No.40 Summer 2013 LASTHENIA NEWSLETTER OF THE DAVIS BOTANICAL SOCIETY

FIELD PROJECTS KEEP THE HERBARIUM BUSY AND FUNDED!

This past year (2013) has been a good We started the Clear Lake State Park one for getting outside into the field, project in January, with a preliminary away from our indoor herbarium visit with the park’s Environmental database projects. In the spring, we of- Scientist, Jim Dempsey, who showed us fered to work (gratis) on a preliminary convenient access routes and pro- list for Clear Lake State Park, vided maps. An undergraduate, Karen which is on the north slopes of Mount Whitestone, came with us to begin an Konocti in Lake County, about two undergraduate research project to map hours (by car) northwest of Davis. We one or more rare in the park. have worked on a number of plant lists Karen and I returned twice in February for the Dept. of Parks and to continue mapping the rare Konocti Recreation since 2006. The department manzanita (Arctostaphylos manzanita doesn’t have much of a budget for this ssp. elegans); Karen presented her find- Ellen Dean and Karen Whitestone at Clear sort of thing, and so they often need a ings on this plant at an undergraduate Lake State Park (with Clear Lake in the back- plant list, and their properties are often research symposium in April. ground). Photo: J. Dempsey a convenient place to take students to In March, I traveled to the park complete their spring plant collection. with a group of Davis Botanical Society volunteers and Collection Manager Jean Shepard for a general collecting expedi- tion. Then I took two more trips in April THE CONSERVATORY’S PLANT LADY and May with 16 UC Davis students (15 interns and one volunteer, Marisol The Botanical Conservatory’s Staff Gonzalez). Horticulturalist, Marlene Simon, has By the end of the quarter, we had col- become locally famous! In September lected nearly 240 specimens in the park 2013 Marlene began a regular stint on and documented three rare plants. With the television show Good Day, Sacra- the help of herbarium volunteer Kate mento as The Plant Lady, the show’s Mawdsley and undergraduate Nathan resident horticultural advisor. She has Gonzalez, we combined our collection been a regular addition to the show Sat- urday mornings between 7 am and 11 continued on page 4 am, answering viewer questions about problems with indoor and outdoor cul- tivated plants. She is an articulate plant IN THIS ISSUE expert, able to field many types of plant Field Projects...... 1 questions, even when she isn’t given the questions ahead of time. If you want to find previous episodes of The Plant The Plant Lady...... 1 Lady on the internet, here are two links you can try. Hopefully, they will still Society Profiles...... 3 work by the time this issue of Lasthenia is in your hands. Field Trips...... 5 http://gooddaysacramento.cbslocal.com/video/9209962-plant-questions- answered/ The Cedars...... 6 http://gooddaysacramento.cbslocal.com/video/9234553-ask-the-plant-lady- Student Grants...... 7 pt-3/ 1 CONSERVATORY UPDATE Where does the money go? If you’ve ever wondered how the annual Conservatory Wish List allocation from Davis Botanical Society member dues, interest from the Label-Making Needs: Conservatory Endowment, and gifts to Software ($500) current operations funds have helped Desktop Computer ($2000) the Botanical Conservatory, read on. Printer ($1000) And while you are doing so, keep in mind that the Conservatory has new BOG Project Needs: curatorial needs that are explained at Soil amendments ($4000) the end of this article and summarized And more BOG materials….. in the wish list to the right. ($4000)

