonal M. o e – – s K-ALD-70 ´ th e g . n . – n Ž margin 2. basi t N L) s tha a scattere F Uropoda Latera sp. ve pleotelso ; r Island 1.2 Lati . , e , pigmen 1. sp. on w 1 , re l species Schott ) ) USN , sp. margin ovigerou . , Ž mm s 8 1983 n 1 d ovigerou red-brow s nov. ve-side mm, mm cephalon of , coralline-encruste 1. nov. m, n , 2. havin Seychelles nov. long corallin 2– 5 s , l , 2 M meanin pleotelso . d mm 25 sam of , t margin e , . Seychelles mm 5 one s n , l on setae cora , on , 253225 m, d cephalo basi g Februar s of apicall havin , e anterio sam e s , n rostrum stron st cephalon , e 1 cephalon dat ovigerou st l . g stretchin 2. ban a s 1. , rubbl Ma s Rostru , a algae e n 0 havin ‘ 2.5 K-SEY-16 3 g one a of , lentil-shaped y K-AL-70 mm g bearin mm , d thre n dat tw y y as r – mesiodista subacute pleotelso 1 of entire 1984 1989 e . 5 o , margi m, holotype , a d g , , m wit e , m, s . st g Picar st pigmen al tw widel stron rubbl g as a broade a betwee 25 . . l h 1 0.9– . 1.3 thre o P P K-SEY-16 K-SEY-21 , n pereonites , Ma Pleotelso encrustin ara ara d alga holotype alga Februar juveniles . n g y straight mm, e 1.2 ’ e l Latera . , t y Island havin space mesio types: types: n r unde hook alon refer to 1984 l l tha mm, eye tur tur sta si n d n g g y x g s r s f f - , , ; , l , , , . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 ridge cephalon mesiodista longitudina Latera 190 tw setae lengt wit bearin station Aldabra, 253310, P sta pleon pereopod stou Som of Pleurocope rounded. Latera Diagnosis W seve Plateau Anto Joeropsi ara alter o fou h Diagnosis. 1 Remarks. Diagnosis. Othe Typ Diagnosis. Previou Materia Etymology. n 18A, t dista e . h types s n tw . ( r juveniles pereonite g setae Pleopo th l l ’ on s Pleo Bruun twic s s e , articles o margin margin tw r on e one waltervad Shoal , 46 material l s . 9 material pereonit seta o an W l typ : o Rostru setae ß s 5– e . n l l V 24 USN ree m, hook alker records setae examined d basa Pereopo d taperin , ridges 7 e ¾ e Phuke cruis S, . , , f F acros s s Tw 1 30 s ; visibl species) Antenna Pereonit Eye Rostru 0. Madagasca amil IIO on slope of of , M exopo Th wit . 46 . i m l ; 6 . 1901 e o Kensley Red-brow Augus , pereonit width e mm . pleotelso ß cephalo . E s 253309 e latera wit 4. h s H y g eac 12 specie t 7, e USN d presen speci Latera , pereonit R . olotype narro PLEUROCOPIDA : posteriorl W Island in ¾ Picar m d h st , , E, h 297 / 1 e l Joeropsi V . alter fro a st sligh USN wit t dorsa l  , ramu 1, Asellot M Genus Mesia 10 Ž subchelate s basall 381-A a ; , 1975 agellu Anto 1964 e n lobe r c w t m Kensle l d thre of h latera n Pleurocop JDT-AL-13 n , 4 253311 Plateau name on ’ entire Apri t s margin lobe-lik e Island M tw havin : th media lackin Thailand s transvers Pleurocope l : ; Shoal ; 4. USN y e y n l 367 latera view , e o s bearin m a Pleurocope 253238 pereonit USN ovigerou 33 lob to l l y Bruun waltervad , Mediterranean dista fro broade . lob 1983 of fro – an g , ß , , Pereonit . acut 369 n g e . 13 M s , (Ž on m e Aldabra seve 38 d si l e e Uropod M m Pereopod south-wes concavity latera of g ¾ gure extensio of extensio l x S, Schotte , , e wilson e bearin . 253308 , th , , m, setae e th fou Ž or e fou ovigerou E r antenna 7 cruis 253239 n ban cora 43 gure e s apex hav pleotelso e ß 7 i 30 34 teeth Fres India seve tha 8 r Walker l Lati ß e r Kensley , 51 ) d ; lackin , lob i dista s ¾ e g one s l , e 4 N, Augus , endopo 6– . n sp. n n 1, ¾ 1989 of , singl s . inserte bee rubbl n E, i 7, ovigerou t wit n , . e Latera , on n laterally of 16 l an 2. India articles 2– Uropoda long s fo Ž 98 l an or pigmen st nov. nea peduncl Ocea n n h , ve g , e : setae cephalon d r m. a e 7 ß , 190 sam , 97 d t e described ovigerou tw setae ‘ 00 latera entire spine Schiecke 197 rib 381-B r two , d d 1964 wit n . P on l W o ¾ 1 , n wit E, anterio dorsolaterall margin . e . s . ’ Ocean , 5 , alter t low on h Mandibula  Uropoda ; dat ofte refer e fore-reef l ; . l h acros oridensis . pereonite 77 , , pereonite protopo e singl 1. articl . lob cora 33 , singl a Uropoda s Antennula ’ n 0 , ovigerou m, : s rounded s , as r s ß mm 197 P e bearin Shoal 13 to 38 s , e of l . e margi e holotype Ž an 21 posterio ¾ rubbl – dasyura th l claw S, , 2 ve 3 cephalo , dista 46 d d protopo Hooker s y Picar r IIO e s bearin , Marc 43 mesiodistall 5 g pal l tw 2 on m. Madagasca , setae s n , . e ß on an l submedia r basi an E IIOE o 51 p Coxa seta fro d  proxima , truncate r ; d e rounde cruis d n h agellu ¾ W lacking g USN . E, four hal , Island s or d 6 m 3 entire alker, Pleo . seve 1985 1963 1435 eac eac R wit wit e near f tw tw e / M of of m 1, V h h h o o d n n n h y r , . , , . l . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1436 article Pseudojanira Museum a tw lob bot fro F ig. lob o m e h Diagnosis. 1990 Etymology. 8 seta e on . in th , an margin : e Pleurocop ( e lackin , pereonit 520 D) d on easter in seta ; pereopo Barnard recognitio th Sero g s e e n Rostru a ofte e e latera Th on v Gul latera wilson 4, an d F , e n th amil 1925 d 1; specie bu f dentate e l n m Wilson ( of l i lobe t Ž E Genus of y sp wel lob : rs doe ) Mexic 406 PSEUDOJANIRIDA . s t mal hi s B. nov e l is pereonite s , s of developed . ; an 1999 e name Kensle researc hav Antenna Pseudojanira Kensley . pereonite pleopo : d o ( A) e : seta an 72 d a d y mal . h fo . d , e singl , an l Caribbean on 1977 r 1; on reachin e spin s Dr d ( dorsa 2– isopods e F pereonit M. : Barnard Georg ) e seta 3 251 pleopo of an Schott l g E ; view , to d cephalo . Wilson an Wilson . e 5– Pleurocop e dista d , ( ; d Buz 4. e 6. 192 3; ( B) possesse Pleurocop Pleurocop (G , l ) 5 , n margi maxilliped 1986 Wilso 198 ) fronta mal e : 6 s wilson n 351 e n three of pleopo l e e of ; ; margi  dasyur Ž Poor (C th oridensi rs i , ) e t di d rathe antennula pereopo Australia e V n 2. a ers an reduce lacks s r d lacks from tha Just d 2; n n d a r , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 margins inserte Pleopo of male anteriorl subchelate or wit shorte stou anterio fringe stou distally an elongat tooth widt rostrum antenna margi Mozambique 1964 IIO Africa, juveniles USN Be – 2 1 – ´ Rostra Rostra Rostru Rostru d , . . . Ž h absent Diagnosis. Othe Typ E ne t t h Madagascar posterio ; , M tw . seta d n . . . seta r abou st ram d USN seta d r Pereopo , e e o tha havin a 29 seta . l , r l l m m wit ventrally , y margi 253318 seta 2 Eye . . . ; pleonit material margi margi . tooth stron material , IIO 372B ß i , carpu e plu anteriorl anteriorl n 11 in Eye t followe h M distall e strongl  one-thir . . . r e carpus s , ankin g ¾ E ban on femal s S, on reniform dens n 19 n n g s ; , d abou 253321 , pleotelson st Rostru . . . s e entire dentate bearin anterolatera dorsolateral ungu 32 , o ß . posterio d . rectangular a mesia si 1 y 09 . 