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Folia conchyliologica , n°17 , Mars 2012 Weyrauch’s type localities: a clarification; with illustrations of types of Orthalicoidea (Mollusca, Gastropoda, Stylommatophora) in the Tucumán museum 1 by Abraham S.H. BREURE Abstract. — Additional, more precise locality data are provided for the species described by Wolfgang K. Weyrauch from South America. Most of the holotypes in the Tucumán museum pertaining to the Orthalicoidea are illustrated. Additionally, a list is provided of new species described by various authors and based on Weyrauch’s material, illustrating the importance of the Weyrauch collection. Key words . — Weyrauch, Argentina, Bolivia, Brazil, Ecuador, Peru, georeferenced localities. Introduction Wolfgang K. Weyrauch worked for many years in Peru and Argentina. He collected many shells, especially in central and northern Peru; based on his collection he described 197 new taxa, which have been listed by BARBOSA et al. (2008). However, the type localities mentioned in that paper only give “C. Peru”, etc., and are too vague to be useful for further references. During revisionary work I found that a more precise (geo)reference to these localities is often needed to interprete the status of these taxa. The following list is therefore an addition to the publication mentioned; only those (164) taxa are herein mentioned for which Weyrauch published type localities (thus excluding new names and taxa above the species level). A best guess has been added in notes (bold numbers in parenthesis; see last section) whenever I was unable to locate the type locality with certainty, e.g. when Weyrauch used local names for places that are not included in the gazetteers used. The localities as given by Weyrauch are all translated into English. Additonally, most of the holotypes of the Orthalicoidea (sensu BREURE & R OMERO , in press) present in the collection of the Tucumán museum are illustrated (referred to by a figure number in parenthesis at the end of the data for the taxon); these species were classified with the families Amphibulimidae and Orthalicidae by BARBOSA et al. (2008). For the types in the Senckenberg collection, see NEUBERT & J ANSSEN (2004). The Weyrauch collection remains an important source of material for our understanding of Neotropical snails, especially species from Peru, as is shown by the section listing 55 (sub)species based on his material but described by others, and other papers which made use of his collection ( e.g. , PILSBRY , 1944a-c, 1945, 1949; HAAS , 1947, 1948, 1951, 1952, 1955, 1959, 1966; ZILCH , 1954; LOOSJES , 1957; LOOSJES & L OOSJES -VAN BEMMEL , 1966; BREURE & E SKENS , 1977, 1981; BREURE , 1978; SALAS -OROÑO , 2007; PIZÁ & C AZZANIGA , 2009). Moreover, Weyrauch has distributed many shells under manuscript names to various museums in the world. Some of this material has been revised (BREURE , 2008; BREURE & N EUBERT , 2008). In the Tucumán museum additional, unpublished material has been found which awaits a further, critical review. Coordinates are according the GNS gazatteer (http://geonames.nga.mil/ggmagaz/ ); in some cases the Fuzzy Gazetteer (http://isodp.hof-university.de/fuzzyg/query/ ) has been used. Elevations, if not given by the author, have