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Solifugae, Eremobatidae)
1998. The Journal of Arachnology 26:113-116 RESEARCH NOTE THE EFFECTS OF REPRODUCTIVE STATUS ON SPRINT SPEED IN THE SOLIFUGE, EREMOBATES MARATHONI (SOLIFUGAE, EREMOBATIDAE) Costs associated with reproduction are de- gravid females of the solifuge Eremobates lineated by trade-offs between the current re- marathoni Muma 1970 . To my knowledge, no productive capacity of an animal and the prob- previous data on sprint speed or the relation- ability of its future survival and reproductive ship between sprint speed and reproductive success (Williams 1966) . The successful anal- status exist for the Solifugae . ysis of life history parameters depends on our Eremobates marathoni is a common inhab- ability to identify proximate mechanisms by itant of the Big Bend region of Trans Pecos which such costs are mediated . Documented Texas (Punzo 1997), which lies within the costs associated with reproduction include de- northern confines of the Chihuahuan Desert. I creased survivorship resulting from physio- collected gravid (G) and nongravid (NG) fe- logical or behavioral changes that accompany males by hand at night with the aid of a head reproduction (Hirshfield & Tinkle 1975 ; Bell lamp as they wandered over the surface of the 1980). For example, if escape from a predator ground, or through the use of pitfall traps as depends on the speed or endurance of a po- described previously (Punzo 1994a) . All so- tential prey organism, then any reduction in lifuges were collected within a 3 km radius of the locomotor performance of gravid females Marathon, Texas (Brewster County) during could increase the risk of predation . Loco- July 1996 . A detailed description of the ge- motor performance has been correlated with ology and dominant vegetation of this area is survivorship in many species of vertebrates given by Tinkam (1948) . -
Arachnida, Solifugae) with Special Focus on Functional Analyses and Phylogenetic Interpretations
HISTOLOGY AND ULTRASTRUCTURE OF SOLIFUGES Comparative studies of organ systems of solifuges (Arachnida, Solifugae) with special focus on functional analyses and phylogenetic interpretations HISTOLOGIE UND ULTRASTRUKTUR DER SOLIFUGEN Vergleichende Studien an Organsystemen der Solifugen (Arachnida, Solifugae) mit Schwerpunkt auf funktionellen Analysen und phylogenetischen Interpretationen I N A U G U R A L D I S S E R T A T I O N zur Erlangung des akademischen Grades doctor rerum naturalium (Dr. rer. nat.) an der Mathematisch-Naturwissenschaftlichen Fakultät der Ernst-Moritz-Arndt-Universität Greifswald vorgelegt von Anja Elisabeth Klann geboren am 28.November 1976 in Bremen Greifswald, den 04.06.2009 Dekan ........................................................................................................Prof. Dr. Klaus Fesser Prof. Dr. Dr. h.c. Gerd Alberti Erster Gutachter .......................................................................................... Zweiter Gutachter ........................................................................................Prof. Dr. Romano Dallai Tag der Promotion ........................................................................................15.09.2009 Content Summary ..........................................................................................1 Zusammenfassung ..........................................................................5 Acknowledgments ..........................................................................9 1. Introduction ............................................................................ -
Giant Whip Scorpion Mastigoproctus Giganteus Giganteus (Lucas, 1835) (Arachnida: Thelyphonida (=Uropygi): Thelyphonidae) 1 William H