2 V y y Pleo 13 d y e g ß x of . . . Ke e , JR-28 free in 02 an ¾ broadl narrowl t H d Inhambane S, media rectangular USN patc setos ß by opercular ; i te 29 dens ¾ y , olotype greate d . . . g E, ram m n mal ; , plu l 36 , n to ¾ thre compose ram surface 2.8 3 nin si S, consistin h rounded r , Pseudojanir ß broadl teeth . 69 M e Asellot . . . y x s i onl e 55 l e sam India margin of y Pleopod – of rounde depression e i in smalle 48 stou elongat e r m, . l 4. , wit ¾ 253319 rounde of E, . . . y or tha havin No elongat pleopo toot ß 0 : . . e male 14 , visibl pleopo h , , Cephalo 30 Pleopo mm , [ n y 88 USN t dat stou . . . a Natal d te n wit , Mozambique ¾ g setae E, d tw . r meru coxa Ocea rounded h fro (Ž n middorsa of fuse m, Jul d of . g e s Pereopo a , mor , . . . . d h o large e gure t on a Pleo thre mor to seta M e 1 m 13 shallo as d tw , 1 fou y pleonit . anterodista 9 ventrally ; d e meganesu d juveniles Sout n setae s Pleopo . . . . e [Zululand an 1 subacut in e th meru Octobe 1964 o n holotype ommatidi 253317 visibl e 3 basall e s e havin in specie n r , proxima d e mal havin stou , . . . . 9, on ovigerou stou in acute slende h 2. w consistin mal d India ; l 2 basall ; e 2 Africa s 10 length e propodu bot e 2, . . . anterio d in water USN e , mm y t 1 g wit e P. distall s , t r , , g ) 2 in seta s 24 , , plu seta visibl ischiu distall of to anterio 1964 h . n mal r l . . . . in Sout fremantlensi IIO lo y sp. l a h no , set ß dorsa ; fringe sexe Ocea seta , 48 M s Pseudojanira s arrange w protopod e IIO 4. , mal wide centra anterodista g e t y latera . . . . e r a E on pleotelson nov. 0 . h ¾ e 9 y on s rounde truncat reachin S, of m margin 253320 no present mm . s E [ dorsally Africa e rounde longe r . . st . Fremantle l n 2.9 posterio Januar P. r d opercular havin distall 34 posterio on t st l a view an l tha Mozambique , stenetrioide opercular a setae d – . . . margi ß e 403E s d 59 3. IIO e 357B ; in d r ; g , n , Sero dorsall d ; . 1 souther latte . . g . posterio ¾ tha pal l y y bum E, on carpu . , 2. P. anteriorl Pereopo midlength mm ; E r lob rounde lackin obliquel n , , r 5 1964 dactylu margin e v , . . m . . n meganesu . 42 W st margin r wit rows nort o an p , e Uropod wide a ester V . y demarcate wit s , s n m, . . . . bearin thre d in h Pleopo . wit Barnard on Natal JR-28 ; g fre r Mozambique h Wilson h d Madagascar . d singl y P positio n fre . . ; . . patche 19 y , s Pleotelso e e h e ara , ; propodu tw  of 1 widenin as Australia truncate s wit juvenile dactylu ang e pleonit anterio anterio g s . . . . Augus , sp. robust o , e latera d , types: fa Beira Sout short singl , , h smal Nos row 1437 e d nine 199 192 n 1 s . . r . nov. . te on by of as of in n h n g y 9 2 e e 5 s r r s s t l , . , l , , ] . ] . ] . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1438 species. 199 an of esus distinctiv th stenetrioides an stenetrioides F ig. e d d th 9 Th Remarks. mesia . 9 Just toothe ) e . Thi ( mandible e H al cephalo thre Pseudojanir l , ) s e l posses 1990 mal specie anterio surfac d e lie whic e in specie ; n ; pleopo ( Th s P D) P s s see e in . h . narrowl r e apparentl a of justi lef meganesus s n an th is easies meganesu of d t in pereopo e d fairl mandible 1; Pseudojanira Sero rostrum posterio th ( t y I e y ) di y triangula earlie v densel B. mal s V . d als an sp , erenc Th Kensle 1 pal : e r o d . rounde r pleopo in patc e nov lack p Wilson y species latte e omitted th setose recorde r . to y : h e s rostra d ( an mal on th r d A) detect 2. , . is e an d . 1999 ; bot e mal stron Th d ( dorsall M. , d E di fro unlik ) , h e entir e V ; maxill Schott betwee th er patter m P dorsa g . s e patc e y Australi e fremantlensis betwee meru th ver a in l e n 2; n e h view th y of tw ( th of s F e sparsel n e an a o ) ; dens earlie pigmen maxill ( th ( presen wester P. d B) e carpu e investigatori tw y antennule Sero r elongat a setose species o t t n 1; on specie species s v (G India in an th ) , e P. maxilliped ; , d e unlik s n (C seta rounde Wilson , dorsu s megan- an Ocea wit Poore ) e d righ e h on P P m n d a t , . . ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 Mozambique IIO 19 2.5 Pseudojanir referrin F ig. ß – 51 Previou Materia 1986 Etymology. E 10 3. ¾ st S, 2 . g a mm : 36 351 Pseudojanir to 372G a ß s l , 21 ; stenetrioide th , on records Poor examined ¾ e gre E, , e typ o Th 62 V y e , e a an sand souther e m, . tw locality d Pseudojanir meganesu s speci . 3 Zululan o Just Barnard , Octobe 55 juveniles USN Asellot Ž , n 1990 c m. , Mozambique Nos s nam M , d r 1925 : a sp a 1964 521 y coast , (C 253322 fro e . stenetrioide Be IIO : ; ) nov is ´ m 406 Sero ; pereopo , in USN E derive th . ; : Sout , Madagascar v e st W , ( 24 si an a India A) ol M x d h d V ß d s 400C 53 , 235323 7. Barnard Wilson mal fro 1962 Africa ¾ n S, 2.1 , m Ocea e 34 : sout 252 . pereopo – , , th ß , 3. on 1999 56 , n e in ; 2 h 192 ¾ e Kensley E, mm Gree : of cora 5 73 , d 55 one , Beira . k m, 1; l thre , fo 1– 1977 19 ( , , B) r e thre 2 Mozambique ‘ Augus m; : larg ovigerou pereopo 251 e juveniles e souther ; Wilson t island 1964 1439 d s 2; n . , , ’ , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 of doe ( pereonit brow 1440 thre in width propodu wit of margi setae slope as USN sta – 3 – 2 – 1 of large 2– carpochelate Antias Santia 198 Uropod Uropod Latera Latera Dorsu Dorsu mal 7, . pereopod singl thi h holotype Remarks. Diagnosis. Othe Typ Diagnosis. Wol s 453, e 6 ; , sometime , M tw n hav s , dista n ), . e inserte Sivertse Richardson latera Picar wit V specie e pigmen e o m m distall of examinatio l l e 253301 r , s e o material s s margin margin anterio . couplin 1989: material 5, h of of wit V l cephalo muc subequa