been extracted from Google Earth (v5.2.1.1588, http://earth.google.com ). Abbreviations used for type depositories are: ANSP, Academy of Natural Sciences, Philadelphia, U.S.A.; FML, Fundación Miguel Lillo, Tucumán, Argentina; FMNH, Field Museum of Natural History, Chicago, U.S.A.; MACN, Museo Argentino de Ciencias Naturales, Buenos Aires, Argentina; MRCN, Museu Rio-Grandense de Ciências Naturais, Porto Alegre, Brazil; SMF, Senckenberg Natur-Museum, Frankfurt am Main, Germany. Taxa belonging to the Orthalicoidea are re-classified according to the systematic arrangement presented in BREURE & R OMERO (in press). Shell heights in the legends are abbreviated with H. 1 A.S.H. Breure, Netherlands Centre for Biodiversity Naturalis, P.O. Box 9517, NL-2300 RA Leiden, the Netherlands ([email protected] ) 4 Folia conchyliologica , n°17 , Mars 2012 Published names by Wolfgang K. Weyrauch acobambensis, Drymaeus WEYRAUCH , 1967b: 482, pl. 3 figs 24-25. Type locality: Peru, [Dept. Junín] Río Tarma valley, between Tarma and Arcobamba, Huailahuichán [11°21'30" S 075°36'11" W], 2900 m. Holotype SMF 155694. Bulimulidae. aequistriata, Gracilinenia WEYRAUCH , 1956a: 110, pl. 6 fig. 2. Type locality: Peru, [Dept. Junín] Chanchamayo valley, on the road from Hacienda Naranjal [11°08'37" S 075°24'00" W] near the village of San Ramón to the mine “Pichita-Caluga” (1) ; elevation not measured, but probably about 1300 m. Holotype FML 1531. agitata, Pseudoglandina WEYRAUCH , 1967b: 486, pl. 4 fig. 53. Type locality: Peru, [Dept. Junín] Chanchamayo valley, between La Merced [11°03'21" S 075°19'43" W] and San Ramón [11°07'17" S 075°21'11" W], 1100 m. Holotype FML 1066 (2) . Bulimulidae. agueroi, Bostryx (Peronaeus) WEYRAUCH , 1960c: 126, pl. 12 figs 39-41. Type locality: Peru, [Dept. Lima] right margin of Río Cañete, road from Cañete to Yauyos, Fundo Yacca [12°30'41" S 075°55'04" W], 2300 m. Holotype SMF 162150. Bulimulidae. aguilari, Bostryx (Bostryx) WEYRAUCH , 1967a: 349, pl. 1 figs 2-6. Type locality: Peru, [Dept. Lima] ‘lomas’ de Amancaes [12°00'38" S 077°02'25" W], 200-250 m. Holotype SMF 162163. Bulimulidae. albicolor, Peruinia WEYRAUCH , 1957a: 13, pl. 1 fig. 11. Type locality: Peru, [Dept. Huánuco] left margin of Río Tulumayo, 23 km NE Tingo Maria, near bridge in the road to Pucallpa, 670 m [09°11'25" S 075°57'26" W (3) ]. Holotype SMF 155719. albocostata, Temesa (Temesa) WEYRAUCH , 1964b: 270, pl. 1 figs 1-2. Type locality: Peru, [Dept. Lima] right margin of Río Alis, near Alis [12°16'43" S 075°47'13" W], 3300 m. Holotype SMF 162108. altispira, Euglandina WEYRAUCH , 1960a: 25, pl. 4 fig. 21. Type locality, Peru, [Dept. Cajamarca,] 25 km NE Cajamarca [sic, (4) ], on path from Sócota [06°18' S 078°44' W] to San Andrés [de Cutervo; 06°13' S 078°40' W], 2400 m. Holotype SMF 162003. altorum, Mesembrinus (Mormus) expansus WEYRAUCH , 1958: 129, pl. 7 fig. 12-13. Type locality: Peru, [Dept. Junín] Chanchamayo valley, right side of Río Chanchamayo, between La Merced and San Ramón, above Puente Herrería, Quimiri Sur [11°05'01" S 075°17'24" W, 1430 m], 1100-1300 m. Holotype SMF 156295. Bulimulidae. altorum, Systrophia (Systrophia) WEYRAUCH , 1967a: 428, pl. 6 fig. 80. Type locality: Peru, [Dept. Junín] Chanchamayo valley, Pichita Caluga (1) , 2200 m. Holotype FML 