square footage, the multi-level benches The new bench arrangement in the succulent have brought our plants further into room. Photo: E. Sandoval the third dimension and lower, so more of our visitors, especially the young, the need and their ability to build the can see more of the plants and less of solutions. Ian was also supported by the floor. I am no longer afraid visitors Gifford Cycad Garden donations, and will succumb to claustrophobia! We many visitors have noted his skilled have also used some of this funding for pruning of the plants around Storer Hall increased student staff to transplant, and the Sciences Laboratory Building. prune, propagate, and maintain the The juniper bushes at the west end of collection. Storer are now interesting sculptures Two of our skilled student rather than view blocking blobs! The succulent room in spring 2013. The walls between the old rooms are long gone employees, Federico Lopez Borghesi, What’s next? We’ve completed about and the benches are almost finished. a recent Plant Biology undergrad, 85% of the grading and recontouring of Photo: E. Sandoval and Ian D. R. Baker, International Ag. an area northwest of the Conservatory, Development, were paid in part by DBS- slated to become the Biological Orchard The Conservatory has seen a originated funds to accomplish major and Gardens (BOG, for short). This steady 6% increase in the number of overhauls of the benches in our cool project will include a large area of fruit tours and participants over the last fern rooms and what used to be two and berry trees and bushes, as well as couple of years, and we expect more desert rooms. The improvements were areas for Mediterranean and California as enrollment across campus increases. a collaboration between what I saw as native plants. This past year we led more than 4,100 Needed soon are additional irrigation individuals on 157 tours, guided mostly equipment and soil amendment by Marlene Simon, staff horticulturalist, materials, with an estimated cost and myself. (I could interject here of $4,000 to $7,000. We hope to that we would be ecstatic to find some be able to plant in late 2014, if the major donations for a new and bigger infrastructure is complete. Dan Isidor, Conservatory, but in the meantime... ). in the development office of the College In order to accommodate this growth, of Biological Sciences, has helped especially as a result of increased with funding for the BOG, but more is enrollment in Intro Biology 2C, we’ve needed. used the funding sources listed earlier Inside the Conservatory, a new to make a variety of improvements software package to produce plant to the ways plants are displayed and labels, with the computer and printer to changes to the bench layouts in and use it, is at the top of the wish list. I’d be around our growing facilities. happy to talk with anyone who would One of the biggest improvements like more information about either of inside the Conservatory occurred in these projects. You can contact me at our two succulent rooms, now one [email protected] or 530-752- continuous room since we removed 0569. the former dividing wall. The aisles are now wider and traffic flow is much E. Sandoval & K. Mawdsley improved thanks to the replacement Interns Marisol Gonzalez and Surbhi Cho- of the wooden benches with tiered phla enjoying displaying an black plastic benches in a figure eight. Amorphophallus corm, Spring, 2013. Although we’ve lost some bench top Photo: E. Sandoval 2 RECENT GIFTS Conservatory Endowment Herbarium Operations Davis Botanical Society Student Eric Conn Bureau of Land Management Grants Fund Ernesto Sandoval California Native Plant Society Michael Barbour Maxine Schmalenberger Friends of the Davis Library & Valerie Whitworth Katherine Mawdsley Michelle Barefoot Herbarium Endowment In Memory of James Neilson: & Luis Perez-Grau Chris Bronny Harold & Roberta Bacheller Eric Conn Mick Canevari Griffin Greenhouse Supplies, Inc. Brenda Grewell & Steve Kidner Eric Conn William & Jean Heflin E. Eric Grissell Joseph DiTomaso Dianne McQuaid Marie Jasieniuk & Frank Roe Lewis Feldman James & Rosella McQuaid Terence & Judith Murphy Brenda Grewell Henri & Dianne Pellissier Thomas & Ann Rost Gordon & Delia Harrington Maxine Schmalenberger Charles, Jessica, & Henry Hughes Conservatory Gifts in Kind Roger Willmarth Charlotte Kimball Maurice Levin Julie Knorr Marjorie March (in memory of June Herbarium Gifts in Kind McCaskill) Gerald Dickinson Sue Nichol Marcel Rejmanek Robert Preston Estate of Barbara & John Hopper Maxine Schmalenberger John Randall Kenneth and Shirley Tucker Roger Willmarth Carol Witham Thank you for Conservatory Operations Kelly Ratliff your support!