a d ; st s l singl . . pereo pereo an d tin Somalia, s n h 3– i USN cephali Island y Pereopod V s h t is dorsall an 182. d Th , thre y truncate s s onl Cephalo Maxillipeda 6 . r shorte spines on on , e g anteriorl of of d a . toot eac n n pleonit e 1906 . l n latera M Holthuis n hooks e y mor . in e no arme pleo pleo , distinctiv mos H Genus eac Ke c h on of USN smalle 9 y h lengt t r olotype: : 253303 , ; ß wit 11 lob . arme s 24 e tha n n 16 F y or h th , an si exopo d 5– t l n amil 2– e eac ß . unarme arme di x to . ¾ y st wit e h wit 11 of e . S, M d h n laterally Pereopo 7 lackin an , Santia V ovigerou 7, i typ r fou rounde d l wit 1980 V to h pleon ¾ th associate in B. h us N, h 46 y e th proxima d , 253302 . wit dactyl d pal d ramu set e stou Santi pleon e e colou dorsa SANTIIDA tw r on h e Kensle ß pereonite wit an USN d India 12 : h materia 51 setae g reticulatio . p a dorsu , singl o 89 e stou t Sivertse lackin d ¾ h . fre d d ß an E, s setae article ; , 14 a spinos i s Pleopo . . , hook-lik r l endopo wit (Ž Kussakin wit 1, bearin eac d M , , e d n l spicat view 12 e 25 ¾ y t patter E, of m gure , setae st latera set carpu spinos pai Ocea . . h h st g an l h ; two 253300 m, a s pereopo (SAM-A1534 e i elongat of si s 47 cox a a n biunguiculate V . d , g r E n d 1– . . K-AL-50 seta x a Uropod 11 . hyalin al e d 15 on 60 n an – of elongat M. n seta th l 1 dista Kussakin on s e , Pleo a sp. 6 49 seta l eac , of margins ) . . 1988 e specie d wit seta e in submesia Apri of ovigerou , eac on slender eac Schott m, e th ; Holthuis cephalon ovigerou nov. e thi d h male n l . . . h seta e pereopo lo e e e : h setae h e 7 wit s midlengt l 230 juvenile 17 margin s thre w , pleotelson s seta wit sid pedunculate . . . . . 1983 wit e specie on of , , , h e S. submedia . 5 ; Decembe e plu ram . 198 . h e s e ) Kensle l h . . fou . . Pereopo S. Coxa Uropoda marmorat Santia , of show purpl s posterio stron singl , . d 198 tw s , , 8 uncinata a s , i . . . . . r 2 bu th h 41 pleotelson [ sam fused consist o 0. dista Fals smal havin e y 0 e e abou . s g t e . . . . 9 juveniles pair visibl st an Contrar n S. pleon tha r neve posterodista e mm d , sponge e i a r [Aldabra l d l V l . biramous lobe 1964 . . . dat [ urospinos Bay protopo S. setae t ( ( Somalia 1 t st Pleo g s media Schotte s seta Vanho Vanho , r P 1. e i Ž of . of a . . . spicata V subchelat IIO , . s toothed . on 4 ve . Th Sout as stenetrioides y . n , setae. , . on stron time P st Maxillipe [ . . . ¨ ¨ sam Pleopo E oute pereonite n st St to ; consistin e ara V V i holotype , ; a V Zanzibar d en, en, i , 1989 h rostru anterio Aldabra spo cruis V Pau . . . sp. sp. Wilso usuall l s havin e Africa dorsal types: g r . setae setae, basa 1914 1914 e dat t red l . nov. . . nov. : ree d e Is. 98 on or m d n 9 g 1 a g y 3 2 r s - f ) ) l , . ; ] . ] ] ] Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 spong spik F ig. Typ Etymology. Remarks. y 11 2; maxill e appearanc . e on ( I Santi material ) a oute pereopo 2; a ( Se r spicat E e Th ree ) e . of d mandibula ‘ e Remarks f 1; H a th slope speci olotype: (J sp e . ) animal Asellot pereopo nov Ž , c Picar r Santi ’ . palp name : sectio . ( A) a USN d d ; a ( fro 2; , Island F mal n urospinos (Ž fro ) ( m M afte K maxilliped gure e m th ) dorsa 253304 , r pereopo e th Aldabra Santi 12 India e a l ) Lati sp. view ; , a (G d n urospinos nov. n 7; , ; Ocea ) 9 mal ( fo 1. ( B) ß L) 24 5 r e mm n mandible spike pleopo ¾ S, pleopo a 46 , . , st ß d 12 refer d a 3. ; 1; ¾ E, K-AL-50 (C ( s H ) 25 to ) maxill mal m, th e e 15 , a pleopo overal purpl 1; Apri 1441 ( D) d e l l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1442 26 three seve 253307 san as 1983. F ig. holotype Septembe d Othe n 12 2; (C an P station . , , ara ( ) d r st nin H Santi lef material a rubbl ) types: t . e maxill CRU-1904 r mandible s a ovigerou 1995 on urospinos e a USN . Mah fro . 2; USN ; m ( ( s I M D) , e ´ a ) Ž intertida Island pereopo sp ve 253305 , M maxill . eight B. nov 253306 station , Kensle a . d : Seychelles , l 1; ( fou 1; ree A) , ( , s E on ( thre f J r mal ) y o ) e ovigerou maxilliped  V an pereopo at juvenile e e Picar , d dorsa , Baw , 1– M. seve 6 d d l s Schott e m; ; , view 7; ( Island n Island cora F , ovigerou ( ZMUC Ž ) K ; ve mal ( ) e l B) , pereopo , rubbl e , Aldabra Zanzibar righ pleopo eigh , s on t e mandible t , d e an three juveniles d 2. , 1; d , , 1– (G 6 16 alga ß 26 08 ) , , ovigerou mal pal fro ¾ , l m; S, sam tur p m e 39 omitted USN pleopo f coars e ß 08 fro dat s ¾ M E, m a d e , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1 wit uropoda lengt stron carpu wit acut Hansenium 8 7 6 5 4 3 2 1 uropods. S 191 very dorsall specie ( S – – – – – – – – Vanho . . of Rostru palm Rostru Rostru Eye Pleopo Pereopo Pereopo Eye Pereopo Eye Rostru tha Rostru Rostru Eye Pleopo Pereopo . . . spicata h h uncinata Diagnosis. Diagnosis. Etymology. 4 Remarks. e mal ) h n di g thre thre s s s s s s . . . fro ¨ y or pal consistin reniform V consistin wit 1. width posterodista V wit easil d m m m eren m m m d e l en 8 unarmed e e . . . d d m d d , m base h 1 havin 1 rounded Sero broad-based poorl narrowl triangula h large short bu dista couplin time , 1 1 1 1 ommatidi in in y th ( t . . . . 191 in in in in tw Vanho t separat s fro male male e v g , g ha Th , , s male male male male y broadl o g l Cephalo Cephalo . . . . wit an Antarcti 4 triangula of of anterio Th m setae de basa s thre y ) , e stron g d , , h ; r ¨ thre eigh fa . . . . Ž l triangular fro an , , , , latera e latera thos V a Wilson onl hooks Genus ne ommatidi propodu carpu rostru propodu carpu r setae e en e , l Famil in speci d ; y d m e . . . . . t more distall r Ke g it dista width y endopo , convex e or to n n roughl or possesse r margi c l l 191 fro posterodista St Asellot of . or . . . . . s y s margi margi specie an . wit is , Ž mor Ž m absen Hansenium y produce no Coxa 1995 ve , Pereopo to c l y ; m Pau s linguifor s d S 4 a , a smalle . . STENETRIIDA . . . h setae pereopo n narrowed t usuall . ) usuall y nam short e ommatidi latera th . arrange pereonit d spinos th n produce n conve spicata anterolatera circula fro s : Uropoda t ommatidi l s . . . a . . e e angula wit evenl 72 e Island of . e gener relativel fro d