10668. andivagus, Naesiotus WEYRAUCH , 1956b: 56, pl. 1 fig. 2. Type locality: Peru, [Dept. Lima] Río Huaura valley, between Huaura and Cerro de Pasco, Cabracancha [10°49'55" S 076°53'23" W], 2200 m. Holotype FML 1461 (2) . Bulimulidae. angelmaldonadoi, Bostryx (Bostryx) modestus WEYRAUCH , 1960a: 31, pl. 3 figs 11-12. Type locality: Peru, [Dept. Lima] left side of Río Lurin, ‘lomas’ E of Pachacamac [12°15'14" S 076°47'38" W], 450 m. Holotype SMF 155595. Bulimulidae. angiportus, Newboldius WEYRAUCH , 1960b: 53, pl. 8 figs 5-6. Type locality: Peru, [Dept. Junín] tributary of Río Chanchamayo, 2 km of Pichita Caluga Mine (1) , 2200 m. Holotype SMF 162045. Bulimulidae. angispira, Bostryx (Bostryx) obliquiportus WEYRAUCH , 1960c: 123, pl. 11 fig. 19. Type locality: Peru, [Dept. Lima] left side of Río Cañete, 5 km of Laraos, [Cerro] Quichao [12°20'45" S 075°48'53" W], 3450 m. Holotype SMF 162183. Bulimulidae. angustus, Bulimulus (Bulimulus) vesicalis WEYRAUCH , 1966: 45, fig. 5. Type locality: Brazil, Edo. Rio Grande do Sul, São Leopoldo, near “Clube Airmore” stadium [29°46'52" S 051°09'25" W]. Holotype MRCN 1576a. Bulimulidae. araozi, Bulimulus (Bulimulus) WEYRAUCH , 1956c: 149, pl. 11 fig. 8. Type locality: Peru, [Dept. Huánuco] left margin of Rio Huallaga, near Tingo Maria [09°19'17" S 076°00'55" W], 670 m. Holotype SMF 155304. Bulimulidae. argentinus, Pupoides (Ischnopupoides) chordatus WEYRAUCH , 1964a: 37, fig. 1. Type locality: Argentina, Prov. Jujuy, near Tilcara [23°34' S 065°22' W], Garganta del Diablo, 2900 m. Holotype FML 522a. bambamarcaensis, Naesiotus (Naesiotus) WEYRAUCH , 1960a: 37, pl. 6 fig. 38. Type locality: Peru, [Dept. Cajamarca] Río Llaucan valley, 4 km NW Bambamarca [06°41' S 078°32' W], Cerro Machaipungo, 3000 m. Holotype SMF 156220. Bulimulidae. 5 Folia conchyliologica , n°17 , Mars 2012 basiplanata, Epiphragmophora WEYRAUCH , 1960a: 43, pl. 6 fig. 39. Type locality: Peru, [Dept. Pasco] road from Cerro de Pasco to Huánuco, valley of Río Payanchan, 3800 m (5) . Holotype SMF 162033. beltrani, Bostryx (Peronaeus) agueroi WEYRAUCH , 1964a: 52, figs. 9-10. Type locality: Peru, [Dept. Lima] left side of Río Cañete, between Laraos [12°20'48" S 075°47'11" W] and Tintín [12°18'10" S 075°48'40" W], 3450 m. Holotype SMF 162152. Bulimulidae. bequaerti, Drymaeus WEYRAUCH , 1956c: 154, pl. 11 figs 12-14. Type locality: Peru, [Dept. Huánuco] near Tingo Maria, right side of Río Monzón near its confluence with Rio Huallaga [09°19'17" S 076°00'55" W], 670 m. Holotype SMF 155309. Bulimulidae. bequaerti, Temesa (Neniatracta) WEYRAUCH , 1957: 25, pl. 1 fig. 12. Type locality: Peru, [Dept. Cajamarca] 1 km NE Bambamarca [06°41' S 078°32' W], 2800 m. Holotype SMF 155720. bermudezae, Bostryx (Pseudoperonaeus) WEYRAUCH , 1958: 111. pl. 9 figs 38-40. Type locality: Peru, [Dept. Lima] Rio Cañete valley, Laraos [12°20'48" S 075°47'11" W], 3600 m. Holotype SMF 156350. Bulimulidae. bicolor, Naesiotus (Naesiotus) WEYRAUCH , 1967a: 408, pl. 6 fig. 83. Type locality: Peru, [Dept. Cuzco] Sicuani [14°16'10" S 071°13'34" W, 3500 m], 3200 m. Holotype FML 3987 (Pl. 2 figs 17-18) . Bulimulidae. birabenorum, Bostryx (Lissoacme) WEYRAUCH , 1965a: 71, figs 1-3. Type locality: Argentina, Prov. Tucumán, near Tafí del Valle [26°51'10" S 065°42'35" W], El Infernillo, 3042 m.