EENY493 Giant Whip Scorpion Mastigoproctus giganteus giganteus (Lucas, 1835) (Arachnida: Thelyphonida (=Uropygi): Thelyphonidae) 1 William H. Kern and Ralph E. Mitchell2 Introduction shrimp can deliver to an unsuspecting finger during sorting of the shrimp from the by-catch. The only whip scorpion found in the United States is the giant whip scorpion, Mastigoproctus giganteus giganteus (Lucas). The giant whip scorpion is also known as the ‘vinegaroon’ or ‘grampus’ in some local regions where they occur. To encounter a giant whip scorpion for the first time can be an alarming experience! What seems like a miniature monster from a horror movie is really a fairly benign creature. While called a scorpion, this arachnid has neither the venom-filled stinger found in scorpions nor the venomous bite found in some spiders. One very distinct and curious feature of whip scorpions is its long thin caudal appendage, which is directly related to their common name “whip-scorpion.” The common name ‘vinegaroon’ is related to their ability to give off a spray of concentrated (85%) acetic acid from the base of the whip-like tail. This produces that tell-tale vinegar-like scent. The common name ‘grampus’ may be related to the mantis shrimp, also called the grampus. The mantis shrimp Figure 1. The giant whip scorpion or ‘vingaroon’, Mastigoproctus is a marine crustacean that can deliver a painful wound giganteus giganteus (Lucas). Credits: R. Mitchell, UF/IFAS with its mantis-like, raptorial front legs. Often captured with shrimp during coastal trawling, shrimpers dislike this creature because of the lightning fast slashing cut mantis 1. -
The Phylogeny of Fossil Whip Spiders Russell J
Garwood et al. BMC Evolutionary Biology (2017) 17:105 DOI 10.1186/s12862-017-0931-1 RESEARCH ARTICLE Open Access The phylogeny of fossil whip spiders Russell J. Garwood1,2*, Jason A. Dunlop3, Brian J. Knecht4 and Thomas A. Hegna4 Abstract Background: Arachnids are a highly successful group of land-dwelling arthropods. They are major contributors to modern terrestrial ecosystems, and have a deep evolutionary history. Whip spiders (Arachnida, Amblypygi), are one of the smaller arachnid orders with ca. 190 living species. Here we restudy one of the oldest fossil representatives of the group, Graeophonus anglicus Pocock, 1911 from the Late Carboniferous (Duckmantian, ca. 315 Ma) British Middle Coal Measures of the West Midlands, UK. Using X-ray microtomography, our principal aim was to resolve details of the limbs and mouthparts which would allow us to test whether this fossil belongs in the extant, relict family Paracharontidae; represented today by a single, blind species Paracharon caecus Hansen, 1921. Results: Tomography reveals several novel and significant character states for G. anglicus; most notably in the chelicerae, pedipalps and walking legs. These allowed it to be scored into a phylogenetic analysis together with the recently described Paracharonopsis cambayensis Engel & Grimaldi, 2014 from the Eocene (ca. 52 Ma) Cambay amber, and Kronocharon prendinii Engel & Grimaldi, 2014 from Cretaceous (ca. 99 Ma) Burmese amber. We recovered relationships of the form ((Graeophonus (Paracharonopsis + Paracharon)) + (Charinus (Stygophrynus (Kronocharon (Charon (Musicodamon + Paraphrynus)))))). This tree largely reflects Peter Weygoldt’s 1996 classification with its basic split into Paleoamblypygi and Euamblypygi lineages; we were able to score several of his characters for the first time in fossils. -
Nine New Species of the Eremobates Scaber Species Group of the North American Camel Spider Genus Eremobates (Solifugae, Eremobatidae)
Zootaxa 4178 (4): 503–520 ISSN 1175-5326 (print edition) http://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2016 Magnolia Press ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4178.4.3 http://zoobank.org/urn:lsid:zoobank.org:pub:20E72076-A0E4-427F-96D5-74A6DD4C9E06 Nine new species of the Eremobates scaber species group of the North American camel spider genus Eremobates (Solifugae, Eremobatidae) PAULA E. CUSHING1 & JACK O. BROOKHART2 Denver Museum of Nature and Science, 2001Colorado Blvd., Denver, CO 80205 USA. E-mail: [email protected]; [email protected] Abstract Nine new species of the Eremobates scaber species group of the solifuge genus Eremobates Banks 1900 are described, eight of them from Mexico. These new species are: E. axacoa, E. bonito, E. cyranoi, E. fisheri, E. hidalgoana, E. jalis- coana, E. minamoritaana, E. zacatecana, and E. zapal and together increase the size of this species group to 23. A key to all species in the E. scaber species group is also provided. Key words: Camel spider, solifugid, revision, taxonomy Introduction Solifugae, commonly known as camel spiders, is a poorly studied order of arachnids. It currently includes 12 families and over 1,100 described species. The phylogeny of the order and of all but one family is currently unresolved. In recent years, our lab has focused on the phylogeny, biology, and natural history of the North American family Eremobatidae Kraepelin 1899 (Brookhart & Cushing 2002, 2004, 2005; Cushing et al. 2005, 2014, 2015; Conrad & Cushing 2011; Cushing & Casto 2012). Cushing et al. (2015) produced a backbone molecular phylogeny of the Eremobatidae that supported the monophyly of several genera and species groups within the family. -
