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BIOLOGICAL SCIENCES

The biology major provides a unified In the College o f Human Ecology, the natural 5. Enrollees in those courses in the curriculum for undergraduates enrolled in sciences distribution requirement is for at least 6 biological sciences in which animal use either the College of Agriculture and Life credits selected from BIO G 109-110, 101 and is a component may, at the professor’s Sciences or the College o f Arts and Sciences. 103 plus 102 and 104, 105-106 or 107-108 or discretion, be asked to sign copies of Courses in biological sciences are integral to from specified courses in chemistry or physics. this statement (USE OF ANIMALS . . .) many disciplines and are basic requirements at the first meeting o f the course.” Switching between BIO G 109-110 and either in many schools and colleges at Cornell. BIO G 101-104 or 105-106 at midyear may Graduate study in the biological sciences is not be possible because o f variation in administered by more than 15 specialized presentation o f topics. Students must receive fields within the Graduate School, as permission o f the instructor to switch ADVANCED PLACEMENT described in the Fields o f Study catalog at sequences. Taking sequences in reverse order For information on credit for advanced www.gradschool.cornell.edu/?p=38. is strongly discouraged in BIO G 101-104 but placement in biological sciences, see allowed in BIO G 105-106. www.biology.cornell.edu/advising/ap.html.

ORGANIZATION Many different departments participate in the USE OF ANIMALS IN THE BIOLOGICAL THE MAJOR biology major. SCIENCES CURRICULUM: CORNELL The major of biological sciences is available to Student services are provided by the Office of UNIVERSITY students enrolled in either the College o f Undergraduate Biology (OUB), www.biology. Agriculture and Life Sciences or the College of cornell.edu. Located in Stimson Hall, the Students wishing to enroll in biology ( “BIO”) Arts and Sciences. The undergraduate program professional and student advisors provide courses should know and understand the is coordinated for students in both colleges by academic and career advising, as well as help following criteria relative to the use o f animals the Office of Undergraduate Biology. By undergraduates find research opportunities on in the teaching program, as passed by the completion o f the sophomore year, all campus. Advisors in the OUB also follow the faculty o f the Division o f Biological Sciences students who intend to major in biological progress o f biology majors and work closely in 1988, and reaffirmed in 1997: sciences must declare the major and a with faculty advisors. Additional services and 1. “Live animals will be used for teaching program o f study through the Office o f resources o f the Biology Center include in certain courses in the biological Undergraduate Biology, in 216 Stimson Hall. tutoring, lecture tapes, examination files, and sciences. Some animals will require Whenever possible, students should include extensive information on summer research humane euthanasia after they have the introductory biology, chemistry, and opportunities and graduate programs. The been used for teaching. mathematics sequences in their freshman center has comfortable areas for studying and 2. Courses bearing the “BIO” description schedule and complete the organic chemistry relaxing. conform to the rules for the care o f lecture course in their sophomore year. The Shoals Marine Laboratory, a cooperative such animals as outlined in Guiding Biology majors should regularly monitor their venture with the University o f New Principles in the Care and Use of progress in the major, and should assess as Hampshire, is located on Appledore Island in Animals (as approved by the Council realistically as possible the likelihood o f the Gulf o f Maine. Its base office in Stimson of the American Physiological Society), achieving at a level that is consistent with Hall provides academic advising for students the Guide for the Care and Use o f their academic and personal goals. Weak interested in the marine sciences and Laboratory Animals (DHEW publication performance in core courses, particularly after administers the SEA Semester program for 86-23, revised 1996; see p. 7, Courses the freshman year, may indicate a need to Cornell students pursuing studies at Woods o f Study), the Animal Welfare Act, and reevaluate aptitude and genuine interest in the Hole, Mass., or aboard the schooner Robert C. the N ew York State Public Health Law. major. Students with questions, particularly Seamans or brigantine Conuith Cramer. Within these regulations, and in with concerns about their ability to complete keeping with the principle of Academic the major, are encouraged to consult with Freedom o f the Faculty, the use o f their biology advisor and to take advantage of animals to aid in teaching any the advising and counseling resources o f the DISTRIBUTION REQUIREMENT biological sciences discipline is at the Office of Undergraduate Biology as well as discretion o f the professor in charge. those o f the university and their college. In the College o f Agriculture and Life Sciences, the Physical and Life Sciences 3- Each course, as well as research The requirements for the biological sciences distribution requirement is a minimum o f 18 projects, in which animals are used major are listed below. Requirements 1-9 must credits, including at least 6 credits o f receives a formal review annually by the be taken for a letter grade. Courses taken for introductory biology satisfied by BIO G 109- Cornell University Institutional Animal the program o f study should be taken for a 110, 105-106, or 101 and 103 plus 102 and Care and Use Committee (IACUC). letter grade unless the course is offered for 104, or 107-108. S-U grades only or if the student’s advisor 4. Any concerns regarding the use of live grants permission. For Students in the College o f Arts and animals in teaching should be Sciences, all biology ( “BIO”) courses can be addressed first to the faculty member 1. Introductory biology for majors used toward fulfillment o f the biological responsible for that course. He or she (one year): BIO G 101 and 103 plus sciences distribution requirement. Please see is required to be in compliance with all 102 and 104, or 105-106. BIO G 107- the Arts and Sciences “Distribution applicable regulations and guidelines. 108, offered during the eight-week Requirements” section o f this catalog for Alternatively, students may choose to Cornell summer session for 8 credits, further details. The following courses are address their concerns to the director also satisfies the introductory biology especially suitable for the distribution o f the Cornell Center for Research requirement for majors. requirement because they have no Animal Resources, Dr. Michele Bailey, 2. General chemistry: CHEM 207-208 or prerequisite: BIO G 101-104, 105-106, 107-108, at 253-3520. The director may initiate 215. Students who, via advanced 109-110, 170; BIOAP 212; BIOEE 154, 207, discussion with the faculty member placement, take only CHEM 208 or 275; BIOGD 184; BIOMI 192; BIONB 111; responsible for a particular course only 215 should be aware that some BIOPL 240, 241. without involving the student if he or professional and graduate schools she would prefer to remain anonymous. require 8 credits o f general chemistry. 156 BIOLOGICAL SCIENCES - 2 I ill] 8

These students may wish to take both programs o f study and their requirements are in biochemistry should take PHYS 207- CHEM 215 and 208 or 215 and 216. listed below: 208 and should consider taking CHEM Students may wish to consult with their 389-390 and its prerequisites. They should 1. Animal Physiology: BIOAP 311 faculty advisor or advisors in the Office be sure to complete CHEM 207-208 or Introductory Animal Physiology, BIOAP o f Undergraduate Biology for further 215-216 during their freshman year. 316 Cellular Physiology, plus a minimum clarification. o f 7 credit hours selected from the • Biology majors in the College o f 3. College mathematics (one year): one following lecture and laboratory courses, Agriculture and Life Sciences who select semester o f calculus (MATH 106, 111, o f which at least 4 credit hours must be a the biochemistry program are allowed to 191, or their equivalent) plus one laboratory course. take up to 61 credit hours in the endowed semester selected from the following: colleges because of the high number of a. Lecture courses: BEE 454 Physiological required endowed courses for this a. a second semester o f calculus Engineering; AN SC 300 Animal program o f study. (MATH 112, 192, or their Reproduction and Development; equivalents). AN SC 410 Nutritional Physiology and 3. Computational Biology: Computation Metabolism; AN SC 427 Fundamentals has become essential to biological b. a course in finite mathematics o f Endocrinology; BIO G 305 Basic research. Genomic databases, protein (MATH 105). Immunology; BIOAP 214 Biological databanks, MRI images o f the human c. a course in statistics (BTRY 301, Basis o f Sex Differences; BIOAP 458 brain, and remote sensing data on MATH 171, AEM 210, ILR 212, Mammalian Physiology; BIOAP 475 landscapes contain unprecedented PSYCH 350, PAM 210, ECON 319, Mechanisms Underlying Mammalian amounts o f detailed information that are ECON 321, SOC 301). Developmental Defects; BIOAP 489 transforming almost all o f biology. Mammalian Embryology; BIOBM 407 4. Organic chemistry: CHEM 257 and Problems investigated by computational Nature o f Sensing and Response: 251, or 357-358 and 251, or 357-358 biologists include topics as diverse as the Signal Transduction in Biological and 301, or 359-360 and 251, or 359- genetics o f disease susceptibility; Systems; BIOBM 437 Eukaryotic Cell 360 and 301. comparing entire genomes to reveal the Proliferation; BIOGD 385 evolutionary history o f life; predicting the 5. Physics: PHYS 101-102, 207-208,* or Developmental Biology; BIOGD 400 A structure, motions, and interactions o f 112-213.* Those who take PHYS 112- Genomics Approach to Studying Life; proteins; designing new therapeutic 213 are advised to complete PHYS 214 BIOGD 401 Genomic Analysis; BIOGD drugs; modeling the complex signaling as well. 483 Molecular Aspects o f mechanisms within cells; predicting how Development; BIOGD 610 Genomes 6. Genetics: BIOGD 281. ecosystems will respond to climate as Chromosomes; BIOGD 612 change; and designing recovery plans for 7. Biochemistry: BIOBM 330, or 331 and Overview of Model Genetic endangered species. The computational 332, or 333. Organisms; BIONB 322 Hormones and biologist must have skills in mathematics, Behavior; BIONB 325 Neurodiseases- 8. Evolutionary biology: BIOEE 278 or statistics, and the physical sciences as well Molecular Aspects; BIONB 326 The BIOPL 448. Note: BIOPL 241 Botany is as in biology. A key goal in training is to Visual System; BIONB 492 Sensory a prerequisite course to BIOPL 448. develop the ability to relate biological Function; NS 331 Physiological and processes to computational models. 9. A program of study selected from the Biochemical Bases o f Human Cornell faculty work primarily in four outline below. Nutrition. subareas o f computational biology: Although not required for the biological b. Laboratory courses: BEE 454 biomolecular structure, bioinformatics and sciences major, a course in statistics is Physiological Engineering; AN SC 301 data mining, ecology and evolutionary recommended for all biology students. Animal Reproduction and biology, and statistical and computational Students should consult their faculty advisors Development; BIO G 401 Introduction methods for modeling biological systems. when choosing appropriate courses in to Scanning Microscopy; BIO G 403 Specific topics o f study include DNA statistics. Transmission Electron Microscopy for databases, protein structure and function, Note: Core courses cannot count toward the Biologists; BIOAP 413 Histology: The computational neuroscience, program o f study requirements. Biology o f the Tissues; BIOAP 319 biomechanics, population genetics, and Animal Physiology Laboratory; BIOAP management o f natural and agricultural 416 Cellular Physiology and Genomics systems. Programs of Study and Requirements Laboratory; BIOBM 440 Laboratory in Beyond core skills in mathematics, As noted in the list o f requirements above, Biochemistry and Molecular Biology; physical sciences, and biology, the students accepted into the biological sciences BIONB 491 Principles of computational biology program o f study major must choose a program o f study. Neurophysiology. Whereas the core requirements o f the biology requires additional course work in curriculum provide the common foundation 2. Biochemistry: BIOBM 440 Laboratory in mathematics and computer programming, deemed essential for all biology majors, the Biochemistry and Molecular Biology; a “bridging” course aimed at connecting role o f the program o f study is to provide physical chemistry (CHEM 287-288 or biology to computation, and an advanced either a concentration in a particular area o f 389-390 or 389-288); 6 credits o f organic course where the theoretical/ biology or, in the case o f the general biology chemistry (CHEM 357-358 or CHEM 359- computational component o f one aspect program o f study, a survey o f biology that is 360); plus one o f the following two o f biology is studied. Students should broad but not superficial. The program o f alternatives. Students wanting to maximize enroll in the more rigorous courses in the study requirement can be met by taking 13 to their background in chemistry should take physical and mathematical sciences and 15 credit hours o f courses chosen by the CHEM 300 Quantitative Chemistry and may wish to take additional courses in student in consultation with his or her biology CHEM 301 Honors Experimental these areas. Chemistry I. Students wanting to gain advisor. Programs o f study for particular Computational biology has applications as further depth in biochemistry and related subject areas are designed by faculty members broad as biology itself. The problems o f disciplines should take BIOBM 432 Survey specializing in the subject. Typically, the interest and the tools available to study of Cell Biology and either CHEM 300 program o f study consists o f one or more them are constantly evolving, so students Quantitative Chemistry or CHEM 252 courses that provide foundation in the subject are encouraged to gain fundamental skills Elementary Experimental Organic and a list o f optional courses from that area or that will serve them throughout their Chemistry. Students choosing the first related areas, many o f which are at an careers. There is great, and increasing, alternative are encouraged to take BIOBM advanced level (300 or higher). Because demand for research scientists and 432. biology is an experimental science, most technical personnel w ho can bring programs o f study require one or more Notes: mathematical and computational skills to laboratory courses. The laboratory the study o f biological problems. The • CHEM 288 is designed for biologists. Five requirement in some programs o f study can program is also an excellent preparation hours o f biochemistry are recommended be met by participation in the independent for graduate study in any area o f biology (331 and 332, or 330 and 334 or 333 and research course (BIO G 499). The possible or computational biology. 334). Students interested in graduate work THE MAJOR 157

Required Courses for Program of recommended for bridging course CS 473 Ecology o f Agricultural Systems; Study in Computational Biology 321/BIOBM 321. MATH 221 Linear NTRES 420 Forest Ecology; BIOSM 413 Algebra and Calculus, MATH 231 Research in Marine Biology; NTRES a. One course in computer programming Linear Algebra, or MATH 420 422 and 423 Wetland Ecology and (CS 100 Introduction to Computer Differential Equations and Dynamical Management, lec and lab; BIOMI 418 Programming or BEE 151 Introduction Systems is recommended for bridging Microbial Ecology; CSS/HORT 466 Soil to Computing) course BIOEE 460. Ecology; BIOP1/ENTOM 440 b. One additional course in mathematics Phylogenetic Systematics; BIOPL 447 • Courses may not be used to (MATH 221 Linear Algebra and Molecular Systematics; BIOPI/ENTOM simultaneously satisfy two different Calculus; or MATH 231 Linear Algebra; 453 Principles and Practice o f requirements. For example, BTRY 408 or MATH 294 Engineering Mathematics Historical Biogeography; BIOEE/EAS can be used to satisfy either II; or MATH 420 Differential Equations 479 Paleobiology; BIOGD 484 requirement (2) or requirement (4), and Dynamical Systems; or BTRY 408 Molecular Evolution. but not both. Theory of Probability; or BTRY 421 Note: Students also are encouraged to Matrix Computation) • Students who use BTRY 408 to fulfill gain experience in some aspect of the additional mathematics c. A bridging course, i.e., a course in field biology through course work at a requirement should not use OR&IE mathematical modeling applied to biological field station and can apply 360 Engineering Probability and biology (BIOEE 362 Dynamic Models up to 6 credits in the place o f courses Statistics II to fulfill the requirement in Biology; BIOEE 460 Theoretical from lists (b) or (c). For example, for an advanced course. Ecology; CS 321/BIOBM 321, students may apply 6 credits from the Numerical Methods in Computational • Biology majors in the College o f following courses taken at the Shoals Molecular Biology; BTRY 382 Agriculture and Life Sciences who Marine Laboratory (BIOSM): BIOSM Introduction to Statistical Genomics select this program o f study are 308 Marine Microbial Ecology; BIOSM and Bioinformatics, BTRY 494 allowed to take up to 61 credit hours 309 Climates and Ecosystems; BIOSM Quantitative Genetics, or CS 428 in the endowed colleges due to the 310 Marine Symbiosis; BIOSM 374 Introduction to Computational high number of required endowed Field Ornithology; BIOSM 377 Biophysics) courses for this program o f study. Diversity o f Fishes; BIOSM 477 Marine d. One course from the following list o f 4. Ecology and Evolutionary Biology Vertebrates; and BIOSM courses in lists b and c. The Ecology and Evolutionary advanced courses: BIOEE 261 Ecology and the Environment. Effective fa ll semester 2005, new students Biology program offers a specialization AN SC 420 Quantitative Animal in Marine Biology and Oceanography must also complete 10 credits from the Genetics; BIOBM 631 Protein Structure following lists: (a) Principles, (b ) (for a description, see “Courses in and Function; BIOGD 481 Population Marine Science”). The Organization for Organisms, and (c ) Ecological and Genetics; BIOGD 484 Molecular Evolutionary Processes. One course must Tropical Studies (OTS) offers an Evolution; BIOGD 487 Human be chosen from list (a) and a second Undergraduate Semester Abroad Genomics; BIOEE 460 Theoretical Program, featuring two courses either from list (b ) o r (c). The remaining Ecology; BIONB 330 Introduction to credits can be satisfied with courses from (Fundamentals in Tropical Biology and Computational Neuroscience; BIONB Field Research in Tropical Biology) all three lists. Students are encouraged to 422 Modeling Behavioral Evolution; that can count as two 3-credit courses take at least one course from each list. BIOPL 440 Phylogenetic Systematics; toward the concentration. Six credits BTRY 408 Theory o f Probability; BTRY a. Principles: BIOEE 453 Speciation; can be applied from the 15-week fall 482/682 Statistical Genomics; BTRY BIOEE 458 Community Ecology; “Semester in Environmental Science” 494 Quantitative Genetics; BTRY 652 BIOEE 460 Theoretical Ecology; BIOEE program offered by the Woods Hole Computationally Intensive Statistical 464 Macroevolution; BIOEE 480/ Marine Biological Laboratory. Inference; CIS 409 Data Structures and ENTOM 470 Ecological Genetics; Algorithms for Computational Science; BIOEE 478 Ecosystem Biology; NTRES 5. G en era l B iology: The program o f study in general biology requires a minimum o f 13 CS 211 Computers and Programming; 310 Applied Population Ecology. CS 426 Introduction to Computational credit hours in addition to courses counted b. Organisms: BIOEE 274 The Biology; CS 626 Computational toward requirements 1-9 on pages 155-156. Vertebrates: Structure, Function, and Molecular Biology; CS 627 These 13 credits must include: Evolution; BIOEE 373 Biodiversity and Computational Biology: The Machine a. One course from each o f three Biology o f the Marine Invertebrates or Learning Approach; MATH 420 BIOSM 376 Marine Invertebrate different programs o f study in biology. Differential Equations and Dynamical Zoology; BIOEE 450 and 451 Only those courses specifically listed Systems; NTRES 310 Applied Mammalogy, lec and lab; BIOEE 470 as fulfilling a program o f study Population Ecology; NTRES 411 and 472 Herpetology lec and lab; requirement are acceptable without Quantitative Ecology o f Fisheries; BIOEE 475 Ornithology; BIOEE 476 permission o f advisor. NTRES 412 Wildlife Population Biology o f Fishes; ENTOM 212 Insect b. A course with a laboratory. Analysis: Techniques and Models; Biology; BIOPL 241 Introductory NTRES 670 Spatial Statistics; OR&IE c. A minimum o f two upper-level (300 Botany; BIOPL 448 Plant Evolution and 360 Engineering Probability and and above) courses of 2 or more the Fossil Record; PL PA 309 Fungi; Statistics II; OR&IE 361 Introductory credits each. BIOSM 449 Seaweeds, Plankton, and Engineering Stochastic Processes Seagrass. 100-level courses are not acceptable for Notes meeting any o f these requirements. c. Ecological and Evolutionary Processes: BIO G 498 may not be used to fulfill • It is strongly recommended that BIOEE 263 Field Ecology; BIOEE 265 the requirements o f this program o f students complete the Core physics Tropical Field Ecology and Behavior; study. BIO G 499 (minimum o f 2 requirement using the PHYS 207-208 NS/ANTHR 275 Human Biology and credits, but no more than 3 credits) option. Evolution; BIOEE/BIONB/ENTOM 369 may count as one o f the upper-level Chemical Ecology; BIOEE/EAS 350 • It is strongly recommended that courses, and may count as the Dynamics o f Marine Ecosystems; students complete the Core organic laboratory course with approval o f the BIOEE/MATH 362 Dynamic Models in chemistry requirement using the advisor, but it cannot count as a course Biology; BIOEE 446 Plant Behavior— CHEM 257/251 option and that the representing a program o f study. Induced Plant Responses to Biotic time saved be used to take either CS Stresses; BIOEE/ENTOM 455 Insect 211 or a second mathematics course Note: It is possible to use a single course Ecology; BIOEE/NTRES 456 Stream from the list above to fulfill more than one requirement. For Ecology; BIOEE 457 and 459 example, BIOAP 413 Histology could • MATH 221 Linear Algebra and Limnology: Ecology o f Lakes, lec and count in all three areas: as a course in the Calculus, MATH 231 Linear Algebra, lab; BIOEE/EAS 462 Marine Ecology; Animal Physiology program o f study, as MATH 294 Engineering Mathematics II, BIOEE 466 and 468 Physiological Plant an upper-level course, and as a course or BTRY 421 Matrix Computation is Ecology, lec and lab; BIOEE/HORT with a lab. 1 5 8 BIOLOGICAL SCIENCES - 2['I' •Jil8

6. Genetics and Development: A minimum 8. Microbiology: Students in the Microbiology, Lee, and BIONB 222 o f 13 credits, usually chosen from the Microbiology program o f study must Neurobiology and Behavior II: following courses: BIOGD 385 complete BIOMI 290 General Introduction to Neurobiology. Graduate- Developmental Biology; BIOGD 400 A Microbiology, Lee; BIOMI 291 General level courses such as BIOBM 631 Protein Genomics Approach to Studying Life; Microbiology, Lab. At least 8 additional Structure and Function; BIOBM 633 BIOGD 401 Genomics Analysis; BIOGD credits are required, which must include Biosynthesis o f Macromolecules; BIOBM 450 Vertebrate Development; BIOGD 480 at least one o f the following courses: 636 Functional Organization o f Eukaryotic Seminar in Developmental Biology; BIOMI 414 Bacterial Diversity; BIOMI 416 Cells; and BIOBM 639 The Nucleus are BIOGD 481 Population Genetics; BIOGD Bacterial Physiology; BIOMI 418 Microbial also acceptable by permission o f advisor. 482 Human Genetics and Society; BIOGD Ecology; BIOMI 485 Bacterial Genetics. Five hours o f biochemistry are 483 Advanced Developmental Biology; recommended (BIOBM 331 and 332, or Additional approved courses are included BIOGD 484 Molecular Evolution; BIOGD 330 and 334, or 333 and 334). CHEM 207- in the list below. Students are invited to 486 Advanced Eukaryotic Genetics; 208 or 215-216 should be completed in complete their requirements in one o f BIOGD 487 Human Genomics; BIOGD the freshman year. three areas o f interest (these are only 489 Mammalian Embryology; BIOGD 490 recommended areas o f interest; students 10. Neurobiology and Behavior: The two- Manipulating the Mouse Genome; BIOGD can design their own course list as long semester introductory course sequence 610-624 Genomics Modules; BIOGD 640 as they meet the requirements described Neurobiology and Behavior I and II Stem Cell Biology; BIOGD 682 above): (a) Prokaryotic Biology, (b) (BIONB 221 and 222) with discussion Fertilization and the Early Embryo; Molecular Microbiology and section (4 credits per semester) and 7 BIOGD 684 Advanced Topics in Biotechnology, and (c) Pathogenic additional credits. The 7 additional credits Population Genetics; BIOGD 687 Microbiology. Courses acceptable to the must include at least one advanced course Developmental Genetics; BIOGD 689 program o f study that cover topics related from the BIONB offerings. “Topics” Cellular Basis o f Development; BIOMI 420 to a particular area o f interest are: courses (BIONB 420s and 720s), Microbial Genomics; BIOMI 485 Bacterial independent study (BIO G 499), BIONB Genetics; ENTOM 400 Insect Prokaryotic Biology: BIOMI 391 Advanced 321, and PSYCH 423 may be used as Development; BIOAP 475 Mechanisms Microbiology Laboratory; BIOMI 414 supplemental credits but do not qualify Underlying Mammalian Developmental Bacterial Diversity; BIOMI 416 Bacterial as advanced courses. Defects; BIONB 493 Developmental Physiology; and BIOMI 418 Microbial Neurobiology; BIONB 495 Molecular and Ecology. Note: Students who declare the program o f study in Neurobiology and Behavior after Genetic Approaches to Neuroscience; Molecular Microbiology and BIOBM 437 Cell Proliferation and taking BIONB 221 or 222 for only 3 credits Biotechnology: BIOMI 391 Advanced must still take the 1-credit discussion section Senescence; BIOBM 439 Molecular Basis Microbiology Laboratory; BIOMI 416 in BIONB 221 and 222. To arrange this, the o f Human Disease; NS 608 Epigenetics; Bacterial Physiology; BIOMI 420 Microbial BIOBM 633 Biosynthesis o f student should consult the professors in Genomics; BIOMI 485 Bacterial Genetics; charge o f the two courses. Macromolecules; BIOBM 639 The and BIOMI 394 Applied and Food Nucleus; BIOEE 453 Speciation; PL BR Microbiology. 11. Nutrition: NS 331 Physiological and 403 Genetic Improvement o f Crop Plants; Biochemical Bases o f Human Nutrition (4 Pathogenic Microbiology: BIOMI 404 PL BR 606 Advanced Plant Genetics; credits) and at least 9 credits o f additional Pathogenic Bacteriology and Mycology; BIOPL 343 Molecular Biology and Genetic course work in the biological aspects o f BIOMI 409 Viruses and Disease; BIOMI Engineering o f Plants; BIOPL 482 Plant nutrition, such as NS 122 Nutrition and 417 Medical Parasitology; and BIOMI 485 Molecular Biology II; BIOPL 483 Plant the Life Cycle; NS 315 Obesity and the Bacterial Genetics. Molecular Biology I; BIOPL 641 Regulation o f Body Weight; NS 332 Laboratory in Plant Molecular Biology. 9. Molecular and Cell Biology: CHEM Methods in Nutritional Sciences; NS 341 Up to 3 credits for this program o f study 357-358 or 359-360; BIOBM 432 Survey Human Anatomy and Physiology; NS 347 may be chosen from other biological o f Cell Biology; BIOBM 440 Laboratory in Human Growth and Development; NS 361 sciences courses, including BIO G 499 Biochemistry and Molecular Biology, or Biology of Normal and Abnormal Independent Undergraduate Research in BIONB 430 Experimental Molecular Behavior; NS 421 Nutrition and Exercise; Biology, with approval from the faculty Neurobiology; and at least 7 additional NS 441 Nutrition and Disease; NS 452 advisor. credits o f courses that have a cell Epidemiology and Dietary Markers o f biological or molecular biological Chronic Disease; NS 603 (alternate years) 7. Insect Biology: ENTOM 212 Insect orientation. The 7 additional hours should Mineral Nutrition: Metabolic, Health, and Biology plus a minimum o f three include at least two courses from the Environmental Aspects; NS 614 Topics in additional courses totaling at least 9 credits following list: BIOAP 416 Cellular Maternal and Child Nutrition; NS 631 selected from the following two groups. At Physiology and Genomics Laboratory; Micronutrients: Function, Homeostasis, least one o f the three additional courses BIOBM 401 Genomic Analysis; BIOBM and Metabolism; and NS 632 Regulation must be selected from group a. 434 Applications of Molecular Biology; o f Macronutrient Metabolism. Some Group a: ENTOM 322 Insect Morphology; BIOBM 435-436 Undergraduate Seminar courses require NS 115 Nutrition, Health, ENTOM 331 Insect Phylogeny and in Biochemistry; BIOBM 437 Regulation o f and Society, which may be used as part o f Evolution; ENTOM 333 Larval Insect Cell Proliferation, Senescence, and Death; the additional 9 credits. Biology; ENTOM 400 Insect Development; BIOBM 439 Molecular Basis o f Human Note: For students in the College o f ENTOM 455 Insect Ecology; ENTOM 483 Disease; BIO G 305 Immunology; BIOGD Agriculture and Life Sciences, credits in NS Insect Physiology 385 Developmental Biology; BIOGD 400 courses count toward the required 55 CALS A Genomics Approach to Studying Life; Group b: ENTOM 315 Spider Biology; credits. For students in the College o f Arts and BIOGD 46l Development and Evolution; ENTOM 325 Insect Behavior; ENTOM 344 Sciences, NS credits will count toward the 100 BIOGD 484 Molecular Evolution; BIOGD Insect Conservation Biology; ENTOM 352 hours required in A&S if those credits fulfill 486 Advanced Eukaryotic Genetics; Medical and Veterinary Entomology; major requirements. BIOGD 487 Human Genomics; BIOGD ENTOM 369 Chemical Ecology; ENTOM 490 Manipulating The Mouse Genome; 12. Plant Biology: Students choose one area 370 Pesticides, Environment, and Human BIOMI 409 Principles o f Virology; BIOMI o f study from the following two options: Health; ENTOM 394 Circadian Rhythms; 420 Microbial Genomics; BIOMI 485 ENTOM 443 Entomology and Pathology of Option (a ) Botany: Students are required Bacterial Genetics; BIONB 425 Molecular Trees and Shrubs; ENTOM 444 Integrated to take BIOPL 241 Introductory Botany. Neurophysiology; BIONB 495 Molecular Pest Management; ENTOM 452 Herbivores Students should then choose, with the aid and Genetic Approaches to and Plants; ENTOM 453 Principles and o f their faculty advisor, a minimum o f Neurosciences; BIOPL 343 Molecular Practice o f Historical Biogeography; three o f the following courses, for a total Biology and Genetic Engineering of ENTOM 456 Stream Ecology; ENTOM 463 o f at least 10 additional credits, to round Plants; BIOPL 347 Laboratory in Molecular Invertebrate Pathology; ENTOM 470 out their botanical training: BIOPL 242 Biology and Genetic Engineering of Ecological Genetics; ENTOM 477 Biological and 244 Plant Function and Growth, Lee Plants; BIOPL 444 Plant Cell Biology. Control; ENTOM 490 Insect Toxicology and Lab; BIOPL 243 Taxonomy o f Students graduating in spring 2008 or Cultivated Plants; BIOPL 245 Plant earlier may include BIOMI 290 General THE MAJOR 159