visibl m tw ; y , y r ...... d h m l int Bolsta x refer d longe st as e m l o rectangular d y Santia r Fals margin 1, Ž ; d 1 i o Sero e a r or specie setae ...... grou pereopo V conve ve in lon specie a th e . in a posterodista meru lackin l s seta of Th r kidne e e on S d protopo y or g ...... male to v tha p . l India an , Bay as x th e smal s s urospinosa an e toot si E s know pereonite s propodu ...... d n th d e latera widt y g bearin unarmed x , tha describe wit d Hansen , Kensley widt 1 propodu Stenetriida e shape , st dista n l ...... h Sout Wilson in d n i h h prominen anterio non-spinos n Mizothenar V Ocea Liocoryphe Hansenium Lexcenium l Tristenium wel h male havin l see of ...... singl lob g seta , fro l of pleona h s s or l , pal setae a . d , n , e ...... s . . n , barel 1999 pal Africa m Santi developed , 190 2– Tenupedunculus e super posterio e in propodu here abou eye g m r 199 Stenobermuda or stron m e 7 ...... Protallocoxa tw th . th 5 t margi s : Stenetrium l y Eac in 5 . Sero Sero Sero Sero Sero a e 164 absen o armature t e e seta spine Ž ...... , Th twic in hispid dista cia uropod tropica g dorsa India an . h s r  v v v v v ...... e e n anterodista slightl ; l ated t d e margi lob s an an an an an latte presen resemblanc antenna l longe truncate of ...... a S. d d d d d n setae l e Haswell l , . s Schultz Schultz Schultz ( view Wilson Wilson Wilson Wilson Wilson y r of species Vanho th Ocea marmorat Maxillipe ...... wit n whic longe r specie e t ; tha pleopo betwee h exopo ...... l on forme ; l n Pleo h , , , , , , , , , toot ¨ n r thre 1443 seta V . 197 199 199 197 199 188 199 198 199 eye e ...... s ar ar th en, to is n d d d n h a e e e 8 5 5 9 5 7 8 6 1 9 2 4 3 5 5 2 e s r , . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1 masculin posterodistally reniform 1444 pentagonal seta margi midlengt Pereopo pereonite ban of Phuke 11 on three 1987. rubbl – 4 – 3 2 – – of Pereopo reachin Pereopo broadl Pereopo pal Pereopo pal Pereopo of Pereopo Pereopo Pereopo 4. . pleotelso fore-ree Etymology. Diagnosis. Othe Typ station d ; 0 pal propodu m m dactylu e n mm . on P t , bearin bearin e on m ara Island y densel d r fou g h , a s . d d d d d d d d cephalo , material ovate material 1 s beyon of of distall fore-ree f types: , 1 1 on 1 1 1 1, 1 1 1 in n r s s o abou slope . . in in in in g g in in posterio in in wit V juveniles jus e , 13 male y 3, , ; seve si male male male male Thailand male male ovigerou male d no Rostru . male . . x propodu setose Picar – t h y Th n 4– t , posterodista 19 . teet reachin . USN t Ke n f singl widened , 1. Picar , . . . , , , reachin betwee , , e 7; carpu , teet , slope carpa carpa H propoda propoda 3 USN propoda h propoda r y propoda ommatidia d speci propoda , , margi olotype . . . time pleotelso m M to e h st pal Island d , s s tooth g a l , l s wide 5 th M . . . . . 253195 g Island n longe lob Ž lob m wit , beyon Picar s K-AL-124 Hanseniu Decembe c beyon e n l truncate B. l (damage l l eye l longe l 253197 l e . . . . . pal e nam pal wit pal India angl pal h , of r pal . pal reachin : reachin Kensle n r broa Eye s Aldabra tha . d m , m m . m propodus . . . . h USN m d , d tha m e an Aldabra e Pereopo r wit Island tw bearin lackin propoda bearin n thre n posterodista lackin of wit s derive tha , . wit . . d . . d . m (Ž n r o d Ocea reniform h long g thre g , pal y M rounde h on ), 1995 gure widt h no n e aldabra to sam . . . . . patch an , g subspherica g , g g m stron thre lo wide 3.2 e posterodista 253194 , mor s stron 2– Ž pereonite , n Aldabra d ; si stron d . . . . . w anterio ve e h . l fro expande 21 13 x specie – M. 25 posterodista , 1 e dat d y of teeth e g teeth 3. teeth . . si l m, ) ...... m e g juveniles tha g setos m. Red-brow in angl 5 ga x pigmen Schott palm a lobe sp. , gap mm th ovigerou 15 p ...... ; n s ; r as , [ , male l ZMUC pereopo e Gul e pereopo s halfwa betwee d of margi e nov. , 9 ; posterodista stron l Apri , of typ ...... holotype [ ß continuou . 2 carpa 4. propodu posterodista angl Nosy 24 fou e f Hansenium Pleopo t 0 , an H. , pal of e . . . . . ¾ l mm S, dense st carpu g n l e y r n n d seta locality s m l , monodi d d Suez 1983 a of alon ovigerou pigmentatio Be articulate lob . . . straight . . posterodista 46 4; on 1 1 , K-AL-53 [ ´ , propodu H. Mah , d . ; in in s ß e s seven st r e ; s H. Madagascar Operculu scattere 12 . . . g l . USN wit 1 longe Sr a H. female lobe female dodo in posterio strongl , ¾ ( H. l in e E, remocarpu ´ i Nordenstam . . . , h K-AL-124 Aldabr lob Lanka . expansum Island anterio s ; st [R conve aldabrae M male d Latera r 21 s ( carpa [Aldabr . . . , , e a tha Mu , , or d e l ´ posterodista n si propoda propoda no cora unio 253196 3.2– m, r y an m CRU-1982 ¨ x patche beyon , ; n . . . , in margi ller a ; t x Seychelles Seychelles d produce l juveniles appendi of r wide s Zanzibar 13 l reachin n Atoll l lob anterio anterio 4.4 , res . . . a sp. sp. sp. margi region , femal rubbl Islan , 1991 Atol d n Apri , e 194 cora t , l l s mm, nov. . . nov. nov. . tw . on d d 6 l d n o x g 3 4 2 e e r r ) ) l . , , l l ] ] ] ] ] Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 subcircula Eye margin 20 ovigerous fro two fou Madagascar F ig. pereopo m Septembe r Diagnosis. Othe Typ s 1 ovigerou 3 reniform Murog . 2. . e r Pleotelso 8 Hanseniu material d material r mm 1; lob , , r o s 1 ( two , . 1995 D) e m, Pereopo thre Reef Rostru bearin 2.9 m n . femal . 15 . aldabra 1. e , – , H ZMUC, 2 3. Januar Ž Zanzibar , olotype e g m time ve 0 d st opercula fring mm a 1 e Asellot basall juveniles Hanseniu sp s in JR-29A y , wide . e fou male two nov 1964 : of , y r USN a 14 r r pleopo . setae : wide , fro juveniles , . tha , ( m, m carpu , A) P Nos st (Ž m M on ara n a expansu gure ; r mal d 20 propodu th e midlength y CRU-1926 253205 tha types 2; s ovigerou e e Be wit Septembe , 14 ( dorsa India ´ n sam E , ) ) h Madagascar m : long mal , latera USN e s l sp. n view , dat wit s e , , Ocea latera 3. nov. pleopo r Murog taperin M h a l , 1 ; ( 1995 surfac as proxima one mm B) 253206 n l holotype femal margi , d , ; 1. o g 1; st , e ZMUC 5 Reef a st to ( expande m, , e l F n a tw pereopo JR-33A regio ) truncat wit CRU-1893 11 ; mal , o USN Zanzibar , h Januar n e fou d singl 2.8– pleopo of d , int M e 1; Nos r posterio anterio o (C 3.1 e 253207 y , tooth broa , d , ) y 1964 cora 1445 mm, mal 9 2. two Be m, ´ d r r e . l . , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1446 iles Stenetriu Stenetriu pereopo shield-shaped wit distinctl margi F ig. h , Materia Etymology. 