SOCIETY PROFILES Marie Jasieniuk

Marie Jasieniuk, 2013-14 Davis forests of eastern Canada to study Botanical Society president, now Oxalis montana for her doctorate. studies weeds in agroecosystems, Post-doctoral opportunities led her especially invasive plants of first to Manitoba and evolutionary and horticultural origin, but she has genetic studies of herbicide resistance roots, so to speak, in herbaria. As in weeds and then south to Montana an undergraduate at the University State University in Bozeman, where she of Saskatchewan, the herbarium worked on modeling crop yield losses hired her for one of those life- and the ecology of invasive plants in changing summer jobs: she and natural areas.. While recounting this another student took a camper with history, Marie gave a most nuanced a canoe on top and followed the reply to a request for her definition crew building a road into the boreal of a “weed:” it is, she said, “context forest, collecting and documenting dependent.” the plants along the route for the Marie came to UC Davis in 2002, to herbarium’s collection. Her interest what is now the Dept. of Plant Sciences. in plants had begun even earlier as She co-teaches the graduate core course she botanized on her parents’ farm in in Plant Biology as well as Science and Saskatchewan. Society 12, Plants and Society 12, and Marie Jasieniuk in the field. Marie’s master’s degree research Bio Sci 2B, which introduces students to Photo courtesy of the subject. in plant community ecology took her the principles of ecology and evolution. into the Canadian subarctic to study She is an enthusiastic proponent of We’re delighted to welcome her back revegetation after forest fires. Then the Davis Botanical Society’s student to a campus herbarium, even though she moved on to McGill University research grants, believing strongly in her duties as DBS president don’t come in Montreal and hiked the deciduous the training of botanists for the future. with a camper and canoe. K. Mawdsley

3 HERBARIUM FIELD PROJECTS (CONT. FROM PAGE 1)

tion, I had help at the beginning of the project from alumna Allyson Ayalon. My wonderful husband, Tom Starbuck, went out with me numerous Saturdays (and the 4th of July) to help finish up various sites. Since Tom and I argue a lot about almost everything, I am afraid that we were rather noisy at the Stevenson’s Bridge site early in the morning on the 4th of July. Nearby residents must have wondered what the heck was happening. I loved doing the Putah Creek monitoring, because I was able to revisit Volunteer plant collectors at Clear Lake State a habitat (and make collections for the General Creek at Sugar Pine Point State Park with lake in background, March 2013. herbarium) in an area that has been Park. Photo: M. Starbuck Photo: Petra Unger sparsely visited by our herbarium’s bota- data with two older park plant lists nists for the past few decades. We have flora of the wetland areas of Sugar Pine and online data from the Consortium numerous collections made by Kather- Point State Park in the Tahoe basin. We of California Herbaria, updating all the ine Esau, one of our early herbarium di- made two field trips to the park this plant names to current scientific names. rectors, in Putah Canyon or along Putah past summer – one in July and one in The 13-page preliminary list is nearly Creek in the 1920s; Putah Canyon is an August. Kate Mawdsley came to help on completed, but there are still quite a few area close to where the dam now stands both trips, and Tom Starbuck and my unexplored areas to visit and plants to and just to the east. daughter, Margaret Starbuck, came in document at the park, and we will con- August. Students Kaya MacMillen and tinue with the project next spring with a Nathan Gonzalez accompanied us on new set of student interns. the July trip. Our contact at the park, In May we had to give up visiting Svetlana Yegorova, accompanied us on Clear Lake State Park and move on to a both outings to become familiar with more local (and funded!) project: veg- our methods. After four visits, we have etation monitoring along Putah Creek. now collected ca. 450 specimens at Davisites will be aware of a local non- Sugar Pine Point State Park, document- profit organization, the Putah Creek ing some interesting finds including Council, which has obtained funding three-way sedge (Dulichium arundina- for restoring and monitoring various ceum), buckbean (Menyanthes trifoliata), aspects of the biology of Putah Creek. the rarely collected meesia moss (Meesia The creek was dammed in the 1950s to longiseta), as well as several rare plants. create Lake Berryessa, which is located This project, which takes some agility west of Davis and provides water for California broomrape along Putah Creek. (to walk up creek beds, climb over logs, Solano County. Once the creek leaves Photo: Kaya MacMillen and wade through water), will continue Lake Berryessa, most of its water goes in 2014, if you want to join us. into the Solano Diversion Canal; the Like much of the Central Valley most remnant of the natural creek runs east of the current herbaceous flora along the E. Dean toward Davis and then south of Davis to creek is nonnative, with woody plants the Yolo Bypass, supplying agricultural providing most of the native cover. areas on its north and south sides. However, there were excellent native The Putah Creek Council has surprises all along the creek, includ- contracted with the UC Davis Museum ing California broomrape (Orobanche of Wildlife and Fish Biology to do californica) close to the bypass. Another biological monitoring, and they have surprise (besides the number of naked been monitoring the birds and vegeta- men along the creek) was the amount tion for nearly 10 years. This spring we of feral pecan growing along the creek; were awarded the vegetation monitoring no previous herbarium specimens have contract, which required revisiting ca. been collected to document this species 300 plots distributed from near the dam growing wild in Yolo County. One major to near the Yolo Bypass. disappointment was not seeing a river I hired several wonderful students to otter the entire four months. Ellen was help me with field work: Kaya Mac- sad. Millan, Maddy Taylor, Arti Lal, and Our final field project of 2013 is one Nathan Gonzalez (the last two are also we are continuing from the previous Three-way sedge (Dulichium arundinaceum). herbarium student assistants). In addi- two years (but now with funding!) – a Photo: M. Starbuck 4 SPRING 2013 FIELD TRIPS TAKE US TO LOCAL GEMS