Geological History and Phylogeny of Chelicerata
Arthropod Structure & Development 39 (2010) 124–142 Contents lists available at ScienceDirect Arthropod Structure & Development journal homepage: www.elsevier.com/locate/asd Review Article Geological history and phylogeny of Chelicerata Jason A. Dunlop* Museum fu¨r Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt University Berlin, Invalidenstraße 43, D-10115 Berlin, Germany article info abstract Article history: Chelicerata probably appeared during the Cambrian period. Their precise origins remain unclear, but may Received 1 December 2009 lie among the so-called great appendage arthropods. By the late Cambrian there is evidence for both Accepted 13 January 2010 Pycnogonida and Euchelicerata. Relationships between the principal euchelicerate lineages are unre- solved, but Xiphosura, Eurypterida and Chasmataspidida (the last two extinct), are all known as body Keywords: fossils from the Ordovician. The fourth group, Arachnida, was found monophyletic in most recent studies. Arachnida Arachnids are known unequivocally from the Silurian (a putative Ordovician mite remains controversial), Fossil record and the balance of evidence favours a common, terrestrial ancestor. Recent work recognises four prin- Phylogeny Evolutionary tree cipal arachnid clades: Stethostomata, Haplocnemata, Acaromorpha and Pantetrapulmonata, of which the pantetrapulmonates (spiders and their relatives) are probably the most robust grouping. Stethostomata includes Scorpiones (Silurian–Recent) and Opiliones (Devonian–Recent), while -
By Heterophrynus Sp. (Arachnida, Phrynidae) in a Cave in the Chapada Das Mesas National Park, State of Maranhão, Brazil
Crossref 10 ANOS Similarity Check Powered by iThenticate SCIENTIFIC NOTE DOI: http://dx.doi.org/10.18561/2179-5746/biotaamazonia.v10n1p49-52 Predation of Tropidurus oreadicus (Reptilia, Tropiduridae) by Heterophrynus sp. (Arachnida, Phrynidae) in a cave in the Chapada das Mesas National Park, state of Maranhão, Brazil Fábio Antônio de Oliveira1, Gabriel de Avila Batista2, Karla Dayane de Lima Pereira3, Lucas Gabriel Machado Frota4, Victoria Sousa5, Layla Simone dos Santos Cruz6, Karll Cavalcante Pinto7 1. Biólogo (Pontifícia Universidade Católica de Goiás, Brasil). Doutorando em Geologia (Universidade de Brasília, Brasil). [email protected] http://lattes.cnpq.br/6651314736341253 http://orcid.org/0000-0001-8125-6339 2. Biólogo (Anhanguera Educacional, Brasil). Doutorando em Recursos Naturais do Cerrado (Universidade Estadual de Goiás, Brasil). [email protected] http://lattes.cnpq.br/1131941234593219 http://orcid.org/0000-0003-4284-2591 3. Bióloga (Anhanguera Educacional, Brasil). Mestranda em Conservação de Recursos Naturais do Cerrado (Instituto Federal Goiano, Brasil). [email protected] http://lattes.cnpq.br/4328373742442270 http://orcid.org/0000-0003-1578-8948 4. Biólogo (Pontifícia Universidade Católica de Goiás, Brasil). Analista Ambiental da Biota Projetos e Consultoria Ambiental LTDA, Brasil. [email protected] http://lattes.cnpq.br/7083829373324504 http://orcid.org/0000-0001-6907-9480 5. Bióloga (Pontifícia Universidade Católica de Goiás, Brasil). Mestranda em Ecologia e Evolução (Universidade Federal de Goiás, Brasil). [email protected] http://lattes.cnpq.br/0747140109675656 http://orcid.org/0000-0002-2818-5698 6. Bióloga (Centro Universitário de Goiás, Brasil). Especialista em Perícia, Auditoria e Gestão Ambiental (Instituto de Especialização e Pós-Graduação (IEPG)/Faculdade Oswaldo Cruz, Brasil). -
A Guide to Arthropods Bandelier National Monument