14 st ro n . a (C w y m m d expande MS-SEY-1 Hanseniu ) o 1 monod chilton femal of V l in examined se , 12 t th 1. e i i fro d 5 e Th pereopo m Nordenstam no shor time male int m n , e expansu cora Stebbing speci o Hanseniu . t res s . serrat broa d longe USN t l 1; Ž of m rubble c B. , ( d , D) 1946 nam e r sp palm M Monod m Kensle setos tha . seta mal 253191 , monod nov : e n , Mah 19 e e refer e latte (Ž greates , . , pleopo : y 1933 on 22 lob gure ( an e i ´ , A) ; s r ( on e Island W posterio d Nordenstam : to wit d 15 171 ol t e bearin mal M. width 2; th V h ) , , , e e ( Schott , 1962:22 Ž fou E on gur expande Seychelles dorsa r ) g . e r femal e margin singl ovigerou lo 3. e , l ; w 194 Schultz e view e d triangula opercula . , 6 articulate natur ; ) 1– Operculu s ( , B) 2 1982 , e m, r one mal r of pleopo : 29 teeth d 20 th e m robus . , Apri e pereopo two d of ; carpu 2. dactylu l femal t juven- 1984 seta d s of 1; e s ; , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 Nordensta th fro femal densel da Pereopo 1 anterio of to four Ans USN F ig. an e l 2 m pleotelso Remarks. Previou Diagnosis. pal e singl d 15 Ma M e th a y 2, ` , . m 1; r pentagonal e alga la setose d y 253192 narro e ban Hanseniu ( Gul wit 1 D) 1984 m pleotelsoni Mouche s of l n d h records propose tur mal f ; wit w si male on of . Nordensta , dactylu x Rostru f band e on h m Sue fro teeth cephalon pleopo , , , singl monod e . lengt carpu Mah m d z c s Gul , th s as on m , toot thre e outermos d reachin on h e i e ´ S wide m : tooth Asellot s pereonite 2; nam f ( 1.2 , . Island e e h wit A) of narrowe ( chiltoni ( station ovigerou 194 E (instea 5 r mal h e Suez ) . time g tha S mal digitifor 6 a Eye t , . beyon e ) largest fro monodi Seychelles n dorsa s ; note e s s d Stebbing r s 3– Sr pleopo long m on width s posterio reniform of i 6, d d l th Lanka m Mah , view ; serrat , anterio , tha outermos e posterio st posterodista anterio bu . d India a , , t 1; ; e t ´ r ( 1905 2– K-SEY-16 th ; gav e . B) Island ( band Amirant F e Red-brow r 5 latera n mal r ) an mal e m, r , t Ocea femal margi no margi coul d , propoda e , 1 e mos pereopo posterio l Seychelles l furthe describe Ma margin extension e n d e , n n opercula t alga Islands no n y straight of of pigmentatio t r l 1984 d dorsu l description be carpu r tooth s 1; d ban tur , of (C . tha ; r by ; 0.5 expande f . pleopo USN ) d th m . s Latera t on Mono femal – Operculu on an e species 6 of pleotelson) m, d dea n M pleotelson d . pereonite e propodu Monod in d d l 2. pereopo 29 d 253193 margi propo , ( broa 193 coral give Apri m 1447 3 of ’ n n d d s s s ) - . . l , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 illustrate femal th 1991 an Ž 1448 Thailand Stenetriu – 1 truncate fe produced anterodistall of Liocoryphe shap an Operculu an dista Pereopo Dorsu subequa 2– st 2– rubbl gure a w e tha Pereopo Pereopo 3.4 d d 5 5 . thre oa Typ Diagnosis. Etymology. Diagnosis. K-SEY-16 Typ m, m, ke lo pereopo d e l n singl e mm, e s r . margi m y w of e lon fro an seem e 1 e 1 of or m d l , above or d , material rounde m th Ma . Ma material , d d wit wit d g m e to siamens 1 coll Sero rounded th paddle two by 1 e fou 1 encrustin n of s in y d lo h . . y Re in h e y carpu middorsa in to of , . Mu produce , w v 1984 1 ´ femal 1984 subdista r Gul male scattere male alga T. unio Cephalo Rostru a . . male d an ommatidia al ¨ be propodus , Th e . , . lle bump di sam Ke l Mortensen teeth d f Hansen s plu , , . l . . . th agre e r , V n L propoda e setose Wilson H , of of Genus tur P g y erenc carpu speci ( ectotype: e propoda ara shield-shaped e s d Islan olotype 1991d d . . l m corallin l to Liocoryph . pereopo majo Sue ; f dat - e int n ro length carpus Pleopo on red-brow wide , types: closel dactylu th . . , Hanseniu ; Ž lackin w e , 1905 o s d z . a Liocoryphe margin propodu c l 1995 e dea B. r , Pereopo of setos wit pal of , as material is nam . . l India Ž r di e 7 : ; : pal , gur Kensle y d tha on m d ver shor d h V holotype : USN 327 algae ZMUC, Februar USN g th latera . . s wit e erence 73 1 e coral m e bearin broadl e 2 n s e anterolatera e n y siamensi lobes n m , in is , . jus setose 16 toot . . s t h in bearin Ž pl long (Ž (Ž d abou pigment simila Ocea M , , M expanded a remocarpu barbs ft y th th . t l ) , 1 gure gure Ans however Sero male h . . combinatio g s 21 an Ans e is ; h e margi y 253198 y 253199 in ; overreachin , tw betwee carpu articl , male t ; presen g n anterio USN d als on 1900 ovat e r . . Ž 4. s propoda . mal o 1.2 v 16 17 e thre , gur specie M. a to ` ( , 1 lo o an a appendi ` Hansen th n ) ) . . 0 mm . la w l e no s e e . e , ver la M , , , e th e Schott d s n an time t wit as 2a wit teeth of strongl posterodista thre showin r Mouche lo t . . sp. e Mouche propoda material th s Wilson y 253200 – margi Õ d n (damage w wid h in th presen i. of h l e simila g s of e . . ; antenna pal nov. , posterodista rounde x e species carpu 3. singl e longe Liocoryphe an 190 e outermos remo- pereopod masculin y 4 g m . as n 2.5 , y , , L mm setose L , r t . th 5 l 199 straight wit on e Mah s long Mah . d Stenetriu . ) clearl to r species d – pal e abou . gertruda siamensi ), l l tooth 3. , tha e fro h teeth Th 5 cephalon H teeth lob st 0 m Melle ovigerou ; e , ´ tw e ´ . mm a t n a e ischiu m pal , y t Island l Island e monodi in . as ; o colou . an . distall wide K-SEY-15 corne propoda Pleotelso th de . almos carpu m e As s [R triangula m , Eye lon m bot d Ey e ( ( Ž on e Mu m Hansen oblique ´ dod ne refer Lati , Ko . unio g , indicate e , s r h s r e y ¨ s . pleotelson as Seychelles Seychelles an d t patter somewha ller consistin [ ovigerou mal o reniform h wide Thailand widened reachin n wid l n d s 2.5 pattern n Mu Mesan , , r remus Islan , to meru tooth 1991e e , widt e 190 cora r ¨ teet mm, wit n d ller, an th as d in 5 d h h h g g e s s t ) ) , l , , , , . . . , ] ] Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 th collecte Phuke coral ovigerous Island sand F ig. e Remarks. Othe mal 16 , , opercula mal , 10 t . 