The Sutter Buttes are billed as the small- est mountain range in the world, rising ca. 2100 feet above the Sacramento Val- ley just northwest of Yuba City in Sutter County. The Buttes are the remains of an extinct Pleistocene-era volcano, and the oak woodland landscape is littered with huge volcanic boulders, reminis- cent of a different place and time. We enjoyed a day of hiking with Pete, who also gave us permission to collect specimens for the herbarium; we collected 70 species, adding one new species to their plant list. Marcel Wild hyacinth (Dichelostemma multiflorum). The view of the Central Valley from Sunrise was able to cross one more place off his Photo: M. Starbuck Ridge, Sutter Buttes. Photo: M. Starbuck list of “must see” places and enjoyed seeing how the vegetation differed from Participants were able to view a number Although we had just one Davis Botani- that of the Sierra Nevada and Coast of rare plants, including Cobb Moun- cal Society spring field trip scheduled Range foothills; he found the vegeta- tain lupine (Lupinus sericatus), nodding on last year’s calendar, we ended up tion similar but missing certain families, harmonia (Harmonia nutans), Napa with two! In the fall of 2012, after our genera, and species. Everyone enjoyed false indigo (Amorpha californica var. calendar for the year had been set and the extra-large flowers of baby-blue-eyes napensis), serpentine collomia (Collo- sent, Marcel Rejmanek, then our board’s (Nemophila menziesii), the beautiful and mia diversifolia), two-carpellate western Past-President said “Let’s go to the dramatic landscape and views, and be- flax Hesperolinon( bicarpellatum), green Sutter Buttes. I have never been there.” ing together in a special place. monardella (Monardella viridis), marsh He was not alone. A number of us had ( fontanum), not yet been to the Buttes, or we had and swamp larkspur (Delphinium uligi- not been there for decades. We knew nosum). that there would be quite a bit of inter- I first investigated this area in the est in this trip. It just so happened that spring of 2012, when I took UC Davis another member of our board, Member- students to the property to make their at-Large Valerie Layne, is a docent with plant collections (under close super- the Middle Mountain Foundation, a vision). We gave our plant list to the non-profit group that gives tours at the property managers, adding several new Sutter Buttes. Valerie worked with Mar- species to their plant list. This time cel and me to make the trip happen. around, we made another addition to On the first Saturday in April, 22 of their plant list – bull clover (Trifolium us carpooled up the Sacramento Valley fucatum). It was a glorious day, and the serpentine plants were in full, profuse to Yuba City to meet our docents from Poppy-filled slope near Willaston Flat, Sutter the Middle Mountain Foundation and Buttes. Photo: M. Starbuck bloom, giving us the impression we to get directions to the Dean Ranch, were walking in a botanical garden. a private land holding on the Sutter E. Dean Buttes owned by Margit and Pete Sands. Our scheduled field trip to Las Posa- das State Forest in Napa County took place on April 27. I led ten Botanical So- ciety members down Hwy 128 to Napa County, through the beautiful Chiles and Pope Valleys, and then up and over a redwood-shaded ridge to the town of Angwin. Las Posadas is just southwest of Angwin along Las Posadas Rd., and it really is a remarkable botanical hotpot. Typical of Napa County, within a rel- atively small area you can see numerous habitats, including coast redwood forest, Douglas fir forest, oak/pine woodland (with several species of oaks), grassland Pete Sands and Ellen Dean identifying wild- (with shallow volcanic vernal pools), Ellen Dean with UC Davis students collect- flowers on Sunrise Ridge, Sutter Buttes. and chaparral; the soil types include ing at Las Posadas State Forest, April 2012. Photo: M. Starbuck volcanic, sedimentary, and serpentine. Photo: Claire Loughran 5 THE CEDARS - THE MOST SPECTACULAR SERPENTINE SITE IN CALIFORNIA?????