Biology; BIOPL 247 Ethnobiology; BIOPL Systematics o f Tropical Plants; ‘ BIOPL requirements under Option 2 must 248 Taxonomy o f Vascular Plants; BIOPL 454 Systematics o f Tropical Plants: complete a minimum of one program of 340 Methods in Biological and Field Laboratory; ‘ ENTOM 212 Insect study course o f at least 3 credits at Cornell. Biochemical Prospecting; BIOPL 342 and Biology; ENTOM 215 Spider Biology: • All courses for the minor must be taken 344 Plant Physiology, Lee and Lab; BIOPL Life on a Silken Thread; ENTOM 315 for a letter grade unless a course is 343 and 347 Molecular Biology and Spider Biology; ‘ ENTOM 322 offered S-U only. Genetic Engineering o f Plants, Lee and Comparative Insect Morphology; Lab; BIOPL 345 Plant Anatomy; BIOPL •ENTOM 331 Introductory Insect • Applications for the minor are located in 348 The Healing Forest; BIOPL 359 Systematics; ‘ ENTOM 333 Maggots, 2l6 Stimson Hall. See Bonnie Cornelia or Biology o f Grasses; BIOPL 380 Strategies Grubs, and Cutworms: Larval Insect Wendy Aquadro for academic advising and and Methods in Drug Discovery; BIOPL Biology; ‘ ENTOM 631 Systematics o f for certifying completion o f the minor. 404 Crop Evolution, Domestication, and the Coleoptera; PL PA 309 Diversity; BIOPL 422 Plant Development; Introductory Mycology; *PL PA 319 Independent Research and Honors BIOPL 440 Phylogenetic Systematics; Field Mycology. Program BIOPL 442 Current Topics in b. BIOEE 453 Speciation; BIOEE 464 Biology majors are encouraged to consider Ethnobiology; BIOPL 444 Plant Cell Macroevolution; BIOEE 479 participating in individual research under the Biology; BIOPL 447 Molecular Systematics; Paleobiology; ‘ BIOPL 440 Phylogenetic direction o f a Cornell faculty member. BIOPL 448 Plant Evolution and the Fossil Systematics; BIOPL 442 Current Topics Students interested in beginning research Record; BIOPL 449 Green Signals and in Ethnobiology BIOPL 447 Molecular Triggers— The Plant Hormones; BIOPL should contact faculty members who have Systematics; ‘ BIOPL 448 Plant compatible research interests. Information 452/454 Systematics o f Tropical Plants and Evolution and the Fossil Record; about faculty research interests and Field Lab; BIOPL 453 Principles and ‘ BIOPL 453 Principles and Practices o f undergraduate research opportunities is Practice o f Historical Biogeography; Historical Biogeography. available in the Office o f Undergraduate BIOPL 462 Plant Biochemistry; BIOEE 452 Biology, 216 Stimson Hall, and at www. Herbivores and Plants; BIOEE 463 and biology.cornell.edu. 465 Plant Ecology and Population Biology, The Minor in Biological Sciences Lee and Lab; or BIOEE 466 and 468 The minor in biological sciences has been Faculty members may consider the student’s Physiological Plant Ecology, Lee and Lab. designed to provide students with a broad previous academic accomplishments, interests background in biology while allowing them Option (b ) Plant Biotechnology: Students and career goals, and the availability o f space some flexibility to choose courses o f interest. are required to take BIOPL 343 and 347 and equipment when agreeing to supervise a Students must have completed one full year of Molecular Biology and Genetic student in their laboratory. Students introductory biology (or its equivalent) to Engineering o f Plants, Lee and Lab. conducting research for the first time must declare the minor. Students will complete 12 to Students choose, in consultation with their enroll in BIO G 299, an S-U course designed 15 credits by taking either all three biology to introduce students to research. After the faculty advisor, a minimum o f 10 core course requirements (listed below) or two additional credits from the following list: first semester, students enroll in BIO G 499. biology core course requirements and 5 or Registration for both o f these classes is done BIOPL 241 Introductory Botany; BIOPL more biology credits from the lists o f approved 242 and 244 Plant Function and Growth, in the Office o f Undergraduate Biology in 216 program o f study courses (found in Courses o f Lee and Lab; BIOPL 342 and 344 Plant Stimson Hall. Students may work with faculty Study) or from the following list o f courses: in any department on campus as long as the Physiology, Lee and Lab; BIOPL 422 Plant Development; BIOPL 444 Plant Cell AN SC 300, 301, 410, 420, 427 research topic is biological. Students may not Biology; BIOPL 462 Plant Biochemistry; earn credit for research done o ff campus, BEE 454 BIOPL 482, 483 Plant Molecular Biology I unless supervised by a Cornell faculty and II modules; PL BR 401 Plant Cell and BTRY 382, 482/682 member. Up to 3 credits o f research may be used to complete the program o f studies in Tissue Culture; or PL BR 402 Plant Tissue CS 426, 626, 627 Culture Laboratory. general biology, genetics and development, CSS 421 systematic and biotic diversity, as well as 4 13. Systematics and Biotic Diversity: A credits in neurobiology and behavior. minimum o f 13 credits from the following EAS 350, 479 two groups, including at least 7 credits The honors program in biological sciences is ENTOM 212, 213, 215, 315, 322, 325, 331, 333, designed to offer advanced training in life from group a and three from group b and 344, 352, 369, 370, 394, 400, 443, 444, 452, at least two laboratory courses (marked science research through the performance of 453, 455, 456, 463, 470, 471, 477, 483, 490, 631 with *). BIO G 499, Independent an original research project under the direct Undergraduate Research in Biology, with NS 222, 315, 331, 332, 341, 347, 361, 421, 431, guidance o f a member o f the Cornell faculty. approval o f the advisor, can be used in 441, 452, 455, 475, 602, 603, 614 Biology majors planning on graduating with honors must apply to the Biology Honors fulfillment o f up to 4 credits in group (a), NTRES 305, 310, 411, 412, 418, 419, 608, 670 and can count as one laboratory course if Program in the spring o f their junior year. it has a laboratory component o f 2 or PL BR 401, 402, 403, 606 Applications and information are available in the Office o f Undergraduate Biology, 216 more credits. PL PA 309, 319 Stimson Hall, or at www.biology.cornell.edu/ a. ‘ BIOEE 264 Tropical Field Biology core courses research/honors.html. To qualify for the Ornithology; ‘ BIOEE 274 The program, students must have been accepted 1. Biochemistry: BIOBM 330, 333, or 331- Vertebrates: Structure, Function, and into the biological sciences major, have 332; one year o f general chemistry and Evolution; BIOEE 371 Human completed at least 30 credits at Cornell, and organic chemistry lecture (CHEM 257 or Paleontology; ‘ BIOEE 373 Biology of have a cumulative Cornell grade point average 357-358) are prerequisites the Marine Invertebrates; BIOEE 405 (GPA) o f at least 3.0. In addition, students must Biology o f the Neotropics; BIOEE 470 2. Evolutionary biology: BIOEE 278 or have at least a 3.0 cumulative Cornell GPA in Herpetology, Lee; ‘ BIOEE 471 BIOPL 448 all biology, chemistry, mathematics, and physics Mammalogy; ‘ BIOEE 472 Herpetology, courses. (Grades earned in courses in other 3. Genetics: BIOGD 281 Lab; ‘ BIOEE 475 Ornithology; ‘ BIOEE departments that are used to fulfill biology 476 Biology o f Fishes; BIOEE 477 Notes: major requirements are included in this Marine Invertebrates Seminar; BIOMI computation.) In addition, candidates must find • BIO G 499 Independent Research may 290 General Microbiology, Lee; a Cornell faculty member to supervise their ‘ BIOMI 291 General Microbiology, not be used to fulfill any requirement for the minor. No course substitutions are research. An honors candidate usually enrolls Lab; BIOMI 414 Prokaryotic Diversity, for credit in BIO G 499 Undergraduate Lee; ‘ BIOPL 241 Introductory Botany; allowed. With the exception o f transfer and study abroad students, no biology Research in Biology under the direction o f the •BIOPL 243 Taxonomy o f Cultivated faculty member acting as honors supervisor, Plants; ‘ BIOPL 245 Plant Biology; courses taken at other institutions will count toward the minor. although the honors program does not require BIOPL 247 Ethnobiology; ‘ BIOPL 248 enrollment for credit. Students accepted into Taxonomy o f Vascular Plants; BIOPL • External transfer students must complete the honors program are required to participate 348 The Healing Forest; BIOPL 359 the core biology courses at Cornell. in honors research seminars during their senior Biology o f Grasses; BIOPL 452 Students who are fulfilling the minor year; submit an acceptable honors thesis; 160 BIOLOGICAL SCIENCES - 2TTBffiTT 8

complete all major requirements; and maintain Online course credit is not acceptable to dissect several invertebrates during the second a 3.00 Cornell cumulative and science GPA transfer for any biology course. semester. For those students w ho object to through graduation. Recommendation to the animal dissection, alternative materials are faculty that a candidate graduate with honors available for study. However, testing involves and at what level o f honors is the responsibility identification o f important structures in real o f the Honors Program Committee. The GENERAL COURSES (BIO G) organisms. student’s final GPA and quality o f his or her Three introductory biology course sequences BIO G 105-106(1105-1106) Introductory thesis are factors in determining the level o f Biology honors recommended. are taught during the academic year: BIO G 101-104, 105-106, and 109-110; and one 105, fall; 106, spring. 4 credits each Students interested in the honors program are during the eight-week summer session: BIO G semester; 2 credits by permission o f strongly encouraged to begin their research 107-108. BIO G 101-104, 105-106, and 107- instructor. Limited to 200 students. Taking projects in their junior year and to consider 108 are intended for biological sciences majors 105-106 in sequence preferred but not spending the following summer at Cornell and other students needing 8 credits from an required. May not be taken for credit after engaged in full-time research on their honors introductory sequence for majors (e.g., BIO G 101-104 or 109-110. N o admittance project. Students interested in staying for the students in a premedical curriculum). Any o f after first week o f classes. First lec o f fall summer also are encouraged to apply to the these sequences meet the prerequisite for semester R Aug. 23, 9:05; additional study Cornell Hughes Scholars Program. upper-level courses listing “one year o f and lab. D. Campbell. Designed primarily for biology majors, Biology majors who are considering study introductory biology for majors” as a preprofessionals, and other students who abroad and graduating with honors are prerequisite. BIO G 109-110 is a course desire a challenging, broad introduction to encouraged to meet with their academic and sequence intended for nonmajors and meets fundamental concepts of biology. Cell biology, research advisor during their sophomore year the prerequisite for many, but not all, upper- physiology, anatomy, and biochemistry are to carefully plan their academic schedule to level courses listing “one year o f introductory strongly emphasized in the fall semester. The meet the requirements o f the honors program. biology” as a prerequisite. Students can earn a maximum o f 8 credits in introductory biology spring semester covers genetics, development, Application forms, requirements, deadline (including advanced placement credits). ecology, evolution, behavior, and the diversity dates for the honors program and the Hughes o f organisms. Students who plan to Scholars Program, and information pertaining BIO G 101-102(1101-1102) Biological concentrate in anatomy and physiology to faculty research may be obtained at the Sciences, Lectures should consider taking this course because o f Office o f Undergraduate Biology, 216 Stimson 101, fall; 102, spring. 2 credits each the strong emphasis on organismal biology. Hall, and at www.biology.comell.edu. semester. Corequisite: BIO G 103 (fall) or Because some testing involves the use of 104 (spring). Prerequisite: for 102, passing predissected specimens, students w ho object grade (D or better) in 101 or permission o f to dissections should take BIO G 101-104. instructor. May not be taken for credit after The course uses an autotutorial format and CURRICULUM COMMITTEE BIO G 105-106 or 109-110. S-U or letter offers considerable flexibility in scheduling. grades by permission o f instructor. First lec Completion o f the course requires mastery o f Many decisions pertaining to the curriculum o f fall semester, F Aug. 24. N o admittance a group o f core units. Testing on these units is and to the programs o f study are made by the after second week o f classes. Evening primarily by oral examination. Students who Biology Curriculum Committee, which meets prelims: fall, Sept. 20 and Oct. 30; spring, elect to take the course must be able to meet monthly during the academic year. The Feb. 21 and Apr. 1. Fall, staff; spring, staff. deadlines. Four formal laboratory sessions are committee consists o f faculty and elected Designed for students w ho intend to offered each semester; additional laboratory student members and welcomes advice and specialize in biological sciences. The fall work is included in the core units. Evaluation suggestions from all interested parties. semester covers the chemical and cellular is based on written reports on experimental basis o f life, energy transformations, work, practical exams, and a comprehensive physiology, neurobiology, and behavior. The final exam. Web site: instructl.cit.comell.edu/ spring semester covers genetics, development, courses/biogl05. ADVISING evolution, and ecology. Each topic is considered in terms o f modern evolutionary BIO G 107-108(1107-1108) General Students in need o f academic advice are theory, and discussions o f plant and animal Biology encouraged to consult their advisors or see an systems are integrated. Summer, 8-week session; 107, weeks 1-4; academic advisor in the Office o f 108, weeks 5-8. 4 credits each. 107-108 Undergraduate Biology, 216 Stimson Hall. BIO G 103-104(1103-1104) Biological fulfills introductory biology requirement for Sciences, Laboratory Students interested in marine biology should majors and forms suitable introductory 103, fall; 104, spring. 2 credits each visit the Shoals Marine Laboratory Office, G14 biology course sequence for students semester. Corequisite: BIO G 101 (fall) or Stimson Hall. intending to go to medical school. 102 (spring). Prerequisite: for 104, passing Prerequisite: one year of college or Students interested in the multidisciplinary grade (D or better) in 103 or permission of permission o f instructor; for BIO G 108, a program biology and society should see instructor. Students registered for lab grade o f D or better in the prerequisite “Special Programs and Interdisciplinary courses w ho are more than 10 minutes late courses (BIO G 101, 103, 105, or 107). Fee Studies” in the “College o f Arts and Sciences” for first meeting o f lab forfeit registration for weeks 1-4: $25; for weeks 5-8, $25. section o f this catalog. in that course; no admittance after second Staff. week o f classes. S-U or letter grades by Designed for students w ho plan further study permission o f instructor. K.-c. Chen. in biology. 107 covers biological metabolism, Designed to provide lab experience with first at the molecular level and then TRANSFERRING CREDIT major biological phenomena to support an progressively to the organ system level. The understanding o f the important concepts, laboratory work involves an introduction to Biology majors are required to complete all principles, and theories of modern biology. A some major techniques, vertebrate dissection, three biology core courses (biochemistry, second objective is to help students gain and a survey of plant organization. 108 seeks evolution, and genetics) at Cornell. expertise in the methods used by biologists to to integrate the topics o f genetics, External transfer students are limited to construct new knowledge. Students are developmental biology, population biology, transferring one core biology course and one exposed to basic concepts, research methods, and ecology in a general consideration of course o f up to 3 credits toward their program including laboratory and data transformation biological evolution. The laboratory work is a o f study. See the Office o f Undergraduate techniques, and instrumentation in the major continuation o f the material covered in 107 Biology for approval procedures. areas o f biology. First-semester topics include and involves more techniques, a survey of biochemistry, physiology, plant biology, and Students who matriculated to Cornell as animal organization, and the design and behavior. In the second semester, laboratory freshmen and Study Abroad students may performance of a field study. For those experience is provided in genetics, transfer program o f study courses at the students who object to animal dissection, biotechnology, invertebrate diversity, plant and discretion o f their advisor. Study Abroad alternative materials are available for study. animal development, and ecology. During the students must obtain approval from the Office However, testing involves identification of first semester, students dissect a doubly pithed important structures in real organisms. of Undergraduate Biology, Director of frog (pithing is done by the staff). Students Advising, to transfer biology core courses. GENERAL COURSES 161

BIO C 109-110(1109-1110) Biological BIO G 200(2000) Special Studies in Using animal, plant, and microbe materials, Principles Biology this section studies chemical fixtures, 109, fall; 110, spring. 3 credits each Fall, spring, or summer. 1-3 credits. cryofixations, ultrathin sectioning, immunogold semester includes lecture and lab. Limited Prerequisite: written permission from localization, quantitative microscopy, and to 600 students. Nonmajors survey course, Office o f Undergraduate Biology. Students metal shadowing techniques. Students have not appropriate for major in biological must register in 216 Stimson Hall. S-U or two additional weeks to complete laboratory science or premed requirement. Both BIO letter grades by permission o f instructor. assignments at the end o f each section.] G 109 and 110, taken in either order, are Staff. BIO G 404(4040) Planning for Graduate required to fulfill distribution requirement Registration device for students who want to Study in Biology in CALS and Human Ecology. Either course take only a portion o f a regular biological Fall. 1 credit. S-U grades only. fulfills Arts and Sciences distribution sciences course— for example, only the L. E. Southard. requirement. Students with transfer credit lectures or only the laboratory in a course that For students w ho plan to pursue a graduate must consult with course instructors for includes both. Only students who have degree leading to a research career. Selected appropriate course placement. Due to already had training equivalent to the portion topics include information on academic and overlap in content, BIO G 109 may not be o f the regular course that is to be omitted may industrial research careers, selecting taken after BIO G 102 or 106, or register in this manner. This course may not appropriate graduate programs, and options equivalent, and BIO G 110 may not be be substituted for 100-level courses and may for funding. Features faculty, graduate student, taken after BIO G 101, 105, or equivalent. not be used to fulfil college distribution and outside speakers. Students write and Note: This course may not satisfy requirements except by permission from the receive feedback on personal statements. prerequisite for upper-level courses in Office of Undergraduate Biology. biology. Letter grades only. Prelims: fall (in BIO G 299(2990) Introduction to BIO G 408(4080) Presentation Skills for class), Sept. 21 and Oct. 31; spring Research Methods in Biology Biologists (evening), Feb. 21 and Apr. 3. H. Greene, Fall, spring, or summer. Variable credit; Spring. 1 credit. S-U grades only. R. Wayne, E. Balko, and staff. max. 3 suggested. S-U grades only. Prerequisite: research experience. Priority Offers a comprehensive knowledge of biology Recommended for freshmen and given to students accepted into Biology as part o f a general education. Broad goals o f sophomores. Students must register for Honors Program. L. Southard and G. Hess. the course encompass an understanding o f the credit in Office o f Undergraduate Biology, Covers oral and written communication skills potential benefits and limitations o f science, 216 Stimson Hall. Applications available in used in presenting research to other scientists. the complexity and workings o f the natural OUB and at www.bio.comell.edu. Add Topics include organization and writing of world, and the internal machinery o f how our deadline is three days before university scientific papers, presentation tips for research bodies and those o f other animals and plants deadline. Any Cornell faculty member seminars, and preparation o f visual aids using work. Fall: biological diversity, genetics, whose research field is biological in nature Microsoft Power Point. All students present a evolution, ecology, behavior, and conservation may serve as a supervisor for this course. 10-minute seminar on their research and biology; Spring: human physiology, plant Non-Cornell supervisors not acceptable. evaluate other presentations. development, genetic engineering, infectious Intended for students w ho are new to diseases, and human health. Laboratory BIO G 410(4100) Teaching High School undergraduate research. Students enrolled in sessions (6 labs per semester) are used for Biology BIO G 299 may be reading scientific literature, problem-solving experiments, demonstrations, Fall. 3 credits. Prerequisite: one year learning research techniques, or assisting with discussions, and dissections (preserved introductory biology; permission of ongoing research. The faculty supervisor vertebrate, invertebrate, and plant materials); instructor. S-U or letter grades. Offered determines the work goals and the form of for those students who object to dissection, alternate years. L. Southard. the final report. alternative materials are available for study Gives students the opportunity to experience without grade penalty. Testing on dissection BIO G 305(3050) Basic Immunology teaching high school science. Students select labs involves identification o f important Lectures (also VETMI 315(3150]) an important biological concept, then develop inquiry-based teaching plans appropriate for structures in real organisms. Fall. 3 credits. Highly recommended: basic courses in microbiology, biochemistry, and high school students. The first part o f the BIO G 112(1120) Issues in Social genetics. S-U or letter grades. Lee. course consists o f lectures, discussion, and Biology: from Diet to Diseases, DNA laboratory experiments, which familiarize the to Deforestation J. A. Marsh. For description, see VETMI 315. students with the scientific content. Students Spring. 3 credits. S-U or letter grades. then work in teams with high school teachers P. J. Davies. BIO G 400(4000) Undergraduate Seminar to develop their curriculum. The final part of An analysis o f current issues o f biological in Biology the course includes practice presentations and relevance and the biological science behind Fall or spring. Variable credit; 1-3 assigned teaching at regional high schools. these issues. Topics will include issues such as for individual seminar offerings; may be food and nutrition, antioxidants, organic repeated for credit. S-U or letter grades. BIO G 431(4310) Frontiers in Biophysics produce, disease prevention, athletic Staff. Fall, full day o f lec, S, Sept. 15, 9 A.M.-4 p. enhancers, genetic testing, cancer, stem cells Specialized seminars on topics o f interest to M. , Racker Room, Biotechnology Bldg. 0.5 and animal cloning, genetically modified undergraduates presented by biology faculty credit. S-U grades only. G. Feigenson and crops, bacteria and antibiotics, viruses, risk, including visiting faculty. staff. statistics and epidemiology, photosynthesis Overview o f current research in biophysics at and global warming, extinction and [BIO G 401(4010) Introduction to Cornell by faculty from different departments overpopulation, invasive species, resource Scanning Electron Microscopy] across the university. Designed for over-utilization. The topics will vary according [BIO G 403(4030) Transmission Electron undergraduates considering a career in to current issues. Microscopy for Biologists biophysics and for graduate students interested in biophysics research opportunities Fall. 1, 3, or 4 credits (4 credits if student BIO G 125(1250) Biology Seminar at Cornell. Fall and spring. 1 credit. Prerequisite: first- takes both sections). Limited to 8 students; year standing or permission o f instructor. minimum o f 4. Prerequisites: BIOAP 313, BIO G 450(4500) Light and Video S-U grades only. Staff. BIOPL 345 or 443. S-U or letter grades. Microscopy for Biologists A first-year seminar designed for students with Tw o sec: 01, 1 credit, weeks 1-4; 02, 3 Fall. 3 credits. Limited to 12 students. Biology AP credit or a strong interest in credits, weeks 5-12. Students may register Prerequisites: one year introductory research. Students will interact with faculty for one or both sec. Fee may be charged. biology and permission o f instructor. Lec, while learning to read and evaluate scientific Lee, lab. Next offered 2008-2009. Staff. lab. R. O. Wayne. publications on current biological topics. Section 01 covers the principles and use of Students leam the relationship between reality Multiple topics and sections will be offered the transmission electron microscope (TEM), and the image using philosophy, mathematics, each semester. with emphasis on proper operation o f the and physical theory. Next they apply these instrument and interpretation o f images tools theoretically and in practice to understand BIO G 170(1700) Evolution of the Earth obtained. Negatively stained materials are used and become experts at image formation and and Life (also EAS 102(1102]) for viewing with the transmission electron analysis using brightfield, darkfield, phase- Spring. 3 credits. S-U or letter grades. microscope. Section 02 covers the principles contract, fluorescence, polarization, J. L. Cisne. and techniques of preparing biological interference, differential interference and For description, see EAS 170. material for transmission electron microscopy. modulation contrast microscopes. They build 162 BIOLOGICAL SCIENCES - 2 8

upon our knowledge and experience to BIO G 705(7050) Advanced Immunology BIOAP 319(3190) Animal Physiology understand how analog image processors and Lectures (also VETMI 705(7050]) Experimentation digital image processors can influence, enhance Spring. 3 credits. Prerequisite: basic Fall. 4 credits. Limited to 40 students per and analyze the images gathered by the immunology course or permission of lab sec. Pre- or corequisite: BIOAP 311 or microscope. Last they learn about many other instructor. Offered alternate even years. permission o f instructor. For pre-med, pre- kinds o f microscopes, including confocal, near Lee. Coordinator: J. A. Marsh. vet juniors and seniors and graduate field, x-ray, acoustic, nuclear magnetic For description, see VETMI 705. students interested in biomedical science. resonance, infrared, centrifuge, atomic force, E. R. Loew, N. A. Lorr, and staff. [BIO G 706(7060) Immunology of Student-conducted in vitro and in vivo and scanning tunneling microscopes. Infectious Diseases (also VETMI 719(7190]) experiments designed to illustrate basic BIO G 498(4980) Teaching Experience physiological processes, physiological research Spring. 2 credits. Prerequisite: basic Fall or spring. 1-4 credits. Limited techniques, instrumentation, experimental immunology course or permission o f ■ H H enrollment. Prerequisites: previous design, and interpretation o f results. Techniques instructor. S-U or letter grades. Offered odd enrollment in course to be taught or include anesthesia, surgical procedures, alternate years; next offered 2008-2009. equivalent. Note: Arts students may not dissection, and real-time computer recording Coordinator: E. Denkers. count this course toward graduation but and analysis. Experiments with isolated living For description, see VETMI 719-1 may, upon petition (one time only) to their tissues or live anesthetized animals examine class dean, carry fewer than 12 other properties o f membranes and epithelia, blood, credits and remain in good standing. This nerves, skeletal and smooth muscle; would affect Dean’s List eligibility but not cardiovascular, respiratory, renal, and eligibility for graduating with distinction. ANIMAL PHYSIOLOGY (BIOAP) reproductive function and their regulation by S-U or letter grades by permission o f the nervous and endocrine systems. instructor. Staff. [BIOAP 214(2140) Biological Basis of Designed to give qualified undergraduate Sex Differences (also B&SOC BIOAP 413(4130) Histology: The Biology 214(2141), FGSS 214(2140]) students teaching experience through actual of the Tissues Fall. 3 credits. Prerequisite: one year involvement in planning and assisting in Spring. 4 credits. Prerequisite: one year introductory biology. S-U or letter grades. biology courses. This experience may include introductory biology. Recommended: Offered alternate years; next offered 2008- supervised participation in a discussion group, BIOBM 330 or 331, or equivalents. S-U or 2009. J. E. Fortune. assisting in a biology laboratory, assisting in letter grades. S. Suarez and L. Mizer. Examines the structural and functional field biology, or tutoring. Provides students with a basis for differences between the sexes. Emphasizes understanding the microscopic, fine-structural, BIO G 499(4990) Independent mechanisms o f mammalian reproduction. and functional organization o f vertebrates Undergraduate Research in Biology Current evidence on the effects o f gender on (primarily mammals), as well as methods of Fall, spring, or summer. Variable credit. S-U nonreproductive aspects o f life is discussed. analytic morphology at the cell and tissue or letter grades. Note: Arts students may Provides students with a basic knowledge o f levels. Emphasizes dynamic interrelations o f not register for more than 6 credits per reproductive endocrinology and with a basis structure, composition, and function in cells semester with one supervisor or 8 credits for objective evaluation o f sex differences in and tissues. per semester with more than one relation to contemporary life.] supervisor. Students in CALS may use up BIOAP 416(4160) Cell Physiology and to 15 credits o f independent study (BIO G BIOAP 311(3110) Introductory Animal Genomics Laboratory Physiology, Lectures (also VTBMS Spring. 4 credits. Limited to 24 students. 499, 498) toward graduation. Up to 3 346(3460]) credits o f research may be used to For pre-med, pre-vet, juniors, seniors, and Fall. 3 credits. Prerequisites: one year complete programs o f study in General graduate students interested in biomedical college biology, chemistry, and Biology, Genetics and Development, and science. Pre- or corequisite: BIOAP 316 or mathematics. Recommended: previous or Systematics and Biotic Diversity, and 4 BIOBM 432 or permission o f instructor. concurrent physics course. S-U or letter credits o f research in Neurobiology and N. A. Lorr, H.-H. Chuang, and staff. grades by permission o f instructor. Evening Behavior. Prerequisite: one semester o f A laboratory course introducing modern prelims. E. R. Loew. BIO G 299 or equivalent or permission of methods and instrumentation in cell General course in animal physiology instructor and Office o f Undergraduate physiology and genomics. Students learn: emphasizing principles of operation, Biology. (1 ) cell culture and imaging by fluorescent regulation, and integration common to a For students with previous undergraduate microscopy; (2 ) isolation and manipulation o f broad range o f living systems from the cellular experience conducting biological research at RNA and DNA, in vitro transcription; to the organismal level. Structure/function Cornell. Students enrolled for this credit (3 ) transformation, transfection, and relationships are stressed along with should be doing independent work on their microinjection; (4 ) protein electrophoresis, underlying physico-chemical mechanisms. own project. Registration forms are available western blotting, and immunocytochemistry; in OUB and on the web at www.bio.cornell. BIOAP 312(3120) Farm Animal Behavior (5) electrophysiology using Xenopus oocyte edu. Add deadline is three days before (also AN SC 305(3050]) expression system and Ussing chambers; and university deadline. Each student must Spring. 2 credits. Prerequisites: one year (6 ) analysis o f gene expression using RT-PCR, submit proposed research project description introductory biology, and introductory animal real time PCR, and microarray analysis. during course registration. Any Cornell faculty physiology (AN SC 100 and 150 or equivalent Students will conclude the course with an member whose research field is biological in is sufficient or BIOAP 311). Recommended: at experiment o f their own design. nature may serve as supervisor for this course. least one animal production course or BIOAP 425(4250) Gamete Physiology and Non-Cornell supervisors not acceptable. equivalent experience. S-U or letter grades. Fertilization (also AN SC 425(4250]) P. Perry and K. A. Houpt. BIO G 600(6000) Graduate Seminar in Fall. 2 credits. Prerequisite: AN SC 300 or For description, see AN SC 305. Biology equivalent. Offered alternate years. Lee. Fall or spring. Variable credit (1-3 credits BIOAP 316(3160) Cellular Physiology J. E. Parks. assigned for individual seminar offerings). Spring. 3 credits. Pre- or corequisite: For description, see AN SC 425. May be repeated for credit. S-U or letter BIOBM 330 or 331 and 332 or 333. BIOAP 427(4270) Fundamentals of grades. Staff. Evening prelims. A. Quaroni. Endocrinology (also AN SC Specialized seminars on topics o f interest to Comprehensive course covering the general 427(4270]) graduate students presented by biology faculty characteristics o f eukaryotic cells; the structure, Fall. 3 credits. Prerequisite: animal or including visiting faculty. composition, and function o f subcellular human physiology course or permission o f BIO G 663(6630) Nanobiotechnology organelles; and the major signal transduction instructor. Lee. P. A. Johnson. (also A&EP 663(6630]) pathways regulating a variety o f physiological For description, see AN SC 427. cell activities. Among the main subjects covered Fall. 3 credits. Letter grades only. H. Hoch. are absorption and transport processes, For description, see A&EP 663. mechanism o f action o f signaling molecules (hormones), the cell cycle and regulation o f cell proliferation, cell-cell communication, extracellular matrix, and carcinogenesis. BIOCHEMISTRY, MOLECULAR AND CELL BIOLOGY 163