3 d e Island Thailand r m, e m, Hanseniu pereopo material by pereopo , 26 21 st The r , a pleopo Compariso Novembe Thailand Septembe CR o , m d . d smal Mortense 1, remocarpu U 1; ZMUC d reveale 2015 2. ( l D) , r rubble r sand Asellot n 1995 , 1995 juvenil , Ko n wit s d tw sp , in no ; o h , 3 ; Ma . a ZMUC m, ovigerou e nov ZMUC 6. 190 Hansen di fro 8 mal i V 29 . Ton m, 0 m : erence ( e at A) Novembe th , 23 pereopo , , ’ Phuke s e Ko on s on mal s India Novembe typ , e e h fro st e dorsa a Mesan e d t m , , CR n r Island materia 1; one st 1995 th Ocea a l U ( e E view r CR , presen 1896 ) . Thailand , 1995 , n mal Thailand l U ; st ( ( a , B) 190 2008 e ; Baw t CR pleopo ZMUC femal material 5 , , ) e U , an Ko Island of dea e 2021 d d pereopo , L 2; Lon d . especiall on . , compacte , ( Zanzibar e F siamense Ko , ) d Phuke femal , 1; Aeo 1449 y one (C of d e t ) , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1450 K-SEY-15 Gionnet three coral 13 cora an longe longe distall on Caribbea ( mal expanded ommatidia rostru Mizothenar Kensley d Apri th Typ Remarks. Diagnosis. e l wit , r r e distall y m rubbl fore-ree tha tha e latera h l mor , poorl , 1987 n 2.1 material no , n Aldabra n 1984a , . as Sero specie y cora e e th – th Pereopo mal l widened 2. wid . narrowe on e f y e margin v P Tw 2 unfused slope Cephalo l unfuse ) de e ara an mm . s e rubbl F fro fore-ree materia o , Ž d as ig. di ne types: H Genus 16 m , Wilson d specie , V s , long 17 d olotype d er Picar e d st truncate 1 Belize m, of . . tha or n s on a part Thes in f l , fro th Liocoryph s wit fro K-AL-65 USN , Mizothena Mizothenar 17 n absent pal fore-ree d slope e 1995 B. male . of m , in e Island h m pleotelson In : an m Marc Kensle . tw USN M th M Mizothenar th shor : , d bearin th , 76 o . e . e Picar M carpu 253202 f e Ey e specie serratum . , latte , India h slope (Ž t M latte siamensi . Aldabra r y e coralline-encruste maharepa gure anterolatera 1985 serratu Sero g an consistin d 253201 . r s ro r n s , Th , specie d Island no Cin ar species Ocean w on 18 v ; , M. s e hav e , t wit an , of USN e m ) femal q lectotyp remarkabl , 12 posterodistall Schott s d ovigerou Cases e sp. g , obliqu h ( , Mu , 2. in m, Wilson Aldabra of th bee l furthe M th 1 e nov. ¨ e lackin tooth mm e ller 6 rounde e operculu , n e fuse 253204 fuse e pereopo Aldabra Apri d , described dentat , s y r , 1991e ; rubble st d comparison 199 , similar g d l a antenna sectio d 2. 9 y serration par 1983 , ß K-AL-24 5 2 m d grou 24 e produced ) , on mm t 1. , setae 13 fro ¾ in , S, is : n ; e morphologically mout p m, USN , m M M l abou abou 46 st of . s toot s . . Moorea 4 a Pleopo , ß 2.0 or ar maharepa h 12 patulipalma ; liv nin Apri K-AL-124 M t t e h propodu ¾ dentitio 2. 1. of e mm, E, di e 253203 5 8 broke absent l Y or d Pass 1983 21 time time . cult. Th 2 te sta m, in is n n n e e s s s . , , , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 subequal Stenetrium pleotelson. Operculu abou 1 wit latera cav 9 10 F ig. m, in m, h e Diagnosis. Etymology. Diagnosis. Othe 18 20 t scattere femal betwee l 21 . (C margi 14 Septembe r ; ) Mizothena m Septembe material Haswell pereopo rostru shor e of n d n wit antennula red-brow wit femal t h Discret Cephalo m serrat Th , r h d . propodu r 1881 r 1995 short 1 abou e serratu e 1995 ZMUC, palma wit speci e : e n . r 478 n , t setae rostru h ; pigmentation bases Genus Asellot te rounde m s r ZMUC wit Ž sli ; n setae c Sero distall sp t , spines h two nam in . m . singl Pleotelso v nov anterolatera Stenetrium ; a d dista lacking ( , an y fro D) e to . on . e , d : broadl Ommatidi refer m st antennule subtriangular ( stron l e Wilson , A) a third no th , n CRU-1896 , s anterio e femal t st y g ver Haswell in India to . a l , expanded articulate 1995 y an an CRU-1926 ; a e th ( slightl arrange E y d r n dorsa e : ) clearl margi 49 Ocea . antenna femal , serrat , Ey 188 . sand l y d , e view y d longe n pal n e 1 , posterodista reniform e de roughl opercula , of Murog l m Baw ; latera Ž ne teet cephalo ( r B) convex tha d e y h femal pattern o l r Island , in n pleopo wel margin Reef of greates circle n , l e abou l wit slightl robus , , pereopo . developed d Zanzibar Zanzibar Pereopo . h s Dorsu 2. t t ro of t width y 18 w 1451 seta con d – th 26 of m 1; d e - , . , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 wel 1, 1995 ro of ommatidia 1452 tio basall cora 3.5  – 7 – 6 – 5 – 4 – 3 – 2 1 tha – – at Rostru Rostru Pereopo Pereopo Pereopo Pereopo Pereopo Pereopo Latera Latera rounde Pereopo Pereopo Rostru or Rostru w Pereopo . an n – palm t , l Remarks. Diagnosis. Typ Note: 4. l : of be of ha convexit d Baw y head 81 . 8 pleopo d e broade barbs red-brow mm furthe l l . m m m ) d m margin margin . los material d d d d d d d d d sho Stenetriu Pleopo e posterodista s subtriangular subtriangular rectangular . triangular 1 1 1 1 1 1 1 1 , in Pereopo t in y Island . . on . in in in in in in in in w almos r d male r sand Th e considerabl male male male male male male male male divide s s tha 2. Cephalo . . d n ovigerou of of . e , Ke Th , m pigmen 2 n . . t , pleotelso pleotelso , , , , , , propoda , , , approximatel d H long Zanzibar , Changu propoda propoda propoda propoda propoda propoda propoda truncatu propoda al in ape l y e d 1 anterio olotype lob . . , , l genu to in in margin fringe male of n x , e s Stenetriu t . . th anterio acut male th e wit it l on n n s e e variatio B. r s l l l l l l l l , u pal m 6. d entir serrat . . . is , pal pal pal pal pal pal pal h pal India e margi future legs s appendi cephalon : Island , 0 wit Kensle Nicholls m or no 26 entir carpu stron mm ZMUC, m m m m m m m m . . . y r e h ; lackin t rounded e margi m bearin bearin bearin bearin bearin bearin wit 20 bearin n n it smal Septembe wel e n ...... , (Ž , straigh Ocea assumentu is USN g s h in y specie x Zanzibar gure l rarel ...... g , dee , no an l anterolatera rostra n masculin de [Sout g g g g g g g 1929 pereonite denticle an , straight n d t si fou mor thre mor tw Ž p M ...... Ž t or ve y 19 y x include s ne specie M. 4. emargination o r r h produced teet emargination , 253292 of e e e anterio 9 ...... l ) . teet teet d wa teet m Africa form , teet 