A Guide to Arthropods Bandelier National Monument Top left: Melanoplus akinus Top right: Vanessa cardui Bottom left: Elodes sp. Bottom right: Wolf Spider (Family Lycosidae) by David Lightfoot Compiled by Theresa Murphy Nov 2012 In collaboration with Collin Haffey, Craig Allen, David Lightfoot, Sandra Brantley and Kay Beeley WHAT ARE ARTHROPODS? And why are they important? What’s the difference between Arthropods and Insects? Most of this guide is comprised of insects. These are animals that have three body segments- head, thorax, and abdomen, three pairs of legs, and usually have wings, although there are several wingless forms of insects. Insects are of the Class Insecta and they make up the largest class of the phylum called Arthropoda (arthropods). However, the phylum Arthopoda includes other groups as well including Crustacea (crabs, lobsters, shrimps, barnacles, etc.), Myriapoda (millipedes, centipedes, etc.) and Arachnida (scorpions, king crabs, spiders, mites, ticks, etc.). Arthropods including insects and all other animals in this phylum are characterized as animals with a tough outer exoskeleton or body-shell and flexible jointed limbs that allow the animal to move. Although this guide is comprised mostly of insects, some members of the Myriapoda and Arachnida can also be found here. Remember they are all arthropods but only some of them are true ‘insects’. Entomologist - A scientist who focuses on the study of insects! What’s bugging entomologists? Although we tend to call all insects ‘bugs’ according to entomology a ‘true bug’ must be of the Order Hemiptera. So what exactly makes an insect a bug? Insects in the order Hemiptera have sucking, beak-like mouthparts, which are tucked under their “chin” when Metallic Green Bee (Agapostemon sp.) not in use. -
Solifugids (Arachnida, Solifugae) Use Adhesive Organs on Their Pedipalps for Prey Capture
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Eileen Hebets Publications Papers in the Biological Sciences 2011 A sticky situation: Solifugids (Arachnida, Solifugae) use adhesive organs on their pedipalps for prey capture Rodrigo H. Willemart University of Nebraska-Lincoln, [email protected] Roger D. Santer University of Nebraska-Lincoln, [email protected] Andrew J. Spence Royal Veterinary College, University of London, [email protected] Eileen Hebets University of Nebraska - Lincoln, [email protected] Follow this and additional works at: https://digitalcommons.unl.edu/bioscihebets Part of the Behavior and Ethology Commons Willemart, Rodrigo H.; Santer, Roger D.; Spence, Andrew J.; and Hebets, Eileen, "A sticky situation: Solifugids (Arachnida, Solifugae) use adhesive organs on their pedipalps for prey capture" (2011). Eileen Hebets Publications. 39. https://digitalcommons.unl.edu/bioscihebets/39 This Article is brought to you for free and open access by the Papers in the Biological Sciences at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Eileen Hebets Publications by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. Published in Journal of Ethology 29 (2011), pp. 177–180; doi: 10.1007/s10164-010-0222-4 Copyright © 2010 Japan Ethological Society and Springer. Used by permission. Submitted February 23, 2010; accepted May 17, 2010; published online June 12, 2010 Supplementary material for this article is included following the “References.” A sticky situation: Solifugids (Arachnida, Solifugae) use adhesive organs on their pedipalps for prey capture Rodrigo H. Willemart,1 Roger D. Santer,1 Andrew J. Spence,2 and Eileen A. Hebets 1 1. School of Biological Sciences, University of Nebraska–Lincoln, Lincoln, NE, USA 2. -
Fossil Calibrations for the Arthropod Tree of Life
bioRxiv preprint doi: https://doi.org/10.1101/044859; this version posted June 10, 2016. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY 4.0 International license. FOSSIL CALIBRATIONS FOR THE ARTHROPOD TREE OF LIFE AUTHORS Joanna M. Wolfe1*, Allison C. Daley2,3, David A. Legg3, Gregory D. Edgecombe4 1 Department of Earth, Atmospheric & Planetary Sciences, Massachusetts Institute of Technology, Cambridge, MA 02139, USA 2 Department of Zoology, University of Oxford, South Parks Road, Oxford OX1 3PS, UK 3 Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PZ, UK 4 Department of Earth Sciences, The Natural History Museum, Cromwell Road, London SW7 5BD, UK *Corresponding author: [email protected] ABSTRACT Fossil age data and molecular sequences are increasingly combined to establish a timescale for the Tree of Life. Arthropods, as the most species-rich and morphologically disparate animal phylum, have received substantial attention, particularly with regard to questions such as the timing of habitat shifts (e.g. terrestrialisation), genome evolution (e.g. gene family duplication and functional evolution), origins of novel characters and behaviours (e.g. wings and flight, venom, silk), biogeography, rate of diversification (e.g. Cambrian explosion, insect coevolution with angiosperms, evolution of crab body plans), and the evolution of arthropod microbiomes. We present herein a series of rigorously vetted calibration fossils for arthropod evolutionary history, taking into account recently published guidelines for best practice in fossil calibration. -