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    FOLIA Folia Malacol. 24(3): 111–177 MALACOLOGICA ISSN 1506-7629 The Association of Polish Malacologists Faculty of Biology, Adam Mickiewicz University Bogucki Wydawnictwo Naukowe Poznań, September 2016 http://dx.doi.org/10.12657/folmal.024.008 PATTERNS OF SPATIO-TEMPORAL VARIATION IN LAND SNAILS: A MULTI-SCALE APPROACH SERGEY S. KRAMARENKO Mykolaiv National Agrarian University, Paryzka Komuna St. 9, Mykolaiv, 54020, Ukraine (e-mail: [email protected]) ABSTRACT: Mechanisms which govern patterns of intra-specific vatiation in land snails were traced within areas of different size, using Brephulopsis cylindrica (Menke), Chondrula tridens (O. F. Müller), Xeropicta derbentina (Krynicki), X. krynickii (Krynicki), Cepaea vindobonensis (Férussac) and Helix albescens Rossmässler as examples. Morphometric shell variation, colour and banding pattern polymorphism as well as genetic polymorphism (allozymes and RAPD markers) were studied. The results and literature data were analysed in an attempt to link patterns to processes, with the following conclusions. Formation of patterns of intra- specific variation (initial processes of microevolution) takes different course at three different spatial scales. At micro-geographical scale the dominant role is played by eco-demographic characteristics of the species in the context of fluctuating environmental factors. At meso-geographical scale a special part is played by stochastic population-genetic processes. At macro-geographical scale more or less distinct clinal patterns are associated with basic macroclimatic
  • Abstract Volume

    Abstract Volume

    ABSTRACT VOLUME August 11-16, 2019 1 2 Table of Contents Pages Acknowledgements……………………………………………………………………………………………...1 Abstracts Symposia and Contributed talks……………………….……………………………………………3-225 Poster Presentations…………………………………………………………………………………226-291 3 Venom Evolution of West African Cone Snails (Gastropoda: Conidae) Samuel Abalde*1, Manuel J. Tenorio2, Carlos M. L. Afonso3, and Rafael Zardoya1 1Museo Nacional de Ciencias Naturales (MNCN-CSIC), Departamento de Biodiversidad y Biologia Evolutiva 2Universidad de Cadiz, Departamento CMIM y Química Inorgánica – Instituto de Biomoléculas (INBIO) 3Universidade do Algarve, Centre of Marine Sciences (CCMAR) Cone snails form one of the most diverse families of marine animals, including more than 900 species classified into almost ninety different (sub)genera. Conids are well known for being active predators on worms, fishes, and even other snails. Cones are venomous gastropods, meaning that they use a sophisticated cocktail of hundreds of toxins, named conotoxins, to subdue their prey. Although this venom has been studied for decades, most of the effort has been focused on Indo-Pacific species. Thus far, Atlantic species have received little attention despite recent radiations have led to a hotspot of diversity in West Africa, with high levels of endemic species. In fact, the Atlantic Chelyconus ermineus is thought to represent an adaptation to piscivory independent from the Indo-Pacific species and is, therefore, key to understanding the basis of this diet specialization. We studied the transcriptomes of the venom gland of three individuals of C. ermineus. The venom repertoire of this species included more than 300 conotoxin precursors, which could be ascribed to 33 known and 22 new (unassigned) protein superfamilies, respectively. Most abundant superfamilies were T, W, O1, M, O2, and Z, accounting for 57% of all detected diversity.