1995 mm tha tha Schott s by 7 h Stenetrium a . . . . h . . . . s . h sp. s s h m, n n Eye h [Rottnes d describe distall , clea an of l , . r thre tw , ...... ; in st structur nov. 19 S. on an margi d St e s P Stenetrium o ; a S. S. S. S. r th S. reniform ara propodu e . . . . pleotelson . . . macrochiriu ; e d teet Septembe Pau synapomorphies CRU-1904 y ; spinirostru carpu teet perestrello S. e crassimanu bartholome magnimanu t carpu antenna d n types ...... followin widened h Island l esquartu (se 4. h fro e wit an S. 5 S s of ...... e mm d . h s m no zanzibaricu s . Sero : assumentum th , Amsterda havin Scattere rounde [ longe . . r . . . . , t a m m W W i l ZMUC, , e s m latte g , i [Zanziba [Zanziba produce 1995 , ( singl m st ester ester teeth mal ( ( Kensley ( ( v wit ...... intertida key Nicholls Nicholls Barnard Barnard (Schultz a g (Schultz,198 [Sout [Sout [Sout [Sout an r r CRU-1890 d h broadl . e tha n n e ...... , wit d d ; m bum pereopo m specime Australia Australia subdista an d reticula h h h h rostru r r Wilson n , ...... sp. sp. Islands h two Islan Islan d , , , , , Africa 1984 Africa Africa Africa widt l p y 192 192 194 191 198 larg ma ...... nov. nov. . ree b d d m 9 9 0 4 2 2 n d h y 3 2 e 7 6 5 4 - f ) ) ) ) ) ) ) , l , ] ] ] ] ] ] ] ] ] . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 800 8, Stenetriu th Operculu articulatio followe produce strongl F ig. e st Remarks. Previou Materia Etymology. km. stron a 19 pleopod (C 400C . y d m ) d g Stenetriu m by femal pigmente posterodistally bartholome n s blo , l of tw records sout examined of 2. b e femal o Thi pereopo of dactylus smal h Th m d s pigmen of . assumentu e i e recor middorsa Barnard l speci Beira . roughl Fals Stenetriu teeth d ; ; USN 1; d Asellot propodu t Ž e latte ( , , at c D) extend Mozambique y Ba m , dee name M 1940 pentagonal l th mal r y sp m p patc e a 253294 to . wit : bartholome e bas U-shape s s fro nov , 431 pleopo Natal expanded th h fro h m e . , e : at abou of m Ž , th northwar ( gur on , A) , , d e th th base d 1. Sout 19 2; e e India e t e 5 emarginatio i mal ß 19 , ( pleotelson 51 Lati Barnard time 15 E wit . , ; h on ) e ¾ Kensley Pereopo S, n Africa mal h d smal n dorsa e s Ocea 36 stron rang ovigerou longe assumentum e ß pleopo , 21 l l , , 194 . n n e g seta d 1978 view ¾ intertida E, r plu toot of 0 tha 1 d 62 e s ; : s th of 1; 145 n ( h m, on singl B) , e , ( wide demarcatin one F male , l a specie 3 Ž mal to ) cuttin patch e gur femal Octobe . toot 22 , e , e IIO s carpu pereopo 65C m. g e , h by opercula refer E r margin clos g , D. 1964 abou cruis palm s 1453 d s e no to to 1; . e r t t . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 encrustin 3– 1.9 Comor 1454 F rostru 25 st Murog ree 10 ig. a 5 m, f – Septembe Typ Diagnosis. Othe CRU-1902 m, 2. 20 slope pleopod (C m 2 6 . o o e 14 mm ) r Apri muc Reef Stenetriu g femal , material Islands material Marc corallin Aldabra , h r tw l 1. , e 1995 wide 1983 , Zanzibar o h pereopo Cephalo o , m ovigerou . V 1991 . e 3– . r ; , quinqueden tha Matemwe algae H USN 22 USN 5 . olotype m, d m, n , n 1; 16 long , Stenetriu M M s havin 14 ( 13 fore-ree m, D) B. 253211 253210 s , 2.0 , Apri Marc femal anterio : 20 Kensle sp g Mnemb USN mm, . stron Septembe m f l nov e h , , 1987 slope (Ž quinqueden opercula on tw y r M . four 1991 g gure : margi an a o e anterolatera ( ; , A) 253208 Island d ZMUC Picar . r 20 , , M. , n r on mal 1995 P st ) alga pleopo ara straight a e Schott s d , e , K-AL-124 ovigerou sp. ; , Zanzibar types l Island dorsa ZMUC on turf l d nov. 2. tooth e e 2; . 0 , l Latera : , ( Mayotte mm view , E USN Aldabra s , st , ) , , on mal lackin , a 9 ; cora , ß l e alga CRU-1891 one M ( 13 margi B) e ¾ , pleopo , l S, , g 253209 Comor l on mal rubbl 9 antenna , turf ß 35 n e 24 st e of ß ovigerou d a ¾ 10 pereopo , S, e 2; pleotelso , , K-AL-24 o ¾ Mayotte on E, cora 46 ( thre Islands l F tooth ß ) 25 oute 12 mal l e d s ¾ m, on E, 1; n r e , , , , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 subapica copulator of agains width in pleotelson on wit basa margin Ž strongl lob Pereopo reticulatio wide CRU-1926 25 ovigerous Mnemb Zanzibar reef ovigerous two Zanzibar of W as attentio give Whethe style obliqu process Stenetrium ve 5. ester to femal mal the Septembe h e Etymology. Diagnosis. Othe Typ Etymology. Remarks. , cephalon 0 n l t r reachin Zanzibar singl it mm , triangula , width an t e th e male. n s tha y wit . , e tw n r a propodu spinos d r e generi wit e an d l Australia setos Operculu , , , material thes y e material Island is o n roughl n , proces h 1 16 14 st , , degre a d tooth h . wit , Murog juveniles namel style Ž draw a g on 4.5 of long r , pereopo th one m, m, e ve carpu e , CRU-1927 1995 c e anterio midlengt h r e Tw tw male – 10 on cephalo Cephalo , placement palm e stron s y t 20 relativel s 20 5. smal . , Th Th n Zanzibar . o teeth . , y m directe m, Eye o pentagonal of reachin 1 distall , anterio mesia ; o s appear to Septembe e e character Septembe mm th , ZMUC st character H , ZMUC, Reef d . no of l r 21 g specie speci setos variabilit a s sam e Typica olotype th ; darke h 1 hal teet reniform n t CRU-1903 , distall n e femal y l Septembe y d , of propodu on , produced dactylu g r Asellot an surface e . wit Stenetriu s unusua lon , Murog Ž f Zanzibar e laterally h tapere Stenetriu s , c e 9 dat to jus margi posterio d of carpu r tw ß l s is on h r one , r s g nam 13 y e Stenetrium patc pereonites : t hav y abou ar , anterolatera 1995 a separat 1995 an o name pereonit tapere roughl ¾ . tw ZMUC, beyon obliqu S, a s e , as see o d ovigerou l Red-brow n d s e r s e wit , h fro ; o su . m feature , Reef 35 , abou narro t ; 1995 refer havin to o produce . holotype n a m r propodu straight one (Ž 9 d middorsall juveniles Y 1. V ZMUC m h P zanzibaric ß margi simila d e m, d glauert amon 10 gure y cien e fo ara 8 narro Matemwe robus e , thi copulator th s . t proximalmos poorl , time r ¾ pentagonal w Zanzibar 20 g E, specie , s 1, s to e an as types th t s st l pereopo n 21 of n . r 6. India d g Septembe an specie to 25 e a , d ; w abou s on th s t lon , i pleopo pigmen 5.0 Eye 5 of y ) tw posterodistall USN typ CRU-1891 Nicholls specie abou thi st longe set d ove e m, mm justif y s a tubula de : a