BIOAP 458(4580) Mammalian Physiology BIOAP 715(7150) Stress Physiology: To BIOBM 330-332(3300-3320) Principles of Spring. 3 credits. Auditors allowed. Be Discussed as Part of Animal Biochemistry Prerequisite: BIOAP 311 or equivalent. Welfare Introductory biochemistry is offered in three Recommended for biological sciences Fall. 1 credit. Prerequisite: BIOAP 311 or formats: individualized instruction (330) and majors, pre-med and pre-vet students, and equivalent. Offered alternate years. lectures (331 and 332) during the academic beginning graduate students in physiology, K. A. Houpt. year, and lectures (333) during the summer. nutrition, and animal science. Evening Emphasizes physiological assessment o f stress. Individualized instruction is offered to a maxim um o f 250 students each semester. prelims. K. W. Beyenbach. BIOAP 720(7200) Animal Physiology and The course offers a treatment o f selected Lectures are given fa ll semester (3 3 V , spring Anatomy Seminar semester (332), and summer (333). topics in vertebrate and human physiology Spring and fall. 1 credit each semester. that emphasizes concepts and a working Prerequisite: admission to graduate field o f BIOBM 330(3300) Principles of knowledge o f physiology. The first course half physiology. M. Roberson. Biochemistry, Individualized surveys biological design and the functional Designed to train graduate students in the Instruction strategies o f multicellular animals. Topics field of physiology to become professional Fall or spring. 4 credits. Prerequisites: one include mammalian fluid compartments, scientists. Students are required to give a year introductory biology and one year homeostasis, and membrane and epithelial seminar on their research. Advice and general chemistry and CHEM 257 or transport. The second half examines the feedback are provided. Throughout the 357-358 (CHEM 358 may be taken mechanism and the regulation of semester, advice is provided on subjects such concurrently) or equivalent, or permission cardiovascular, gastrointestinal, and renal as preparation o f manuscripts, seminars, and o f instructor. Recommended: concurrent systems. Course concluding lectures aim to grant proposals. registration in BIOBM 334. May not be illustrate the integration o f systems in the taken for credit after BIOBM 331, 332, or regulation o f acid/base balance. Clinical BIOAP 757(7570) Current Concepts in 333. S-U or letter grades. Evening prelims: examples o f dysfunction will underscore the Reproductive Biology fall, Sept. 27 and Oct. 30; spring, Feb. 21 role o f normal function, and some diseases Fall. 3 credits. Limited to 20 students. and Apr. 1. J. E. Blankenship, P. C. Hinkle, will be traced to the deepest roots o f their Prerequisites: undergraduate degree in and staff. molecular etiology. Weekly problem sets count biology and strong interest in reproductive Fourteen units that cover protein structure and 50 percent o f the final grade. biology. S-U or letter grades. Offered function, enzymes, basic metabolic pathways, alternate years. J. E. Fortune, P. A. Johnson, BIOAP 475(4750) Mechanisms DNA, RNA, protein synthesis, and an and staff. introduction to gene cloning. N o formal Underlying Mammalian Team-taught survey course in reproductive Developmental Defects (also NS lectures, autotutorial format. physiology/endocrinology. Lectures by a 475(4750]) number of reproductive biologists on various BIOBM 331(3310) Principles of Spring. 3 credits. Prerequisites: BIOBM 330, aspects o f male reproductive function Biochemistry: Proteins and 331-332, or 333 (may be taken (endocrine regulation, testis function, Metabolism concurrently). Offered alternate years. spermatogenesis, sperm physiology/function); Fall. 3 credits. Prerequisites: one year D. Noden and P. Stover. female reproductive function (endocrinology, introductory biology, one year general Focuses on the causes o f developmental ovarian development and function, oocyte chemistry, and CFIEM 257 or 357-358 (CHEM defects and how genetic changes or physiology/function); fertilization and gamete 257 or 357 should not be taken concurrently) teratogenic insults disrupt developmental transport; pregnancy; parturition; lactation; or equivalent, or permission o f instructor. regulatory and metabolic pathways. aging; reproductive technology. Student May not be taken for credit after BIOBM 330 BIOAP 489(4890) Mammalian Embryology participation in the form o f discussions and/or or 333- S-U grades by permission of (also BIOGD 489(4890]) presentations. instructor. Lee; evening prelim Oct. 18. Spring. 3 credits. Prerequisite: introductory G. W. Feigenson. biology. Offered alternate years. Related Courses in Other Departments The chemical reactions important to biology, D. M. Noden. and the enzymes that catalyze these reactions, Examines the early formation o f the Advanced Work in Animal Parasitology are discussed in an integrated format. Topics mammalian body and placenta, emphasizing (VETMI 737) include protein folding, enzyme catalysis, comparative aspects, and morphogenesis and bioenergetics, and key reactions o f synthesis Animal Reproduction and Development histogenesis o f each organ system. and catabolism. (AN SC 300) [BIOAP 619(6190) Lipids (also NS BIOBM 332(3320) Principles of 602(6020]) Comparative Insect Morphology (ENTOM Biochemistry: Molecular Biology Fall. 2 credits. Next offered 2008-2009. 322) Spring. 2 credits. Prerequisites: one year A. Bensadoun. introductory biology and previous or Developmental Biology (BIOGD 385) For description, see NS 602.] concurrent registration in organic chemistry, or permission o f instructor. May Fundamentals of Endocrinology (AN SC 427) BIOAP 710-718(7100-7180) Special not be taken for credit after BIOBM 330 or Topics in Physiology Research in Marine Biology (BIOSM 413) 333. S-U or letter grades by permission o f Fall or spring. 1 or 2 credits per topic; may instructor. Lee. B. K. Tye. be repeated for credit. Enrollment in each Sensory Function (BI0NB 492) Comprehensive course in molecular biology topic may be limited. S-U or letter grades that covers the structure and properties o f by permission o f instructor. Teaching Experience (BIO G 498) DNA, DNA replication and repair, synthesis Lectures, laboratories, discussions, and and processing o f RNA and proteins, the seminars on specialized topics. Undergraduate Research in Biology (BIO G regulation o f gene expression, and the 499) BIOAP 711(7110) Readings in Applied principles and applications o f recombinant Animal Behavior DNA technologies, genomics, and proteomics. Fall. 1 credit. Prerequisite: BIOAP 311 or BIOBM 333(3330) Principles of equivalent. Offered alternate years. Lee. Biochemistry: Proteins, Metabolism, K. A. Houpt. BIOCHEMISTRY, MOLECULAR AND and Molecular Biology BIOAP 714(7140) Cardiac CELL BIOLOGY (BIOBM) Summer, eight-week session. 4 credits. Electrophysiology BIOBM 132(1320) Orientation Lectures in Prerequisites: one year introductory biology, one year general chemistry, and Fall. 1 credit. S-U grades only. Offered Molecular Biology and Genetics (also CHEM 257, or 357-358, or equivalents, or alternate years. R. Gilmour. BIOGD 132(1320]) permission o f instructor. May not be taken Survey o f cardiac potentials, passive Spring, weeks 1-3- 0 credits. Primarily for for credit after BIOBM 330, 331, or 332. membrane properties, ion channels, and freshmen, sophomores, and transfer H. Nivison. cardiac arrhythmias. Emphasizes nonlinear students. S-U grades only. Lee. Staff. Topics include the structure and function o f dynamic aspects o f cardiac electrophysiology Six professors discuss their research and proteins, enzyme catalysis, metabolism, and and cardiac arrhythmias. promising new areas for research in the the replication and expression o f genes. future. 164 BIOLOGICAL SCIENCES - 2 8

BIOBM 334(3340) Computer Graphics BIOBM 438(4380) The RNA World [BIOBM 443(4430) Experimental and Molecular Biology Spring. 3 credits. Prerequisites: BIOBM 330 Molecular Neurobiology (also BIONB Fall or spring. 1 credit. Prerequisite: or 331/332 or 333, or permission o f 430(4300]) BIOBM 333 or 331-332 (BIOBM 332 may instructor. A. Ke. Spring. 4 credits. Limited to 12 students. be taken concurrently) o r Corequisite: Part o f the excitement about “the RNA world” Letter grades only. Disc, lab. Offered BIOBM 330. J. E. Blankenship, P. C. Hinkle, stems from the recognition that RNA is ancient alternate years; next offered 2008-2009. and staff. and that the evolution o f life as w e know it D. L. Deitcher. Visualization o f complex biomolecules using depended upon RNA evolving both For description, see BIONB 430.] Silicon Graphics computers. Group informational and catalytic capabilities. This presentations on current topics in molecular course explores these ideas but more generally [BIOBM 450(4500) Principles of Chemical Biology (also CHEM 450(4500]) biology. provides a comprehensive introduction to RNA Fall. 3 credits. Prerequisites: CHEM 357- BIOBM 432(4320) Survey of Cell Biology biology. Many o f the most interesting topics in the RNA biology, such as the mechanism o f the 358 or 359-360 or equivalent. Next offered Spring. 3 credits. Prerequisite: BIOBM 330, 2008-2009. T. P. Begley. RNA interference and its widespread 333, or 331, and previous or concurrent For description, see CHEM 450.] applications, will be covered in detail. Other registration in 332, or equivalent. topics require consideration o f essential RNA- Recommended: BIOGD 281. Lee. Evening [BIOBM 483(Sec 04)[4530] Molecular protein complexes such as ribosomes, prelims Mar. 4 and Apr. 17. V. M. Vogt. Aspects of Plant Development I (also spliceosomes, telomerase, and Signal Survey o f a wide array o f topics focusing on the BIOPL 483.4) recognition particles. Classical experiments as general properties o f eukaryotic cells. Topics Fall. 1 credit. 12 lec TBA. Next offered well as up-to-date research are covered in this include methods used for studying cells, the 2008-2009.). B. Nasrallah. course. A portion o f each class is devoted to structure and function o f the major cellular For description, see BIOPL 483 4]. discussion and questions. organelles, and analyses o f cellular processes BIOBM 485(4850) Bacterial Genetics such as mitosis, endocytosis, cell motility, BIOBM 439(4390) Molecular Basis of (also BIOMI/BIOGD 485(4850]) secretion, cell-to-cell communication, gene Human Disease (also BIOGD Fall. 2 credits. Prerequisite: BIOGD 281. expression, and oncogenesis. Some o f the 439(4390]) Recommended: BIOMI 290 and BIOBM material is covered in greater depth in BIOBM Fall. 3 credits. Prerequisites: biochemistry 330 or 331 and 332 or 333. Lec. 437, BIOGD 483, and BIOBM 632, 636, and and molecular biology (e.g., BIOBM 330, J. E. Peters. 639. 331-332, or 333) and genetics (e.g., For description, see BIOMI 485. BIOGD 281) or permission o f instructor. [BIOBM 434(4340) Applications of Recommended: cell biology (e.g., BIOBM BIOBM 631(6310) Protein Structure, Molecular Biology to Medicine, Dynamics, and Function Agriculture, and Industry 432 or BIOAP 316) and physiology (e.g., BIOAP 311 or 458). S-U or letter grades. Fall. 3 credits. Prerequisites: BIOBM 330 or Fall. 3 credits. Prerequisites: BIOBM 330 or Lec. W. L. Kraus. 333 or 331-332 and organic chemistry. 333 or 331/332. Recommended: BIOBM This course examines how changes in the Recommended: physical chemistry course. 432. S-U or letter grades. Lee. Next offered normal expression, structure, and activity o f S-U or letter grades. Lec. L. Nicholson. 2008-2009. S. Ely. gene products caused by genetic mutations, Presentations on the basic principles o f Lecture topics emphasize transgenic animal epigenetic phenomena, and environmental protein structure, dynamics, and function. and plant systems that constitute marketed or agents lead to human diseases. The material Specific topics include protein folding, near-market applications such as production focuses on how these changes lead to stability, dynamics, evolution, folded o f pharmaceuticals in milk, edible and nucleic alterations in normal cellular processes, as conformations, structure prediction, molecular acid vaccines, gene therapy, and high-tech well as the resulting physiological recognition, and basic enzyme kinetics. agricultural products. Additional non- consequences. Topics are selected from transgenic topics will include cancer BIOBM 633(6330) Biosynthesis of hormone insensitivity syndromes, inborn treatments and relevant aspects o f the human Macromolecules errors o f metabolism, gene fusions resulting in genome projects. An overview o f human Fall. 2 credits. Prerequisite: BIOBM 330 or hybrid proteins, gene amplification, gene immunology and its relationship to drug 333 or 331-332. Recommended: BIOGD inactivation, disruption o f signaling pathways, development will be provided. Students will 281. Lec. J. W. Roberts and D. B. Wilson. disruption o f metabolic pathways, and the also explore relevant scientific literature.] Synthesis o f DNA, RNA, and proteins, and molecular actions o f infectious agents and regulation of gene expression. BIOBM 435-436(4350-4360) environmental toxins. Examples o f diseases Undergraduate Biochemistry are selected to emphasize various aspects o f BIOBM 636(6360) Functional Seminar genetics, molecular biology, cell biology, Organization of Eukaryotic Cells 1 credit; may be repeated. Prerequisites: physiology, immunology, and endocrinology Spring. 3 credits. Prerequisites: BIOBM 330 upperclass standing; BIOBM 330, 333, or that have been presented in other courses. In or 333 or 331-332, and 432, or equivalents. 331-332, or written permission o f addition, the methods used to identify the Lec. W. J. Brown. instructor. S-U grades only. D. Wilson. underlying biochemical and genetic basis o f Aims to provide an integrated view of Selected papers from the literature on a given the diseases, as well as possible eukaryotic cell organization as elucidated topic are evaluated critically during 12 one- pharmaceutical and genetic therapies for using biochemical molecular, genetic, and cell hour meetings. treating the diseases, are presented. A portion biological approaches. Major topics include of the lecture periods will be devoted to the cytoskeleton, membrane traffic, and cell BIOBM 437(4370) Regulation of Cell discussion and practice questions. polarity. Together with BIOBM 437, 632, and Proliferation, Senescence, and Death 639 this course provides broad coverage o f (also BIOGD/TOX 437(4370]) BIOBM 440(4400) Laboratory in the cell biology subject area. Spring. Variable credit; students may take Biochemistry and Molecular Biology lec for 2 credits or lec and disc for 3 Fall, spring, or summer (three-week BIOBM 638(6380) Macromolecular credits. Limited to about 20 students per session). 4 credits. Limited enrollment. Interactions and Cell Function disc; priority given to graduate students. Priority given to undergraduate biology Spring. 2 credits. Prerequisite: BIOBM 330 Prerequisite: BIO G 101-102 or 105-106 majors in Biochemistry or Molecular and or 333 or 331-332. Recommended: BIOBM and BIOBM 330 or 331/332. Cell Biology programs o f study and to 631 or 633. S-U or letter grades. Lec. J. Fu. Recommended: BIOGD 281 and BIOBM graduate students with minor in field o f Lectures focusing on the principles o f protein- 432. S-U or letter grades. S. Lee. biochemistry. Prerequisites: BIOBM 330 or protein and protein-nucleic acid interactions Covers a wide spectrum o f issues related to 333 or 331-332 (at least one o f 331-332 that underlie cellular processes such as signal cell proliferation in eukaryotes. Lectures completed but one may be taken transduction, intracellular traffic, gene include various aspects o f the regulation o f concurrently). S. Ely and H. Nivison. regulation, and cell development. The cell division cycle and signal transduction Experiments related to molecular biology emphasis throughout is on the structural basis pathways, with additional topics on (includes PCR, DNA cloning, hybridization o f these processes as related to cell function. oncogenesis, cell aging, and cell death. The analysis, restriction mapping, and DNA Some specific topics are signal amplification, facts as well as concepts and logics behind sequence analysis), protein purification and nuclear import and export, transcription by findings are presented in the lectures. analysis (salt fractionation, ion exchange RNA polymerases, RNA processing and export, Research articles are analyzed and discussed chromatography, affinity chromatography, and translation o f mRNAs. in depth during discussion section. SDS-PAGE, and immunoblotting), and determination o f enzyme kinetic parameters. ECOLOGY AND EVOLUTIONARY BIOLOGY 165

[BIOBM 639(6390) The Nucleus BIOBM 831(8310) Advanced Biochemical ECOLOGY AND EVOLUTIONARY Spring. 2 credits. Prerequisite: BIOBM 330 Methods I or 333 or 331-332, or equivalent. Fall. 6 credits. Requirement for, and limited BIOLOGY (BIOEE) Recommended: BIOGD 281. Lee. Next to, first-year graduate students in field of BIOEE 154(1540) Introductory offered 2008-2009. J. T. Us. biochemistry, molecular, and cell biology. Oceanography, Lectures (also EAS Lectures on topics o f eukaryotic genome S-U grades only. Lab and disc. 154(1540]) organization, chromatin structure, regulation o f Organizational meeting first F o f semester Fall. 3 credits; optional 1-credit laboratory gene expression, RNA processing, the 10:10. T. C. Huffaker. offered as BIOEE/EAS 155. S-U or letter structure and movement o f chromosomes, and The first half o f this course comprises an grades. B. C. Monger. nuclear export and import. Covers the intensive laboratory covering fundamental For description, see EAS 154. structure and function o f the nucleus at the aspects o f modem molecular biology and cell molecular and cell biological levels and, biology. The second half comprises research BIOEE 155(1550) Introductory together with BIOBM 437, 632 and 636, in the laboratory o f a professor chosen by the Oceanography, Laboratory (also EAS provides broad coverage of the cell biology student (see BIOBM 832). Students must 155(1550]) subject area.) enroll separately for each half. Fall. 1 credit. Corequisite: BIOEE/EAS 154. S-U or letter grades. B. C. Monger. BIOBM 641(6410) Laboratory in Plant BIOBM 832(8320) Advanced Biochemical For description, see EAS 155. Molecular Biology (also BIOPL Methods II 641(6410]) Spring. 6 credits. Requirement for, and BIOEE 207(2070) Evolution (also HIST Fall. 4 credits. Prerequisites: BIOGD 281 or limited to, first-year graduate students in 287(2870], S&TS 287(2871]) equivalent, BIOBM 330 or 331 or field o f biochemistry, molecular, and cell Fall or summer (six-week session). 3 credits. equivalent, and permission o f instructor. biology. S-U grades only. Lab. V. Vogt. Intended for students with no background S-U grades by permission o f instructor. Research in the laboratories o f two different in college biology. May not be taken for Lab. M. R. Hanson and K. Van Wijk. professors chosen by the student. credit after BIOEE 278. Does not meet For description, see BIOPL 641. Arrangements are made jointly between the evolutionary biology requirement for director o f graduate studies and the research biological sciences major. S-U or letter BIOBM 730(7300) Protein NMR advisor. grades. W. B. Provine. Spectroscopy (also VETMM Evolution is the central concept in biology. This 707(7070]) BIOBM 833(8330) Research Seminar in course examines evolution in historical and Spring. 2 credits. Prerequisites: CHEM 389 Biochemistry cultural contexts. This course aims to and 390, or 287 and 288, or permission of Fall or spring. 1 credit each semester; may understand the major issues in the history and instructor. S-U or letter grades. Offered be repeated for credit. Requirement for, current status o f evolutionary biology and alternate years. Lee. L. K. Nicholson and and limited to, second-, third-, and fourth- explore the implications o f evolution for culture. R. E. Oswald. year graduate students majoring in field o f Issues range from controversies over Students acquire the tools necessary for biochemistry, molecular and cell biology. mechanisms o f evolution in natural populations understanding multidimensional NMR o f S-U grades only. W. L. Kraus and to the conflict between creationists and proteins. NMR fundamentals and schemes for V. M. Vogt. evolutionists. magnetization transfer, water suppression, Each student presents one seminar per year decoupling, and others are presented. on his or her thesis research and then meets BIOEE 261(2610) Ecology and the with instructors and thesis committee Environment BIOBM 732-737(7320-7370) Current members for evaluation. Fall or summer (three-week session). 4 Topics in Biochemistry credits. Prerequisite: one year introductory Fall or spring. 0.5 or 1 credit for each BIOBM 836(8360) Methods and Logic in biology. S-U or letter grades. Fall: topic; may be repeated for credit. Biochemistry, Molecular and Cell B. F. Chabot, A. A. Dhondt, and staff. Prerequisite: BIOBM 330 or 333 or 331-332 Biology, Part I Summer: one weekend field trip. or equivalent. S-U grades only. Spring. 1 credit. Prerequisite: first-year A. T. Vawter. Lectures and seminars on specialized topics. graduate students majoring in field o f Fall: Explores interactions between the Topics for fall and spring to be announced in biochemistry, molecular, and cell biology. environment and organisms in the context o f the course and time roster published at the S-U grades only. Sem and disc. G. P. Hess. individuals, populations, communities, and beginning o f each semester or the department Seminar with critical discussion by students o f ecosystems. Emphasizes basic ecological mini-courses web site, www.mbg.cornell.edu/ original research papers selected by faculty principles and processes intrinsic to cals/mbg/about/courses/mini-courses.cfm. members o f the field o f biochemistry, understanding the world around us and in molecular and cell biology. BIOBM 738(7380) Macromolecular more advanced studies in the environmental Crystallography (also CHEM BIOBM 838(8380) Methods and Logic in sciences, including management-oriented 788(7880]) Biochemistry, Molecular and Cell disciplines. Major topics include adaptive Fall. 3 credits. Prerequisite: permission of Biology, Part II strategies o f organisms, population dynamics, instructor. Lee. S. E. Ealick. Spring. 2 credits. Prerequisite: second-year species interactions, community structure and For description, see CHEM 788. graduate students majoring in field o f ecosystem function, biodiversity, biochemistry, molecular and cell biology or biogeochemistry, productivity, human BIOBM 751(7510) Ethical Issues and field of genetics and development. S-U influences on ecosystems, and sustainable Professional Responsibilities grades only. D. Shalloway. practices. Spring. 1 credit. Prerequisite: graduate Interactive seminar to develop the general Summer: Introduction to principles o f ecology, students beyond first year. S-U grades only. skills needed to support a career in scientific concerning the interactions between Organizational meeting first W o f semester. research: experimental design, writing organisms and their environment. Deals with Sem. P. Hinkle. scientific papers and grants, oral presentation, both terrestrial and aquatic ecology, drawing Ethical issues in research and the professional basic statistical and computational methods, examples from both plant and animal studies. responsibilities o f scientists are discussed based and managing a research laboratory. Exercises Phenomena that occur at the individual, on readings and occasional lectures. The topics focus on the preparation o f a mock research population, community, and ecosystem levels are intended to cover the requirements for grant proposal. o f organization are examined through ethical training o f graduate students on training classroom lectures and discussion and through grants and follow the recommendations o f the Related Courses in Other Departments a series o f lab and field experiences in natural Office o f Research Integrity. Lipids (BIOAP 619, NS 602) habitats around Ithaca and in the Adirondack BIOBM 830(8300) Biochemistry Seminar Forest Preserve. Ecological principles are Fall or spring. 0 credits. Prerequisite: Molecular Techniques for Animal Biologists applied extensively to current environmental graduate students in Biochemistry, (AN SC 650) problems and issues. Molecular, and Cell Biology. Lee open to Teaching Experience (BIO G 498) BIOEE 263(2630) Field Ecology everyone. V. Vogt. Fall. 3 credits. Pre- or corequisite: BIOEE Lectures on current research in biochemistry, Undergraduate Research in Biology (BIO G 26l. Letter grades only. One weekend field presented by distinguished visitors and staff 499) trip. A. Kessler. members. 166 BIOLOGICAL SCIENCES - 2 il' 8

Field exercises designed to give students topics such as the origin and evolution o f BIOEE 363(3630) Field Methods in direct experience with fieldwork, with various vertebrate groups, sensory systems, Ornithological Research emphasis on developing observational skills, thermoregulation, life history, locomotion, Summer (eight-week session). 5 credits. journal keeping, and a landscape perspective. feeding, size, and scaling. Laboratories include Limited to 15 students. Prerequisites: Topics include plant succession, niche dissections o f preserved vertebrate animals introductory biology or equivalent, interest relationships o f insects, influence o f and noninvasive live animal demonstrations. and ability to spend all day in the field herbivores and competition on plant under variable weather conditions, performance, decomposition of soil litter, BIOEE 278(2780) Evolutionary Biology including intense sun and periods o f rain, foraging behavior, census methods, and use o f Fall or spring. 3 or 4 credits; 4-credit and permission o f instructor. S-U or letter scientific collections. option involves writing component and grades. D. W. Winkler and staff. two disc per week; 4-credit option limited Detailed, hands-on experience with the BIOEE 264(2640) Tropical Field to 20 students per sec each semester. methods commonly used in ornithological Ornithology (Students may not preregister for 4-credit field research, focusing on different Winter, Jan. 3-17, 2008, two-week, full-time option; interested students complete methodologies used to obtain data on course. 3 credits. Limited to 12 students; application form on first day o f class.) organismal structure and function, life history minimum o f 8. Prerequisite: permission o f Limited to 300 students. Prerequisite: one characteristics, and behavior. While being instructor. Intended for students with year introductory biology or permission of immersed in an ongoing, intensive research limited or no bird knowledge. S-U or letter instructor; first-semester freshmen by program in the Ithaca area, students learn grades. Daily fieldwork, disc, reading, and permission o f instructor. S-U or letter about the types of evolutionary and ecological individual project. Cost o f tuition covers grades. One all-day Sat. field trip. Evening questions that can be addressed through use airfare, food, and lodging. A. A. Dhondt. prelims: spring, Feb. 28 and Apr. 1. Fall, o f different research methodologies, Provides students with the opportunity to M. A. Geber; spring, K. R. Zamudio. experimental design, and basic statistical study birds intensively in a neotropical Considers explanations for patterns o f diversity analysis techniques. environment. Students learn observational and and for the apparent good fit o f organisms to field techniques, participate in group research the environment. Topics include the genetic BIOEE 369(3690) Chemical Ecology (also projects and in daily seminars. The group is and developmental basis o f evolutionary BIONB/ENTOM 369(3690]) housed in the Biodiversity Center at Punta change, processes at the population level, the Spring. 3 credits. Prerequisites: one Cana. One or two field trips are taken to theory o f evolution by natural selection, levels semester o f introductory biology for majors national parks in the Dominican Republic. o f selection, concepts o f fitness and or nonmajors and one semester of introductory chemistry for majors or BIOEE 265(2650) Tropical Field Ecology adaptation, modes o f speciation, long-term and Behavior trends in evolution, rates o f evolution, and nonmajors or equivalents, or permission of extinction. Students taking the 4-credit option instructor. S-U or letter grades. A. Kessler, Winter, field course based in Kenya, Africa. read additional materials from the primary A. Agrawal, G. Jander, and J. S. Thaler. 4 credits. Limited to 15 students. literature and write a series o f essays in place Why are chilies so spicy? This course Prerequisites: one introductory biology o f the regular prelims. examines the chemical basis o f interactions course and permission o f instructors. Letter between species and is intended for students grades only. I. J. Lovette and BIOEE 350(3500) Dynamics of Marine with a basic knowledge o f chemistry and D. Rubenstein. Ecosystems (also EAS 350(3500]) biology. Focuses on the ecology and chemistry Gives students a broad hands-on Fall. 3 credits. Limited to 25 students. o f plants, animals, and microbes. Stresses understanding o f tropical biology, ecology, Prerequisites: one year o f calculus and chemical signals used in diverse ecosystems, and behavioral ecology. Students gain semester o f oceanography (i.e., BIOEE/EAS using Darwinian natural selection as a experience with experimental design and data 154), or permission o f instructor. S-U or framework. Topics include: plant defenses, collection, field methods, basic statistics, letter grades. Offered alternate years. microbial warfare, communication in marine interpretation and evaluation o f primary C. H. Greene and R. W. Howarth. organisms, and human pheromones. scientific literature, and scientific paper For description, see EAS 350. writing. Most in-country costs (e.g., ground BIOEE 371(3710) Human Paleontology transportation, room, board) are covered by BIOEE 351(3510) Marine Ecosystems (also AHTHR 371(3710]) course tuition, but students pay separately for Field Course (also EAS 351(3510]) Fall. 4 credits. Limited to 18 students. their international airfare and there may also Spring, full-time, three-week course. 4 Prerequisite: one year introductory biology be a small supplementary laboratory fee. credits. Limited to 25 students. Prerequisites: or ANTHR 101 or permission o f instructor. enrollment in Cornell Abroad Earth and Letter grades only. Occasional field trips. BIOEE 267(2670) Introduction to Environmental Sciences Semester in Hawaii; K. A. R. Kennedy. Conservation Biology one semester o f calculus and two semesters Broad survey o f the fossil evidence for human Fall. 3 credits. Intended for both science o f biology or permission o f instructor. Letter evolution with special attention to skeletal and and nonscience majors. May not be taken grades only. C. H. Greene, C. D. Harvell, and dental anatomy, geological contexts, for credit after NTRES 410. Completion o f B. C. Monger. paleoecology, dating methods, archaeological BIOEE 267 not required for NTRES 410. S-U For description, see EAS 351. associations, and current theories o f human or letter grades. One Sat. a.m. field trip. origins and physical diversity. Offered alternate years. J. W. Fitzpatrick. BIOEE 362(3620) Dynamic Models in Broad exploration of biological concepts and Biology (also MATH 362(3620]) BIOEE 373(3730) Biodiversity and practices related to conserving the earth’s Spring. 4 credits. Prerequisites: two Biology of the Marine Invertebrates biodiversity; integrates ecological, semesters introductory biology (BIO G Fall (but course must be taken previous evolutionary, behavioral, and genetic 101-102, 105-106, 107-108, 109-110 or summer at Shoals Marine Laboratory principles important for understanding equivalent) and completion o f mathematics [SML]), three-week, full-time course. 5 conservation issues o f the 21st century. Topics requirements for Biological Sciences major credits (students enroll for credit during include species and ecosystem diversity, or equivalent. S-U or letter grades. Offered fall semester). Limited to 24 students. values o f biodiversity, causes o f extinction, alternate years. S. P. Ellner and Prerequisites: one year introductory risks facing small populations, simulation J. M. Guckenheimer. biology for majors; permission o f faculty modeling, design o f nature preserves, the Introductory survey of the development, because o ff campus. Letter grades only. Endangered Species Act, species recovery, computer implementation, and applications o f Daily and evening lec, lab, and fieldwork. ecosystem restoration and management, and dynamic models in biology and ecology. Case- Total cost for room, board, and overhead past and future human impacts on the planet. study format covering a broad range o f at SML: $1,200. Offered alternate years. current application areas such as regulatory C. D. Harvell. BIOEE 274(2740) The Vertebrates: networks, neurobiology, cardiology, infectious Introduction to the biology and evolution of Structure, Function, and Evolution disease management, and conservation of the major invertebrate phyla, concentrating on Spring. 4 credits. Prerequisite: one year endangered species. Students also learn how marine representatives. In addition to the introductory biology. Fee: $25. S-U or letter to construct and study biological systems evolution o f form and function, lectures cover grades. B. A. McGuire. models on the computer using a scripting and aspects o f ecology, behavior, physiology, Introductory course in vertebrate organismal graphics environment. chemical ecology, and natural history o f biology that explores the structure and invertebrates. SML exposes students to a function o f vertebrates with an emphasis on wealth o f marine and terrestrial invertebrates trends in vertebrate evolution. Lectures cover in their natural habitats. Regular field ECOLOGY AND EVOLUTIONARY BIOLOGY 167 excursions allow an excellent opportunity to (thermal and mixing), chemical (dissolved [BIOEE 466(4660) Physiological Plant study freshly collected and in situ elements and compounds), and organismal Ecology, Lectures representatives o f most o f the major phyla. dynamics. Lakes are exciting environments for Spring. 3 credits. Limited to 30 students. study in their own right and for gaining Prerequisite: BIOEE 26l or introductory [BIOEE 405(4050) Biology of the perspective on ecological and evolutionary plant physiology. Letter grades; S-U grades Neotropics processes in general. by permission only. Offered alternate Spring. 2 credits. Limited to 18 students. years; next offered 2008-2009. J. P. Sparks.] Prerequisite: BIOEE 26l or permission o f BIOEE 458(4580) Community Ecology instructor. S-U or letter grades. Next Spring. 4 credits. Prerequisites: BIOEE 261, BIOEE 467(4670) Seminar in the History of Biology (also HIST 415[4150], offered 2008-2009. A. S. Flecker.] 278, or permission o f instructor. S-U or letter grades. Offered alternate years. B&SOC/S&TS 447[4471]) Fall or summer (six-week session). 4 credits. [BIOEE 446(4460) Plant Behavior M. A. Geber and A. Agrawal. Induced Plant Responses to Biotic Intersection between ecology and evolution of Limited to 18 students. S-U or letter grades. Stresses species interactions. Covers historical and W. B. Provine and G. Gorman. Spring. 3 credits. Limited to 12 students. current views on community structure and Specific topics change each year. Topic for fall Prerequisite: BIOEE 261 or permission o f diversity. Topics include impacts o f species 2007: Evolution and religion. H ow can instructor. S-U or letter grades. Next interactions on ecology and evolution o f evolutionists prepare for teaching biological offered 2008-2009. A. Kessler.] community players, multispecies webs and evolution in a nation where the great majority of people either disbelieve in evolution or [BIOEE 450(4500) Mammalogy, Lectures natural selection in complex communities. believe that purposive causes direct evolution? Spring. 3 credits. Recommended: BIOEE Approach is empirical and methodological. 274. Letter grades; S-U grades by BIOEE 459(4590) Limnology: Ecology of [BIOEE 468(4680) Physiological Plant permission only. Offered alternate years; Lakes, Laboratory Ecology, Laboratory next offered 2008-2009. B. A. McGuire.] Spring. 2 credits. Pre- or corequisite: Spring. 2 credits. Limited to 15 students. BIOEE 457. Letter grades; S-U grades by Pre- or corequisite: BIOEE 466. Letter [BIOEE 451 (4510) Mammalogy, grades only. Offered alternate years; next Laboratory permission only. One weekend field trip. offered 2008-2009. J. P. Sparks.] Spring. 1 credit. Limited to 16 students. Fee for food on field trip: $15. Offered Pre- or corequisite: BIOEE 450. Letter alternate years. N. G. Hairston, Jr. and staff. [BIOEE 469(4690) Food, Agriculture, and grades; S-U grades by permission only. Laboratories and field trips devoted to studies Society (also B&SOC/S&TS Fee: $15. Travel to Cornell University o f the biological, chemical, and physical 469[4691]) Museum o f Vertebrates (CUMV) at the properties of lakes and other freshwater Spring. 3 credits. Limited to 20 students. Laboratory o f Ornithology is necessary. environments. Exercises focus on Prerequisite: introductory ecology course One all-day field trip may be scheduled. understanding the freshwater environment, on or permission o f instructor. S-U or letter Offered alternate years; next offered 2008- experimentation, and on understanding grades. Next offered 2008-2009. 2009. B. A. McGuire.] ecological processes within lakes. Optional A. G. Power.] vertebrate dissection (fish) during one [BIOEE 453(4530) Speciation laboratory exercise and during a portion of [BIOEE 470(4700) Herpetology, Lectures Spring. 4 credits. Limited to 40 students. the weekend field trip. Spring. 2 credits. Limited to 50 students. Prerequisites: BIOEE 278 and BIOGD 281 Recommended: BIOEE 274 and concurrent or equivalents, or permission o f instructor. [BIOEE 460(4600) Theoretical Ecology enrollment in BIOEE 472. Letter grades; S-U or letter grades. Offered alternate years; Spring. 4 credits. Limited enrollment. S-U grades by permission only. Offered next offered 2008-2009. R. G. Harrison.] Prerequisites: completion o f Biological alternate years; next offered 2008-2009. Sciences mathematics requirement or H. W. Greene.] [BIOEE 455(4550) Insect Ecology (also equivalent, and either one additional ENTOM 455[4550]) semester o f mathematics, statistics, or [BIOEE 472(4720) Herpetology, Fall. 4 credits. Recommended: ENTOM 212 modeling (e.g., BEE 260/453/475, NTRES Laboratory or BIOEE 261 or permission o f instructor. 310/411, BIONB 422) or permission of Spring. 2 credits. Limited to 35 students. S-U or letter grades. Offered alternate instructor. S-U or letter grades. Offered Pre- or corequisite: BIOEE 470. Letter years; next offered 2008-2009. J. S. Thaler.] alternate years; next offered 2008-2009- grades; S-U grades by permission only. S. P. Ellner.] Fee: $30. Occasional field trips and special BIOEE 456(4560) Stream Ecology (also projects. Offered alternate years; next NTRES 456(4560]) [BIOEE 462(4620) Marine Ecology (also offered 2008-2009. H. W. Greene.] Fall. 4 credits. Limited to 40 students. EAS 462(4620]) Prerequisite: BIOEE 26l or permission o f Fall. 3 credits. Limited to 75 students. [BIOEE 473(4730) Ecology of Agricultural instructor. S-U or letter grades. Field Prerequisite: BIOEE 26l. Letter grades; S-U Systems (also HORT 473(4730]) project with lab papers. One Sat. field trip. grades by permission only. Offered Fall. 3 credits. Limited to 45 students. Offered alternate years. A. S. Flecker and alternate years; next offered 2008-2009. Prerequisite: BIOEE 26l or permission of C. E. Kraft. C. D. Harvell and C. H. Greene.] instructor. S-U or letter grades. During first For description, see NTRES 456. six weeks o f class, Thurs. meetings may run BIOEE 464(4640) Macroevolution later because o f field trips. Next offered BIOEE 457(4570) Limnology: Ecology of Spring. 4 credits. Limited to 35 students. Lakes, Lectures 2008-2009- L. E. Drinkwater and Prerequisite: BIOEE 278 or permission o f A. G. Power.] Spring. 3 credits. Prerequisite: BIOEE 261 instructor. Interested graduate students or written permission o f instructor. strongly encouraged to preregister. Letter BIOEE 475(4750) Ornithology Recommended: introductory chemistry. grades; S-U grades by permission only. Spring. 4 credits. Limited to 35 students. Letter grades; S-U grades by permission Offered alternate years. A. R. McCune. Prerequisite: permission o f instructor by only. Offered alternate years. Advanced course in evolutionary biology preregistering in E141 Corson Hall. N. G. Hairston, Jr. centered on large-scale features o f evolution. Recommended: BIOEE 274. Letter grades; Limnology is the study o f fresh waters and Areas o f emphasis include phylogeny S-U grades by permission only. Carpooling other inland, nonmarine environments. This reconstruction, patterns and processes o f to Lab o f Ornithology necessary. Fee: $15. course focuses on lakes and ponds, which are speciation, the origin o f evolutionary novelty, Occasional field trips and special projects. discussed as distinct aquatic environments causes o f major evolutionary transitions, and Offered alternate years. D. W. Winkler. with clear terrestrial boundaries, and within patterns o f diversification and extinction in the Lectures cover various aspects o f the biology which ecological interactions are especially fossil record. Discussion o f these problems o f birds, including anatomy, physiology, evident. In lakes, interactions between involves data and approaches from genetics, systematics, evolution, behavior, ecology, and organisms are often strong and adaptations morphology, systematics, paleobiology, biogeography. Laboratory includes dissection easily recognized. Physical and chemical development, and ecology. o f dead material, studies o f skeletons and properties o f the environment impact plumages, and specimen identification of organisms in important ways and organisms, avian families o f the world and species of likewise, influence physics and chemistry. As a N ew York. result, lakes provide excellent systems for understanding the links between physical 168 BIOLOGICAL SCIENCES - 2007-2008