  • Gastropoda, Clausiliidae)

    Gastropoda, Clausiliidae)

    SOME ANATOMICAL, SYSTEMATICAL AND GEOGRAPHICAL DATA ON NENIINAE (GASTROPODA, CLAUSILIIDAE) by F. E. LOOSJES and A. C. W. LOOSJES-VAN BEMMEL Vossenlaan 4, Wageningen, Netherlands (With 38 text-figures and 1 plate) I. INTRODUCTION Up to the moment more than one hundred species and subspecies of Neniinae are known from Central and South America. They are incorporated into 22 genera and subgenera, almost exclusively based on characters of the shells. Only little is known of the anatomy of the animals (Binney, 1871; Binney & Bland, 1871; Baker, 1961; Hesse, 1925; Scott, 1954; Loosjes, 1957; Pilsbry, 1926; Polinski, 1922; Zilch, 1953, 1959). These authors described the radula and/or the genital organs of one or several species. Thus from eleven species the radula has been described and from five the genital apparatus. By the courtesy of Prof. Dr. W. Weyrauch we were able to study the anatomical features of a relatively large number (38) of Peruvian and Argentine Neniinae. We wish to express our most cordial thanks to him for his generosity and also to Dr. B. Hubendick, who provided us with a sample of Nenia tridens (Chemnitz) from Puerto Rico. In Thiele's (1929/31) classification the American Neniinae are grouped into three genera, Nenia, Peruinia and Temesa, based on characters of the shells and of the radula. It turned out impossible to fit the information we found, into this system as it indicates the presence of many more units. In this survey we will follow Zilch's (1960) classification, which enumerates a number of genera and subgenera, based on conchological characters, although the order will be altered according to the results obtained (conf.
  • Spermatogenesis and Sperm Ultrastructure in the Land Slug Limax Flavus (Gastropoda, Pulmonata) from Egypt

    Spermatogenesis and Sperm Ultrastructure in the Land Slug Limax Flavus (Gastropoda, Pulmonata) from Egypt

    Advances in Biological Research 7 (6): 253-265, 2013 ISSN 1992-0067 © IDOSI Publications, 2013 DOI: 10.5829/idosi.abr.2013.7.6.76164 Spermatogenesis and Sperm Ultrastructure in the Land Slug Limax flavus (Gastropoda, Pulmonata) from Egypt Irene Sameh Gamil Department of Zoology and Entomology, Faculty of Science, Helwan University, Cairo, Egypt Abstract: Spermatogenesis and sperm ultrastructure are examined and described for the first time in the garden slug Limax flavus (Gastropoda, Pulmonata, Stylommatophora) in Egypt. Spermatogonia show round nuclei with patchy heterochromatin. The primary spermatocytes are characterized by the presence of synaptonemal complexes. The secondary spermatocytes are reduced in size and contain less cytoplasm and clustered stacks of Golgian cisternae with small proacrosomal vesicles lying close to them. During spermiogenesis, electron-dense plaques develop at both the future anterior and posterior poles of the nuclear surface. These plaques determine the apparent antero-posterior axis of the spermatid, the distal plaque indicates the future anterior part of the cell and the basal one the posterior part where an abundant number of mitochondria are aggregated. The mature spermatozoon shows the characteristic sperm features: an acrosomal vesicle supported by an acrosomal pedestal; a helically keeled nucleus, a neck region and a complex elongate middle piece featuring paracrystalline and matrix layers sheathing the axoneme, coarse fibers and glycogen helices. Key words: Spermiogenesis Spermatozoa Hermaphrodite snail Mollusca Pulmonate INTRODUCTION study, a detailed description of spermatogenesis process till the formation of mature autospermatozoa The bulk of the land Gastropoda consists of is presented. Studies on the spermatozoa and Pulmonata including the most important order spermatogenesis have been used extensively to Stylommatophora which is one of the most diverse and explore taxonomic and phylogenetic relationships of economically significant groups of living molluscs [1].