Curriculum Vitae
Paula E. Cushing, Ph.D. Senior Curator of Invertebrate Zoology, Department of Zoology, Denver Museum of Nature & Science, 2001 Denver,Colorado 80205 USA E-mail: [email protected]; Web: https://science.dmns.org/museum- scientists/paula-cushing/, http://www.solifugae.info, http://spiders.dmns.org/default.aspx; Office Phone: (303) 370-6442; Lab Phone: (303) 370-7223; Fax: (303) 331-6492 CURRICULUM VITAE EDUCATION University of Florida, Gainesville, FL, 1990 - 1995, Ph.D. Virginia Tech, Blacksburg, VA, 1985 - 1988, M.Sc. Virginia Tech, Blacksburg, VA, 1982 - 1985, B.Sc. PROFESSIONAL AFFILIATIONS AND POSITIONS Denver Museum of Nature & Science, Senior Curator of Invertebrate Zoology, 1998 - present Associate Professor Adjoint, Department of Integrative Biology, University of Colorado, Denver, 2013 - present Denver Museum of Nature & Science, Department Chair of Zoology, 2006 – 2011 Adjunct Faculty, Department of Ecology & Evolutionary Biology, University of Colorado, Boulder, 2011 - present Affiliate Faculty Member, Department of Biology and Wildlife, University of Alaska, Fairbanks, 2009 – 2012 Adjunct Faculty, Department of Bioagricultural Sciences and Pest Management, Colorado State University, 1999 – present Affiliate Faculty Member, Department of Life, Earth, and Environmental Science, West Texas AMU, 2011 - present American Museum of Natural History, Research Collaborator (Co-PI with Lorenzo Prendini), 2007 – 2012 College of Wooster, Wooster, Ohio, Visiting Assistant Professor, 1996 – 1997 University of Florida, Gainesville, Florida, Postdoctoral Teaching Associate, 1996 Division of Plant Industry, Gainesville, Florida, Curatorial Assistant for the Florida Arthropod Collection, 1991 National Museum of Natural History (Smithsonian), high school intern at the Insect Zoo, 1981; volunteer, 1982 GRANTS AND AWARDS 2018 - 2022: NSF Collaborative Research: ARTS: North American camel spiders (Arachnida, Solifugae, Eremobatidae): systematic revision and biogeography of an understudied taxon. -
On the Habitat Use of the Neotropical Whip Spider Charinus Asturius (Arachnida: Amblypygi)
ZOOLOGIA 35: e12874 ISSN 1984-4689 (online) zoologia.pensoft.net RESEARCH ARTICLE On the habitat use of the Neotropical whip spider Charinus asturius (Arachnida: Amblypygi) Júlio M.G. Segovia1,2, Lúcia C. Neco3, Rodrigo H. Willemart1,2,4 1Laboratório de Ecologia Sensorial e Comportamento de Artrópodes, Escola de Artes, Ciências e Humanidades, Universidade de São Paulo. Rua Arlindo Béttio 1000, Ermelino Matarazzo, 03828-000 São Paulo, SP, Brazil. 2Programa de Pós Graduação em Zoologia, Instituto de Biociências, Universidade de São Paulo. Rua do Matão 321, Travessa 14, 05508-900 São Paulo, Brazil. 3Departamento de Psicologia Experimental, Instituto de Psicologia, Universidade de São Paulo. Avenida Professor Mello de Morais, 1721 Butantã, 05508030 São Paulo, SP, Brazil. 4Programa de Pós Graduação em Ecologia e Evolução, Universidade Federal de São Paulo. Rua Professor Artur Riedel 275, 09972-270 Diadema, Brazil. Corresponding author: Júlio M.G. Segovia ([email protected]) http://zoobank.org/51469941-5834-4AA2-A1DB-A424F0377392 ABSTRACT. The non-random occupation of habitats is termed habitat selection. Some species of whip spiders select trees with burrows at their base, while others use substrates such as rocks. Here, we investigated the habitat use by Charinus asturius Pinto-da-Rocha, Machado & Weygoldt, 2002, an endemic species of Ilhabela Island in Brazil. We found that C. asturius is more likely to be found under rocks that cover larger areas of substrate. Our results also suggest the existence of territorialism in C. asturius and show that C. asturius adults may be found again on the same rock a week later. Additionally, our data show that C.