[BIOEE 476(4760) Biology of Fishes [Sec 01 Tropical Field Ecology (BIOEE 675(6750) Current Topics in Plant Fall. 4 credits. Limited to 24 students. Spring. 2 credits. Prerequisite: for Molecular Ecology Recommended: BIOEE 274 or equivalent undergraduates, experience or course work Fall. 1 credit; may be repeated for credit. experience in vertebrate zoology. Letter with terrestrial, marine, or freshwater Limited to 20 students. Prerequisite: grades; S-U grades by permission only. organisms. Extended field trip over winter graduate standing or permission of Small lab fee may be required. Tw o field break. letter grades only. Fee to cover instructor. S-U or letter grades. Offered trips. Offered alternate years; next offered transportation and housing: TBA. Offered alternate years; next offered 2008-2009. 2008-2009. A. R. McCune.] alternate years; next offered 2008-2009. A. Kessler.] C. D. Harvell, J. P. Sparks, and BIOEE 477(4770) Marine Invertebrates BIOEE 750-760(7590-7600) Special Seminar N. G. Hairston, Jr.] Topics in Evolution and Ecology Fall. 1 credit. Prerequisite: BIOEE 373 or Sec 02 Graduate Field Course in Ecology Fall or spring. 1-3 credits; may be repeated permission o f instructor. S-U grades only. Spring. 3 credits. Prerequisite: graduate for credit. Limited enrollment. Letter Offered alternate years. C. D. Harvell and standing. Letter grades only. Fee charged grades; S-U grades by permission only. J. G. Morin. to help cover food and lodging for trip to Staff. Discussions and directed readings center on Florida. Offered alternate years. Independent or group-intensive study of current research themes in invertebrate J. P. Sparks. special topics o f current interest. Content biology. Designed as an on-campus Designed to give graduate students experience varies each semester. companion course to the field-based BIOEE in defining questions and designing field 373 Biology o f the Marine Invertebrates. investigations. The course is based at the BIOEE 761(7610) Microsatellite DNA: Techniques Students write individual research essays Archbold Biological Station in central Florida based on projects done in the field. over spring break and during the following Fall. 1 credit; may be repeated for credit. week. The class visits several ecosystems Limited to 12 students. Prerequisite; permission o f instructor. Primarily for [BIOEE 478(4780) Ecosystem Biology including sand pine scrub, cattle ranches, Spring. 4 credits. Prerequisite: BIOEE 26l cypress swamps, and the everglades. graduate students; undergraduates or equivalent. S-U or letter grades. Offered admitted only under exceptional alternate years; next offered 2008-2009. BIOEE 661(6610) Environmental Policy circumstances. E-mail S. M. Bogdanowicz C. L. Goodale and R. W. Howarth.] (also ALS 661(6610], B&SOC (smb31) by end o f Aug. if interested. S-U 461(4611]) grades only. Fee: TBA. R. G. Harrison and BIOEE 479(4790) Paleobiology (also EAS Fall and spring. 3 credits each semester; S. M. Bogdanowicz. 479(4790]) students must register for 6 credits each Construct and screen genomic DNA libraries Spring. 4 credits. Prerequisites: one year semester since R grade given at end o f fall for microsatellite loci. Lectures and group introductory biology for majors and either semester. Limited to 12 students. discussions regarding microsatellite isolation, BIOEE 274, 373, or permission of Prerequisite: permission o f instructor. Letter characterization, and evolution. Informal instructor. S-U or letter grades. Offered grades only. D. Pimentel. presentations o f student research projects. alternate years. W. D. Allmon. Focuses on complex environmental issues. For description, see EAS 479. Ten to 12 students, representing several BIOEE 764(7640) Plant-Insect disciplines, investigate significant Interactions Seminar BIOEE 480(4800) Ecological Genetics Fall or spring. 1 credit; may be repeated (also ENTOM 470(4700]) environmental problems. The research team for credit. Prerequisite: for undergraduates, Spring. 3 credits. Prerequisite: BIOEE 278 spends two semesters preparing a scientific report for publication in Science or permission o f instructor. S-U grades only. or permission o f instructor. S-U or letter A. Agrawal, J. S. Thaler, and A. Kessler. grades. B. P. Lazzaro. BioScience. Thus far, every study has been published. Group-intensive study o f current research in For description, see ENTOM 470. plant-insect interactions. Topics vary from BIOEE 490(4900) Topics in Marine BIOEE 668(6680) Principles of semester to semester, but include: chemical Biology Biogeochemistry defense, coevolution, insect community Spring. 2 credits; may be repeated for Spring. 4 credits. Limited to 20 students. structure, population regulation, biocontrol, credit. Limited to 15 students. Prerequisite: Prerequisite: solid background in ecology, tritrophic interactions, and mutualism. permission o f instructor. Primarily for environmental chemistry, or related environmental science; for undergraduates, BIOEE 767(7670) Currant Topics in undergraduates. S-U or letter grades. Ecology and Evolutionary Biology Offered alternate years. J. G. Morin and permission o f instructor. S-U or letter grades. Offered alternate years. Fall. 4 credits. Prerequisite: for M. J. Shulman. undergraduates, permission o f instructor. Seminar courses on selected topics in marine R. W. Howarth and C. L. Goodale. Lectures cover the biotic controls on the S-U grades only. P. P. Feeny. biology; may include laboratory or field trips. Critical evaluation and discussion o f theory Topics and time of organizational meeting are chemistry o f the environment and the chemical control o f ecosystem function. and research in ecology and evolutionary shown in departmental course offerings listed biology. Lectures by faculty and student-led on the web site. Emphasis is on cycles o f major elements and minor elements globally and in selected discussions o f topics in areas o f current BIOEE 660(6600) Field Studies in ecosystems, stressing the coupling o f element importance. Ecology and Evolutionary Biology cycles. A comparative approach is used to BIOEE 770-775(7700-7705) Workshop in Fall or spring. Variable credit. Prerequisites: illustrate similarities and differences in element Biogeochemistry BIOEE 261, taxon-oriented course, and cycling among ecosystems. Analysis o f both Fall or spring. 1-3 credits; may be repeated permission o f instructor. Letter grades; S-U theoretical and applied issues, including for credit. Limited to 15 students. grades by permission only. Lee and field global atmospheric changes and factors Prerequisite: BIOEE 668. S-U grades only. trips TBA. Estimated costs: TBA. Staff. controlling the acidification o f lakes and soils. Staff. Provides students with opportunities to learn [BIOEE 671(6710) Palaeoanthropology of Workshop-forum in which graduate students field techniques and new biota by interact with invited world leaders in participating in an intensive series o f field South Asia (also ANTHR 671(6371], ASIAN 671(6671]) biogeochemistry. Workshop topics change exercises. Extended field trips may be each semester. A one-week workshop is scheduled during fall break, intersession, or Fall. 3 credits. Limited to 15 students. Letter grades only. Next offered 2008-2009. preceded by seven one-hour preparatory spring break. The regions visited, trip discussions o f readings. objectives, and other details are announced by K. A. R. Kennedy.] the various instructors at an organizational [BIOEE 673(6730) Human Evolution: BIOEE 780(7800) Graduate Seminar in meeting held at the beginning o f the semester. Concepts, History, and Theory (also Ornithology (also NTRES 780(7800]) Meetings on campus are devoted to ANTHR 673(6373]) Fall or spring. 1 credit; may be repeated orientation and reports on completed projects. Spring. 3 credits. Prerequisite: one year for credit. Prerequisite: for undergraduates, introductory biology or ANTHR 101 or permission o f instructor. S-U grades only. permission o f instructor. Letter grades only. I. J. Lovette, A. A. Dhondt, D. W. Winkler, Next offered 2008-2009. K. A. R. Kennedy.] and J. L. Dickinson. Group intensive study o f current research in ornithology. Topics vary from semester to semester. GENETICS AND DEVELOPMENT 169

BIOEE 899(8990) M.S. Thesis Research Geographic Information Systems (CSS 420) General study o f the fundamental principles o f Fall or spring. 1-15 credits. Prerequisite: genetics in eukaryotes and prokaryotes. Topics admission to field o f ecology and Practicum in Forest Farming as an include gene transmission, gene linkage and evolutionary biology. S-U or letter grades. Agroforestry System (CSS/HORT/ recombination, gene structure, gene and E&EB field faculty. NTRES 426) chromosome mutations, gene expression, and Thesis research conducted by an M.S. student the manipulation o f genes. Soil Ecology (CSS/HORT 466) in the field o f ecology and evolutionary BIOGD 282(2820) Human Genetics biology with advice and consultation o f a Introductory Geological Sciences (EAS 101) Spring. 2 or 3 credits; 2 credits if taken major professor who is a member of the field. after BIOGD 281. Prerequisite: one year Evolution of the Earth System (EAS 301) BIOEE 999(9990) Ph.D. Dissertation introductory biology or equivalent. S-U or Research Introduction to Biogeochemistry (EAS/NTRES letter grades. Lee. M. L. Goldberg. Fall or spring. 1-15 credits. Prerequisite: 303) Designed for nonmajors. Lectures provide the admission to field o f ecology and technical background needed to understand evolutionary biology as Ph.D. student. S-U Insect Biology (ENTOM 212) controversial personal, social, and legal or letter grades. E&EB field faculty. implications o f modem genetics that are Dissertation research conducted by a Ph.D. Insect Behavior (ENTOM 325) discussed in section meetings. student in the field o f ecology and BIOGD 385(3850) Developmental Biology evolutionary biology with advice and Introductory Insect Systematics (ENTOM Fall. 3 credits. Prerequisite: BIOGD 281. consultation o f a major professor w ho is a 331) Lee. K. Liu. member o f the field. Maggots, Grubs, and Cutworms: Larval Introduction to the morphogenetic, molecular Related Courses In Other Departments Insect Biology (ENTOM 333) and cellular, and genetic aspects o f the developmental biology o f animals. Evolutionary Theory and Human Behavior Techniques of Multivariate Analysis (ILRST (ANTHR 375/675) 410) BIOGD 394(3940) Circadian Rhythms (also ENTOM/BIONB/PL PA Ethics and the Environment (B&SOC/S&TS Statistical Analysis of Qualitative Data (ILRST 394(3940]) 206, PHIL 246) 411) Fall. 2-3 credits. Prerequisite: ENTOM 212 or BIOGD 281 or BIONB 221 or 222 or Physical Hydrology for Ecosystems (BEE Human Biology and Evolution (NS/ANTHR permission o f instructor. S-U or letter 371) 275) grades. Lee. K. Lee. This course will explore a fundamental feature Evolution of the Earth and Life (BIO G 170, Environmental Conservation (NTRES 201) o f living organisms found in all kingdoms: EAS 102) Applied Population Ecology (NTRES 310) how the cellular 24-hour biological clock General Microbiology, Lectures (BIOMI 290) operates and influences the biological Global Ecology and Management (NTRES activities. The course will cover fundamental Prokaryotic Diversity (BIOMI 414) 322) properties o f biological rhythms and cellular and molecular structure o f circadian oscillators Microbial Ecology (BIOMI 418) Forest Ecology (NTRES 420) o f model organisms including cyanobacteria, fungi, insects, plants, and mammals. One- Neurobiology and Behavior I: Introduction to Forest Ecology, Laboratory (NTRES 421) credit (optional) lab module offered in Behavior (BIONB 221) conjunction with lec. Wetland Ecology and Management, Lec/Lab Methods in Animal Behavior (BIONB 323) (NTRES 422/423) BIOGD 400(4000) A Genomics Approach to Studying Life Ecology of Animal Behavior (BIONB/BIOSM Fungi (PL PA 309) Fall. 3 credits. Prerequisites: one year 329) introductory biology plus BIOGD 281 or 330 or 333 or 331/332 or permission o f Modeling Behavioral Evolution (BIONB 422) instructor. S-U or letter grades. Lec. Animal Communication (BIONB 426) GENETICS AND DEVELOPMENT J. Schimenti. Introduction to principles underlying the Animal Social Behavior (BIONB 427) (BIOGD) organization of genomes and the methods of BIOGD 132(1320) Orientation Lectures in studying them, emphasizing genome-wide Introductory Botany (BIOPL 241) Molecular Biology and Genetics (also approaches to research. Covers the application BIOBM 132(1320]) o f genomics methodologies for addressing Taxonomy of Vascular Plants (BIOPL 248) Spring, weeks 1-3. 0 credits. Primarily for issues including evolution, complex systems, Phylogenetic Systematics (BIOPL/ENTOM freshmen, sophomores, and transfer genetics and gene: phenotype relationships. 440) students. S-U grades only. Lee. Staff. Includes periodic, in-depth discussions o f For description, see BIOBM 132. landmark or timely genomics papers. Molecular Systematics (BIOPL 447) BIOGD 280(2800) Lectures in Genetics BIOGD 401(4010) Genomic Analysis Plant Evolution and the Fossil Record (BIOPL Fall, spring, or summer (eight-week Spring. 3 credits. Prerequisites: BIOGD 400 448) session). 3 credits. Lee component o f or permission o f instructor. Lec. BIOGD 281. Not open to students majoring T. P. O ’Brien. Principles and Practice of Historical in biological sciences; may not be used to Overview of approaches and tools used in Biogeography (BIOPL/ENTOM 453) fulfill requirements for biological sciences genomics research. Covers experimental and major. Prerequisites: one year introductory computational technologies as well as Field Ornithology (BIOSM 374) biology or equivalent, or permission of theoretical concepts important for the study o f instructor. Lee. Highly recommended: genomes and their function. Topics include Field Marine Biology and Ecology (FMBE) problem-solving sessions. T. D. Fox, high-throughput DNA sequencing and (BIOSM 375) R. J. MacIntyre, and D. Nero. genotyping, genetic mapping of simple and Seaweeds, Plankton, and Seagrass: The For description, see BIOGD 281. complex traits, RNA expression profiling, proteomics, genome modification and Ecology and Systematics of Marine BIOGD 281(2810) Genetics transgenesis, and computational genomics. Plants (BIOSM 449) Fall, spring, or summer (eight-week session). 5 credits. Not open to freshmen Biological Statistics I (BTRY/STBTRY 301, fall semester. Prerequisite: one year NTRES 313) introductory biology or equivalent. Lee, Statistical Genomics (BTRY/STBTRY 482) lab. Highly recommended: problem-solving sessions. T. D. Fox, R. J. MacIntyre, and Soil Science (CSS 260) D. Nero. 170 BIOLOGICAL SCIENCES - 2TB B TTi 8

BIOCD 437(4370) Regulation of Cell BIOGD 452 Molecular Biology of Plant developing basic principles the course Proliferation, Senescence, and Death Organelles (also BIOPL 482, Sec 5) discusses the evolution and organization o f (also BIOBM/TOX 437(4370]) Spring. 1 credit. Prerequisites: BIOPL 483 genomes from microbes to higher eukaryotes Spring. Variable credit; students may take Sec 1 or BIOGD 281 and permission of including humans, and the relationship lec for 2 credits or lec and disc for 3 instructor. S-U or letter grades. between molecular evolution at the sequence credits. Limited to about 20 students per M. R. Hanson and D. B. Stem. level and the evolution of developmental disc; priority given to graduate students. For description, see BIOPL 482, Sec 5. pathways and systems. Prerequisites: BIO G 101-102 and BIOBM 330 or 331/332. Recommended: BIOGD [BIOGD 461(4610) Development and BIOGD 485(4850) Bacterial Genetics 281 and BIOBM 432. S-U or letter grades. Evolution (also BIOMI/BIOBM 485(4850]) S. Lee. Spring. 3 credits. Prerequisites: BIOEE 278, Fall. 2 credits. Prerequisite: BIOGD 281. For description, see BIOBM 437. BIOGD 281, BIOBM 332 or 330 or 333. Recommended: BIOMI 290 and BIOBM Recommended: BIOGD 385. Lec. Offered 330 or 331 and 332 or 333. Lec. BIOGD 439(4390) Molecular Basis of alternate years; next offered 2008-2009. J. E. Peters. Human Disease (also BIOBM M. Wolfner. For description, see BIOMI 485. 439(4390]) This course explores the molecular and Fall. 3 credits. Prerequisites: biochemistry genetic pathways and mechanisms that BIOGD 486(4860) Advanced Eukaryotic and molecular biology (e.g., BIOBM 330, regulate animal development, and how they Genetics 331/332, or 333) and genetics (e.g., BIOGD are modified through evolution to result in the Spring. 4 credits. Enrollment may be 281) or permission o f instructor. dazzling array o f forms and functions seen in limited to 50 students. Prerequisites: Recommended: cell biology (e.g., BIOBM the animal kingdom.] BIOGD 281, BIOBM 330 or 333 or 331 and 432 or BIOAP 316) and physiology (e.g., 332. S-U or letter grades. E. E. Alani. BIOAP 311 or 458). S-U or letter grades. [BIOGD 481(4810) Population Genetics Develops fundamental skills in eukaryotic Lec. W. L. Kraus. Fall. 4 credits. Prerequisite: BIOGD 281, genetic analysis through lectures and by For description, see BIOBM 439. BIOEE 278, or equivalents. Lec, disc. Next reading, analyzing, and presenting research offered 2008-2009. C. F. Aquadro. articles. Concepts are presented within the BIOGD 400(4000) A Genomics Approach Population genetics is the study o f the context o f a well-studied field, such as to Studying Life transmission o f genetic variation through time chromosome segregation. The basic tools that Fall. 3 credits. Prerequisites: one year and space. This course explores how to have been developed to study this field are introductory biology plus BIOGD 281 or quantify this variation, what the distribution o f used to analyze other topics such as 330 or 333 or 331/332 or permission o f variation tells us about the structure o f natural vegetative and meiotic cell cycle control, instructor. S-U or letter grades. Lec. populations, and about the processes that lead embryonic development, pathogen resistance J. Schimenti. to evolution. Topics include the diversity and in plants, and human genetics. Introduction to principles underlying the measurement o f genetic variation, mating and organization o f genomes and the methods o f reproductive systems, selection and fitness, BIOGD 487(4870) Human Genomics studying them, emphasizing genome-wide genetic drift, migration and population Fall. 3 credits. Prerequisite: BIOGD 281. approaches to research. Covers the application structure, mutation, multilocus models, the Lec. A. G. Clark. of genomics methodologies for addressing genetics o f speciation, quantitative traits, and Applies fundamental concepts o f transmission, issues including evolution, cloning, stem cells, the maintenance o f molecular variation. population, and molecular genetics to the complex systems, genetics and gene: Emphasis is placed on DNA sequence variation problem of determining the degree to which phenotype relationships. Includes periodic, in- and the interplay between theory and the data familial clustering o f diseases in humans has a depth discussions o f landmark or timely from experiments and natural populations. genetic basis. Emphasizes the role o f full genomics papers. Specific case studies include the population genome knowledge in expediting this process o f gene discovery. Stresses the role o f BIOGD 440(4400) Stem Cell Biology: genetic issues involved in DNA fingerprinting, Basic Science and Clinical the genetic structure and evolution o f natural statistical inference in interpreting genomic Applications and domesticated populations, and the study information. Population genetics, and the central role o f understanding variation in the Spring. 3 credits. Limited to 30 students. o f adaptation at the molecular level. Examples human genome in mediating variation in Prerequisites: BIOBM 432 or BIOGD 385 are drawn from studies o f animals, plants, and disease risk, are explored in depth. Methods or permission o f instructor. S-U grades by microbes.] such as homozygosity mapping, linkage permission o f instructor. Lec, disc. BIOGD 482(4820) Human Genetics and disequilibrium mapping, and admixture T. Tumbar. Society mapping are examined. The format is a series This course will cover basic aspects o f tissue Fall. 4 credits. Limited to 24 students. o f lectures with classroom discussion. morphogenesis and homeostasis with Prerequisite: biological sciences majors; Assignments include a series o f problem sets emphasis on the biological role of embryonic priority given to seniors studying genetics and a term paper. and adult stem cells in development, and their and molecular and cell biology and possible clinical applications. The focus will biochemistry; BIOGD 281 and BIOBM 330 BIOGD 489(4890) Mammalian be placed on mouse and human stem cells. or 333 or 331 and 332. R. A. Calvo. Embryology (also BIOAP 489(4890]) The discussion will be structured around Presentation o f some o f the science and Spring. 3 credits. Prerequisite: introductory relevant research papers that allow more in- technology o f human genetics, plus discussion biology. Offered alternate years; next depth analysis o f the material taught during o f the ethical, social, and legal implications o f offered 2009-2010. D. M. Noden. lectures. recent advances in the field. Topics include Examines the early formation o f the BIOGD 450(4500) Vertebrate assisted reproductive strategies, eugenics, genetic mammalian body and placenta, emphasizing Development counseling, genetic screening (pre-implantation, comparative aspects, and morphogenesis and histogenesis o f each organ system. Fall. 3 credits. Prerequisites: BIOGD 281, prenatal, neonatal, pre-symptomatic, carrier, and and either BIOBM 432 or BIOGD 385. S-U workplace), wrongful life and wrongful birth, BIOGD 490(4900) Manipulating the or letter grades. M. J. Garcia-Garcia. genetic effects o f abused substances, genetics Mouse Genome (also NS 490(4900]) This course explores the developmental and behavior, human cloning, forensic uses o f Fall. 2 credits. Course meets during first genetics, and therapy for genetic diseases. mechanisms employed by vertebrate half o f semester (R 1:25-3:20; Aug. 23-Oct. Students lead some discussions. There is a major organisms. Topics include the detailed analysis 4) and provides background information writing component to the course. o f the genetic, molecular, and cellular events for VTBMS 701/TOX 701 Mouse Pathology underlying development in frogs, fish, mice, BIOGD 484(4840) Molecular Evolution and Transgenesis, which meets during and humans. Course readings include original Spring. 3 credits. Prerequisites: BIOGD second half. Students interested in both research articles. Students are encouraged to 281. Lec. D. Barbash. courses must register for them separately. participate in class discussions. Explores the various processes by which DNA Prerequisites: BIOGD 281 and BIOBM 330, and protein sequences evolve over time, and 332, or 333 or NS 320. Letter grades only. how this evolution at the molecular level P. D. Soloway. relates to changes in the morphology, Functional genomic analysis has benefited behavior, and physiology of organisms that enormously from experimental manipulation of have occurred over time scales ranging from the genomes o f many organisms. The mouse thousands to billions o f years. After has been the model o f choice for such studies GENETICS AND DEVELOPMENT 171 in mammals. This course explores the tools BIOGD 613(6130) Genomics and Society BIOGD 780(7800) Current Topics in available for experimental manipulation o f the Spring, weeks 10-13. 1 credit. Prerequisite: Genetics and Development mouse genome, including transgenesis, gene BIOGD 281 or BIOGD 400 or permission Fall or spring. 1 credit; may be repeated for targeting, gene trapping, chemical mutagenesis, o f instructor. S-U or letter grades. Disc. credit. Limited to 20 students. Primarily for and cloning by nuclear transplant. Also S. Kresovich and S. Tanksley. graduate students; priority given to majors discussed are use o f recombinant inbred mice A multidisciplinary examination o f four to six in field o f genetics. Prerequisite: for for complex trait analysis. Readings from the selected topics that relate to the applications undergraduates, written permission of scientific literature focus on seminal o f biological insights derived from genomic instructor. No auditors. S-U grades only, by applications o f these methods. analysis. Technical, scientific, ethical, political, permission o f instructor. Seminar TBA. Staff. legal, and/or social aspects o f each topic will BIOGD 781(7810) Problems in Genetics BIOGD 608(6080) Epigenetics (also NS be considered from various perspectives. 608(6080]) and Development Fall. 2 credits. Prerequisites: BIOGD 281 BIOGD 620(6200) Evolutionary Genomics Fall. 2 credits. Prerequisite: first-year and BIOBM 330, 332, or 333 or NS 320. of Bacteria graduate students in field o f genetics and Letter grades only. Planned W F 11:15— Spring. 1 credit. M. Stanhope. development. Disc TBA. Staff. 12:05; occasional evening meetings for Comparative genomics o f bacteria is a Introduction to the research literature in student presentations. P. D. Soloway. valuable approach to deriving information on selected areas through weekly problem sets Epigenetic effects refer to reversible alterations pathogenesis, antibiotic resistance, host and discussions. in chromatin structure that can stably and adaptation, and genome evolution. This course BIOGD 782-783(7820-7830) Current heritably influence gene expression. These provides an evolutionary perspective on Genetics/Development Topics comparative bacterial genomics, focusing in changes include covalent modifications to Spring. 0.5 or 1 credit for each topic; may particular on pathogens o f human and DNA itself or to proteins bound to DNA as be repeated for credit. S-U grades only. Lee agricultural importance. The course will well as noncovalent remodeling o f chromatin. and sem on specialized topics. Staff. This course examines selected epigenetic include lectures, discussion o f relevant phenomena described in several eukaryotes, scientific literature, and hands-on BIOGD 784(7840) Introduction to mechanisms regulating these effects, and their bioinformatics sessions. Quantitative Analysis Fall. 1 credit. Letter grades only. E. Alani phenotypic consequences when normal [BIOGD 638(6380) Filamentous Fungal regulation is lost. Reading materials are from Genomics and Development (also (organizer), A. Clark, C. Bustamante, and current literature, and participation in class PL PA 638(6380]) M. Goldberg. discussion is required. The goal o f this course is to introduce Spring, last four weeks o f semester. bioinformatic and probability/statistical tools at 1 credit. S-U or letter grades. Prerequisite: BIOGD 610(6100) Genomes as an intuitive level that will be meaningful to BIOGD 281 or equivalent, or permission of Chromosomes first-year graduate students in Genetics and instructor. Lee. Offered alternate years; Fall. 3 credits. Limited to 15 students. Development. Prerequisites: BIOGD 281 and BIOBM 330 next offered 2008-2009. B. G. Turgeon. or 333 or 331/332 or equivalent by For description, see PL PA 638.] BIOGD 786(7860) Research Seminar in Genetics and Development permission o f instructor. Letter grades only. BIOGD 682(6820) Fertilization and the T. P. O ’Brien and P. E. Cohen. Early Embryo Fall and spring. 1 credit. Requirement for, The eukaryotic genome is partitioned into and limited to second-, third-, and fourth- Spring. 2 credits. Prerequisites: BIOGD discrete structural units, the chromosomes. year graduate students in genetics and 281; BIOBM 332, 330 or 333; and BIOGD The course examines how chromosome development. S-U grades only. Staff. 385 or permission o f instructor. Lee. organization is related to chromatin structure, Each graduate student presents one seminar Offered alternate years. M. F. Wolfner. gene expression, DNA replication, repair and per year based on his or her thesis research. We explore the latest molecular/cell/genetic stability. Special emphasis is placed on how The student then meets with the thesis findings about the biology o f gametes, the linear arrangement o f sequence features committee members for an evaluation o f the fertilization and early development— and their along the chromosome, such as genes and presentation. application to fertility modulation, “cloning” regulatory modules, relate to the functional and stem cells. BIOGD 787(7870) Seminar in Genetics organization o f the genome in the nucleus. and Development Experimental and computational approaches [BIOGD 687(6870) Developmental Fall and spring. 1 credit. Prerequisite: used to address chromosome structure and Genetics graduate students in Genetics and function are studied. Fall. 2 credits. Limited to 20 students. Development. S-U grades only. Sem, TBA. Prerequisites: BIOGD 281 and 385 or BIOGD 611(6110) Genome Maintenance Staff. equivalents. S-U or letter grades. Lee TBA. Mechanisms Seminars in current research in genetics and Offered alternate years; next offered 2008- Fall. 1 credit. Meets only during second half developmental biology conducted by 2009. K. J. Kemphues. o f semester beginning Oct. 15. Limited to 25 distinguished visitors and staff. Course focuses on methods o f genetic analysis students. Prerequisites: BIOGD 281, as well o f fruitflies, nematodes, mice, and fish to as BIOBM 330, or 333, or 331/332 (or understand mechanisms o f development. No Related Courses in Other Departments equivalents). S-U or letter grades. R. Weiss. text. Lectures and problems from literature.] The course focuses on the molecular Advanced Plant Genetics (PL BR 606) mechanisms utilized by eukaryotic cells to [BIOGD 689(6890) Cellular Basis of Biosynthesis of Macromolecules (BIOBM preserve genomic integrity. Topics to be Development 633) discussed include endogenous and exogenous Fall. 2 credits. Limited to 20 students. sources o f mutation, DNA repair pathways, Prerequisites: BIOGD 281, 385, and either Current Topics in Biochemistry (BIOBM 732- and cell cycle checkpoint mechanisms. Also BIOBM 330 or 331-332. Lee. S-U or letter 737) addressed will be how genome maintenance grades. Next offered 2010-2011. J. Liu. impacts genome plasticity and evolution, as Focuses on the integration o f different cellular Evolutionary Biology (BIOEE 278) well as the relationship between genomic processes in various developmental contexts. instability and disease, especially cancer. Topics include cell polarity, cell migration, cell Laboratory in Molecular Biology and Genetic adhesion and fusion, cell growth and Engineering of Plants (BIOPL 347) BIOGD 612(6120) Overview of Model Genetic Organisms proliferation, cell-cell communication, and cell death. Students are required to read current Laboratory in Plant Molecular Biology (BIOPL Spring. 1 credit. Limited to 20 students. literature and participate in discussions in 641) Prerequisites: BIOGD 281 or 400 or class.] permission o f instructor. S-U or letter Molecular Biology and Genetic Engineering grades. J. Schimenti and staff. of Plants (BIOPL 343) Presents the features o f various model organisms and their relative merits for Plant Cytogenetics (PL BR 446) conducting various types o f genomics/genetics Plant Genome Organization (PL BR 483.3) research. Model systems discussed include: yeast, Arabidopsis, Drosophila, C. elegans, Plant Molecular Biology I (BIOPL 483) zebrafish, and mice. 172 BIOLOGICAL SCIENCES - 2007-2008