g pereonite o antenna – propodus Ž hav , n a sp. s at s ZMUC e y t 5. s ve Ž , CRU-1896 r 25 as Mnemb , specie Ocea demarcatin M locality d ne t reniform style 2 8 fro , r , bas mos d y s e 19 st t nov. mm propoda 1. m, 1 wide st t Septembe tha d r abou r a in a 2 a 253293 of m , 6 a in 1995 propoda e palm distall 1929 t in separat apex n t l 20 CRU-1893 n , s broa , time CRU-1902 , smal almos of s th wit teet Stenetrium , ; of thre a cora on , by wide t th y wit 2– latte Septembe e ; Zanziba . Island pleotelson , ; h 1. , e e , ZMUC dorsu h s , male d l int l y fro implyin Red-brow 7, dactylu e g san 6 h l male it wit on ovigerou teet r t longe low . e rathe l truncat fro r s palm o seta al Pleopo a 1995 m time an , genu tooth e longe d h subapica , , l broadl Ž h m ; m transvers , wit . Rottnes d Zanzibar on ve othe , rostru r . , e stron r r on Murog r , s o Murog s ; g of on Island 4. ove s di Nevertheless tha . h latte V . juveniles 1995 e ZMUC r whe Baw longe som d 5 s on is Pereopo n pereopo V r e Operculu it copulator pleotelson tha Matemwe, y mm r n 2 use g s ovigerou specie m doubtful scattere l r n rounde t strongl mos in . e 5.0 e greates . o e n spinos spinos o cuttin basall Island havin , r , folde Islan bands doub palm male wide 15 Reef Reef , thre 1455 tha mm, , t s on d d st m, of of m n d d d d y y 1 g g y a 1 e e e e s t t , , . , , , , , . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 1456 – 1 sperm ro male shorter Stenetrigus Stenobermuda F ig. bearin Pereopo bearin Pereopo w Diagnosis. 21 , of pleopod an (C carpu . tube. ; g g d ) pectinat rostru Stenetriu abou d abou d mal propoda Schultz 1 1 s in e Schultz in no 2. t t m pereopo male fou 10 Cephalo male e t Ke m narrow , l produced seta setae 1982 r palm , y , zanzibaric seta propoda , 1979 e propoda to d : . on ; e 58 , 1 n th Genus ( Pleopo on : triangular E . dactylu cuttin 904 e wit ; ) l cuttin India B. propodu a femal pal l ; h pal sp Sero Kensle Stenobermuda g m d anterolatera s . g edg m n e bearin 2 an nov v edg opercula . Ocea bearin e in an d Ey s . y e : propoda abou d g . mal e an ( n Wilson abou A) of g d specie . . e r 10 t thre l M. mal pleopo wit twic t toot – Schultz . . l fou 12 , Schott palm e e 1995 h s e . . r or h obliqu of dorsa dista obliqu d longe wel ; : fou . . 2; Stenobermuda , 77 ( e 197 D) l l ( l e . r . . F r e developed view fringe pectinat ommatidia ) 9 pectinat femal tha S. mal . ; n syzygus ( e . d e B) palm e pleopo pereopo e lamell . setae setae ; mal S. . antenna , ( Pereopo Barnard latte e brucei ; [Sout d ; a dactylu dactylu d pereopo 1; an [ 1 r Zanzibar] (G h d bearin dactylu sp. l , Africa d ) toot shor 194 s s mal nov. d 1 0 in 1; h g e s t ) ] Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 anterolatera acute Baw F fringe demarcatin propodu anterio merus lengt ig. 2.5 Typ Diagnosis. 22 e h , mm, d , Island . subequa r surpassin e seta carpu s margins Stenobermud material subcircular one g e l , on tooth palm s Zanzibar l an Cephalo g cuttin to ; 2. . d , carpu antenna . 2 basa Ey latte propodu H mm a , g olotype wit , e bruce 25 l margin s , n weakl r (C width h sam Asellot havin m, havin l wit Stenobermud thre ) i tooth s pereopo 21 sp e h y : , havin . e dat . g pigmented g wit ZMUC, Pereopo antenna Septembe a stou nov cluste ; fou a fro h rostru d g as (Ž . t singl : m r 2; fringe stou gure r ( holotype slende a l th ( A) d of D) m e bruce toot r , e 2, t 3. consistin latera 1995 22 mal India Ž mal d elongat carpu narrowl 1 ve r h ) seta mm i e e fringe . . sp. low stou l pleopo n P dorsa e s tooth , ara Ocea on 3. e st nov. g , y t d a 5 of subacute seta types: fringe l posterio triangular d time setae CRU-1894 . view n fou 1. Pereopo e s r on d ; ; ZMUC longe ommatidia ( dactylu seta , r B) mesia anterolatera margin , d e , reachin mal r 1 san tha posterodistally , male l s on e surfac d , n havin . pereopo e robus Pleotelso wide fro g , ovigerou ischium beyon l m g e , toot t 1457 nea reef wit fou d set 1; n h h d a r r s ; , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:54 24 July 2008 S bot recorde an th wes margin tw 1458 F B B B B B References Atol Island Ž th Institution Sciences Richmond acknowledg Queenslan Michell o Dr Museum th Nationa Clar Madagascar we additio Acknowledgements specie of ve V . resi olst eddard arnard arnard arnard e e e d o s brucei Wate h Remarks Thi Etymology. ar t Smithsonian Kuwai India Director collectin k l robus Stephe , possesse India ad e W Asellota 1873 Africa identi addition addition to posses an s s E., W s n ; d ver e alon ashington , Foundatio r th l , d , dactylu , , , , . wor alker K. to n va fro Museu F. an K. – K. 1973, K. e Universit Copenhagen t n in d ’ , y t Ž 76 n s n Ocean marin n seta th catio e g s S. d . E. , Ocea m an Museum g , an gratefu k H. Museu O ) H. s . s s th H. Grab durin wit de e wit muc , , Nationa Atmospheri and Par fro season to Y woul d to Bermud a d , Stenobermud e , 1886 , anonymou e Ms s Maresi , r ’ n m 1914 1940 ce h 1925 s e graniti h Th m th 112 narrowe n on biunguiculate t th Saud Merw . th h of n Aldabr K Isopoda thank m e 2, g of severa e Thi y at of e , l d Madang e e Lind ensley narrowe , fo ( , hithert lis posterio an Repor , th , s Repor , to 1) of 32 Natura specie T generou no Stil Contribution a Contribution th Contribution l , r t i s Australia on e a anaidacea d , c e wh , of barringtoni Museu th logisti is Arabi t e s IIOE 164 a Da ( 381– l l of Seychelles Joh a r S. , c hav th e Aldabr s , forme t Bay t Isopoda other o a o W Annal , Researc propodu B. s on collector – Research a of r following e th acutirostrat B. recorde Papu r contribute muc r 174 n l ard 543. is , e ; s assistanc es a c e syzygu th History , margin propod th 1999 m , Dr Kensle McCai possibl Sout s , name , provisio Sout suppor . e r s wit , e durin h a Salaa a wh describe Isopod s Voyag , a of of s Assistan Isopod s al Kristia , h Ne Annal large to d Atol . h to n. to s l Tw o s h th h s Natura , Program marin d : ; smal w th of of i g , of y n e th Africa mad W e m th e Africa g. propodu Mr ( d t o Smithsonia e fo of a e Sout l Barnard a an r e Guinea n wh s th give withou a e ellington an th ric of th ne durin pereopo crustacea n. 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