environments, and (so it’s speculated) perhaps on other planets where peridot- ite and water occur. Our knowledge of The Cedars owes a great deal to one person, Roger Raiche, who fell in love with the site in 1981. His many years of botanical ex- ploration and advocacy were rewarded in 2011 when the central part of the site was acquired with conservation funds and transferred to the Bureau of Land Management. Access is by a private road and is therefore generally only possible on organized trips. The Davis Botanical Society’s May 2014 visit should be ideal for seeing the endemic Calochortus, among other delights. For more information, see Roger’s Fremontia article: http://www.cnps.org/cnps/publications/ fremontia/Fremontia_Vol37-No2.pdf

Susan Harrison

View of Sargent’s Cypress growing on the cliffs of The Cedars. Photo: S. Harrison Editor’s Note: Susan Harrison, Professor of Environmental Science and Policy, is our new The Cedars in western Sonoma County Member-at-Large on the Davis Botancial Society Board. She will lead the May 2014 is a 7500-acre island of raw serpentine field trip to The Cedars. Susan was named a and peridotite rock in a sea of coastal Fellow of the Ecological Society of America mixed evergreen forest. Its angular cliffs in 2013. and gnarled Sargent’s Cypress forests combine with coastal fog to give it the look of classical Japanese landscape painting. The Cedars may be the single most botanically important serpentine out- crop in California. It is the unique home (the only location of) a fairy lantern (Calochortus raichei), buckwheat (Er- iogonum cedrorum), and creambush (Ho- Lady Slipper Orchid. Photo: S. Harrison lodiscus dumosus ssp. cedrorum), as well as the “Serpentine Night” giant stream orchid beloved of Arboretum sale-goers Geological features of The Cedars (Epipactis gigantea form rubrifolia). It are as significant as its botanical wealth. is one of very few locales for a fleabane Seeps emerge along its rocky stream (Erigeron serpentinus), manzanita (Arcto- channels with a strange chemistry that staphylos bakeri ssp. sublaevis), jewel- includes high pH, high calcium, and flower Streptanthus( glandulosus ssp. a whiff of hydrocarbons, and these hoffmanii), and other serpentine rarities. oozings produce elaborately layered Other species reach their southern range calcium carbonate deposits known as limits at The Cedars, including white- the Wedding Cake and the Mineral leaf manzanita (Arctostaphylos viscida), Falls. Traces of DNA extracted from cotton grass (Eriophorum criniger), these mysterious waters provide clues The Cedars Wedding Cake formation. and lady-slipper orchid (Cypripedium to the identities of microbes that can Photo: S. Harrison californicum). live in some of the earth’s more extreme 6 2013 STUDENT GRANT RECIPIENTS The Davis Botanical Society awarded The final two grants are for a record five student research grants projects on more familiar territory. for 2013-14, a tribute both to the Kyle Christie, a doctoral student in the excellence of the projects proposed by Population Biology graduate group, UC Davis students and the generosity will use the Jack Major award to of DBS members and other donors investigate modes and mechanisms of who make the program possible. The speciation in the California flora using projects demonstrate the diversity of mountain jewelflower Streptanthus( plant-related research on the campus, tortuosus) and several near relatives as as well as an impressive range of field study subjects. He will use genomic locations. techniques to test hypotheses that Javier Jauregui Lazo, a Master’s then tap into a botanical observation different speciation mechanisms have student in the Horticulture and network across the nation to study the operated for individual species in the Agronomy graduate group, will use extent of its spread since introduction jewelflower group, acquiring tissue the Eric Grissell award to conduct about 50 years ago. Field experiments samples and seed (as well, of course, phylogenetic, biogeographic, and at the park will test several hypotheses as herbarium specimens) from across taxonomic analyses of Acaena, in to explain the success of the invasion, the geographic and environmental the Rose family. California has only including reduced insect herbivory, range of each species. one native species of Acaena, but increased animal-mediated seed Elaine Chow’s project will there are many in South America, dispersal, and allelopathy. study competition between a rare Australia, New Zealand, and on Albania is the final exotic locale for perennial endemic, Eureka Valley subantarctic islands—all parts of this year’s awardees: Courtney Jallo, evening-primrose, and an invasive ancient Gondwana. Javier will collect who spent three years there as a Peace annual, Russian thistle, in Death and analyze material from most of the Corps volunteer, will return for an Valley National Park to understand species from Chile and New Zealand ethnobotanical study of comparative the mechanisms underlying species to test the hypothesis that current traditional knowledge of medicinal and interactions. Her early studies species result from multiple migration aromatic plants in Communist and post- suggest that Russian thistle is an events over time from southern Communist generations. Albanian herbs “opportunistic invader” rather than South America to Australia and New such as sage, oregano, and thyme are a strong competitor in Death Valley; Zealand. an important export product because it occurs in much greater densities Christopher Dillis, a student of their quality and unique subspecies, along disturbed roadsides than in in the Animal Behavior Graduate but development pressure may threaten wilderness plots less than 20 meters group, is using a Davis Botanical their habitats if younger generations do from the road. Chow will conduct Society research award to study not recognize their value. Jallo used her ex situ experimental plantings of the the distribution and ecology of the Davis Botanical Society research award two species over two years to test the invasive neotropical tree Bellucia to travel to Albania during summer effect of factors such as herbivory pentamera in Indonesia. Limited to 2013 to conduct interviews during on the growth of evening-primrose disturbed forest patches in its native harvest season for the plants in four in competition with Russian thistle. Latin America, the plant is spreading regions of the country. She will prepare The Larry Mitich award is helping to in shaded primary forest as an a paper for publication as part of her support her dissertation research. invader. He will map the tree’s current Master’s degree program in International Congratulations to all the 2013-14 distribution in detail at Ganung Agricultural Development. She award recipients. We look forward to Palung National Park to record its anticipates the research may contribute hearing the results of the studies at a ecological diversity in Indonesia and to conservations efforts in both future Fall program meeting! ecological and anthropological fields. K. Mawdsley OUR HERBARIUM PHOTOGRAPHER LEAVES FOR COLLEGE