Plant Molecular Biology II (BIOPL 482) structure and function. Students are macromolecule sequence-based methods to encouraged to take this course during their assess community diversity and microbial The Nucleus (BIOBM 639) third year o f study. activity in a variety of ecosystems. Other Undergraduate Research in Biology (BIO G topics discussed include bacterial growth and BIOMI 394(3940) Applied and Food survival, population biology, and microbial 499) Microbiology (also FD SC 394(3940]) interactions. Fall. 2-3 credits. Prerequisites: BIOMI 290- Molecular Neurobiology BIONB 420/720 291. C. A. Batt. [BIOMI 420(4200) Microbial Genomics (also BIOBM 435/735) For description, see FD SC 394. Spring. 2 credits. Prerequisites: BIOMI 290, [BIOMI 397(3970) Environmental BIO G 281, BIOBM 330, or equivalent. Microbiology (also CSS 397(3970]) Offered alternate odd-numbered years; next offered 2008-2009. J. P. Shapleigh and Spring. 3 credits. Prerequisites: BIOEE 26l J. D. Helmann. MICROBIOLOGY (BI0MI) or BIOMI 290 or CSS (SCAS) 260 or Genomic information is revolutionizing BIOMI 290(2900) General Microbiology permission o f instructor. Offered alternate biology. This course discusses the impact o f Lectures odd-numbered years; next offered 2008- genomic information on the study o f microbial Fall, spring, or summer (six-week session). 2009. E. L. Madsen. physiology, evolution, and biotechnology. 3 credits. Prerequisites: one year Discusses the biological properties, evolution, Topics include both techniques (automated introductory biology for majors and one and behavior o f microorganisms in natural DNA sequencing, assembly, annotation, DNA year college chemistry, or equivalent. systems in relation to past and present chips) and applications (genom e-wide analysis Highly recommended: concurrent environmental conditions on Earth and other o f transcription, functional genomics).] registration in BIOMI 291. W. C. Ghiorse. living planets. Also considers the functional Comprehensive overview of the biology of role o f microorganisms in ecologically and BIOMI 431(4310) Medical Parasitology microorganisms, with emphasis on bacteria. environmentally significant processes through (also VETMI 431(4310]) Topics include microbial cell structure and discussion o f specific topics such as nutrient Fall. 2 credits. Prerequisites: zoology and function, physiology, metabolism, genetics, and toxic elemental cycles, transformation o f biology courses. D. Bowman. diversity, and ecology. Some material may pollutant chemicals, wastewater treatment, For description, see VETMI 431. overlap with BIOGD 281 and BIOBM 330. environmental biotechnology, and Also covers applied aspects of microbiology astrobiology] BIOMI 482(4640)(Sec 02) Molecular such as biotechnology, the role of Plant-Microbe Interactions (also BIOMI 404(4040) Pathogenic Bacteriology BIOPL 482, Sec 02, PL PA 464(4640]) microorganisms in environmental processes, and Mycology (also VETMI 404(4040]) Spring. 1 credit. Prerequisites: BIOGD 281, and medical microbiology. 4-credit option Spring. 2 or 3 credits; 3 credits with lec BIOBM 330 or 331 or 333, and BIOPL 483 involves one discussion per w eek led by and sem. Prerequisites: BIOMI 290 and (sec 01) or equivalents. S-U or letter faculty in the Department o f Microbiology and 291; for undergraduates, permission o f grades. Offered alternate even years. will involve readings and a writing instructor. Highly recommended: BIO G S. C. Winans. assignment. Students may not pre-register for 305. Offered alternate even years. For description, see BIOPL 482, Sec 02. 4-credit option: Interested students complete D. Debbie. an application form on first day o f class For description, see VETMI 404. BIOMI 485(4850) Bacterial Genetics (enrollment will be limited to 20 students). (also BIOGD/BIOBM 485(4850]) BIOMI 291(2910) General Microbiology BIOMI 409(4090) Principles of Virology Fall. 2 credits. Prerequisite: BIOGD 281. Laboratory (also VETMI/PL PA 409(4090]) Recommended: BIOMI 290 and BIOBM Fall. 3 credits. Prerequisites: BIOMI 290, Fall or spring, summer (six-week session). 330 or 331 and 332 or 333. J. E. Peters. 291 or permission o f instructor. Students gain a detailed understanding o f how 2 credits. Pre- or corequisite: BIOMI 290. Recommended: BIOBM 330-332, 432. bacteria maintain and pass on genetic S. M. Merkel. Letter grades only. G. R. Whittaker and information with a strong focus on the Study o f the basic principles and techniques S. G. Lazarowitz. bacterium Escherichia coli. They discover the o f laboratory practice in microbiology, and For description, see VETMI 409. processes by which bacteria evolve through fundamentals necessary for further work in different mutations and the exchange o f the subject. [BIOMI 414(4140) Prokaryotic Diversity genetic information. The course explores how BIOMI 292(2920) General Microbiology Spring. 3 credits. Prerequisites: BIOMI 290 genes are regulated efficiently through Discussion and 291. Recommended: BIOBM 330 or 331 negative and positive regulation and by global or 333. Offered alternate odd-numbered Spring. 1 credit. Pre- or corequisite: BIOMI regulatory mechanisms. Upon completion o f 290. S-U grades only. Staff. years; next offered 2008-2009. S. H. Zinder. the course students should understand the Consideration of the evolutionary biology, Series o f discussion groups in specialized tools used to manipulate bacterial genomes physiology, ecology, genetics, and practical areas o f microbiology to complement BIOMI for the understanding o f bacteria and other potential o f important groups o f prokaryotes. 290. living organisms. Topics include prokaryotic phylogeny, the BIOMI 331(3310) General Parasitology evolution of diverse mechanisms of energy BIOMI 610(6100) Introduction to (also VETMI 331 [3310]) conservation, fixation of carbon and nitrogen, Chemical and Environmental Spring. 2 credits. Prerequisites: one year and adaptation to extreme environments.] Toxicology (also TOX 610(6100]) introductory biology. D. Bowman. Fall. 3 credits. Prerequisite: graduate Introduction to the basic animal parasites, BIOMI 416(4160) Bacterial Physiology standing in field or permission o f stressing systematics, taxonomy, general Spring. 3 credits. Prerequisites: BIOMI 290, instructor. Letter grades. A. Hay. biology, ecological interactions, and behavior 291, and BIOBM 330 or 331, or Introduction to the general principles of o f nonmedically important groups. Introduces equivalents. Offered alternate even years. toxicology including the sources, mechanisms, the major animal parasites: protozoan, J. P. Shapleigh. and targets o f toxic agents. Gives special nematode, platyhelminth, acanthocephalan, Focuses on physiological and metabolic attention to the interaction between toxic annelid, and arthropod. functions o f bacteria. Consideration is given to agents and biological systems at both the chemical structure, regulation, growth, and organismal and ecological level. The effects of BIOMI 391(3910) Advanced Microbiology energy metabolism. Special attention is given both anthropogenic and natural toxins are Laboratory to those aspects o f bacterial metabolism not examined with respect to genetic and Fall. 3 credits. Prerequisites: BIOMI 290, normally studied in biochemistry courses. developmental toxicity as well as 291, and BIOBM 330 or 331 or 333. carcinogenesis and specific organ toxicity. Priority given to biological sciences BIOMI 418(4180) Microbial Ecology students in microbiology program o f study. Spring. 3 credits. Prerequisites: BIOMI 290 W. C. Ghiorse, J. P. Shapleigh, and and 291, or 398 and permission of S. H. Zinder. instructor, and BIOBM 330 or 331 and 332. Illustrates basic principles o f experimental E. R. Angert. microbiology. The course is organized into Understanding the role o f microorganisms in four modules that last three weeks each: (1) natural environments is one o f the greatest ecology, (2) physiology, (3) genetics, and (4) challenges facing microbiologists. This course introduces current biochemical and NEUROBIOLOGY AND BEHAVIOR 173

[BIOMI 651(6080) Genomics of BIOMI 740(6430) Veterinary Perspectives Immunology of Infectious Diseases (BIO G Bacterium-Host Interactions (also on Pathogen Control in Animal 706, VETMI 719) PL PA 608(6080]) Manure (also VTMED/BEE 740(6430]) Fall, second half o f semester. 1 credit. Spring, eight weeks. 2 credits. Prerequisite: Introduction to Scanning Electron Microscopy Prerequisites: BIOMI 290 or equivalent or third- and fourth-year veterinary students. (BIO G 401) permission o f instructor. S-U or letter Letter grades only. D. D. Bowman. Fungi (PL PA 309) grades. Offered alternate even years; next For description, see VTMED 740. offered 2008-2009. A. Collmer and Light and Video Microscopy for Biologists BIOMI 791(7910) Advanced Topics in S. Winans. (BIO G 450) Microbiology For description, see PL PA 608.] Fall or spring. 1 credit; may be repeated Limnology: Ecology of Lakes, Lectures BIOMI 690 Prokaryotic Biology for credit. Prerequisite: graduate standing (BIOEE 457) Fall and spring. 4 weeks/8 lec. 1 credit per in microbiology. S-U grades only. Sec 01 Magical Mushrooms, Mischievous Molds Bacterial Genetics, S. C. Winans; Sec 02 sec to be offered. (PL PA 201) Environmental Microbiology, E. R. Angert. Sec 1 Microbial Structure and Function Reading and presentation by graduate students Microbiology for Environmental Engineering Fall. J. P. Shapleigh. o f current literature in selected areas of (CEE 451) Discusses those macromolecules and modern microbiology. assemblages o f macromolecules that together Plant Virology (PL PA 645) define the structure o f the prokaryotic cell. BIOMI 796(7960) Current Topics in Principles of Biogeochemistry (BIOEE 668) This includes external structures, such as cell Microbiology wall, flagella, pili, and peptidoglycan and Fall and spring. 0.5 or 1 credit for each The Soil Ecosystem (CSS 366) internal structures such as specialized vesicles topic; may be repeated for credit. Primarily and other large complexes. for graduate students in microbiology. Prerequisite: upper-level courses in Sec 2 Environmental Microbiology microbiology. S-U grades only. Lec. Staff. Fall. E. L. Madsen. Lectures and seminars on special topics in NEUROBIOLOGY AND BEHAVIOR Core course o f concepts, methods, and current microbiology. literature that reveals the multidisciplinary (BIONB) nature o f environmental microbiology and its BIOMI 797(7970) Scientific BIONB 111(1110) Brain Mind and relationship to prokaryotic biology. Discusses Communication Skills Behavior (also PSYCH/COGST the crucial roles that microorganisms play in Fall and spring. 1 credit each semester. 111(1110]) catalyzing biogeochemical reactions Requirement for graduate students in Spring. 3 credits. Prerequisite: none. throughout the biosphere. graduate field of microbiology for first two Intended for freshmen and sophomores in humanities and social sciences; not open Sec 3 Microbial Physiology/Diversity semesters; third semester optional. S-U to juniors and seniors. Not recommended Fall. S. H. Zinder. grades only. Staff. for psychology majors; biology majors may Reviews the major energy-conserving modes The ability to communicate effectively is not use for credit toward major. Letter o f metabolism and their phylogenetic essential for success as a scientist. The primary grades only. Planned M W F 9:05. distributions among both bacteria and archaea. goal o f this course is to provide students with E. Adkins-Regan and R. R. Hoy. Topics include phylogenetic analysis, an opportunity to develop self-confidence and For description, see COGST 111. fermentation, respiration, photosynthesis, refine their formal oral presentation skills. pathways o f carbon and nitrogen fixation, and Students are asked to present topical seminars BIONB 221(2210) Neurobiology and evolution o f the three domains o f life. that are critically evaluated by the instructor. Behavior I: Introduction to Behavior Feedback for improving the presentation and Fall. 3, 4, or 5 credits; 4 credits with one Sec 4 Microbial Genetics peer evaluations are emphasized. Spring. J. D. Helmann. disc per week; 5 credits with two disc per week and participation in Writing in the Reviews the fundamental concepts o f BIOMI 798(7980) Graduate Research Majors program; 4- or 5-credit option microbial genetics including mutations and Seminar in Microbiology their analysis, plasmids, conjugation, Fall and spring. 1 credit each semester. required o f students in neurobiology and behavior program o f study. Limited to 15 transformation, transduction, transposition, Requirement for graduate students in recombination, repair, and mutagenesis. graduate field o f microbiology. S-U grades students per 4-credit disc. Priority given to only. Staff. students studying neurobiology and Sec 5 Microbial Pathogenesis All graduate students in the field o f behavior. Limited to 12 students in 5-credit Spring. S. C. Winans. microbiology are required to attend and option (students may not preregister for Introduction to the fundamental concepts o f present a seminar concerning their research at 5-credit option; interested students bacterial pathogenesis including the normal least once each year. complete application form on first day of flora, pathogen entry and colonization, the class). Not open to freshmen. Prerequisite: production and regulation o f toxins, horizontal BIOMI 799(7990) Microbiology Seminar one year introductory biology for majors. transfer o f pathogenesis determinants, and the Fall and spring. Requirement for all May be taken independently of BIONB roles o f both specific and nonspecific host graduate students in graduate field o f 222. S-U or letter grades. Planned M W F defenses. Examples include bacterial microbiology. Open to all who are 12:20; disc TBA. P. W. Sherman and staff. pathogens o f both animals and plants. interested. Staff. General introduction to the field o f animal BIOMI 699(6990) Toxicology Journal Club behavior. Topics include evolution and Sec 01— Environmental Toxicology Related Courses in Other Departments behavior, behavioral ecology, sociobiology, (TOX 6990) Advanced Food Microbiology (FD SC 607) chemical ecology, communication, orientation Spring. 1 credit. Required for toxicology and navigation, and hormonal mechanisms of students until post A exam. A. G. Hay. Advanced Immunology Lectures (BIO G/ behavior. VETMI 705) BIOMI 725(7250) Mechanisms of BIONB 221(2210) Neurobiology and Microbial Pathogenesis (also VETMI Advanced Work in Bacteriology, Virology, or Behavior I: Introduction to Behavior 725(7250]) Immunology (VETMI 707) Summer, six-week session. 3 or 4 credits; 4 credits with one disc per week. Limited to Spring. 3 credits. Prerequisites: for Phytobacteriology Research Updates (PL PA 30 students. Prerequisite: one year undergraduates, written permission o f 647) instructor; BIOMI 404, 409, 417, or introductory college biology. S-U or letter equivalent. Highly recommended: Basic Immunology, Lectures (BIO G 305, grades. Course fee: none. Planned M-F completion o f two o f the three courses. VETMI 315) TBA. Staff. D. Debbie, M. Hesse, H. Marquis, J. Parker, General introduction to the field o f animal Current Topics in Oomycete Biology (PL PA behavior. Topics include evolution and M. Scidmore, and G. Whittaker. 644) For description, see VETMI 725. behavior, behavioral ecology, sociobiology, Food Microbiology, Laboratory (FD SC 395) chemical ecology, communication, orientation and navigation, and hormonal mechanisms o f Food Microbiology, Lectures (FD SC 394) behavior. 174 BIOLOGICAL SCIENCES - 2 8

BIONB 222(2220) Neurobiology and [BIONB 324(3240) Biopsychology Covers the basic ideas and techniques Behavior II: Introduction to Laboratory (also PSYCH 324(3240]) involved in computational neuroscience. Neurobiology Fall. 4 credits. Limited to 18 students. Surveys neural dynamics o f networks o f cells, Spring. 3 or 4 credits; 4 credits with disc Prerequisites: junior or senior standing; neural coding, learning, memory models, and written projects; 4-credit option PSYCH 223 or, BIONB 221 or 222, and sensory coding.] required o f students studying neurobiology permission o f instructor. Planned T R 1:25— and behavior. Limited to 15 students per 4:25. Letter grades only. Next offered BIONB 331(3310) Human Sociobiology disc; priority given to students studying 2008-2009. T. J. DeVoogd. Spring. 3 credits. Limited to 100 students. neurobiology and behavior. Not open to For description, see PSYCH 324.] Prerequisite: BIONB 221 or PSYCH 223 or freshmen. Prerequisites: one year permission o f instructor. S-U or letter introductory biology for majors and one [BIONB 325(3250) Insect Behavior (also grades. Planned M W 2:55-4:10. P. Barclay. year o f chemistry. May be taken ENTOM 325(3250]) Lecture-based course drawing on research in independently o f BIONB 221. S-U or letter Spring. 3 credits. Limited to 55 students. evolutionary biology and animal behavior to grades. Planned M W F 12:20; disc TBA. Prerequisite: ENTOM 212 or BIONB 221. investigate various aspects o f human social C. D. Hopkins and staff. Intended for juniors, seniors, and behavior. Findings are presented from areas General introduction to the field o f cellular beginning graduate students. S-U or letter such as evolutionary psychology, and integrative neurobiology. Topics include grades. Planned T R 10:10-11:25. Offered anthropology, human behavioral ecology, and neural systems, neuroanatomy, developmental alternate years; next offered 2008-2009. evolutionary game theory. Topics may vary neurobiology, electrical properties of nerve L. S. Rayor. slightly from year to year, but include mating, cells, synaptic mechanisms, neurochemistry, For description, see ENTOM 325.] cooperation (with kin and nonkin), conflict motor systems, sensory systems, learning, and BIONB 326(3260) The Visual System and aggression, parental behavior, costly signaling, and culture. memory. Some discussion sections include Spring. 4 credits. Limited to 25 students. dissections o f preserved brains. Prerequisite: BIONB 222 or BIOAP 311 or BIONB 369(3690) Chemical Ecology (also BIONB 231(2310) Sophomore Seminar: permission o f instructor. S-U or letter BIOEE/ENTOM 369(3690]) Topics in Cognitive Studies (also grades. Planned M W F 10:10; disc TBA. Spring. 3 credits. Prerequisites: one PSYCH 231/531(2310/6331], COGST Offered alternate years. H. C. Howland. semester o f introductory biology for majors 531(6331]) The visual systems o f vertebrates are or nonmajors and one semester of Spring. 4 credits. 231 limited to 15 discussed in breadth and depth as well as introductory chemistry for majors or students. Planned M W 2:55-4:10. some aspects o f invertebrate vision. Topics nonmajors or equivalents, or permission of S. Edelman. include the optics and anatomy o f eyes, instructor. S-U or letter grades. Planned M For description, see PSYCH 231. retinal neurophysiology, structure and function W F 11:15. A. Agrawal, G. Jander, o f higher visual centers, ocular motility, and A. Kessler, and J. Thaler. BIONB 321(3210) State of the Planet ocular and visual system development. For description, see BIOEE 369. Spring. 2 or 3 credits (3-cr. option includes disc sec). Prerequisites: none. S-U grades [BIONB 327(3270) Evolutionary BIONB 392(3920) Drugs and the Brain only. Planned M W 8:40-9:55; disc TBA. T. Perspectives on Human Behavior Fall. 4 credits. Limited to 90 students. Eisner and M. L. Zeeman. Fall. 3 credits. Limited to 20 students. Prerequisites: BIONB 222 or equivalent This interdisciplinary course is intended for Prerequisites: BIONB 221 and permission course in neurobiology by permission of any student with concern for the global crises o f instructor. Letter grades only. Planned instructor. Recommended: knowledge of we collectively face. During this course you M W 2:55-4:10. Next offered 2008-2009. biochemistry. S-U or letter grades. Planned T will be introduced to current data and S. T. Emlen. R 10:10-11:25; disc TBA. Offered alternate engaged in analysis o f those data, establish a Socratically taught, discussion-based course years. R. M. Harris-Warrick. global context for your specialized education, dealing with evolutionary perspectives on Introduction to neuropharmacology, with an and be provided with a toolbox to bring human behavior.] emphasis on the neural mechanisms o f awareness, analysis and action to your lives [BIONB 328(3280) Biopsychology of psychoactive drugs. Topics include a brief and careers beyond graduation. Learning and Memory (also PSYCH introduction to neuropharmacology and a 332(3320]) discussion o f the major neurotransmitter BIONB 322(3220) Hormones and families. The rest o f the course covers the Behavior (also PSYCH 322(3220]) Spring. 3 credits. Limited to 65 students. major psychoactive drugs, including cocaine, Fall. 3 credits. Limited to 60 students. Prerequisites: one year o f biology and heroin, psychedelics, marijuana, and alcohol, Prerequisites: junior or senior standing; any either a biopsychology course or BIONB as well as pharmaceuticals for the treatment o f one o f the following: PSYCH 223 or 222. S-U or letter grades. Graduate students, see PSYCH 632. Planned M W F anxiety, schizophrenia, and depression. BIONB 221 or 222 or one year Includes a term paper in the form o f a grant introductory biology plus psychology 11:15. Next offered 2008-2009. T. J. DeVoogd. proposal to study a current problem in course. Tw o lec plus sec in which students neuropharmacology. read and discuss original papers in the For description, see PSYCH 332.] field, give oral presentation, and write term [BIONB 329(3290) Ecology of Animal BIONB 394(3940) Circadian Rhythms paper. Letter grades only. Graduate Behavior (also BIOSM 329(3290]) (also BIOGD/ENTOM/PL PA 394(3940]) students, see PSYCH 722. Planned M W F Summer. 4 credits. Limited to 12 students. Fall. 2 or 3 credits. Prerequisite: 200-level 11:15. E. Adkins-Regan. Prerequisite: one year introductory college biology course. S-U or letter grades. For description, see PSYCH 322. biology. Recommended: ecology, Planned T 10:10-11:00; additional lab R psychology, or behavior course. S-U or BIONB 323(3230) Methods in Animal 1:25-4:25 for 3-cr. option. K. Lee. letter grades. Special two-week course Behavior For description, see PL PA 394. Fall. 4 credits. Limited to 24 students. offered at Shoals Marine Laboratory (SML), Prerequisite: BIONB 221. Letter grades located on an island o ff Portsmouth, N.H. [BIONB 396(3960) Introduction to only. Offered alternate years. Planned M W For more details and an application, Sensory Systems (also PSYCH 1:25-4:25. S. L. Vehrencamp and contact SML office, G14 Stimson Hall. Daily 396/696(3960/6960]) C. Botero. lec, lab, and fieldwork for two weeks. Next Spring. 4 credits. Limited to 25 students. This course provides hands-on experience offered summer 2009. SML faculty. Prerequisites: introductory biology or with modern methods for studying animal For description, see BIOSM 329 ] biopsychology, plus second course in behavior both in the field and in the [BIONB 330(3300) Introduction to behavior, biopsychology, cognitive science, laboratory. In-depth class projects will be Computational Neuroscience (also neuroscience, or perception; knowledge of complemented with a series o f shorter PSYCH/COGST/BME 330(3300]) elementary physics, chemistry, and behavior. S-U or letter grades. Planned T R workshops and demonstrations. Topics include Fall. 3 or 4 credits; 4 credits includes lab 10:10-11:25. Offered alternate years; next animal sound recording, field videography providing additional computer simulation offered 2008-2009. B. P. Halpem. radio-tracking, mapping, animal color analysis, exercises. Limited to 25 students. For description, see PSYCH 396.] capture/marking methods, odor analysis, Prerequisites: BIONB 222 or permission o f measuring dominance hierarchies, and instructor. S-U or letter grades. Offered behavioral statistics. alternate years; next offered 2008-2009. Planned M W 2:55-4:10. C. Linster. NEUROBIOLOGY AND BEHAVIOR 175

BIONB 413(4130) Molecules of Social BIONB 423(4230) Cognitive [BIONB 427(4270) Animal Social Behavior and Emotion Neuroscience (also PSYCH Behavior Spring. 3 credits. Prerequisite: permission 425/625(4250/6250]) Fall. 4 credits. Limited to 30 students. o f instructor. Letter grades only. Planned Fall. 4 credits. Limited to 20 students Prerequisites: BIONB 221 and BIOEH 26l T R 10:10-11:25. Offered alternate years. Prerequisites: introductory biology; or 278, and permission o f instructor. Letter D. P. McCobb. biopsychology or neurobiology (e.g., grades only. Planned T R 2:30-4:25. Active-learning course with specific topics to PSYCH 223 or BIONB 221); and Offered alternate years; next offered 2008- be determined by students. Focuses on introductory course in perception, 2009. P. W. Sherman. molecular, neural, and endocrine mechanisms cognition, or language (PSYCH 102, 209, Writing-intensive advanced course for upper- underlying innate and learned behavior 214, or 215 essential). S-U or letter grades. division students. Lectures, discussions, patterns, and their relationship to social, Graduate students, see PSYCH 625. student presentations examine topics in ecological, and evolutionary context. Planned M W F 9:05. One lab in sheep animal behavioral ecology, human Neurotransmitters, hormones and receptors brain dissection. Offered alternate years. sociobiology, Darwinian medicine.] governing such behaviors as parental care, B. L. Finlay. territoriality, cooperativity, courtship, and For description, see PSYCH 425. [BIONB 428(4280) Clinical Neurobiology stress responses are examples o f topics o f Fall. 3 credits. Limited to 20 students. BIONB 424(4240) Neuroethology (also Prerequisites: two courses from BIONB interest. Format includes library research, oral PSYCH 424(4240]) and written presentations, teamwork, and peer 222, BIOGD 281, BIOBM 330 or 331; co­ Fall. 3 credits. Limited to 50 students. review. registration in one o f the two is acceptable Prerequisites: BIONB 221 or 222, or one year by permission o f instructor. Open to BIONB 420(4200) Topics in Neurobiology introductory biology for majors and advanced undergraduates. S-U or letter and Behavior permission o f instructor. S-U or letter grades. grades. Planned M W 2:30-4:25. Offered Fall or spring. Variable credit; may be Planned MWF 10:10. Offered alternate years. alternate years; next offered 2008-2009. repeated for credit. Primarily for C. D. Hopkins. R. Booker. undergraduates. S-U or letter grades. Staff. Neuroethologists take a comparative and This course focuses on the etiology, Courses on selected topics in neurobiology evolutionary approach to study the nervous epidemiology, cellular and molecular basis, and behavior; can include lecture and seminar system. They ask, how do brains o f animals and strategies for treating a number of courses. See department office (W363 Mudd compare and how did they evolve? How are neurodiseases (e.g., Alzheimer’s disease, Hall) for offerings. neural circuits adapted to species-typical depression, ADHD).] behavior? What is the hope and interest in the BIONB 421(4210) Effects of Aging on study o f a large diversity o f animals, compared [BIONB 429(4290) Olfaction and Taste: Sensory and Perceptual Systems with a specialized look at just a few Structure and Function (also PSYCH (also PSYCH 431/631(4310/6310]) mammalian species? Can w e hope to 429(4290]) Fall. 3 or 4 credits; 4-credit option involves understand how animals with specialized Spring. 3 or 4 credits; 4-credit option term paper or creation o f relevant web behaviors have specialized nervous systems? requires term paper. Priority given to site. Limited to 35 students. Prerequisites: What is the sensory world o f a real animal junior and senior psychology and biology introductory course in biology or and how does it vary from species to species? majors and graduate students. Graduate psychology, plus second course in These and other questions derive this students, see PSYCH 629. Prerequisite: one perception, neuroscience, cognitive introductory survey of neuroethology, 300-level course in biopsychology or science, or biopsychology. Planned T R including exotic senses, amazing motor equivalent. Planned T R 10:10-11:25. 10:10-11:25. B. P. Halpern. programs, and surprising integration. Offered alternate years; next offered 2008- For description, see PSYCH 431. 2009. B. P. Halpern. BIONB 425(4250) Molecular For description, see PSYCH 429.1 BIONB 422(4220) Modeling Behavioral Neurophysiology Evolution Fall. 3 credits. Limited to 20 students. [BIONB 430(4300) Experimental Fall. 4 credits. Limited to 25 students. Prerequisite: BIONB 222 or permission o f Molecular Neurobiology (also BIOBM Prerequisites: BIONB 221, one year o f 443(4430]) instructor. S-U or letter grades. Planned T calculus, course in probability or statistics, Spring. 4 credits. Limited to 12 students. R 2:55-4:10. Offered alternate years. and permission o f instructor; advanced Corequisite: BIOBM 440 lab. Letter grades D. P. McCobb. undergraduates and graduate students. S-U Focuses on ion channels, the primary proteins only. Planned T all day; disc TBA. Offered or letter grades. Planned T R 2:55-4:10; generating cellular electrical signals in nerve alternate years; next offered 2008-2009. computer lab TBA. Offered alternate years. cells and other excitable cells (e.g., muscle, D. L. Deitcher. H. K. Reeve. Experiments include PCR, cloning o f DNA heart, glands). Reviews the latest Intensive lecture and computer lab course on fragments, RNA purification, restriction digests, electrophysiological and molecular genetic modeling strategies and techniques in the bacterial transformation, DNA sequencing, and experiments. Diversity of electrophysiology study o f behavioral evolution. Population- deriving from channel structure and protein interactions. Emphasis on molecular genetic (including quantitative-genetic), static expression patterns is considered in the techniques to study neurobiological problems.] optimization, dynamic programming, and contexts o f behavior and behavioral plasticity game-theoretic methods are emphasized. BIONB 431(4310) Genes and Behavior (learning), neural development, and channel These approaches are illustrated by Spring. 3 credits. Limited to 50 students. evolution. Format includes written and oral application to problems in optimal foraging, Prerequisite: BIONB 222. S-U or letter presentations, reviewing scientific literature in sexual selection, sex ratio evolution, animal grades. Planned T R 2:55-4:10. Offered selected areas, and proposing new communication, and the evolution o f alternate years. J. R. Fetcho. experiments. cooperation and conflict within animal social Our genes influence how we behave. This groups. Students learn to critically assess BIONB 426(4260) Animal Communication lecture course explores the current recent evolutionary theories o f animal Spring. 4 credits. Limited to 50 students. understanding of how genes influence the behavior, as well as to develop their own Prerequisite: BIONB 221. Letter grades behavior o f a variety o f animals, including testable models for biological systems o f only. Planned T R 2:55-4:10; disc TBA. humans. Topics include the genetic basis of interest or to extend pre-existing models in Offered alternate years. J. W. Bradbury and hearing, movement, learning, memory, novel directions. The Mathematica software S. L. Vehrencamp. intelligence, sexual behavior, aggression, program is used as a modeling tool in the Communication is the “glue" that holds sleep, and diseases o f behavior. The focus is on the unprecedented insight that modern accompanying computer lab (no prior societies together. This course examines how molecular and genetic tools are providing into experience with computers required). and why animals communicate. Topics include the genetic basis o f behavior. the role o f the environment in shaping animal signals, whether animals tell the truth to each BIONB 442(4420) Instrumentation for other, why some bird songs are simple and Biology others complex, and what kinds o f signals Fall. 4 credits. Limited to 12 students. might be exchanged between species. Prerequisite: one semester o f calculus. Letter grades only. Planned T R 8:40-9:55, lab W 1:25-4:25. B. R. Land. 176 BIOLOGICAL SCIENCES • 2 8

Introduction to biological electronics, nerve cells differentiate both morphologically Related Courses in Other Departments computer interfacing techniques, and data and biochemically? The role of cues such as Evolutionary Perspectives on Behavior reduction. Gives a basic understanding o f the hormones and developmental genes in neural (PSYCH 535) techniques for using electrical recording development is discussed. Readings are taken combined with a computer in a biological from original journal articles. Biopsychology of Normal and Abnormal context. Examples are drawn from practical BIONB 494(4940) Brain Evolution and Behavior (PSYCH/NS 361) neurobiological instrumentation problems and Behavior the electronic basis o f neural circuitry. Developmental Biopsychology (PSYCH 422) Spring. 3 credits. Limited to 50 students. [BIONB 444(4440) Neural Computation Intended for juniors, seniors, and graduate Evolution of Human Behavior (PSYCH 326) Spring. 3 credits. Limited to 10 students. students. Prerequisite: BIONB 222 or Prerequisites: BIONB 222 or permission o f equivalent. S-U or letter grades. Planned T R Topics in Biological Anthropology (ANTHR instructor. S-U or letter grades. Planned T 2:55-4:10. Offered alternate years. A. H. Bass. 490) Organization and evolution of R 2:30-3:20, lab T R 3:35-4:25. Offered Primate Behavior and Ecology (ANTHR 390) alternate years; next offered 2008-2009. neuroanatomical pathways as substrates for T. A. Cleland. species-typical vertebrate behaviors. The Teaching Experience (BIO G 498) Lecture and computer lab course covering the course is divided into three major sections: biophysical mechanisms underlying neural development, general principles o f brain The Brain and Sleep (PSYCH 440/640) computation and information coding by organization, and co-evolution o f vertebrate neurons and networks.] brain and behavior. Undergraduate Research in Biology (BIO G 499) BIONB 470(4700) Biophysical Methods [BIONB 495(4950) Molecular and Genetic (also A&EP/VETMM 470(4700]) Approaches to Neuroscience OTS Undergraduate Semester Abroad Fall. 3 credits. Prerequisites: solid Fall. 3 credits. Limited to 25 students. Programs knowledge o f basic physics and Prerequisites: junior, senior, or graduate mathematics through sophomore level. standing; BIONB 222 and BIOBM 330 or Shoals Marine Laboratory Program Recommended: knowledge of cellular 332. Letter grades only. Planned T R 2:55- Spider Biology (ENTOM 215) biology. Letter grades only. Planned M W 4:10. Offered alternate years; next offered 2:55-4:10. M. Lindau. 2008-2009. D. L. Deitcher. Navigation, Memory, and Context: What Focuses on how molecular and genetic For description, see A&EP 470. Does the Hippocampus Do? (PSYCH approaches have led to major advances in 423/623) BIONB 491(4910) Principles of neuroscience. Lectures, student presentations, Neurophysiology (also BME and discussions examine research articles.] 491(4910]) Spring. 4 credits. Limited to 20 students. [BIONB 496(4960) Bioacoustic Signals in Prerequisite: BIONB 222 or written Animals and Man PLANT BIOLOGY (BIOPL) permission o f instructor. S-U or letter Fall. 3 credits. Limited to 12 students. grades for graduate students by permission Prerequisites: junior, senior, or graduate BIOPL 240(2400) Green Wortd/Blue o f instructor. Planned M W 10:10; lab standing; one year introductory biology, Planet planned M or T 12:20-4:25. B. R. Johnson. PHYS 101-102 or 207-208, and permission Fall. 3 credits. S-U or letter grades. Lee. Laboratory-oriented course designed to teach o f instructor. S-U or letter grades. Planned T. Silva. the theory and techniques o f modern cellular M W 9:05; lab TBA. Offered alternate Focuses on helping individuals understand neurophysiology including computer years; next offered 2008-2009. C. W. Clark how scientific information relates to the issues acquisition and analysis o f laboratory results. and R. R. Hoy. they face as citizens, in management decision Lecture time is used to introduce laboratory Teaches students about animal acoustic making, and in public policy. To what extent exercises and discuss results, to supplement signaling by introducing them to various should genetic engineering o f crop plants be laboratory topics, and to discuss primary animal acoustic systems.] permitted? Should w e place limits on fossil research papers. Extracellular and intracellular fuel consumption as a means o f limiting BIONB 720(7200) Seminar in Advanced global warming and global climate change? recording and voltage clamp techniques Topics in Neurobiology and Behavior explore motor neuron and sensory receptor Must human endeavors be restricted in certain Fall or spring. Variable credit; may be tiring properties, and examine the cellular basis areas to maintain diversity? The format o f this repeated for credit. Prerequisite: graduate for resting and action potentials and synaptic course is interactive, with lectures and standing or permission o f instructor. S-U or transmission. Invertebrate preparations are used discussions about how w e as a society deal letter grades. Staff and students. as model systems. See instructl.cit.comell.edu/ with controversial issues. Designed to provide several study groups courses/bionb491/index.html. each semester on specialized topics. A group BIOPL 241(2410) Introductory Botany [BIONB 492(4920) Sensory Function may meet for whatever period is judged Fall. 3 credits. Lee, lab. K. J. Niklas. (also PSYCH 492/692(4920/6920]) adequate to enable coverage of the selected Introductory botany for those interested in the Spring. 4 credits. Limited to 25 students. topics. Ordinarily, topics are selected and plant sciences. Emphasizes structure, Prerequisite: 300-level course in circulated during the preceding semester. reproduction, and classification o f biopsychology, or BIONB 222, or BIOAP Discussion o f current literature is encouraged. angiosperms and the history o f life on earth. 311, or equivalent; knowledge of Suggestions for topics should be submitted by Laboratory emphasizes development o f skills elementary physics, chemistry, and faculty or students to the chair o f the in handling plant materials, including behavior. S-U or letter grades. Graduate Department of Neurobiology and Behavior. identification. First and second weeks o f students, see PSYCH 692. Planned M W F laboratory are field trips, starting with the first BIONB 721(7210) Introductory Graduate day o f classes. Those who register fo r an 10:10. Offered alternate years; next offered Survey in Neurobiology and Behavior 2008-2009. H. C. Howland and evening laboratory are still required to attend Fall. 2 credits. Requirement for graduate B. P. Halpern. the afternoon field trips. students majoring in neurobiology and For description, see PSYCH 492.] behavior. Concurrent registration in BIONB BIOPL 242(2420) Plant Function and BIONB 493(4930) Developmental 221 and 222 not required. S-U grades only. Growth Lectures Neurobiology Planned W 4:00-6:00. D. L. Deitcher and Spring. 3 credits. Primarily for Fall. 3 credits. Limited to 20 students. staff. undergraduates in agricultural sciences but Prerequisite: BIONB 222 or permission o f Lectures, readings, and discussion introduce first- also for any biological sciences students instructor. S-U or letter grades by year graduate students to the research activities wanting to know about plant function; permission o f instructor. Planned M W o f the faculty in the graduate field of suitable as second-level course for nonmajors 2:55-4:10. Offered alternate years. neurobiology and behavior. Class meets weekly to satisfy biology distribution requirement. R. Booker. for two hours. Students also prepare a research Prerequisites: one year introductory biology Lectures covering the development o f the proposal on a potential topic for their thesis and/or BIOPL 241. Corequisite for plant nervous system, taking examples from both research (in the format o f an NSF or NIH grant). science undergraduates (and highly vertebrates and invertebrates. Emphasis is on This proposal is prepared in consultation with recommended for other science majors): cellular and molecular issues, that is, how do one or more relevant faculty members. BIOPL 244. Recommended: one year PLANT BIOLOGY 177

introductory chemistry. May not be taken for BIOPL 248(2480) Taxonomy of Vascular BIOPL 347(3470) Laboratory in Molecular credit after BIOPL 342 except by written Plants Biology and Genetic Engineering of permission o f instructor. S-U or letter grades. Spring. 4 credits. Prerequisite: one year Plants Evening prelims. P. J. Davies. introductory biology. May not be taken for Spring. 2 credits. Limited to 24 students. H ow plants function and grow. Examples deal credit after BIOPL 243. S-U or letter grades. Prerequisite: BIOPL 343 or permission of with crop plants or higher plants where Lee, lab. Offered alternate years; not instructor. Recommended: concurrent possible, though not exclusively. Topics offered 2008-2009. J. I. Davis. enrollment in BIOPL 343. S-U or letter include cell structure and function; plant Introduction to the classification o f vascular grades. Lab. M. E. Nasrallah. metabolism, including photosynthesis; light plants, with attention to the goals o f Companion to BIOPL 343 with laboratory relations in crops; plant-water relations; water taxonomy, the processes o f plant evolution, activities that focus on the practice o f plant uptake, transport, and transpiration; irrigation and the means o f analyzing evolutionary biotechnology. Students transfer genes to o f crops; sugar transport; mineral nutrition; relationships among plants. The laboratory plants by a variety o f methods and analyze growth and development— hormones, presents an overview o f vascular plant their expression in the host genome by use of responses to light, flowering, fruiting, diversity, with particular attention to the reporter gene assays and by the preparation dormancy, and abscission; stress; tissue flowering plants. and analysis o f nucleic acids. culture; and genetic engineering o f plants. BIOPL 342(3420) Plant Physiology, BIOPL 348(3480) The Healing Forest [BIOPL 243(2430) Taxonomy of Lectures Spring. 2 credits. Prerequisites: introductory Cultivated Plants (also HORT Spring. 3 credits. Prerequisites: one year biology or plant biology or permission o f 243(2430]) introductory biology. Corequisite: BIOPL instructor. Lec/disc. Offered alternate years. Fall. 4 credits. Prerequisite: one year 344 or written permission o f instructor. E. Rodriguez. introductory biology or written permission o f May not be taken for credit after BIOPL Ethnobotanical and ethnopharmacological instructor. May not be taken for credit after 242 unless written permission obtained consideration o f the role o f plants, fungi, and BIOPL 248. Lee, lab. Offered alternate years; from instructor. Lee. T. G. Owens. insects in traditional and western medicine. next offered 2008-2009. M. A. Luckow. Integrated and interdisciplinary study o f the Studies o f indigenous and lay societies Study o f ferns and seed plants, their processes that contribute to the growth, illustrate the ecological, systematic, relationships, and their classification into competition, and reproduction o f plants. biochemical, and cultural aspects o f herbal families and genera, emphasizing cultivated Topics include, but are not limited to, plant- medicines and are placed in the broader plants. Particular emphasis is placed on water relations, membrane properties and context o f such interdependent themes as the gaining proficiency in identifying and processes, photosynthesis, plant respiration, conservation o f biological and cultural distinguishing families and in preparing and mineral and organic nutrition, stress diversity, human health, bioprospecting, using analytic keys. Attention is also given to physiology, control o f growth and compensation for indigenous knowledge, and the economic importance o f taxa, to the basic development, and responses to the sustainable development. taxonomic literature, and to the elements o f environment. Emphasis is on the relationship nomenclature.] between structure and function from the [BIOPL 359(3590) Biology of Grasses molecular to the whole-plant level. Spring 2 credits. Prerequisite: one year BIOPL 244(2440) Plant Function and introductory biology or course in plant Growth, Laboratory BIOPL 343(3430) Molecular Biology and systematics or permission o f instructor. S-U Spring. 2 credits. Limited to 14 students per Genetic Engineering of Plants or letter grades. Lee. Lab. Offered alternate sec. Corequisite: BIOPL 242. May not be Spring. 2 credits. Prerequisite: one year years; next offered 2008-2009. J. I. Davis. taken for credit after BIOPL 344. Disc and general biology or permission o f instructor. Systematics and related aspects o f the biology of lab; students must take lab and disc on S-U or letter grades. Lee. M. E. Nasrallah. the graminoid plant families (grasses, sedges, same day. T. Silva. Introduction to current studies involving and rushes), with the principal emphasis on Experiments exemplify concepts covered in recombinant DNA technology and its grasses. Major topics include phylogenetics, BIOPL 242 and offer experience in a variety of application to the improvement o f plants. taxonomy, physiology, reproductive biology, biological and biochemical techniques, from Emphasizes genetic transformation speciation, and biogeography. The roles of the cellular to whole plant level. methodology, gene expression systems, and graminoid plants in natural and human- strategies for increasing productivity. The BIOPL 245(2450) Plant Biology disturbed environments are discussed, as are the course is directed toward undergraduates who origins o f cultivated species.] Summer, six-week session. 3 credits. wish to become familiar with the theory and Limited to 24 students. Lee, lab. T. Silva. practice of plant biotechnology. BIOPL 380(3800) Strategies and Methods Introductory botany, including plant in Drug Discovery identification. Emphasizes structure, BIOPL 344(3440) Plant Physiology, Spring. 2 credits. Prerequisite: one year reproduction, and classification o f flowering Laboratory introductory biology and organic chemistry plants. Much o f the laboratory work is Spring. 2 credits. Corequisite: BIOPL 342. course or permission o f instructor. S-U or conducted outdoors taking advantage of May not be taken for credit after BIOPL letter grades. M. A. Aregullin. several outstanding natural areas available for 244. Similar to BIOPL 244 but at more Covers strategies and methodologies in study. Those who lack college-level biology are advanced level. Lab, disc. T. Silva. chemotaxonomy, chemical ecology, and expected to work closely with the instructor on Experiments exemplify concepts covered in ethnobotany, as they are used in chemical supplemental instructional materials. BIOPL 342 and offer experience in a variety of prospecting for new pharmaceuticals. biological and biochemical techniques, from Discusses the biosynthesis and distribution of [BIOPL 247(2470) Ethnobiology the cellular to whole plant level, with plant secondary metabolites, the use o f Fall. 3 credits. S-U or letter grades. Lee, emphasis on experimental design. techniques in isolation and structure disc. Offered alternate years; next offered elucidation o f natural products, and biological 2008-2009. Staff. BIOPL 345(3450) Plant Anatomy assays in the discovery o f chemicals with Consideration o f the principles, methods, and Fall. 4 credits. Limited to 15 students. pharmacological activity. issues o f ethnobiology. Emphasis is on the Prerequisite: one year introductory biology past and present ecological, evolutionary, or a semester o f botany. Lee, lab. BIOPL 404(4040) Crop Evolution, economic, and cultural interrelationships o f A. Gandolfo. Domestication and Diversity (also humans in traditional and lay societies with Descriptive course with equal emphasis on PL BR/IARD 404(4040]) their plants and animals, as a means o f development and mature structure. Lecture, Fall. 2 credits. Prerequisites: BIOGD 281 or understanding the place and future o f humans laboratory, and reading are integrated in a PL BR 225 or permission o f instructor. S-U in the biosphere. Traditional medicines, study guide. The laboratory offers the or letter grades. Lee. S. Kresovich. underutilized organisms, resource opportunity to develop the practical skills For description, see PL BR 404. management, and ownership o f nature, and required to make anatomical diagnoses and to methodology are among the topics covered.] write anatomical descriptions. 178 BIOLOGICAL SCIENCES - 2 8

BIOPL 422(4220) Plant Development nuclear and organellar genomes is described senescence, defense mechanisms, amino acid Fall. 2 credits. Lee. Prerequisites: course from the standpoint o f their suitability for biosynthesis, and small molecule transport. work in molecular biology (e.g., BIOBM systematic and evolutionary studies. Genomics-based experimental tools such as proteomics and metabolomics are discussed. 330, 331/332, or 333), and genetics (e.g., [BIOPL 448(4480) Plant Evolution and BIOGD 281), or permission o f instructor. the Fossil Record BIOPL 482 Plant Molecular Biology II S-U or letter grades. J. Hua. Spring. 3 credits. Prerequisite: BIOPL 241 Spring. 1-6 credits; 1 credit per sec. Introduction to plant development, studying or equivalent, or permission o f instructor. Prerequisites: BIOGD 281 and BIOBM 330 the mechanisms o f morphogenesis and cell Lee, lab. Offered alternate years; next or 332, or equivalents. Recommended: fate determination at the organismal, cellular, offered 2008-2009. K. J. Niklas and BIOBM 331. S-U or letter grades. and molecular levels. W. L. Crepet. Series o f four-week modules on specialized [BIOPL 440(4400) Phylogenetic Introduction to evolution, surveying major topics. Coordinator: J. B. Nasrallah. changes in plants from the origin o f life to the Systematics (also ENTOM 440(4400]) Sec 01 Molecular Plant-Pathogen present. Emphasizes plant form and function, Interactions I and II (also PL PA 462.1) adaptations to particular ecologic settings, and 1 credit. 12 lec. A. Collmer, S. G. Lazarowitz, Spring. 4 credits. Limited to 24 students. evolutionary theory as it relates to plants.] G. Martin, and B. G. Turgeon. I Prerequisite: introductory biology or Examination o f the molecular and cellular permission o f instructor. Lee, lab. Offered BIOPL 449(4490) Green Signals and Triggers— The Plant Hormones (also factors that control pathogen-plant interactions alternate years; next offered 2008-2009. HORT 449(4490]) from the perspectives of pathogen biology K. C. Nixon. Spring. 2 credits. Prerequisite: one year and plant responses to pathogen infection. Basic and advanced theory and methods o f introductory biology and plant physiology Beginning spring 2004, alternate years focus phylogenetic analysis. Introduces students to (BIOPL 242 or 342) or permission of on: (1 ) plant perception o f microbial cladistic analysis using parsimony and gain instructor. S-U or letter grades. Offered pathogens and the interplay o f plant defenses experience with computer-aided analysis o f alternate years. P. J. Davies. and pathogen counterstrategies that result in taxonomic data, including both morphological Study o f plant hormones and how they resistance or susceptibility to disease and molecular data sources. Topics include regulate plant growth and development. production, with topics including the genetic applications o f phylogenetic methods to Topics include the discovery, role in growth nature o f dominant and recessive resistance, biogeography and evolutionary studies.] and development, mode o f action, and induction o f pathogen defense genes, BIOPL 442(4420) Currant Topics In practical uses o f the plant hormones auxin, apoptotic responses that limit infection, and Ethnobiology gibberellins, cytokinins, abscisic acid, ethylene, RNA interference; and (2) the genetic and Fall. 2 credits. Limited to 12 students. and brassinosteroids. molecular mechanisms of microbial Prerequisite: permission o f instructor. S-U pathogenesis, with an emphasis on fungal and or letter grades. Lec/disc. Offered alternate BIOPL 452(4520) Systematics of Tropical bacterial virulence proteins, toxins, and their Plants years. Staff. deployment systems. Fall. 3 credits. Prerequisite: BIOPL 243 or Explores the interrelationships o f plants and 248. Letter grades only. Lee, lab. Offered Sec 02 Molecular Plant-Microbe animals with humans from a wide range o f every three years. K. C. Nixon. Interactions (also BIOMI 482, Sec 2, PL perspectives. Topics considered are The families o f plants encountered solely or PA 464) contemporary issues, theory, and methodology chiefly in tropical regions are considered in a 1 credit. S-U or letter grades. 12 lec. of ethnobotany and ethnobiology, and the role phylogenetic context in lectures, discussions, Offered alternate years. S. C. Winans. o f plants and animals in human lives, in and laboratory, with the aim o f providing Focuses on the interactions o f Agrobacteria subsistence and exchange, and in thought. basic points o f recognition for, and an and Rhizobia with plants. Topics on BIOPL 443(4430) Topics and Research understanding of, diversity and relationships in Agrobacterium-plant interactions include Methods in Systematics these families. plant-microbe recognition mechanisms, T-DNA Fall or spring. 1-2 credits; 1 credit per sec. transfer process, oncogenesis, and use o f Prerequisite: written permission o f BIOPL 453(4530) Principles and Practice Agrobacterium to produce transgenic plants. of Historical Biogeography (also instructor. S-U or letter grades. K. C. Nixon. Topics on Rhizobium-plant interactions ENTOM 453(4530]) Series o f 1-credit modules on specialized include regulation o f nitrogenase activity and Fall. 3 credits. Prerequisite: systematics topics in systematics. Topics and instructors expression, organization and function of the course or permission o f instructors. S-U or vary each semester. May not be taught every sym plasmid, nodule development, and plant letter grades. Lee, lab. Offered alternate semester. Topics and instructors are listed in genetics involved in plant-microbe interaction. years. J. K. Liebherr and M. A. Luckow. the division’s catalog supplement issued at the For description, see ENTOM 453- Sec 03 Light Signal Transduction in beginning o f the semester. Plants [BIOPL 444(4440) Plant Cell Biology BIOPL 454(4540) Systematics of Tropical 1 credit. S-U or letter grades. 12 lec. Plants: Field Laboratory Fall. 4 credits. Limited to 24 students. Offered alternate years. T. Brutnell. Spring. 1 credit. Limited to 15 students. Prerequisites: one year introductory In addition to providing plants with energy for Prerequisite: BIOPL 452 or permission o f biology or permission o f instructor. Lee, photosynthesis, light plays an essential role in instructor. Letter grades only. For more lab. Next offered 2008-2009. R. O. Wayne. the development o f higher plants. Light quality details and application, contact L. H. Bailey Uses evidence from microscopy, physiology, and intensity is carefully monitored by the Hortorium, 467 Mann Library. Offered biochemistry, and molecular biology to try to plant to avoid neighboring vegetation, set the every three years. K. C. Nixon. unravel the mystery o f the living cell. Studies circadian clock, and adjust photosynthesis Intensive orientation to families o f tropical the dynamics o f protoplasm, membranes, and rates. This course focuses on recent studies flowering plants represented in forests o f the the various organelles. The mechanisms o f cell that have illuminated the molecular basis o f American Tropics. Emphasis is on field growth and division, the relationship o f the light signal transduction networks in higher identification combined with laboratory analysis cytoskeleton to cell shape and motility, the plants. Readings are assigned from current o f available materials in a “whole-biology” interaction o f the cell with its environment, literature with an emphasis on those that use context. Two-week field trip over winter break. and the processes that give rise to genomics tools such as microarray analysis to multicellular differentiated plants are BIOPL 462(4620) Plant Biochemistry address fundamental questions in red/far-red and blue light signal transduction. investigated.] Spring. 3 credits. Prerequisites: BIOPL 242 BIOPL 447(4470) Molecular Systematics or 342 or equivalent and BIOBM 330 or [Sec 04 Plant Gene Evolution and 331 or equivalent or permission of Phylogeny Fall. 3 credits. Prerequisites: BIOEE 278 or instructor. Letter grades only. Lee. J. Rose 1 credit. 12 lec. Offered alternate years; BIOGD 281 or BIOBM 330, or BIOBM 332, and K. Van Wijk. next offered 2008-2009. J. J. Doyle. or written permission o f instructor. Lee. Focuses on biochemistry of plant specific Practical applications o f molecular systematics/ Offered alternate years. J. J. Doyle. processes, with the aim to obtain an evolution for plant molecular biologists and Theory and practice o f using molecular integrative overview o f plant biochemistry. other non-systematists. The course focuses on evidence, particularly DNA sequence data, for Examples include processes such as cell wall two basic issues: methods and principles for addressing diverse systematic and evolutionary biochemistry, pigment biosynthesis and inferring relationships among genes and the questions. Emphasis is on phylogeny degradation, secondary metabolism, use o f data to hypothesize relationships reconstruction, particularly in eukaryotic systems. The organization and evolution o f PLANT BIOLOGY 179 among plants. Evolutionary patterns and BIOPL 483 Plant Molecular Biology I Sec 05 Molecular Breeding (also PL BR processes o f genes and gene families are Fall. 1-5 credits; 1 credit per sec. 483.5) discussed, as well as rates o f sequence Prerequisites: BIOGD 281 and BIOBM 330 1 credit. 12 lec. Offered alternate years. evolution, paralogy and orthology, the effects or 332, or equivalents. Recommended: S. D. Tanksley. of recombination and concerted evolution of BIOBM 331. S-U or letter grades. Application o f DNA markers to the gene phylogenies, and the implications o f Coordinator: J. B. Nasrallah. identification, manipulation, and isolation o f using gene or allele phylogenies to infer Series o f four-week modules on specialized genes important to plant and animal organismal evolutionary patterns.] topics. productivity using molecular genetic Sec 05 Molecular Biology of Plant Sec 01 Concepts and Techniques in Plant techniques. Students learn how to design and Organelles (also BIOGD 452) Molecular Biology (also PL PA 463.01, execute experiments to identify quantitative 1 credit. S-U grades by permission o f PL BR 483.01) trait loci (QTLs), as well as how to apply instructor. 12 lec. M. R. Hanson and 2 credits. Lec. S. R. McCouch, molecular markers to plant and animal D. B. Stern. J. Giovannoni, and J. Rose. breeding programs. Plants contain three different genomes— in the Introductory module that provides a broad Sec 06 Plant Senescence (also HORT nucleus, chloroplasts, and mitochondria. This overview o f molecular biology concepts relevant 625.2) course examines the organization, expression, to the plant sciences. Serves as a prerequisite to 1 credit. S-U or letter grades. (12 lec). and evolution o f plant organelle genomes. other modules in the BIOPL 483 (fall) and S. Gan. Special topics include RNA editing and stability, BIOPL 482 (spring) series. The course is divided Introduces molecular, genetics, and genomics effects o f organelle mutations on plant into three sections: (1) Gene discovery: covers approaches in plant senescence and reproduction and photosynthesis, and genetic, molecular, and genomics approaches to postharvest research. Topics include gene chloroplast transformation for expression o f the isolation o f plant genes; (2) Gene expression, regulation, and function associated useful foreign proteins. characterization: covers DNA sequence analysis, with physiological and biochemical changes in assessment o f gene expression, functional Sec 06 Plant Biotechnology (also PL BR senescing, maturing, and/or ripening plants or genomics approaches, and production of 482.6, PL PA 462.2) parts. Genetic manipulation o f senescence/ transgenic plants; (3) Analysis and 1 credit. S-U or letter grades. 12 lec. ripening processes are also discussed. characterization o f proteins and metabolites: E. D. Earle. includes proteomics approaches to the analysis BIOPL 641(6410) Laboratory in Plant Deals with production and uses o f transgenic o f plant proteins, protein-protein interactions, Molecular Biology (also BIOBM plants for agricultural and industrial purposes. and metabolic profiling through emerging 641(6410]) Topics include procedures for gene metabolomic techniques. This course consists of Fall. 4 credits. Prerequisites: BIOGD 281 or introduction and control o f gene expression, two lectures and one day o f discussion per equivalent, BIOBM 330 or 331 or as well as strategies for obtaining transgenic week. Course material is coordinated with equivalent, and permission o f instructor. plants that are resistant to insects, diseases, BIOPL 641 (lab). Emphasis is on understanding S-U grades by permission o f instructor. and herbicides, produce useful products, or techniques and approaches appropriate for Lab. M. R. Hanson, H. Wang, T. Brutnell, have improved nutritional and food processing different experiments and objectives. G. Jander, J. Hua, M. Scanlon, and characteristics. Regulatory and social issues K. van Wijk. relating to plant biotechnology are discussed. Sec 02 Proteomics in Plant Biology Includes selected experiments on gene 1 credit. S-U or letter grades. 12 lec. Sec 07 Plant Cell Walls: Structure to expression, biolistic transformation, confocal K. van Wijk. Proteome microscopy, laser capture microdissection, Introduction to proteomics and mass 1 credit. S-U or letter grades. 12 lec. microarray analysis, genetic mapping and spectrometry and its application in plant Offered alternate years. J. Rose. mutant analysis, transposon tagging, biology. Includes discussion o f protein Examines the structure and function o f plant proteomics, and metabolite analysis. separation, protein tagging and visualization cell walls, exploring their dynamic nature and techniques; principles o f biological mass [BIOPL 642(6420) Plant Mineral Nutrition fundamental contribution to numerous aspects spectrometry and interpretation o f spectra; (also CSS 642(6420]) o f plant growth and development. Topics bioinformatics tools in proteomics; comparative Spring. 3 credits. Prerequisite: BIOPL 342 include wall biosynthesis; wall structure and proteomics; phosphorylation mapping. or equivalent. Lec. Offered alternate years; composition; regulation of cell expansion and Discusses limitations and possibilities o f next offered 2008-2009. L. V. Kochian and differentiation; defense against pathogens and proteomics on plants for which little sequence R. M. Welch. signaling; the apoplast as a metabolically information is available and experimental Detailed study of the processes by which active subcellular compartment; and analytical papers involving plant proteomics. plants acquire and use mineral nutrients from techniques: from biochemistry to proteomics. the soil. Topics include the uptake, [Sec 03 Plant Genome Organization and [Sec 08 Plant Imaging translocation, and compartmentation o f Function (also PL BR 483.3) 1 credit. 4 weeks. Limited to 12 students. mineral elements; root-soil interactions; the 1 credit. 12 lec. Offered alternate years; Priority given to graduate students. S-U or metabolism o f mineral elements; the next offered 2008-2009. S. D. Tanksley. letter grades. Offered alternate years; next involvement o f mineral nutrients in various Covers the structure and variation o f plant offered 2008-2009. R. Turgeon. physiological processes; and the nutrition o f nuclear genomes, including changes in At some point, most research projects involve plants adapted to extreme environmental genome size, centromere/telomere structure, structural analysis. This may necessitate hand stresses (e.g., acid soils). Specific mineral DNA packaging, transposable elements, sectioning, in situ hybridization, electron elements are emphasized to illustrate these genetic and physical mapping, positional gene microscopy, confocal imaging, or any of a topics.] cloning, genomic sequencing and comparative host o f other possibilities. The key to success genomics.] BIOPL 647(6470) Systematic Biology is having a proper foundation so that the Journal Club correct choice of procedures can be made. [Sec 04 Molecular Aspects of Plant Fall or spring. 1 credit; may be repeated This course provides an overview o f the Development I (also BIOBM 483.4) for credit. Intended for graduate students theory o f practical light and electron 1 credit. 12 lec. Next offered 2008-2009. and advanced undergraduates in systematic microscopy focusing on plant tissue J. B. Nasrallah. biology. S-U grades only. Disc TBA. Bailey preparation, hand and microtome sectioning, Focuses on the molecular genetics o f plant Hortorium staff. staining, optical techniques, histochemistry, development with an emphasis on plant Discussions led by staff, visitors, and students and the localization o f macromolecules. reproductive biology. Current approaches to on topics o f current importance to systematic Students will gain an appreciation o f the the elucidation o f the molecular signals and biology. potentials and limitations o f available methods pathways that lead to the establishment o f the and learn how these methods can be differentiated state o f floral cells and organs [BIOPL 649(6490) Solute Transport in combined to answer specific research are discussed. Topics include cell-cell Plants (also BEE 649(6490]) questions. The course will consist o f six signalling in the establishment o f pattern and Fall. 3 credits. Letter grades only. Lec. lectures and four two-hour labs, plus a short functional differentiation o f specialized cell Offered alternate years; next offered 2008- project.] types, and the control o f developmental 2009- R. M. Spanswick. pathways by endogenous and external cues. For description, see BEE 649.] The module is a companion to BIOPL 482, Sec 02 (Molecular Plant-Microbe Interactions).] 180 BIOLOGICAL SCIENCES - 2 8