This issue of Lasthenia marks the end of an era, as it is probably the last issue with photos by our student photographer, Margaret Starbuck. She has graduated from high school and deserted the herbarium for college! I have heavily depended on Margaret for photographs of our many field excursions over the past seven years. It has been a wonderful thing to have someone with a good eye snapping photos so that I could collect plants and/or lead a field trip. Thank you to the many Davis Botanical Society members who have encouraged Margaret in her school and ex- tracurricular activities. You have attended plays, choir concerts, art shows, donated money to drama troupes, and you took the time to talk with her on field trips. The herbarium baby is now all grown up. This photo is the only photo that she took of herself during last Spring’s Sutter Buttes field trip – looks like she was having fun. E. Dean 7 DAVIS BOTANICAL SOCIETY LASTHENIA LASTHENIA, the Newsletter of the Davis Botanical Society, is published in collaboration with the staff of the UPCOMING 2014 EVENTS UC Davis Botanical Conservatory and Center for Plant Diversity. Lichen Workshop - Saturday, January 11 Editor: Kate Mawdsley Issue Contributors: E. Dean, E. Sando- Botanical Tea in the Herbarium - Wednesday, January 22 val, K. Mawdsley, M. Starbuck, S. Harri- son, C. Loughran, A. Wight, M. Simon, J. Dempsey, P. Unger, K. MacMillen, M. Museum Day - Saturday, February 8 Jasieniuk Design: Susan Gloystein Field Trip to San Pedro Valley Park - Saturday, February 22 Layout: Ellen Dean DBS OFFICERS, 2013-2014 Horticultural Workshop - Saturday, March 1 President: Marie Jasieniuk President-elect: Brenda Grewell Membership Vice President: Picnic Day - Saturday, April 12 Pat McGuire & Kate Mawdsley Secretary: Marlene Simon Spring Meeting - Thursday, May 8 Treasurer: Robert Rhode Past President: Joe DiTomaso Field Trip to The Cedars - Saturday, May 17 Members at Large: Susan Harrison, Valerie Layne Student Member at Large: For more information, see http://herbarium.ucdavis.edu Jorge Perez Zabala or for horticultural workshop email [email protected] Ex officio: Dan Potter, Ernesto

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