BIOPL 651(6510) Water Transport in Seminar with student presentations o f current 2. BIOSM 364 Field Marine Science or Plants (also BEE 647[6470]) topics in systematics. BIOSM 375 Field Marine Biology and Fall. 2 credits. Letter grades only. Lee. Ecology and at least one 400-level BIOSM Offered alternate years. R. M. Spanswick. BIOPL 746(7460) Seminar in Systematic field course at the Shoals Marine Botany: Student Research For description, see BEE 647. Laboratory Spring. 1 credit. Prerequisite: graduate BIOPL 654(6540) Botanical standing or permission o f instructor. Letter 3. BIOEE 462 Marine Ecology Nomenclature grades only. Sem. Bailey Hortorium staff. Undergraduate Specialization in Ocean Fall. 1 credit. Prerequisite: written Student-led seminar presentation based on his Sciences permission o f instructor. S-U grades only. or her thesis research or a related topic. Lee and disc. Offered alternate years. Science o f Earth Systems majors have the M. A. Luckow. BIOPL 749(7490) Graduate Research in option o f specializing their program o f study Botany in the area o f ocean sciences. This Fall or spring. Variable credit; may be interdisciplinary specialization is intended for repeated for credit. S-U or letter grades. students with interests in understanding the Staff. I Analysis o f the International Code o f Botanical interaction o f biological, chemical, geological, Similar to BIO G 499 but intended for Nomenclature and its application to various and physical processes in ocean systems. In graduate students w ho are working with plant groups. addition to fulfilling the Science o f Earth faculty members on an individual basis. Systems general requirements (see the SES BIOPL 656(6560) Topics in Plant program description in Interdisciplinary Evolution Related Courses in Other Departments Centers, Programs, and Studies section o f Spring. 1 credit. Prerequisite: BIOPL 448 or catalog), students in ocean sciences are equivalent background in evolution, or Current Topics in Plant Molecular Ecology required to take four advanced courses from written permission o f instructor. Lab and (BIOEE 675) the following list to fulfill their major disc. Offered alternate years. K. J. Niklas. Fungi (PL PA 309) requirements: Series o f selected topics to provide a background in plant evolution, paleobotanical Seaweeds, Plankton, and Seagrass: The 1. BIOEE 373 Biology o f the Marine literature, and evolutionary theory. Among the Ecology and Systematics of Marine Invertebrates topics discussed are the origin o f a terrestrial Plants (BIOSM 449) 2. BIOEE 457 Limnology flora, the evolution o f the seed plants, and the Fungal Biology (PL PA 649) 3. BIOEE 462 Marine Ecology origin and adaptive radiation o f the angiosperms. Physiological Plant Ecology, Lectures and 4. BIOEE 478 Ecosystem Biology Laboratory (BIOEE 466/468) BIOPL 740(7400) Plant Biology Seminar 5. BIOEE 490 Topics in Marine Biology Fall and spring. 0 credits (no official Plant Behavior-Induced Plant Responses to 6. BIOSM 377 Diversity o f Fishes registration). Requirement for graduate Biotic Stresses (BIOEE 446) students doing work in plant biology. 7. BIOSM 310 Marine Symbiosis Plant Cytogenetics Laboratory (PL BR 446) W. L. Crepet. 8. BIOSM 309 Climates and Ecosystems Lectures on current research in plant biology, Teaching Experience (BIO G 498) presented by visitors and staff. 9. BIOSM 329 Ecology o f Animal Behavior Undergraduate Research in Biology (BIO G 10. BIOSM 364 Field Marine Science BIOPL 741(7410) Problems in Plant Cell 499) and Molecular Biology 11. BIOSM 365 Underwater Research Spring. 2 credits. Prerequisite: first- and 12. BIOSM 374 Field Ornithology second-year graduate students in Plant Cell and Molecular Biology Program. Disc. 13. BIOSM 375 Field Marine Biology and Introduction to the research literature in plant COURSES IN MARINE SCIENCE Ecology molecular and cellular biology through weekly Cornell offers an extensive listing o f 14. BIOSM 413 Research in Marine Biology problem sets and discussions. undergraduate courses in marine science. 15. BIOSM 449 Seaweeds, Plankton and Undergraduates interested in pursuing studies BIOPL 742(7420) Current Papers in Plant Seagrass Biology in marine science are encouraged to explore Fall or spring. 1 credit. Limited enrollment. the undergraduate specialization in marine 16. BIOSM 376 Marine Invertebrate Zoology Primarily for graduate students, with biology offered through the Office of (Note: Not same as BIOEE 373) Undergraduate Biology, the undergraduate priority given to majors or minors in plant 17. BIOSM 477 Marine Vertebrates molecular biology. Prerequisite: for specialization in ocean sciences offered undergraduates, written permission o f through the Science o f Earth Systems Program, 18. EAS 375 Sedimentology and Stratigraphy instructor. S-U grades only. Sem. Staff. and the summer program o f courses offered 19. EAS 455 Geochemistry by the Shoals Marine Laboratory. Further BIOPL 743(7430) Faculty Research in information on these programs can be found 20. EAS 475 Special Topics in Oceanography Plant Cell and Molecular Biology at the Cornell Marine Programs Office, G14 21. EAS 479 Paleobiology Fall. 1 credit. Prerequisite: graduate Stimson Hall, or at www.sml.cornell.edu. standing or written permission from Sea Semester member o f Plant Cell and Molecular Undergraduate Specialization in Marine BIOSM 366 SEA: Introduction to Biology Program or from coordinator for Biology and Oceanography Oceanography undergraduates. Disc TBA. Staff. Biological sciences majors in the Ecology and Introduction for graduate students to the Evolutionary Biology program of study have BIOSM 367 SEA: Introduction to Maritime research being conducted by Cornell faculty in the option o f specializing their program o f Studies the Plant Cell and Molecular Biology Program. study in the area o f Marine Biology. This BIOSM 368 SEA: Introduction to Nautical BIOPL 744(7440) Graduate Research in specialization is intended for students with Science Plant Cell and Molecular Biology interests in understanding the unique aspects BIOSM 369 SEA: Practical Oceanography I Fall or spring. 1 credit. Requirement for, o f organismal biology in the marine and limited to, second-, third-, and fourth- environment. In addition to fulfilling the major BIOSM 370 SEA: Practical Oceanography II and the ecology and evolutionary biology year graduate students in Plant Cell and BIOSM: 372 SEA: Practical Oceanography III Molecular Biology. Sem. Staff. program o f study requirements, students in Each student presents one seminar per year marine biology are encouraged to enroll in BIOSM 378 SEA: Oceans and Climate: Oceans on his or her thesis research and then meets the following courses: in the Global Carbon Cycle with the thesis committee members for 1. BIOEE 154 The Sea: An Introduction to BIOSM 379 SEA: Ocean Science and Public evaluation. Oceanography Policy BIOPL 745(7450) Seminar in Systematic BIOSM 380 SEA: Oceanographic Field Botany Methods Fall. 1 credit. Prerequisite: graduate standing or permission o f instructor. S-U grades only. Sem. Bailey Hortorium staff. SHOALS MARINE LABORATORY 181

BIOSM 381 SEA: Independent Research in program offered in cooperation with the Sea details or an application, contact SML Oceans and Climate Education Association (SEA). office, G14 Stimson Hall. Daily lec, lab, and fieldwork for two weeks. BIOSM 362 SEA: Maritime History and Culture The following marine sciences courses are currently administered by the Cornell Marine Allows students w ho are not biology majors to BIOSM 371 SEA: Marine Environmental experience the breadth o f the marine sciences Programs Office. (Not all o f these courses are History under field conditions at an island laboratory. offered each semester; consult the SML web Aspects o f biology, geology, earth science, Students in both marine science specializations site for current offerings: www.sml.comell.edu.) are exposed to an integrated program o f chemistry, and physics are included. Specific study, emphasizing a natural progression o f BIOSM 111(1110) A Marine Approach to topics include beach, salt marsh, tidal mud formal course work combined with ample Introductory Biology flat, tide pool, and benthic offshore opportunities for practical field experience. Summer. 8 credits. Prerequisite: score o f 4 environments; identification o f marine plants or higher on AP Biology Exam (which and animals; chemical and physical These courses must be taken concurrently. fulfills introductory biology requirement for oceanography; marine geology; and ecology Special program run by the Sea Education biology majors). W. E. Bemis and o f kelp beds and urchin barrens. Association. Contact the Marine Programs J. B. Heiser. office (255-3717) for more details. A four-week course for pre-freshmen at Shoals BIOSM 162(1620) Marine Environmental Science Marine Lab on Appledore Island in the Gulf o f Maine including daily fieldwork, boat trips, Summer. 4 credits. Prerequisite: open to outdoor adventure, and practices for high school students who have successfully completed two high school science SHOALS MARINE LABORATORY sustainable living. Intensive lectures, laboratory, and fieldwork occur in a learning environment courses. Special two-week course offered (BIOSM) emphasizing individual skill building, project at Shoals Marine Laboratory (SML), located G14 Stimson Hall, 255-3717 design and execution, and collaboration with on an island o ff Portsmouth, N.H. For faculty and peers. We explore four major more details or an application, contact SML The objective o f the Shoals Marine Laboratory themes— ecology, behavior, development, and office, G14 Stimson Hall. Daily lec, lab, (SML) is to provide undergraduates and other and fieldwork for 14 days. SML faculty. genomics— and connect them using examples interested adults a unique opportunity to explore Environmental studies have become an from evolution. Social issues discussed range marine sciences in an island setting noted for its from global environmental change to integral component o f high school programs biota, geology, and history. SML has established a sustainable fisheries, emerging infectious all around the country; however, opportunities national reputation for excellence and has diseases, and stem cell research. to apply this course work to the marine become North America’s largest marine field environment are limited. Marine station focusing on undergraduate education. BIOSM 123(1230) Ocean Sciences Environmental Science focuses on coastal The summer population o f Appledore Island Summer. 4 credits. Prerequisite: permission marine habitats, with an emphasis on issues as is limited to about one hundred people at any o f instructor. Letter grades only. D. Taylor. they relate to global habitats and concerns. one time. Participants and faculty members An inquiry-based, hands-on introduction to Laboratory exercises and fieldwork include can literally and figuratively immerse marine biology using the rocky intertidal and explorations along Appledore Island’s rocky themselves in their explorations, free from open sea as a natural laboratory. Students will intertidal zone and excursions to neighboring distractions common to most academic learn the skills necessary for success in islands to observe harbor seal and seabird institutions. Because SML is a residential science courses: understanding disciplinary colonies. Offshore cruises include facility, a sense o f community develops that discourse; reading scientific papers critically, oceanographic sampling exercises and field makes courses and seminars at SML generating scientific hypotheses; designing trips to seabird and whale foraging grounds. outstanding educational and intellectual experiments and interpreting data using basic Lectures and discussions expose MES students experiences. Participants learn from and statistics; and constructing, presenting and to topics ranging from fishes to fisheries, exchange ideas with a wide range of understanding data in graphs and tables. seaweeds to lobsters, and plankton to whales. specialists whose primary interests are marine Students will learn and apply these skills to Fundamental scientific research methods and but whose perspectives often differ, providing field exercises centered on textbooks, primary equipment are introduced, and each student fertile ground for lively discussions. literature, lectures, seminars and discussions. has the opportunity to be involved in group research projects. Credit courses at Shoals Marine Laboratory are BIOSM 160(1600) The Oceanography of full-time, intensive learning experiences. the Gulf of Maine BIOSM 204(2040) Biological Illustration Courses may be taken sequentially, but not Summer. 4 credits. Limited to 24 students. Summer. 2 credits. Special one-week concurrently. A typical day combines lecture Special two-week course offered aboard a course offered at Shoals Marine Laboratory sessions, laboratory and field work, field trips SEA vessel and at Shoals Marine (SML), located on an island o ff Portsmouth, to nearby islands and the mainland, and Laboratory (SML), located on an island off N.H. For more details or an application, collecting and research excursions aboard the Portsmouth, N.H. For more details or an contact SML office, G14 Stimson Hall. Daily laboratory’s 47-foot research vessel, John M. application, contact SML office, G14 sessions for one week. SML faculty. Kingsbury, or the 36-foot research vessel, John Stimson Hall or Sea Education Association General discussion o f scientific publishing, B. Heiser. Field experience is an integral office, P.O. Box 6, Woods Hole, MA 02543. illustration labeling, color techniques, and component o f all courses, using Appledore’s Daily lec, lab, and fieldwork for two printing processes. Provides the scientist or extensive intertidal and subtidal zones, wading weeks. SML faculty. science student a chance to experience several bird rookeries, and seabird colonies. Faculty, Exciting opportunity to explore the offshore illustration techniques with the goal o f obtaining drawn from Cornell University, the University and near-coastal environments o f the Gulf o f an overview o f scientific and wildlife illustrations. of New Hampshire, and other leading Maine for advanced high school students. The student may choose a single technique to academic institutions, are selected based not Students spend 10 days aboard the Sea explore in depth. Course size is limited so that only on their academic excellence but also on Education Association’s sailing vessels round individual attention can be emphasized. trip between Woods Hole, Mass., and the Isles their teaching ability in the field. In addition, BIOSM 210(2100) Boats for Biologists there are numerous guest lecturers including o f Shoals via Georges Bank and the Gulf o f Summer. 2 credits. S-U or letter grades. engineers, coastal planners, and specialists Maine. Besides operating the ship, students W. E. Bemis. from private industry, government, and the study the many characteristics o f this unique Marine and freshwater biologists rely on academic community. ocean environment. Following the sea component, students spend seven days at the boats— small and large— for transportation, The Ithaca campus functions o f the Shoals Shoals Marine Laboratory collecting data environmental sampling, data collection, and Marine Laboratory are centered in the Cornell characteristic o f the Isles o f Shoals coastal other tasks. This course offers biologists at all Marine Programs office, G14 Stimson Hall. The environment. stages o f their careers a chance to learn basic office serves as an advising center for students boat handling, piloting, navigation, and interested in the marine sciences, maintains a BIOSM 161(1610) Introduction to Marine common sampling techniques to enhance browsing library with updated information on Science their opportunities for research and education. graduate study and career opportunities as well Summer. 4 credits. S-U or letter grades. This course meets all certification as on marine programs at other institutions, Special two-week course offered at Shoals requirements for basic small boat handling by and administers the SEA Semester, a 17-credit Marine Laboratory (SML), located on an state-specific agencies and National island o ff Portsmouth, N.H. For more Association o f Boating Law Administrators. 182 BIOLOGICAL SCIENCES - 2 li 8

BIOSM 225(2250) Sustainability in the student learns to use a variety o f light biological oceanography, ichthyology, and 21 st Century microscopic techniques, and contributes, fisheries. Attention is also given to Summer. 4 credits. W. E. Bemis. through fieldwork and by written reports, to a introductory physical and chemical Students will consider the challenges o f comprehensive survey o f symbiotic oceanography and marine geology. Marine institutionalizing sustainability. Through guest associations on and around Appledore Island. ecology and the effects o f human activity on lectures and fieldwork, they will grapple with the marine environment are included. Students challenges inherent to the field, devising BIOSM 312(3120) Biology of the Lobster apply this knowledge by conducting a transect strategies to enhance sustainability on Summer. 2 credits. Prerequisite: one year study toward the end o f the course. FMS Appledore Island. Topics include: systems o f college level biology. J. Factor. places emphasis on ichthyology, fisheries thinking, food and fisheries, engineering and An introduction to the biology o f the biology, general oceanography (biological, energy, and climate science. American lobster, Homarus americanus. The physical, and chemical), and marine geology. course will include an overview of this FMBE (BIOSM 375) places an additional BIOSM 276(2760) Seabird Conservation ecologically and economically important emphasis on ecology, especially in the Summer. 2 credits. J. Ellis. species, as w ell as cover several major topics intertidal zone; ecological, evolutionary and Combines lectures from specialists (e.g., in depth, each taught by a lobster biologist physiological adaptations o f marine organisms; ecologists and wildlife veterinarians) with a expert in that field. Topics may include life and field experiments. variety o f field-based activities related to history, larval development and seabird ecology and conservation. Topics will metamorphosis, anatomy, physiological BIOSM 365(3650) Underwater Research include: seabird identification, behavioral adaptation, fisheries and fishing methods, Summer. 4 credits. Prerequisites: one year studies, census techniques, population threats feeding mechanisms, ecology, and behavior. college-level biology, recognized SCUBA (e.g., fisheries bycatch, pollution), and Course will include lecture, laboratory, certification, and medical exam. S-U or restoration. A field trip to a nearby seabird discussion, and the natural field environment letter grades. Special two-week course restoration island will be included. o f Appledore Island. offered at Shoals Marine Laboratory (SML), located on an island o ff Portsmouth, N.H. BIOSM 308(3080) Field Microbial Ecology BIOSM 320(3200) Functional Morphology For more details or an application, contact Summer. 4 credits. Prerequisite: of Marine Organisms SML office, G14 Stimson Hall. Daily lec introductory biology or permission of Summer. 4 credits. Prerequisite: one year and fieldwork for two weeks. instructor. E. Zettler and L. A. Zettler. introductory biology or one semester Covers the philosophy o f research, hypothesis The microbial world dominates the biosphere introductory biology, general zoology, and testing and experimental design, sampling in terms o f biomass, diversity, and metabolic general botany. F. Fish. methods, various underwater techniques, flexibility. This course will introduce students A study o f the structure, form, and function o f diving physics and physiology, and use of to collecting, observing, and identifying live morphological adaptations in marine plants dive tables. Emphasizes subtidal ecological representatives o f these fascinating microbial and animals as examined through a research. Requirements include critical organisms including bacteria, protists, fungi, mechanical and ecological perspective. The evaluation o f several journal articles and and microscopic animals. Taxonomy and course will investigate the biomechanics of production o f a research proposal. ecology o f the basic groups o f microorganisms marine organisms through lecture, laboratory will be covered while students learn to collect demonstrations, and independent research BIOEE 373(3730) Biology of the Marine in the field for observation, experimentation projects. Invertebrates and isolation. Fall (but course must be taken previous BIOSM 329(3290) Ecology of Animal summer at Shoals Marine Laboratory BIOSM 309(3090) Coastal Ecology and Behavior (also BIONB 329(3290]) [SML]), three-week, full-time course. 5 Bioclimates Summer. 4 credits. Prerequisite: one year credits (students enroll for credit during Summer. 4 credits. Prerequisite: one year introductory college biology. fall semester). Limited to 24 students. college-level biology; background Recommended: course work in ecology, Prerequisites: one year introductory preferred in physics/physical geography. psychology, or behavior. S-U or letter biology for majors; permission o f faculty S-U or letter grades. Special two-week grades. Special two-week course offered at because o ff campus. Letter grades only. course offered at Shoals Marine Laboratory Shoals Marine Laboratory (SML), located on Daily and evening lec, lab, and fieldwork. (SML), located on an island o ff Portsmouth, an island o ff Portsmouth, N.H. For more Offered alternate years. C. D. Harvell and N.H. For more details or an application, details and an application, contact SML J. G. Morin. contact SML office, G14 Stimson Hall. Daily office, G14 Stimson Hall. Daily lec, lab, and lec, lab, and fieldwork for two weeks. SML fieldwork for two weeks. SML faculty. BIOSM 374(3740) Field Ornithology faculty. The ecological significance of behaviors of Summer. 4 credits. Prerequisite: one year Study o f the fundamentals o f organism- coastal organisms, with emphasis on field and college-level biology. S-U or letter grades. environment interaction developed through laboratory research methods. Lectures and Special two-week course offered at Shoals defining and measuring abiotic factors readings address the major subareas o f Marine Laboratory (SML), located on an including solar radiation, temperature, behavior (communication, orientation, social island o ff Portsmouth, N.H. For more atmospheric moisture, precipital wind, and behavior, foraging, predator avoidance, and details or an application, contact SML currents. On-site exploration o f the dynamics sensory mechanisms). Each student engages in office, G14 Stimson Hall. Daily lec and of meteorology and the role of abiotic and short-term behavioral observation and fieldwork for two weeks. SML faculty. biotic factors in the life o f coastal and marine prepares a research proposal for studying a Introduction to field ornithology focusing on plants and animals including humans. problem within the course subject area. the biology, ecology, and behavior o f the avifauna on the Isles o f Shoals. Focuses on BIOSM 310(3100) Marine Symbiosis BIOSM 364(3640) Field Marine Science fieldwork designed to observe and study Summer. 4 credits. Prerequisite: one full (FMS) many concepts frequently taught in the year college-level biology. Recommended: Summer. 8 credits. Prerequisite: one year classroom setting including territoriality, background in microbiology or cell college biology. S-U or letter grades. breeding biology, and survivorship. Students biology. S-U or letter grades. Special two- Special four-week course offered twice learn and apply numerous ornithological field week course offered at Shoals Marine each summer at Shoals Marine Laboratory methods including various census techniques, (SML), located on an island o ff Portsmouth, Laboratory (SML), located on an island o ff territory mapping, banding, behavioral Portsmouth, N.H. For more details or an N.H. Students may not take FMS after observations, and creating a field notebook. application, contact SML office, G14 taking FMBE (BIOSM 375). For more Stimson Hall. Daily lees and fieldwork for details or an application, contact SML BIOSM 375(3750) Field Marine Biology two weeks. SML staff. Office, G14 Stimson Hall. Daily lec, lab, and Ecology (FMBE) Introduction to the concepts of symbiosis as and fieldwork for four weeks. Three core Summer. 8 credits. Prerequisites: one full applied to marine organisms, with an faculty members assisted by up to 15 year college-level biology. S-U or letter emphasis on microbial symbionts. Students visiting lecturers, including representatives grades. Four-week course offered at Shoals develop the ability to analyze symbioses using o f governmental agencies. SML faculty. Marine Laboratory (SML), located on an a comprehensive set o f criteria, including Designed for the student who desires an initial island o ff Portsmouth, N.H. For more duration, propagation, specificity, integration overview o f the marine sciences, this course details or an application, contact SML and modes o f interaction. Morning lectures emphasizes living material in natural habitats. office, G-14 Stimson Hall. Daily lec, lab, are followed by afternoons collecting, Most of the course work is concerned with and fieldwork for four weeks. SML faculty. preparing and studying live specimens. Each the biology o f intertidal plants and animals, SHOALS MARINE LABORATORY 183

Designed for students seeking an introduction to BIOSM 413(4130) Research In Marine make up the group Chondrichthyes. This the marine sciences and marine ecology; FMBE Biology course will cover advanced topics in the emphasizes fieldwork in natural habitats. Summer. 6 credits. Prerequisite: one year evolution, diversity, anatomy, functional Examines aspects o f the biology and ecology o f college-level biology. Recommended: morphology, neurobiology, sensory systems, marine organisms, including intertidal plants and experience in ecology or physiology. S-U or behavior, reproduction, development, and invertebrates, fishes, marine mammals and birds, letter grades. Special three-week course conservation o f cartilaginous fishes. biological oceanography, and human impacts on offered at Shoals Marine Laboratory (SML), the marine environment. FMBE places a special located on an island o ff Portsmouth, N.H. BIOSM 472(4720) Molecular Systematics emphasis on the ecology o f the intertidal zone For more details or an application, contact Summer. 4 credits. Prerequisite: one year and ecological, evolutionary, and physiological SML office, G14 Stimson Hall. Daily lec, lab, introductory biology; genetics and/or cell adaptations o f marine organisms. Students may and fieldwork for three weeks. SML faculty. biology with laboratory components. not take FMBE after taking FMS (BIOSM 364). Introduction to the physiological ecology and Recommended: evolutionary biology, functional morphology o f marine plants and ecology vertebrate/invertebrate zoology. [BIOSM 376(3760) Marine Invertebrate animals, with emphasis on selected algal and A. Shedlock. Zoology invertebrate examples from the Gulf o f Maine. An introduction to basic concepts in Summer. 6 credits. Prerequisite: one year Topics include photosynthesis in the marine systematic biology, evolutionary genetics, introductory biology and permission o f environment; respiration in intertidal molecular ecology, and conservation biology instructors. Students may not take BIOSM organisms; carbohydrates, proteins, and lipids emphasizing the natural history o f marine 376 after taking BIOEE 373. S-U or letter as nutrients in the sea; acclimation and organisms. Students integrate field sampling grades. Special three-week course offered at tolerance of tide-pool biota; and biological techniques, taxonomy, and curation o f Shoals Marine Laboratory (SML), located on responses to competition and grazing. Field specimens from Appledore Island and an island o ff Portsmouth, N.H. For more and laboratory exercises explore principles surrounding waters with molecular diagnostics details or an application, contact SML office, and procedures used to characterize the completed in the laboratory. Standard methods G14 Stimson Hall. Daily lec, lab, and physical, chemical, and biotic environment o f for DNA purification, amplification, fieldwork for three weeks. Offered alternate intertidal and shallow subtidal organisms, sequencing, and genotyping are used to years; offered summer 2008. SML faculty. including determination o f temperature, light, address questions about population structure, Introduction to the biology and evolution of salinity, oxygen and nutrient levels, and in kinship, and species phylogeny. the major invertebrate phyla, concentrating on vivo functional analyses o f metabolic BIOSM 477(4770) Marine Vertebrates marine representatives. Emphasizes the phenomena. The process o f scientific Summer. 6 credits. Prerequisites: vertebrate evolution o f form and function, and the investigation is the predominant theme o f the biology course. S-U or letter grades. Special ecology, behavior, physiology, chemical course. ecology, and natural history o f invertebrates. three-week course offered at Shoals Marine Appledore Island’s unique location provides BIOSM 445(4450) Forensic Science for Laboratory (SML), located on an island off an excellent venue for the study o f freshly Marine Biologists Portsmouth, N.H. For more details or an collected and in situ representatives o f most Summer. 2 credits. Prerequisite: satisfactory application, consult SML office, G14 o f the major phyla.] completion of college-level course in Stimson Hall. Daily lec, lab, and fieldwork biology, ecology, or marine science. for three weeks. SML faculty. BIOSM 377(3770) Diversity of Fishes W. Lord. Topics in marine vertebrate biology Summer. 6 credits. Prerequisite: one full Forensic science represents the unique emphasizing laboratory studies, field year college-level biology. Recommended: merging o f scientific insight and the law. collections or observations, and readings from background in vertebrate biology. S-U or Forensic Science for Marine Biologists the current literature. Topics include letter grades. Special two-week course provides a field-oriented introduction to the systematics o f fishes o f the Gulf o f Maine; offered at Shoals Marine Laboratory (SML), forensic science domain and the utilization o f elasmobranch physiology; interpretation o f life located on an island o ff Portsmouth, N.H. marine biology within the justice system. history and parameters from otolith For more details or an application, contact Students receive comprehensive instruction microstructure; teleost skeletomuscular SML office, G14 Stimson Hall. Daily lees concerning the recognition, documentation, structure and function; population biology and and fieldwork for two weeks. SML staff. collection, and preservation of physical the contemporary Gulf o f Maine fishery; Intensive lecture, laboratory, and field course. evidence. Additionally, students develop Mesozoic marine reptiles; the biology o f sea Lectures cover the basic anatomy and practical incident response, scene turtles in cold water; coloniality in sea birds; physiology of fishes with examples drawn management, and forensic teamwork skills. avian adaptations to life at sea; evolution and from a wide variety o f fishes from throughout systematics o f marine mammals; diving the world. The course emphasizes the BIOSM 449(4490) Seaweeds, Plankton, physiology; and ecology and conservation of diversity o f fishes in two aspects, diversity o f and Seagrass: The Ecology and existing marine mammal populations. Systematics of Marine Plants evolutionary solutions to problems faced by Dissection o f vertebrate animals is a part o f Summer. 4 credits. Prerequisite: BIOSM 364 fishes and the great diversity o f different types one or more laboratory sessions. o f fishes that inhabit the world. Laboratory or one year introductory biology. S-U or exercises cover the anatomy and osteology of letter grades. Special two-week course BIOSM 495(4950) Research Methods In teleost fishes and identification o f local offered at Shoals Marine Laboratory (SML), Marine Biology species. Each student selects a different local located on an island o ff Portsmouth, N.H. Summer. 1 credit. Corequisite: BIOSM 499 species o f teleost fish to study and dissect and For more details or an application, consult or permission o f instructor. Primarily for prepares a comprehensive paper on its SML office, G14 Stimson Hall. Daily lec, undergraduates. Special eight-week course morphology, soft anatomy, and osteology. lab, and fieldwork for two weeks. SML offered at the Shoals Marine Laboratory faculty. (SML). For more details or an application, BIOSM 410(4100) The Herring Gull’s Overview o f the major marine algal groups, contact SML office, G14 Stimson Hall. World including aspects o f anatomy morphology, Weekly sem for eight weeks. SML faculty. Summer. 2 credits. Prerequisite: development, life histories, physiology, and use. Seminar course on research methodology, introductory biology. T. Seeley P. Sherman, Laboratories and fieldwork emphasize experimental design, statistical analyses, and and J. Shellman Sherman. relationships between distribution and major scientific writing. The course is designed to An introduction to the study o f animal environmental parameters and involve student assist students in the research they are behavior in the field by focusing on the projects. conducting while enrolled in BIOSM 499. behavior o f the herring gulls nesting on Appledore Island. Topics will include natural BIOSM 465(4650) Sharks: The Biology, BIOSM 499(4990) Research In Biology selection and behavior, levels o f analysis, Evolution, and Conservation of Summer. Variable credit; 2 credits per animal communication, territoriality, kin Sharks and Their Allies seven days on site. For more details and recognition, orientation, and mating systems. Summer. 2 credits. Prerequisite: vertebrates an application, contact SML office, G14 The Herring G ull’s World, the classic book o f or comparative anatomy and ichthyology Stimson Hall. or permission o f SML director. J. Morrissey Nobel Laureate Niko Tinbergen, will be read Section A: Independent Biological and W. E. Bemis. and discussed. Methods o f measuring behavior Research: Independent study with a member The last 30 years have produced an explosion and designing experiments will be taught, and o f the Shoals Marine Laboratory core faculty, o f new information on the biology o f the students will conduct individual research based on student faculty interest and available approximately 1,000 living species of sharks, projects. facilities. A short proposal o f research must be skates, rays, and chimaeras, which collectively sent with application materials. 184 BIOLOGICAL SCIENCES - 2fiTiCTCTifil 8

Research Experiences for Undergraduates For more information, contact Shoals Marine BIOSM 372(3720) SEA Practical (REU) Laboratory office, G14 Stimson Hall, or call Oceanography III 0 credits. The National Science Foundation SEA directly at 800-552-3633. Program costs Summer. 3 credits. Prerequisites: BIOSM (NSF) Research Experiences for are to be paid in place of regular Cornell 366, 367, and 368. Undergraduates (REU) program provides tuition and fees. Theories and problems raised in class are tested in the practice o f oceanography at sea. support for undergraduates to pursue Instructors for the SEA Semester include supervised, independent research projects During lectures and watch standing, students faculty o f the Sea Education Association and are instructed in the operation o f basic at the Shoals Marine Laboratory. Nine the Woods Hole Oceanographic Institution students are selected from a competitive, oceanographic equipment, in the and others. national pool to participate in the eight- methodologies involved in the collection, week summer program. For more analysis, and reduction o f oceanographic data, information and an application, contact SEA Basic Semester and in the attendant operations o f sailing an SML office, G14 Stimson Hall, or view BIOSM 366(3660) SEA Introduction to oceanographic research vessel. Group research SML’s web site at www.sml.cornell.edu. Oceanography projects are completed. 3 credits. Corequisites: BIOSM 367 and 368. I BIOSM 650(6500) Field Marine Ecology Survey o f the characteristics and processes o f SEA Oceans and Climate Semester and Environmental Science for the global ocean. Introduces oceanographic Teachers BIOSM 368(3680) SEA Introduction to concepts and develops them from their bases Nautical Science Summer. 2 credits. Prerequisites: one year in biology, physics, chemistry, and geology. college-level biology. Recommended: Provides a broad background in BIOSM 378(3780) SEA Oceans and teaching experience. A special one-week oceanography with special attention to areas Climate: Oceans in the Global course offered at Shoals Marine Laboratory pertinent to the subsequent cruise. Guest Carbon Cycle (SML), located on an island o ff Portsmouth, lecturers from the Woods Hole research Fall, spring. 4 credits. Prerequisite: 3 lab N.H. For more details or an application, community interpret current trends and science courses or permission o f instructor. contact SML office, G-14 Stimson Hall. activities in this rapidly evolving field. SML faculty. Daily lec and fieldwork for one week. SML Students develop individual projects to be This course examines the role of the oceans faculty. carried out at sea. in the climate system, addressing topics such Intended for teachers o f grades 6-12 but also as the global carbon cycle, the thermohaline open to undergraduate junior and senior BIOSM 367(3670) SEA Introduction to circulation, and aspects o f global change students interested in teaching. Teachers Maritime Studies including warming and sea level rise. develop hands-on, experiential approaches to 3 credits. Corequisites: BIOSM 366 and 368. the marine sciences, with an emphasis on Interdisciplinary consideration of our BIOSM 379(3790) SEA Ocean Science coastal and environmental issues. Extensions to relationship with the marine environment. and Public Policy (HA) freshwater ecology also are included. Fieldwork Covers the elements o f maritime history, law, Fall, spring. 3 credits. SEA faculty. is emphasized, with numerous excursions to literature, and art necessary to appreciate our This course seeks to provide students with a the rocky intertidal and with off-shore ocean marine heritage and to understand the fundamental understanding o f the intersection sampling. Lectures focus on biodiversity, political and economic problems of between climate change and governmental adaptations, predator-prey interactions, contemporary maritime affairs. policy and the interrelationship between environmental sustainability, and how to science and governmental policy. After an BIOSM 368(3680) SEA Introduction to introduction to the development o f maritime engage and motivate students with aquatic Nautical Science projects. law and sovereignty on the high seas, students 3 credits. Corequisites: BIOSM 366 and 367. will examine why societies funded oceanic BIOSM 699(6990) Research in Biology for An introduction to the technologies o f research. Teachers operation at sea. The concepts o f navigation Summer. 2 credits per week. Prerequisite: (piloting, celestial and electronic), naval BIOSM 380(3800) SEA Oceanographic BIOSM 650. Special course offered at architecture, ship construction, marine Field Methods Shoals Marine Laboratory (SML), located engineering systems, and the physics o f sail Fall, spring. 3 credits. Prerequisite: on an island o ff Portsmouth, N.H. For are taught from their bases in astronomy, successful completion o f BIOSM 378. SEA more details or an application, contact SML mathematics, and physics. Provides the faculty. office, G14 Stimson Hall. theoretical foundation for the navigation, This course introduces students to all aspects Opportunity for teachers who have taken seamanship, and engineering that students o f oceanographic fieldwork. Students learn BIOSM 650 to return to Shoals to pursue in employ at sea. practical skills in the operating principles and safe deployment of oceanographic greater depth a topic o f their choosing under BIOSM 369(3690) SEA Practical the direction o f the BIOSM 650 faculty. instrumentation. Skills acquired enable Oceanography I students to carry out an independent research 4 credits. Prerequisite: BIOSM 366. project. SEA Semester Theories and problems raised in the shore In cooperation with the Sea Education component are tested in the practice o f BIOSM 381(3810) SEA Independent Association (SEA), the Shoals Marine oceanography at sea. Students are introduced Research in Oceans and Climate Laboratory offers a semester-length sequence to the tools and techniques o f the practicing Fall, spring. 4 credits. Prerequisite: of courses designed to provide college oceanographer. During lectures and watch successful completion o f BIOSM 378. SEA undergraduates with a thorough academic, standing, students are instructed in the faculty. scientific, and practical understanding o f the operation o f basic oceanographic equipment; This course provides upper-level study sea. This sequence is repeated approximately in the methodologies involved in the collection, focused on oceanography and climate, once every two months throughout the year. reduction, and analysis o f oceanographic data; including the design and completion of an Students spend the first half o f SEA Semester and in the attendant operations o f a sailing independent research project that is (a six-week shore component) in Woods Hole, oceanographic research vessel. comparable in scope to an undergraduate Mass., receiving instruction in oceanography, senior research thesis. BIOSM 370(3700) SEA Practical nautical science, and maritime studies. The Oceanography II second half o f SEA Semester (a six-week sea SEA: Documenting Change in the Caribbean 4 credits. Prerequisites: BIOSM 368 and 369. component) is spent at sea aboard the SSV Semester Building on the experience o f Practical Robert C. Seamans or the SSV Corwith Oceanography I, students assume increasing BIOSM 362(3620) SEA Maritime History Cramer. Enrollment is open to both men and responsibility for conducting oceanographic and Culture (CA) (HA) women judged capable of benefiting from SEA research and overseeing operations o f the Fall, spring. 4 credits. SEA faculty. semester; a student must have successfully vessel. The individual student is ultimately In this course, students will explore political, completed at least one college-level responsible directly to the chief scientist and cultural and social changes in the Caribbean laboratory science course (or its equivalent) the master o f the vessel for the safe and since just before Europeans arrived at the end in order to be admitted to SEA Semester or orderly conduct o f research activities and o f the 15th century. Student will see how the SEA Summer Session. No prior sailing related operations o f the vessel. Each student physical nature o f the region has influenced experience is necessary. Cornell students undertakes an individual research project patterns o f settlement and development and enrolled in the SEA Semester must take the designed during the shore component. the impact o f European expansion. entire sequence. FACULTY ROSTER 185

BIOSM 366(3660) SEA Practical Feeny, Paul P., Ph.D., Oxford U. (England). Morin, James G., Ph.D., Harvard U. Prof., Oceanography I Prof, Ecology and Evolutionary Biology Ecology and Evolutionary Biology BIOSM 368(3680) SEA Introduction to Fitzpatrick, John W., Ph.D., Princeton U. Prof., Mortlock, Robert P., Ph.D., U. o f Illinois. Prof. Nautical Science Ecology and Evolutionary Biology/ Emeritus, Microbiology Laboratory of Ornithology Nasrallah, June B., Ph.D., Cornell U. Prof., BIOSM 370(3700) SEA Practical Flecker, Alexander S., Ph.D., U. o f Maryland. Plant Biology Oceanography II Assoc. Prof., Ecology and Evolutionary Nasrallah, Mikhail E., Ph.D., Cornell U. Prof., Biology Plant Biology BIOSM 371(3710) SEA Marine Fox, Thomas D., Ph.D., Harvard U. Prof., Environmental History (CA) (LA) Naylor, Harry B., Ph.D., Cornell U. Prof. Molecular Biology and Genetics Emeritus, Microbiology Fall, spring. 4 credits. SEA faculty. Fu, Jianhua, Ph D., U. Pittsburgh. Asst. Prof., Niklas, Karl J., Ph.D., U. o f Illinois. Prof., Plant This course is designed to explore the Molecular Biology and Genetics Biology interaction o f ecological factors in ocean, Ghiorse, William C., Ph.D. Rensselaer Nixon, Kevin C., Ph.D., U. o f Texas, Austin. coastal and island environments; the impact o f Polytechnic Inst. Prof., Microbiology Assoc. Prof., Plant Biology (Bailey human actions on those environments; and Gibson, Jane, Ph.D., U. o f London (England). Hortorium) the need for local, regional and international Prof. Emeritus, Molecular Biology and Owens, Thomas G., Ph.D., Cornell U. Assoc. responses and strategies to mitigate and Genetics Prof., Plant Biology manage that impact. Goldberg, Michael L., Ph.D., Stanford U. Prof., Paolillo, Dominick J., Jr., Ph.D., U. o f Molecular Biology and Genetics California. Davis. Prof. Emeritus, Plant Hanson, Maureen R., Ph.D., Harvard U. Prof., Biology Molecular Biology and Genetics/Liberty Parthasarathy, Mandayam V., Ph.D., Cornell U. FACULTY ROSTER Hyde, Prof., Plant Biology Prof. Emeritus, Plant Biology Harrison, Richard G., Ph.D., Cornell U. Prof., Peters, Joseph, Ph.D., U. o f Maryland. Asst. New York State College of Agriculture Ecology and Evolutionary Biology Prof, Microbiology and Life Sciences Harris-Warrick, Ronald M., Ph.D., Stanford U. Raguso, Robert A., Ph.D., U. o f Michigan. Prof., Neurobiology and Behavior Assoc. Prof, Neurobiology and Behavior Adler, Kraig K., Ph.D., U. o f Michigan. Prof., Harvell, C. Drew, Ph.D., U. o f Washington. Reeve, H. Kern, Ph.D., Cornell U. Prof., Neurobiology and Behavior Prof., Ecology and Evolutionary Biology Neurobiology and Behavior Agrawal, Anurag, Ph.D., U. o f California, Hay, Anthony, Ph.D., U. o f California. Assoc. Roberts, Jeffrey W., Ph D., Harvard U. Robert J. Davis. Assoc. Prof., Ecology and Prof, Microbiology Appel Professor o f Cellular and Molecular Evolutionary Biology/Entomology Helmann, John D., Ph.D., U. o f California, Biology, Molecular Biology and Genetics Alani, Eric E., Ph.D., Harvard U. Assoc. Prof, Berkeley. Prof., Microbiology Rodriguez, Eloy, PhD., U. o f Texas. Prof, Molecular Biology and Genetics Henry, Susan, Ph.D., U. o f California, Berkeley. Plant Biology (Bailey Hortorium) Anderson, John M., Ph.D., New York U. Prof. Prof., Molecular Biology and Genetics and Root, Richard B., Ph.D., U. o f California, Emeritus, Molecular Biology and Genetics Dean CALS Berkeley. Prof. Emeritus, Ecology and Angert, Esther R., Ph.D., Indiana U. Asst. Prof., Hopkins, Carl D., Ph.D., Rockefeller U. Prof., Evolutionary Biology/Entomology Microbiology Neurobiology and Behavior Rose, Jocelyn, Ph.D., U. o f California, Davis. Barbash, Daniel A., Ph.D., U. o f California, Howarth, Robert W., Ph.D., Massachusetts Inst, Asst. Prof., Plant Biology Berkeley. Asst. Prof., Molecular Biology and of Technology/Woods Hole Oceanographic Russell, James B., Ph.D., U. o f California, Genetics Institution. David R. Atkinson Professor o f Davis. Prof., Microbiology Bates, David M., Ph.D., U. o f California, Los Ecology and Environmental Biology, Scanlon, Michael, Ph.D., Iowa State U. Assoc. Angeles. Prof. Emeritus, Plant Biology Ecology and Evolutionary Biology Prof., Plant Biology (Bailey Hortorium) Hua, Jian, Ph.D., California Inst, o f Seeley, Jr., Harry W., Ph.D., Cornell U. Prof. Bemis, William E., Ph.D., U. o f California, Technology. Asst. Prof., Plant Biology Emeritus, Microbiology Berkeley. Prof., Ecology and Evolutionary Ingram, John W., Ph.D., U. o f California, Shalloway, David I., Ph.D., Massachusetts Inst, Biology/Shoals Marine Laboratory Berkeley. Prof. Emeritus, Plant Biology o f Technology. Greater Philadelphia Prof., Bruns, Peter J., Ph D., U. o f Illinois. Prof. (Bailey Hortorium) Molecular Biology and Genetics Emeritus, Molecular Biology and Genetics Jagendorf, Andre T., Ph.D., Yale U. Liberty Shapleigh, James P , Ph.D., U. o f Georgia. Cade, Thomas J., Ph.D., U. o f California, Los Hyde Bailey Professor of Plant Physiology Assoc. Prof., Microbiology Angeles. Prof. Emeritus, Ecology and Emeritus, Plant Biology Stinson, Harry T., Ph.D., Indiana U. Prof. Evolutionary Biology Kemphues, Kenneth J., Ph D., Indiana U. Prof., Emeritus, Molecular Biology and Genetics Calvo, Joseph M., Ph.D., Washington State U. Molecular Biology and Genetics Tye, Bik-Kwoon, Ph.D., Massachusetts Inst, of William T. Keeton Professor in Biological Kessler, Andre, Ph D., Max Planck Inst, for Technology. Prof., Molecular Biology and Sciences, Molecular Biology and Genetics Chemical Ecology/Friedrich Schiller U. o f Genetics Chabot, Brian F., Ph D., Duke U. Prof., Jena (Germany). Asst. Prof., Ecology and Uhl, Charles H., Ph D., Cornell U. Prof. Ecology and Evolutionary Biology Evolutionary Biology/Boyce Thompson Inst, Emeritus, Plant Biology Clayton, Roderick K., Ph.D., California Inst, o f for Plant Research Uhl, Natalie W., Ph D., Cornell U. Prof. Technology. Prof. Emeritus, Plant Biology Kingsbury, John M., Ph.D., Harvard U. Prof. Emeritus, Plant Biology (Bailey Hortorium) Crepet, William L., Ph.D., Yale U. Prof., Plant Emeritus, Plant Biology Van Wijk, Klaas J., Ph.D., Groningen U. (The Biology (Bailey Hortorium)* Kraus, W. Lee, Ph.D., U. o f Illinois. Asst. Prof., Davies, Peter J., Ph.D., U. o f Reading Netherlands). Assoc. Prof, Plant Biology Molecular Biology and Genetics Vogt, Volker M., Ph.D., Harvard U. Prof., (England). Prof., Plant Biology Lis, John T., Ph.D., Brandeis U. Prof, Davis, Jerrold I., Ph.D., U. o f Washington. Molecular Biology and Genetics Molecular Biology and Genetics Walcott, Charles, Ph.D., Cornell U. Prof., Assoc. Prof., Plant Biology (Bailey Lovette, Irby J., Ph.D., U. o f Pennsylvania. Neurobiology and Behavior Hortorium) Assoc. Prof., Ecology and Evolutionary Wayne, Randy O., Ph.D., U. o f Massachusetts. Dhondt, Andre A., Ph.D., Ghent State U. Biology/Laboratory of Ornithology Assoc. Prof, Plant Biology (Belgium). Edwin H. Morgens Professor of Luckow, Melissa A., Ph D., U. o f Texas, Austin. Winans, Stephen C., Ph D., Massachusetts Inst. Ornithology, Ecology and Evolutionary Assoc. Prof., Plant Biology (Bailey of Technology. Prof., Microbiology Biology/Laboratory of Ornithology Hortorium) Winkler, David W., Ph.D., U. o f California, Dondero, Norman C., Ph.D., Cornell U. Prof. MacDonald, Russell E., Ph.D., U. o f Michigan. Berkeley. Prof., Ecology and Evolutionary Emeritus, Microbiology Prof. Emeritus, Molecular Biology and Biology Doyle, Jeffrey J., Ph.D., Indiana U. Prof., Plant Genetics Wu, Ray, Ph.D., U. o f Pennsylvania. Prof., Biology (Bailey Hortorium) MacIntyre, Ross J., Ph.D., Johns Hopkins U. Molecular Biology and Genetics Dress, William J., Ph.D., Cornell U. Prof. Prof., Molecular Biology and Genetics Zahler, Stanley A., Ph.D., U. o f Chicago. Prof. Emeritus, Plant Biology (Bailey Hortorium) Madsen, Eugene L., Ph.D., Cornell U. Assoc. Emeritus, Molecular Biology and Genetics Eisner, Thomas, Ph.D., Harvard U. Jacob Prof, Microbiology Zinder, Stephen H., Ph D., U. o f Wisconsin. Gould Schurman Professor Emeritus, Marks, Peter L., Ph.D., Yale U. Prof. Emeritus, Prof., Microbiology Neurobiology and Behavior/Ecology and Ecology and Evolutionary Biology Evolutionary Biology* McCune, Amy R., Ph.D., Yale U. Prof., Ecology Emlen, Stephen T., Ph.D., U. o f Michigan. and Evolutionary Biology Jacob Gould Schurman Professor, Neurobiology and Behavior 186 BIOLOGICAL SCIENCES - 2 8

Other Teaching Personnel Wang, Haiyang, Adjunct Asst. Prof., Boyce Kennedy, Kenneth A. R., Ph.D., U. of Blankenship, James E., M.S., Cornell U. Sr. Thompson Inst./Plant Biology California, Berkeley. Prof. Emeritus, Ecology Lee., Molecular Biology and Genetics and Evolutionary Biology Calvo, Rita A., Ph.D., Cornell U. Sr. Lee., College of Arts and Sciences Lee, Siu Sylvia, Ph.D., Bailor Coll, o f Medicine. Molecular Biology and Genetics Adkins-Regan, Elizabeth, Ph.D., U. o f Asst. Prof., Molecular Biology and Genetics Chen, Kuei-chiu, Ph.D., N ew York U. Lee., Pennsylvania. Prof., Neurobiology and Leonard, Samuel L., Ph.D., U. o f Wisconsin. Neurobiology and Behavior Behavior/Psychology Prof. Emeritus, Molecular Biology and Ely, Susan, Ph.D., Tufts U. Lee., Molecular Aquadro, Charles F., Ph.D., U. o f Georgia. Genetics Biology and Genetics Prof., Molecular Biology and Genetics/ Linster, Christine, Ph.D., Pierre and Marie Hester, Laurel, Ph.D., U. Michigan. Lee., Ecology and Evolutionary Biology Curie U. Assoc. Prof., Neurobiology and Neurobiology and Behavior Bass, Andrew H., Ph.D., U. o f Michigan. Prof., Behavior S H l Lorr, Nancy, Ph.D., U. o f Oregon. Lee., Neurobiology and Behavior Liu, Jun Kelly, Ph.D., Cornell U. Asst. Prof., BBS Physiology Blackler, Antonie W., Ph.D., U. o f London Molecular Biology and Genetics N £ | McGuire, Betty A.. Ph.D., U. o f Massachusetts. (England). Prof., Molecular Biology and McCobb, David, Ph.D., U. o f Iowa. Assoc. Sr. Lee., Ecology and Evolutionary Biology Genetics Prof., Neurobiology and Behavior Merkel, Susan, M.S., Cornell U. Sr. Lee., Booker, Ronald, Ph.D., Princeton U. Assoc. MacDonald, June M. Fessenden, Ph.D., Tufts Microbiology Prof., Neurobiology and Behavior U. Assoc. Prof. Emeritus, Molecular Biology Nivison, Helen T., Ph.D., U. o f California, Bretscher, Anthony P., Ph.D., Leeds U. and Genetics/Program on Science, Davis. Lee., Molecular Biology and Genetics (England). Prof., Molecular Biology and Technology, and Society Rehkugler, Carole M., M.S., Cornell U. Sr. Lee., Genetics Nicholson, Linda, Ph.D., Florida State U. Microbiology Brown, William J., Ph.D., U. o f Texas Health Assoc. Prof., Molecular Biology and Silva, Thomas, Ph.D., Cornell U. Sr. Lee., Plant Science Center, Dallas. Prof., Molecular Genetics Biology Biology and Genetics Power, Alison G., Ph.D., U. o f Washington. Southard, Laurel E., M.S., Tulane U. Lee., Clark, Andrew G., Ph.D., Stanford U. Prof., Prof., Ecology and Evolutionary Biology/ Undergraduate Biology Molecular Biology and Genetics/Ecology Science and Technology Studies and Evolutionary Biology Provine, William B., Ph.D., U. o f Chicago. Joint Appointees Chuang, Huai-hu, Ph.D., U. o f California. Asst. Charles A. Alexander Professor o f Biological Baldwin, Ian T., Adjunct Prof., Max Planck Prof., Molecular Physiology Sciences, Ecology and Evolutionary Inst, for Chemical Ecology/Ecology and Deitcher, David, Ph.D., Harvard Medical Biology/History Evolutionary Biology School. Assoc. Prof., Neurobiology and Schimenti, John C., Ph.D., U. o f Cincinnati. Bloom, Stephen E., Prof., Veterinary/ Behavior Prof., Molecular Biology and Genetics Microbiology and Immunology Ellner, Stephen P., Ph.D., Cornell U. Prof., Seeley, Thomas D., Ph.D., Harvard U. Prof., Bradbury, Jack, Prof., Neurobiology and Ecology and Evolutionary Biology Neurobiology and Behavior Behavior/Library o f Natural Sounds Feigenson, Gerald W., Ph.D., California Inst, o f Shaw, Kerry L., Ph.D., Washington U., St. Brutnell, Thomas, Prof., Plant Breeding/Plant Technology. Prof., Molecular Biology and Louis. Prof., Neurobiology and Behavior Biology Genetics Sherman, Paul W., Ph.D., U. o f Michigan. Prof., Doyle, Jeffrey J., Prof., Plant Biology (Bailey Fetcho, Joseph R., Ph.D., U. o f Michigan. Prof., Neurobiology and Behavior Hortorium)/Plant Biology General Neurobiology and Behavior Sparks, Jed P., Ph.D., Washington State U. Asst. Foote, Robert H., Jacob Gould Schurman Prof. Finlay, Barbara, Ph.D., Massachusetts Inst, o f Prof., Ecology and Evolutionary Biology Emeritus, Animal Science/Physiology Technology. Prof., Psychology Tumbar, Tudorita, Ph.D., U. o f Illinois, Urbana- Giovannoni, James G., Adjunct Asst. Prof., Garcia Garcia, Maria J., Ph.D., U. Autonoma Champaign. Asst. Prof., Molecular Biology USDA Science and Education de Madrid Spain. Asst. Prof., Molecular and Genetics Administration/Plant Biology Biology and Genetics Turgeon, Robert, Ph.D., Carleton U. (Canada). Hanson, Maureen, Prof., Molecular Biology Geber, Monica A., Ph.D., U. o f Utah. Assoc. Prof., Plant Biology and Genetics/Plant Biology Prof., Ecology and Evolutionary Biology Wallace, Bruce, Ph.D., Columbia U. Prof. Hrazdina, Geza, Prof., Food Science and Gibson, Quentin H., Ph.D./D.Sc., Queen’s U. Emeritus, Molecular Biology and Genetics Technology Geneva/Plant Biology Ithaca (Northern Ireland). Greater Philadelphia Wilson, David B., Ph.D., Stanford U. Prof., Jahn, Margaret M., Assoc. Prof., Plant Professor Emeritus in Biological Sciences, Biochemistry, Molecular Biology and Breeding/Plant Biology Molecular Biology and Genetics Genetics Jander, Georg, Adjunct Asst. Prof., Boyce Goodale, Christine L., Ph.D., U. o f New Wolfner, Mariana F., Ph.D., Stanford U. Prof., Thompson Inst./Plant Biology Hampshire. Asst. Prof., Ecology and Molecular Biology and Genetics Kochian, Leon V., Adjunct Prof., USDA Science Evolutionary Biology Zamudio, Kelly R., Ph.D., U. o f Washington. and Education Administration/Plant Biology Greene, Harry W., Ph.D., U. o f Tennessee. Assoc. Prof., Ecology and Evolutionary Korf, Richard P., Prof. Emeritus, Plant Prof., Ecology and Evolutionary Biology Biology Pathology/Plant Biology (Bailey Hortorium) Hairston, Nelson G., Jr., Ph.D., U. o f Kresovich, Stephen, Prof., Plant Breeding/Plant Washington. Frank H. T. Rhodes Professor Other Teaching Personnel Biology of Environmental Science, Ecology and Balko, Elizabeth A., Ph.D., College o f Liebherr, James K., Assoc. Prof., Entomology/ Evolutionary Biology Environmental Science and Forestry. Sr. Plant Biology (Bailey Hortorium) Halpern, Bruce P., Ph.D., Brown U. Prof., Lee., Plant Biology McClure, Polley A., Prof., Information Neurobiology and Behavior/Psychology Johnson, Bruce R., Ph.D., Boston U. Sr. Lee., Technologies/Ecology and Evolutionary Heppel, Leon A., Ph.D., U. o f California, Neurobiology and Behavior Biology Berkeley. Prof. Emeritus, Molecular Biology Land, Bruce R., Ph,.D., Cornell U., Sr. Lee., McCouch, Susan R., Assoc. Prof., Plant and Genetics Neurobiology and Behavior Hess, George P., Ph.D., U. o f California, Breeding/Plant Biology Joint Appointees Pimentel, David, Prof. Emeritus, Entomology/ Berkeley. Prof., Molecular Biology and Levin, Simon A., Adjunct Prof., Princeton U./ Ecology and Evolutionary Biology Genetics Ecology and Evolutionary Biology Rossman, Michael J., Adjunct Prof., Purdue U./ Hinkle, Peter C., Ph.D., N ew York U. Prof., Likens, Gene E., Adjunct Prof., Institute o f Molecular Biology and Genetics Molecular Biology and Genetics Ecosystem Studies/Ecology and Stem, David B., Adjunct Prof., Boyce Howland, Howard C., Ph.D., Cornell U. Prof., Evolutionary Biology Thompson Institute/Plant Biology Neurobiology and Behavior/Biomedical Tanksley, Steven, Prof., Plant Breeding/Liberty Sciences College of Veterinary Medicine Hyde Bailey Prof., Plant Biology Hoy, Ronald R., Ph.D., Stanford U. Merksamer Alcaraz, Ana, D.V.M., Ph.D., Cornell U. Lee., Thaler, Jennifer S., Assoc. Prof., Entomology/ Prof., Neurobiology and Behavior Biomedical Sciences Ecology and Evolutionary Biology Huffaker, Tim C., Ph.D., Massachusetts Inst, of Beyenbach, Klaus W., Ph.D., Washington State Thompson, John F., Adjunct Prof., USDA Technology. Prof., Molecular Biology and U. Prof., Biomedical Sciences Science and Education Administration/Plant Genetics Bezuidenhout, Abraham, D.V.Sc., U. o f Biology Ke, Ailong, Ph.D., Johns Hopkins U. School o f Pretoria (South Africa), Sr. Lee., Biomedical Vehrencamp, Sandra, Prof., Neurobiology and Medicine. Asst. Prof., Molecular Biology and Sciences Behavior/Library o f Natural Sounds Genetics Buckles, Elizabeth, D.V.M., Ph.D., U. o f Weiss, Robert S., Ph.D., Baylor Coll, o f Wisconsin. Asst. Prof., Pathology Medicine. Asst. Prof., Biomedical Sciences Catalfamo, James, M.S., Ph.D., Union Coll. Sr. Wootton, John F., M.S., Ph.D., Cornell U. Prof., Res. Assoc., Population Medicine and Biomedical Sciences Diagnostic Services Xin, Hong-Bo, Ph.D., Beijing Medical U. Chuang, Huai-hu, Ph.D., U. o f California. Asst. (People’s Republic o f China). Asst. Prof., Prof., Molecular Physiology Biomedical Sciences Cohen, Paula, Ph.D., U. of London (England). Yen, Andrew, Ph.D., Cornell U. Prof, Asst. Prof., Genetics Pathology and Director, Graduate Studies in Famum, Cornelia E., D.V.M., Ph.D., U. of Environmental Toxicology, Biomedical Wisconsin, Madison. Prof., Biomedical Sciences Sciences FitzMaurice, Mamie C., V.M.D., Ph.D., U. o f College of Engineering Pennsylvania. Instr., Biomedical Sciences Fortune, Joanne E., Ph.D., Cornell. U. Prof., Joint Appointees Biomedical Sciences Cisne, John L., Assoc. Prof., Geological Gilmour, Robert F., Ph.D., SUNY Upstate Sciences/Biological Sciences Medical Center. Prof., Biomedical Sciences Webb, Watt W., Prof., Applied and Engineering Gleed, Robin, BVSc, MRCVS, U. o f Liverpool Physics/Biological Sciences (England). Assoc. Prof., Clinical Sciences Gunn, Teresa M., Ph.D., U. o f British Columbia Biological Sciences (Canada). Asst. Prof., Biomedical Sciences Hermanson, John W., M.S., Ph.D., U. o f Florida Joint Appointees Gainesville. Assoc. Prof., Biomedical Snedeker, Suzanne M., Asst. Prof., Center for Sciences the Environment/Biological Sciences Houpt, Katherine A., V.M.D., Ph.D., U. o f Pennsylvania. Prof., Clinical Sciences Division of Nutritional Sciences Kotlikoff, Michael I., Ph.D., U. o f California, Joint Appointees Davis. Prof., Biomedical Sciences Lin, David, Ph.D., U. o f California, Berkeley. Arion, William J., Prof., Nutritional Sciences/ Asst. Prof., Biomedical Sciences Molecular Biology and Genetics Loew, Ellis R., Ph.D., U. o f California, Los Bensadoun, Andre, Prof., Nutritional Sciences/ Angeles. Prof., Biomedical Sciences Physiology Lorr, Nancy, Ph.D., U. o f Oregon. Lee., Kazarinoff, Michael N., Assoc. Prof., Nutritional Biomedical Sciences Sciences/Molecular Biology and Genetics Ludders, John, D.V.M., Washington State U. Wright, Lemuel D., Prof. Emeritus, Nutritional Prof., Clinical Sciences Sciences/Molecular Biology and Genetics Maza, Paul, D.V.M., U. o f St. Kitts (West Indies). Lee., Biomedical Sciences •Joint appointment with College o f Arts and McDonough, Sean, D.V.M., Ph.D., U. of Sciences Pennsylvania. Assoc. Prof., Biomedical fjoint appointment with College of Veterinary Sciences Medicine Meyers-Wallen, Vicki, V.M.D., Ph.D., U. o f fjoint appointment with College of Agriculture Pennsylvania. Assoc. Prof., Biomedical and Life Sciences Sciences §Joint appointment with College of Minor, Ronald. V.M.D., Ph.D., U. o f Engineering Pennsylvania. Prof., Biomedical Sciences Mizer, Linda, D.V.M., Ph.D., Ohio State U. Sr. Lee., Biomedical Sciences Nikitin, Alexander Yu, M.D., Ph.D., Petrov Research Inst, o f Oncology (Russia). Asst. Prof, o f Pathology, Biomedical Sciences Njaa, Bradley, L., D.V.M., MVSc, U. o f Saskatchewan. Asst. Prof., Biomedical Sciences Noden, Drew M., Ph.D., Washington U. (St. Louis). Prof., Biomedical Sciences O ’Brian, Timothy, Ph.D., U. o f Illinois. Prof., Biomedical Sciences Oswald, Robert, Ph.D., Vanderbilt U. Prof., Molecular Medicine Quaroni, Andrea, Ph.D., U. o f Pavia (Italy). Prof., Biomedical Sciences Rawson, Richard E., D.V.M., Ph.D., U. o f Minnesota. Sr. Lee., Biomedical Sciences Roberson, Mark, Ph.D., U. o f Nebraska. Assoc. Prof., Biomedical Sciences Schimenti, John, Ph.D., U. o f Cincinnati. Prof., Biomedical Sciences Schlafer, Donald H., D.V.M., Ph.D., U. o f Georgia. Prof., Biomedical Sciences Shepard, Laura, D.V.M., Cornell U. Instr., Biomedical Sciences Suarez, Susan, Ph.D., U. o f Virginia. Prof., Biomedical Sciences Summers, Brian, B.V.Sc., Ph.D., Prof., Biomedical Sciences Travis, Alexander J., V.M.D., Ph.D., U. of Pennsylvania. Asst. Prof., Biomedical Sciences